taxonID	type	description	language	source
03E6755EA3527716C33EFF46FC5FFE34.taxon	description	(Figs 2 A – I)	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA3527716C33EFF46FC5FFE34.taxon	materials_examined	Type locality. Central Vietnam, Th ừa Thiên-Hu ế Prov., B ạch Mã National Park. Type material (15 spec.). HOLOTYPE: ♂ (VNMN), ‘ C VIETNAM: Thua Thien Hue | Prov., Bach Ma National Park | (16.193 ° N 107.853 ° E) 1250 m [p] || 28. V – 1. VI. 2017 L. Bartolozzi | E. Orbach, V. Sbordoni, | S. Bambi & A. Bandinelli leg. | (numero Mag. 3089) [p] ’. PARATYPES: 1 ♂ (ZMPC), ‘ Dambri. B ảo Lâm. | Lâm Đ ồng, Vietnam. | March, 2017, | local collector leg. [p] ’; 3 ♂♂ 2 ♀♀ (CZPC), ‘ Dambri. B ảo Lâm. | Lâm Đ ồng, Vietnam. | March, 2018, | local collector leg. [p] ’; 4 ♂♂ 4 ♀♀ (CZPC), ‘ Dambri. B ảo Lâm. | Lâm Đ ồng, Vietnam. | April, 2018, | local collector leg. [p] ’.	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA3527716C33EFF46FC5FFE34.taxon	description	Description of holotype (♂). Body shape. Elongate ovoid, weakly convex. Color. Ground color blackish-brown with strong dark green metallic luster; antennomeres 1 – 6, maxillary palpi, labial palpi and lateral teeth of protibia dark orange, antennal club black; elytra black with two arcs of rather small yellow spots: 6 anterior spots encircling scutellum and 8 additional spots of which the most anterior is situated beneath humeral umbone and the most posterior lies in interstice 1 and costal interval 2 at ca. 2 / 3 of elytral length. Head. Clypeus subtrapezoidal, disc very densely, transversely rugopunctate, anterior corners rounded; anterior margin weakly reflexed; frons rugopunctate and with very shallow impression at middle, confluently punctate at sides; vertex finely and sparsely punctate at middle, coarsely punctate laterally; ratio of interocular width / width of head approximately 0.64; antennal club longer than antennomeres 1 – 6 combined. Pronotum approximately 1.56 times wider than long, with oblique impression posteriorly of middle on each side and median longitudinal furrow, with steep decline along lateral margin; sharply defined area posteriorly of anterior angle with deep, coarse, confluent punctures, area of lateral impression with coarse, separate punctures, remaining surface smooth with scarce micropunctures; with sparse, erect setae at lateral margin; anterior angles acute and strongly produced; posterior angles rectangular; sides of pronotum slightly converging anteriad in posterior half, strongly converging in anterior half; basal marginal line present only near posterior angle; all other marginal lines complete. Scutellum nearly semicircular, broader than long, finely and very sparsely punctate. Elytra regularly striate; costal intervals and interstices similarly slightly convex; strial punctures distinct, coarse, subsutural interstice with a secondary stria irregularly doubled anteriorly and disappearing in posterior quarter; vague secondary striae also present in anterior part of interstice 2 (anterior 1 / 5) and 3 (anterior half); elytral surface with sparse micropunctation; humeral umbone and apical protuberance prominent; opaque area at apical curvature narrow; lateral margin with flat paramarginal area between humerus and middle of elytra; epipleuron broad near humerus, ending slightly posteriorly of middle of elytron; epipleuron with soft, white setae continuing along lateral margin to apical curvature; another 2 – 3 large white setae near and at apicosutural angle; posterior margins evenly, separately rounded. Propygidium with dense fringe of white, rather short setae along posterior margin. Pygidium strongly convex; apex broadly rounded; punctation densely punctate-imbricate, transversely confluent; with scattered white setae near base; apex with several long, erect brownish setae. Ventral thoracic surface densely covered with soft, long, white setae. Meso-metaventral process short, compressed between mesocoxae, projecting slightly downward in lateral view, anteriorly vertical and straight; apex subrectangular, somewhat rounded; bulbiform in ventral view. Abdominal ventrites with transverse band of rather sparse white setae (usually interrupted in middle) and irregular white setae on each side of anterior half of ventrites 2 – 4; ventrites 1 – 2 and anterior half of ventrite 3 carinate laterally. Legs. Mesofemur and metafemur with two bands of long white setae, one along anterior margin, the second emerging from a transverse row of punctures parallel to posterior margin. Protibia bidentate, rather slender, approximately 3.9 times longer