taxonID	type	description	language	source
03C53526FFCFFFC7FEC1F9B8BA39F7F6.taxon	description	(Figs 1 – 2)	en	Yin, Zi-Wei, Kurbatov, Sergey A., Cuccodoro, Giulio, Cai, Chen-Yang (2019): Cretobrachygluta gen. nov., the first and oldest Brachyglutini in mid-Cretaceous amber from Myanmar (Coleoptera: Staphylinidae: Pselaphinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 101-106, DOI: 10.2478/aemnp-2019-0008, URL: http://dx.doi.org/10.2478/aemnp-2019-0008
03C53526FFCFFFC7FEC1F9B8BA39F7F6.taxon	type_taxon	Type species. † Cretobrachygluta laurasiensis sp. nov., here designated.	en	Yin, Zi-Wei, Kurbatov, Sergey A., Cuccodoro, Giulio, Cai, Chen-Yang (2019): Cretobrachygluta gen. nov., the first and oldest Brachyglutini in mid-Cretaceous amber from Myanmar (Coleoptera: Staphylinidae: Pselaphinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 101-106, DOI: 10.2478/aemnp-2019-0008, URL: http://dx.doi.org/10.2478/aemnp-2019-0008
03C53526FFCFFFC7FEC1F9B8BA39F7F6.taxon	diagnosis	Diagnosis. Head with small, nude vertexal foveae, lacking sulcus connecting foveae; antennae with club formed by apical three antennomeres; gular region with broad median ridge. Pronotum with small, nude median and lateral antebasal foveae. Each elytron with four basal foveae; subhumeral foveae and marginal striae present; discal striae absent. Metacoxae contiguous, moderately projecting posteriorly; tarsomere 2 distinctly shorter than tarsomere 3; tarsi each with single pretarsal claw. Abdomen with tergite 1 (IV) much longer than 2 (V); tergites 1 – 4 (IV – VII) with broad paratergites; tergite 1 (IV) with two basolateral foveae, lacking mediobasal fovea; sternite 1 (III) as long as 2 (IV) along midline.	en	Yin, Zi-Wei, Kurbatov, Sergey A., Cuccodoro, Giulio, Cai, Chen-Yang (2019): Cretobrachygluta gen. nov., the first and oldest Brachyglutini in mid-Cretaceous amber from Myanmar (Coleoptera: Staphylinidae: Pselaphinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 101-106, DOI: 10.2478/aemnp-2019-0008, URL: http://dx.doi.org/10.2478/aemnp-2019-0008
03C53526FFCFFFC7FEC1F9B8BA39F7F6.taxon	description	Description. Body length slightly over 1.8 mm; habitus (Fig. 1) strongly convex dorso-ventrally, in dorsal view strongly broadened posterior to pronotum. Head (Fig. 2 B) roundly quadrate, slightly wider than pronotum; vertex raised, with pair of small and nude vertexal foveae (Fig. 2 A; vf); frons shallowly and broadly impressed, impression flanked laterally by widely separated antennal tubercles, and demarcated anteriorly by broad frontal-clypeal ridge; ocular-mandibular carinae (Fig. 2 D; omc) distinct; gular region broadly impressed at middle, two close but clearly separated foveae (Fig. 2 C; gf) in impression; broad, longitudinally oval median gular ridge present anterior to foveae and extending to mouthparts, demarcated laterally by sulci. Antennae (Fig. 2 A) eleven-segmented; club well-developed, formed by apical three antennomeres, with sub-oval and basally truncate antennomeres 11 (Fig. 2 A; a 11) much wider than preceding antennomeres. Maxillary palpi four-segmented; palpomere 1 minute, elongate, palpomere 2 pedunculate in basal half, strongly broadening toward apex, palpomere 3 subtriangular, approximately as wide and half as long as palpomere 4, palpomere 4 (Fig. 2 B; mp 4) largest, sub-oval, with elongate apical palpal cone. Pronotum (Fig. 2 B) slightly elongate, widest at apical third; lateral margins rounded, sides convergent toward apex at apical two-fifths and narrowing toward base at basal third; disc with nude median (Fig. 2 B; maf) and lateral antebasal (Fig. 2 B; laf) foveae subequal in size, lacking additional carinae, sulci or tubercles; paranotal carinae either short, or lacking; lateral procoxal foveae present, moderately separated. Elytra (Fig. 2 B) slightly wider than long, widest point posterior