than wide; proximal tooth short, situated close to the rather long, weakly curved apical tooth; inner spur short, positioned at level of space between proximal and apical teeth. Metatibia fusiform. Protarsomere 5 (without claws) slightly longer than protarsomeres 1 – 4 combined; inner protarsal claw approximately 3 / 4 as long as protarsomere 5, strongly curved and deeply incised apically, upper branch spiniform, lower branch broad, obliquely truncated; outer mesotarsal claw approximately as long as combined length of mesotarsomeres 1 – 4, curved, deeply incised at apex, upper branch spiniform; metatarsal claws very unequal, outer claw approximately twice as thick and distinctly longer than inner. Aedeagus as in Figs 2 F – H. Female. Protibia slender, apical tooth of protibia long and spatulate; protarsus articulated slightly basally of level of proximal tooth; inner spur long, articulated between 1 / 2 and 2 / 3 of tibial length; protarsus very slender, protarsomere 5 (without claws) shorter than tarsomeres 1 – 4 combined; modified claws of pro- and mesotarsus shorter, two apical branches more equal than in males; antennal club slightly longer than antennomeres 2 – 6 combined. Measurements. Total body length 10.4 – 11.9 mm (HT 11.9 mm), total body width 6.0 – 7.2 mm (HT 7.2 mm). Morphological variation. All six paratypes have the sides of the pronotum suffused with orange and dark orange femurs and tibiae with the knee joints and apices of tibiae dark metallic green. The size of the yellow elytral spots is variable. Some spots become divided in specimens with generally smaller spots. Number of elytral setae near apico-sutural angle varies between two and five. Shape of aedeagus remains consistent.	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA3527716C33EFF46FC5FFE34.taxon	diagnosis	Differential diagnosis. Glenopopillia albopilosa sp. nov. is clearly separated from all other species in this genus by several characters: the setae on the elytral margin and epipleuron are soft and white, not rigid and dark as in all other species. The pronotum has only one oblique lateral impression (none in G. klossi, two in all other species). The coarsely punctate areas of the pronotum are sharply defined (more gradually changing in the other species). The legs are generally more slender than in all other species; the apical tooth of the protibia is laterally protruding beyond the proximal tooth (not surpassing the proximal tooth in all other species). The antennal club is very long in males, longer than the combined length of the remaining antennomeres (longer than antennomeres 2 – 6 combined in the other species). The elytra are predominantly black in G. albopilosa sp. nov. whereas larger parts are light colored in the other species.	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA3527716C33EFF46FC5FFE34.taxon	etymology	Etymology. The specific epithet (adjective in the nominative singular case) references the color of the setae on the lateral elytral margin, which is white and not blackish- brown as in all the other species of this genus. Collecting circumstances. The holotype was collected with a beating sheet at the forest edge (L. Bartolozzi, pers. comm.).	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA3527716C33EFF46FC5FFE34.taxon	distribution	Distribution. Central Vietnam, Th ừa Thiên-Hu ế and Lâm Đ ồng Provinces.	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA3527705C108FF0EFD46FBF7.taxon	type_taxon	Type species. Glenopopillia maculata Lin, 1980, by original designation.	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA3527705C108FF0EFD46FBF7.taxon	description	Generic characters. Scarabaeidae, Rutelinae, Anomalini, Popilliina. Body elongate ovoid, dorsum weakly convex. Length 10.0 – 14.0 mm. Body with strong greenish or reddish metallic luster, except for elytra which are contrastingly orange brown or maculate black-yellow and legs which are partly orange. Clypeus weakly reflexed. Pronotum with two large, deep oblique impressions near each lateral margin (except for Glenopopillia albopilosa sp. nov. and G. klossi), with anterior angles acute and strongly produced and posterior angles obtuse; sides strongly converging in anterior third; basal margin of pronotum straight before scutellum, basal marginal line interrupted medially; surface finely and sparsely punctate. Elytron with prominent humeral umbone and apical protuberance; posterior margin evenly rounded; apicosutural angle dentiform; surface with regular punctate striae; all intervals slightly convex; apical curvature lined by a more or less extensive opaque area; elytral lateral margin with row of usually dark, rigid setae (all setae white and soft only in G. albopilosa sp. nov.), which are present also at apico-sutural angle, except for G. klossi. Base of mesepimeron not exposed beyond elytral base in dorsal view. Meso-metaventral process short (previously known as mesosternal process), compressed between mesocoxae, anteriorly vertical, straight; apex subcircular, bulbiform in ventral view. Posterior margin of propygidium exposed by elytra, with fringe of white setae, except for G. klossi. Pygidium strongly convex in males. Protibia bidentate. Metatibia strongly fusiform. Proximal abdominal ventrites laterally carinate, all abdominal ventrites with transverse band of whitish setae. Sexual dimorphism present as follows: protibia wide with approximated teeth (terminal tooth short and pointed) and spur short in males vs. protibia more slender with teeth more distant (terminal tooth long and spatulate) and spur long in females; protarsus somewhat thicker and protarsomeres 1 – 4 shorter in males, while protarsus very slender and protarsomeres 1 – 4 longer in females; inner protarsal claw and outer mesotarsal claw very long, cleft, with upper branch spiniform in males, shorter and less unequal in females; metatarsal claws very unequal in males, more equal in females.	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA3527705C108FF0EFD46FBF7.taxon	diagnosis	Diagnosis. The character of the long, rigid or soft setae along the lateral margin of the elytra (soft only in G. albopilosa sp. nov.), with similar setae usually present at the apico-sutural angle, is unique within the subtribe Popilliina.	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA3527705C108FF0EFD46FBF7.taxon	discussion	Identification key to the Glenopopillia species	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA354771AC3CEFE46FDCBFB14.taxon	description	(Figs 3 A, G; 4 A, G, M; 5 A, G, M)	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA354771AC3CEFE46FDCBFB14.taxon	materials_examined	Type locality. India, Meghalaya, 3 km E of Tura, 25 ° 30 ’ N 90 ° 14 ’ E, 500 – 1150 m a. s. l. Type material (21 spec.). HOLOTYPE: ♂ (NMPC), ‘ NE INDIA; MEGHALAYA; 1999 | 3 km E of Tura; 500 – 1150 m; | 25 ° 30 ’ N 90 ° 14 ’ E; 1. – 8. v. | Zd. Koštál lgt [p] ’. PARATYPES: 3 ♂♂ 12 ♀♀ (CZPC), same data as holotype; 1 ♂ 2 ♀♀ (IZAS), same data as holotype; 1 ♀ (CZPC), ‘ NE INDIA; MEGHALAYA; 1999 | 3 km E of Tura; 500 – 1150 m; | 25 ° 30 ’ N 90 ° 14 ’ E; 15. – 22. iv. | Zd. Koštál lgt [p] ’; 1 ♀ (PPCB), ‘ NE INDIA, Meghalaya state | West Garo Hills reg., TURA | 5 – 7. V. 1996, alt. 700 ± 100 m, | GPS N 25 ° 30.7 ’ E 90 ° 13.9 (WGS 84) | E. Jendek & O. Šauša leg. || Coll. P. Pacholátko | Brno | Merhautova 68 | Czech republic [p] ’.	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA354771AC3CEFE46FDCBFB14.taxon	description	Description of holotype (♂). Body shape. Elongate ovoid, weakly convex. Color. Ground color blackish-brown with strong green to red metallic luster; legs dark orange except for mesoand metatarsus, meso- and metafemur, and apical part of metatibia being dark brown with green metallic luster; clypeus and antenna light brown; elytra including all margins, epipleura and suture blackish-brown with weak metallic sheen; a large oblique reniform yellow spot between humerus and costal interval 1 posteriorly not reaching middle of elytron, one small elongate yellow spot in anterior part of costal interval 5, two small yellow spots slightly posteriorly of middle: one elongate in costal interval 3, another transverse extending from costal interval 4 to 5, and a subquadrate yellow spot in the apical 4 / 5 extending from costal interval 1 to 2. Head. Clypeus subrectangular, disc very densely, transversely, confluently punctate; anterior corners rounded; anterior margin weakly reflexed; frons very shallowly impressed, laterally punctate like clypeus, separately punctate at middle; vertex very sparsely punctate; ratio of interocular width / width of head approximately 0.67; antennal club slightly longer than antennomeres 2 – 6 combined. Pronotum approximately 1.5 times wider than long, with two deep, oblique impressions on each side (posterior impression larger than anterior) and deep median longitudinal furrow, with steep decline along lateral margin; punctation of pronotum distinct between lateral impressions, extremely shallow and sparse only in very middle, punctures becoming gradually larger laterally; sparse erect setae present near anterior angles and along lateral margin; anterior angles acute