to middle; each elytron with four small but distinct basal foveae (Fig. 2 B; bef); sutural striae (Fig. 2 B; ss) complete; discal striae absent; subhumeral foveae (Figs 2 B, 2 D; shef) present; short marginal striae (Figs 2 B, 2 D; ms) extending from subhumeral foveae to basal third of elytral length; posterolateral clefts (Fig. 2 B; plc) broad and shallow. Hind wings fully developed. Thorax with narrowly separated median mesoventral foveae (Fig. 2 C; mmsf); lateral mesoventral foveae (Fig. 2 D; lmsf) simple; lateral mesocoxal foveae (Fig. 2 C; lmcf) and lateral metaventral foveae (Fig. 2 C; lmtf) present; lateral sutures separating meso- and metaventrite well-marked, fully developed; posterior margin of metaventrite with narrow split at middle. Abdomen (Figs 2 A – C) dorsoventrally flattened, with generally rounded lateral margins; segments telescoping. Five tergites (IV – VIII) (Figs 2 A, 2 B; tIV – VIII) visible, with broad paratergites on segments 1 – 4 (IV – VII); tergite 1 (IV) much longer than tergite 2 (V), sides narrowing toward base, lacking mediobasal foveae, with pair of basolateral foveae (Fig. 2 B; blf), short triangular discal carinae (Fig. 2 B; dc), and broad, shallow transverse impression at base. Six sternites (III – VIII) (Fig. 2 C; sIII – VIII) visible; sternites 1 (III) and 2 (IV) subequal in length along midline; sternite 2 (IV) lacking mediobasal foveae and basal impression, with small basolateral foveae. Foveal pattern of remaining abdominal segments barely visible based on available specimen, but one pair of basolateral foveae putatively present on each of tergites 2 – 4 (V – VII). Legs moderately elongate; all three pairs of coxae contiguous; metacoxae (Fig. 2 C; mtc) moderately projecting posteriorly; dorsal margin of all trochanters short, base of femora thus close to coxal-trochanteral articulation. Tarsi three-segmented (Fig. 2 box; t 1 – 3), with short tarsomere 1, and longer tarsomeres 2 and 3; tarsomere 2 distinctly shorter than tarsomere 3; tarsi each with one elongate claw (Fig. 2 box; tc). Male mesotrochanters and mesotibiae modified on ventral margin and at near apex, respectively; fore and hind legs simple.	en	Yin, Zi-Wei, Kurbatov, Sergey A., Cuccodoro, Giulio, Cai, Chen-Yang (2019): Cretobrachygluta gen. nov., the first and oldest Brachyglutini in mid-Cretaceous amber from Myanmar (Coleoptera: Staphylinidae: Pselaphinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 101-106, DOI: 10.2478/aemnp-2019-0008, URL: http://dx.doi.org/10.2478/aemnp-2019-0008
03C53526FFCFFFC7FEC1F9B8BA39F7F6.taxon	etymology	Etymology. The generic name is a combination of the prefix ‘ Creto - ’ derived from the Cretaceous, and Brachygluta Thomson, 1859, type genus of Brachyglutini. The gender is feminine.	en	Yin, Zi-Wei, Kurbatov, Sergey A., Cuccodoro, Giulio, Cai, Chen-Yang (2019): Cretobrachygluta gen. nov., the first and oldest Brachyglutini in mid-Cretaceous amber from Myanmar (Coleoptera: Staphylinidae: Pselaphinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 101-106, DOI: 10.2478/aemnp-2019-0008, URL: http://dx.doi.org/10.2478/aemnp-2019-0008
03C53526FFCCFFC1FC78FA52BA97FE3D.taxon	materials_examined	Type material. HOLOTYPE (SNUC-Paleo- 0076): a well-preserved male in an approximately 7.2 × 5.0 mm clear, yellowish amber piece; upper Albian to lower Cenomanian, Hukawng Valley, northern Myanmar (SNUC).	en	Yin, Zi-Wei, Kurbatov, Sergey A., Cuccodoro, Giulio, Cai, Chen-Yang (2019): Cretobrachygluta gen. nov., the first and oldest Brachyglutini in mid-Cretaceous amber from Myanmar (Coleoptera: Staphylinidae: Pselaphinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 101-106, DOI: 10.2478/aemnp-2019-0008, URL: http://dx.doi.org/10.2478/aemnp-2019-0008