and strongly produced; posterior angles obtuse; sides of pronotum converging anteriad in posterior two thirds, then strongly curved, strongly converging, and slightly sinuate in anterior third; basal marginal line interrupted before scutellum between level of elytral interstice 2 on each side; all other marginal lines complete. Scutellum nearly semicircular, broader than long, finely and sparsely punctate. Elytra regularly striate; three inner costal intervals (1, 2 and 3) slightly more convex than interstices; strial punctures distinct, subsutural interstice with secondary stria being irregularly doubled anteriorly and disappearing close to posterior elytral margin; vague secondary striae also present in very anterior part of interstice 2 (slightly longer than scutellum) and anterior 1 / 3 of interstice 3; elytral surface with sparse micropunctation; humeral umbone and apical protuberance prominent; opaque area at apical curvature narrow; lateral margin with wide flat paramarginal extension between humerus and middle of elytra; epipleuron broad near humerus, ending slightly posteriorly of elytral midlength; epipleuron with numerous soft short white setae near humerus and stout spiniform black setae beginning posteriorly of humerus and extending along lateral margin to apical curvature (becoming gradually larger apicad); one additional large spiniform seta present near apico-sutural angle; posterior margin evenly rounded. Propygidium with dense fringe of white setae along posterior margin, covering approximately 1 / 2 of propygidial length. Pygidium very convex; apex broadly rounded; punctation rather sparse, transverse on disc (except across vague midline), transforming into concentrically arranged dense striolation at sides and base; with two small spots of white setae near base; apex with several long, erect brownish setae. Ventral thoracic surface densely covered with soft, long, white setae. Meso-metaventral process short, compressed between mesocoxae, projecting slightly downward in lateral view, anteriorly vertical and straight; apex subrectangular, somewhat rounded; bulbiform in ventral view. Abdominal ventrites with dense transverse band of long, white setae in posterior half (broadly interrupted in middle) and irregular white setae on each side of anterior half of ventrites 2 – 4; ventrites 1 – 2 and anterior half of ventrite 3 carinate laterally. Legs. Mesofemur and metafemur with two bands of long white setae, one along anterior margin, the second emerging from a transverse row of punctures parallel to posterior margin. Protibia bidentate, broadened, rather long, approximately 3.9 times longer than wide; proximal tooth short, situated close to the rather short, weakly curved apical tooth; inner spur short, positioned at level of space between proximal and apical tooth. Metatibia fusiform; protarsus slender; protarsomere 5 (without claws) slightly longer than tarsomeres 1 – 4 combined; inner protarsal claw approximately 3 / 4 as long as protarsomere 5, deeply incised apically, upper branch spiniform, lower branch broad, obliquely truncated; outer mesotarsal claw approximately as long as combined length of mesotarsomeres 1 – 4, curved, deeply incised at apex, upper branch spiniform; metatarsal claws very unequal, outer claw approximately twice as thick and 1 / 3 longer than inner. Aedeagus as in Figs 5 A, G, M. Female. Protibia slender, apical tooth of protibia long and spatulate; protarsus articulated slightly basally of level of proximal tooth; inner spur long, articulated between 1 / 2 and 2 / 3 of tibial length; protarsus very slender, protarsomere 5 (without claws) shorter than tarsomeres 1 – 4 combined; modified claws of pro- and mesotarsus shorter, two apical branches more equal than in males; antennal club as long as antennomeres 2 – 6 combined. Measurements. Total body length 10.3 – 11.8 mm (HT 11.6 mm), total body width 5.6 – 6.6 mm (HT 6.4 mm). Morphological variation. Elytral yellow spots vary slightly in shape and size. Number of elytral spiniform setae near apico-sutural angle varies between two and five. Shape of parameres very constant.	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA354771AC3CEFE46FDCBFB14.taxon	diagnosis	Differential diagnosis. Glenopopillia forceps sp. nov. is most similar to G. maculata Lin, 1980 and G. nagaii (Sabatinelli, 1997). It differs from these species in the following characters: protibia more slender in males; yellow lateral spot of elytron divided; secondary stria on interstice 2 present only anteriorly; abdominal ventrites with distinct irregular white setae on each side of the anterior half of ventrites 2 – 4. Moreover, in G. forceps sp. nov. the aedeagus is rather short with asymmetric parameres (Figs 5 A, G, M).	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA354771AC3CEFE46FDCBFB14.taxon	etymology	Etymology. The specific epithet refers to the forceps-like shape of the parameres of the new species (noun in nominative case, standing in apposition).	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA354771AC3CEFE46FDCBFB14.taxon	distribution	Distribution. India (Meghalaya, Garo Hills).	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA3587718C09AFB26FF6CFC54.taxon	description	(Figs 6 A – I)	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA3587718C09AFB26FF6CFC54.taxon	materials_examined	Type locality. ‘ Tonkin, Chapa [= Vietnam, Lào Cai Province, Sapa] ’. Type material studied (2 spec.). SYNTYPES: 1 ♂ (BMNH), ‘ Co- | type [yellow] [p] || 1930.275 [hw] | TONKIN [p] | Chapa | 20. VI. [hw] 191 [p] 8 [hw] | JEANVOINE [p] || Strigoderma ♂ | fossulata cotype Bend [hw] | E. Benderitter, det. [p] ’; 1 ♂ (IRSB), “ TONKIN [p] | Chapa | 11. VII. [hw] 191 [p] 7 [hw] | JEANVOINE [p] || Strigoderma ♂ | fossulata cotype Bend [hw] | E. Benderitter, det. [p] || R. I. Sc. N. B. 16.117 | L. Burgeon, coll. Et det. [p] ”.	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA3587718C09AFB26FF6CFC54.taxon	description	Redescription of syntype (♂ from BMNH). Body shape. Elongate ovoid, weakly convex. Color. Ground color blackish-brown with strong green to red metallic luster; legs dark orange except for mesoand metatarsus, meso- and metafemur, and apical part of metatibia which are dark brown with green metallic luster; clypeus and antenna light brown; elytra predominantly light brown with weak metallic sheen; sutural margin narrowly blackened; some vague, pale yellow spots as follows: one posteriorly of scutellum; two near middle of outer margin: inner spot in costal interval 3, outer one slightly larger than inner and spanning costal interval 4 to 5; one subquadrate spot approximately between 2 / 3 and 3 / 4 of elytral length, from costal interval 1 to 2. Head. Clypeus subrectangular, disc very densely, transversely, confluently punctate; anterior corners rounded; anterior margin weakly reflexed; frons very shallowly impressed, laterally punctate like clypeus, separately punctate at middle; vertex very sparsely and finely punctate; ratio of interocular width / width of head approximately 0.67; antennal club longer than antennomeres 2 – 6 combined. Pronotum approximately 1.5 times wider than long, with two deep, oblique impressions on each side (posterior impression larger than anterior) and deep median longitudinal furrow; with steep decline along lateral margin; disc extremely finely and sparsely punctate, punctures becoming gradually larger laterally; sparse erect setae present near anterior angles and along lateral margin; anterior angles acute and strongly produced; posterior angles obtuse; sides of pronotum weakly converging anteriad in posterior two thirds, then strongly curved, strongly converging, and slightly sinuate in anterior third; basal marginal line interrupted before scutellum between level of elytral costal interval 3 on each side; all other marginal lines complete. Scutellum nearly semicircular, broader than long, finely and sparsely punctate. Elytra regularly striate; three inner costal intervals (1, 2 and 3) slightly more convex than interstices; strial punctures large, distinct, subsutural interstice with secondary stria being irregularly doubled anteriorly and almost reaching the posterior elytral margin; distinct but discontinuous secondary striae also present on interstice 2 and 3; elytral surface with sparse micropunctation; humeral umbone and apical protuberance prominent; opaque area at apical curvature broad, including interstice 4 laterally; lateral margin with wide flat paramarginal extension between humerus and middle of elytra; epipleuron broad near humerus, ending slightly posteriorly of elytral midlength; epipleuron with numerous soft short white setae near humerus and stout spiniform black setae beginning posteriorly of humerus and extending along lateral margin to apical curvature (becoming gradually larger apicad); three additional large spiniform setae present near apico-sutural angle; posterior margin evenly, separately rounded. Propygidium with dense fringe of white setae along posterior margin covering approximately 1 / 3 of propygidial length. Pygidium very convex; apex broadly rounded; punctation transverse, rather sparse on disc (except