03C53526FFCCFFC1FC78FA52BA97FE3D.taxon	diagnosis	Diagnosis of male. As for the genus (vide supra), plus the following: body length 1.85 mm; mesotrochanters protuberant at ventral margin; mesotibiae with small, triangular preapical spur.	en	Yin, Zi-Wei, Kurbatov, Sergey A., Cuccodoro, Giulio, Cai, Chen-Yang (2019): Cretobrachygluta gen. nov., the first and oldest Brachyglutini in mid-Cretaceous amber from Myanmar (Coleoptera: Staphylinidae: Pselaphinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 101-106, DOI: 10.2478/aemnp-2019-0008, URL: http://dx.doi.org/10.2478/aemnp-2019-0008
03C53526FFCCFFC1FC78FA52BA97FE3D.taxon	description	Description. Habitus (Fig. 1) rather stout; total body length (combined length of head, pronotum, elytra, and abdomen) 1.85 mm. Body reddish brown, with mouthparts and tarsi paler, integument of dorsal surface densely covered with suberect fine setae. Head slightly broader than long; length from anterior margin of clypeus toward base 0.30 mm, width across eyes 0.41 mm; eyes roundly prominent, each composed of approximately 75 facets. Length of antenna 0.86 mm; antennomere I (scape) cylindrical, about 1.65 times as long as wide, antennomere II (pedicel) roundedcylindrical, much narrower than scape, 1.2 times as long as wide, and about 0.6 times as long as scape, antennomeres 3 – 7 almost moniliform, slightly elongate, antennomere 8 distinctly transverse, antennomeres 9 – 11 obconical, successively wider apically, antennomere 11 about twice as long as antennomere 10, truncate at base, broadest at near middle, and narrowing toward apex, with inner margin broadly emarginate in apical half (observable in Fig. 1 A, right antenna). Maxillary palpi as in Figs 2 B – D; length of palpomere 1 0.03 mm, palpomere 2 0.12 mm, palpomere 3 0.06 mm, palpomere 4 0.12 mm. Pronotum slightly longer than wide, length along midline 0.41 mm, maximum width 0.37 mm; sides roundly expanded laterally. Elytra transverse, length along suture 0.58 mm, maximum width 0.66 mm, anterior margin broadly impressed, posterior margin truncate. Legs moderately elongate, densely setose; ventral margin of mesotrochanters projecting in conspicuous process; mesotibiae bearing small, triangular preapical spur. Abdomen broadest at tergite 1 (IV), length along midline 0.66 mm, maximum width 0.57 mm; tergites and sternites densely covered with suberect, posteriorly directed setae; tergite VIII with deep apical notch at middle of posterior margin.	en	Yin, Zi-Wei, Kurbatov, Sergey A., Cuccodoro, Giulio, Cai, Chen-Yang (2019): Cretobrachygluta gen. nov., the first and oldest Brachyglutini in mid-Cretaceous amber from Myanmar (Coleoptera: Staphylinidae: Pselaphinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 101-106, DOI: 10.2478/aemnp-2019-0008, URL: http://dx.doi.org/10.2478/aemnp-2019-0008
03C53526FFCCFFC1FC78FA52BA97FE3D.taxon	discussion	Remarks. The protuberant mesotrochanters, spinose mesotibiae, and apically notched tergite VIII are interpreted as male sexual characters based on our experience in the extant brachyglutine fauna.	en	Yin, Zi-Wei, Kurbatov, Sergey A., Cuccodoro, Giulio, Cai, Chen-Yang (2019): Cretobrachygluta gen. nov., the first and oldest Brachyglutini in mid-Cretaceous amber from Myanmar (Coleoptera: Staphylinidae: Pselaphinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 101-106, DOI: 10.2478/aemnp-2019-0008, URL: http://dx.doi.org/10.2478/aemnp-2019-0008
03C53526FFCCFFC1FC78FA52BA97FE3D.taxon	etymology	Etymology. The specific epithet refers to the occurrence of this species on the supercontinent Laurasia some 99 million years ago. Adjective.	en	Yin, Zi-Wei, Kurbatov, Sergey A., Cuccodoro, Giulio, Cai, Chen-Yang (2019): Cretobrachygluta gen. nov., the first and oldest Brachyglutini in mid-Cretaceous amber from Myanmar (Coleoptera: Staphylinidae: Pselaphinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 101-106, DOI: 10.2478/aemnp-2019-0008, URL: http://dx.doi.org/10.2478/aemnp-2019-0008