across vague midline), transforming into concentrically arranged dense striolation at sides and base; with two small spots of white setae near base and another two vague spots in a small depression at lateral margin; apex with several long, erect brownish setae. Ventral thoracic surface densely covered with soft, long, white setae. Meso-metaventral process short, compressed between mesocoxae, projecting slightly downward in lateral view, anteriorly vertical and straight; apex subrectangular, somewhat rounded; bulbiform in ventral view. Abdominal ventrites with a transverse band of dense, long, white setae in posterior half (broadly interrupted in middle) and irregular white setae on each side of the anterior half of ventrites 2; ventrites 1 – 3 carinate laterally. Legs. Meso- and metafemur with two bands of long white setae, one along anterior margin, the second emerging from a transverse row of punctures parallel to posterior margin. Protibia bidentate, rather long, broadened, approximately 4.3 times longer than wide; proximal tooth short, situated close to the rather short, curved apical tooth; inner spur short, at level of proximal tooth. Metatibia fusiform; protarsus slender; protarsomere 5 (without claws) slightly longer than tarsomeres 1 – 4 combined; inner protarsal claw approximately 3 / 4 as long as protarsomere 5, deeply incised apically, upper branch spiniform, lower branch broad, obliquely truncated; outer mesotarsal claw approximately as long as combined length of mesotarsomeres 1 – 4, curved, deeply incised at apex, upper branch spiniform; metatarsal claws very unequal, outer claw approximately twice as thick and 1 / 3 longer than inner. Aedeagus as in Figs 6 F – H. Female. Unknown. Measurements. Total body length 12.5 – 12.9 mm, total body width 7.0 – 7.2 mm.	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA3587718C09AFB26FF6CFC54.taxon	diagnosis	Differential diagnosis. Glenopopillia fossulata is very similar to G. r. rufipennis Lin, 1980 and G. skalei sp. nov. in having the same light brown color type of the elytra. Glenopopillia skalei sp. nov. is clearly separated by the partly blackened costal interval 1, and the wider protibia in males. The two syntypes have only one transverse band of setae on each abdominal ventrite while the anterior part is more or less glabrous. In G. r. rufipennis, there are white setae present in the anterior half of abdominal ventrites 2 – 4. Punctures in abdominal ventrites are fewer and smaller; setae at the metafemur are distinctly shorter and less dense; spiniform setae in metatibia are not strong as G. r. rufipennis. The shape of the parameres is very specific. In contrast to G. r. rufipennis and G. skalei sp. nov., the parameres of G. fossulata are symmetrical with the apex strongly curved but simple, without subapical denticle (Figs 6 F – H).	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA3587718C09AFB26FF6CFC54.taxon	distribution	Distribution. Vietnam (Lào Cai Province) (BENDERITTER 1929).	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA35A771EC088FBE6FD49FB94.taxon	description	(Figs 7 A – I)	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA35A771EC088FBE6FD49FB94.taxon	materials_examined	Type locality. ‘ Selangor, Bukit Kutu, 3000 – 3460 F’ [Malaysia, Selangor State, Bukit Kutu mt., ca. 3 ° 32.6 ’ N, 101 ° 43.2 ’ E] ’.	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA35A771EC088FBE6FD49FB94.taxon	materials_examined	Type material studied (1 spec.). LECTOTYPE: ♂ (present designation, ZMHB), ‘ Bukit Kutu | Selangor | April 1915 [p] | 3000 – 3460 [hw] || Zool. Mus. | Berlin [p] || Spilota | klossi | Type Ohs. [red] [hw] ’.	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA35A771EC088FBE6FD49FB94.taxon	materials_examined	Additional material studied (8 spec.). MALAYSIA: PERAK: 1 ♀ (CZPC), W. Perak, 30 km SE of Ipoh, Cameron Highlands, Ringlet, 900 m, 25. iv. – 5. v. 2001, P. Čechovský lgt.; 1 ♂ 2 ♀♀ (MSPC), Malaysia, Cameron Highlands. MYANMAR: TANINTHARYI REGION: 2 ♂♂ (CZPC), Tenasserim, 20. iv. 1995; 2 ♂♂ 1 ♀ (CZPC), Tenasserim, 23. iv. 1996, local collectors.	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA35A771EC088FBE6FD49FB94.taxon	description	Redescription of lectotype (♂). Body shape. Elongate ovoid, weakly convex. Color. Ground color blackish-brown with strong green metallic luster; all legs except tarsi dark orange; all tarsi, proximal half of femur, and apical part of meso- and metatibia dark brown with green metallic luster; antenna light brown; elytra including all margins, epipleura and suture blackish-brown with weak metallic sheen; a large oblique reniform yellow spot between humerus and costal interval 1 posteriorly not reaching middle of elytron; three interconnected yellow spots forming an oblique transverse band just posteriorly to middle of elytron; inner spot larger and subquadrate, spanning costal interval 1 to 2, middle spot rather small, spanning interstice 2 and costal interval 3, outer spot extending from interstice 3 to costal interval 5. Head. Clypeus subrectangular, disc very densely, transversely, confluently punctate; anterior corners rounded; anterior margin weakly reflexed; frons very shallowly impressed medially, densely punctate, medial impression sparsely punctate; vertex very sparsely punctate; ratio of interocular width / width of head approximately 0.61; antennal club longer than antennomeres 2 – 6 combined. Pronotum approximately 1.4 times wider than long, with small impression on each side; disc extremely finely and sparsely punctate, punctures becoming gradually larger laterally; sparse erect setae present near anterior angles and along lateral margin; anterior angles acute and strongly produced; posterior angles obtuse; sides of pronotum very weakly converging anteriad in posterior two thirds, then strongly curved, strongly converging, and slightly sinuate in anterior third; basal marginal line interrupted before scutellum between level of elytral costal interval 3 on each side; all other marginal lines complete. Scutellum nearly semicircular, longer than wide, finely and sparsely punctate. Elytra regularly striate; two inner costal intervals (1 and 2) slightly convex, all other costal intervals and interstices almost flat; strial punctures distinct; subsutural interstice with a secondary stria being irregularly doubled anteriorly and disappearing in posterior third; vague secondary striae also present in anterior half of interstices 2 and 3; elytral surface with sparse micropunctation; humeral umbone and apical protuberance prominent; opaque area at apical curvature very narrow, laterally ending at apical protuberance; lateral margin with moderately wide flat paramarginal extension between humerus and middle of elytron; epipleuron broad near humerus, ending slightly posteriorly of elytral midlength; epipleuron with several short white setae near humerus and rigid but rather short spiniform black setae beginning posteriorly of humerus and extending along lateral margin to apical curvature (becoming gradually larger apicad), apico-sutural angle without setae; posterior margin evenly rounded. Propygidium glabrous. Pygidium strongly convex; apex broadly rounded; punctation dense, transverse; apex with several long, erect brownish setae. Ventral thoracic surface densely covered with soft, long, white setae. Meso-metaventral process short, compressed between mesocoxae, projecting slightly downward in lateral view, anteriorly vertical and straight; apex subcircular; bulbiform in ventral view. Abdominal ventrites with transverse band of dense, long, white setae in posterior half (broadly interrupted in middle); ventrites 1 – 2 and anterior half of ventrite 3 carinate laterally. Legs. Meso- and metafemur with several irregular bands of long white setae. Protibia bidentate, broadened, approximately 4.0 times longer than wide; proximal tooth short, situated close to the rather short, curved apical tooth; inner spur short, at level of space between proximal and apical tooth. Metatibia strongly fusiform; protarsus slightly thickened; protarsomere 5 (without claws) longer than tarsomeres 1 – 4 combined; inner protarsal claw approximately 3 / 4 as long as protarsomere 5, deeply incised apically, upper branch spiniform, lower branch broad, obliquely truncated; outer mesotarsal claw as long as mesotarsomere 5, incised at apex, upper branch spiniform; metatarsal claws very unequal, outer claw approximately twice as thick and 1 / 3 longer than inner. Aedeagus as in Figs 7 F – H. Female. Protibia slender, apical tooth of protibia long and spatulate; protarsus articulated slightly basally of level of proximal tooth; inner spur long, positioned between 1 / 2 and 2 / 3 of tibial length; protarsus very slender, protarsomere 5 (without claws) shorter than tarsomeres 1 – 4 combined; modified claws of pro- and mesotarsi shorter, two apical branches more equal than in males; antennal club as long as antennomeres 2 – 6 combined. Measurements. Total body length 10.8 – 12.5 mm (LT 11.3 mm), total body width 5.2 – 7.4 mm (LT 6.2 mm). Morphological variability. Elytral spots vary slightly in shape and size, the preapical transversal band sometimes divided into two spots. Color of legs variable, in some specimens all legs dark brown with green metallic luster. Secondary stria on subsutural interstice sometimes only present in anterior half of elytra, sometimes almost reaching the posterior margin. Shape of parameres very constant.	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA35A771EC088FBE6FD49FB94.taxon	diagnosis	Differential diagnosis. This species is peculiar and differs from all other species of this genus by the following characters: pronotum more or less even, with only one small impression on each side; apico-sutural angle of elytron without spiniform setae, setae along lateral margin less strong and shorter compared to all other species except G. albopilosa sp. nov. with short soft setae; propygidium without long white setae at posterior margin; pygidium smooth, without white long setae; modified claws of proand mesotarsi of males more robust, the upper branch extremely small. Moreover, the general shape of the aedeagus differs significantly from the shape found in all other species: The parameres are very elongated, without a subapical tooth as found most other species, and the ventral plate has a long extension reaching the apical curvature of the parameres (Figs 7 F – H).	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA35A771EC088FBE6FD49FB94.taxon	distribution	Distribution. Malaysia (Perak, Selangor) (OHAUS 1926). First country record from Myanmar (Tanintharyi Region).	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA35A771EC088FBE6FD49FB94.taxon	discussion	Remarks. Because OHAUS (1926) did not state how many syntypes were included in the original description of this species, and it is not entirely certain that the type series was monospecific; therefore, a lectotype was designated by us.	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA35C771CC141FBA6FEFFFA94.taxon	description	(Figs 3 B, H; 4 B, H, N; 5 B, H, N; 8 A – J)	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA35C771CC141FBA6FEFFFA94.taxon	materials_examined	Type locality. ‘ Guangxi, Pingxiang, Daqingshan’. Type material studied (1 spec.). HOLOTYPE: ♂ (GIABR): ‘ ťı ± 4 m [= Guangxi, Mt. Daqing Shan] | 1974.5.26 ~ 31 | ffl 罗ήι † × ṣ [= leg. Lin Ping, Luo Yuliang & Li Yaoquan] || HOLOTYPE [red] [p] || Glenopopillia | maculata sp. nov. [hw] | ※ + à [p] ffl 1978 [hw] [= det. Lin Ping 1978] || 477 [p] ’ (Fig. 8 I). Additional material studied (4 spec.). CHINA: GUANGXI ZHUANG A. R.: 1 ♂ (CCPC), Jiuwanshan Mts, Huanjiang County, Hechi City, Guangxi, China, 1100 m, 1. vi. 2015, Y. - Q. Lu lgt.; 1 ♂ (CCPC), Jiuwanshan Mts, Huanjiang County, Hechi City, Guangxi, China, 1100 m, 29. v. 2015, Y. Q. Lu lgt. VIETNAM: B ẮC GIANG PROVINCE: 1 ♂ (CZPC) (Figs 3 B, H; 4 B, H, N; 5 B, H, N), Tay Yen Tu Nat. Res., Thanh So’n, 21 ° 12.812 Nʹ 106 ° 45.846 Eʹ, 86 m, 18. v – 21. v. 2015, A. Skale lgt.; 1 ♀ (CZPC), Tay Yen Tu Nat. Res., Van Danh (bei Dong Ri), 21 ° 09.96 Nʹ, 106 ° 49.56 ʹE, 415 m, 20. v. 2015, A. Weigel lgt.	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA35C771CC141FBA6FEFFFA94.taxon	description	Redescription of holotype (♂). Body shape. Elongate ovoid, weakly convex. Color. Ground color blackish-brown with strong green to red metallic luster; legs dark orange except for mesoand metatarsus, meso- and metafemur, and apical part of metatibia being dark brown with green metallic luster; clypeus and antenna light brown; elytra including all margins, epipleura and suture blackish-brown with weak metallic sheen; with following maculae: a large oblique reniform yellow spot between humerus and first costal interval extending posteriorly but not reaching middle of elytron, one small elongate yellow spot in anterior part of costal interval 5, two small approximated yellow spots slightly posteriorly of middle: one elongate in costal interval 3, second transverse, spanning costal interval 4 to 5, and a subquadrate yellow spot in apical 4 / 5 spanning costal interval 1 to 2. Head. Clypeus subsemicircular, disc very densely, partly transversely confluently punctate, anterior corners widely rounded; anterior margin weakly reflexed; frons shallowly impressed, laterally punctate like clypeus, punctures separate at middle; vertex sparsely punctate; ratio of interocular width / width of head approximately 0.65; antennal club longer than antennomeres 2 – 6 combined. Pronotum approximately 1.5 times wider than long, with two deep, oblique impressions on each side (posterior impression larger than anterior) and a shallow median longitudinal furrow, with steep decline along lateral margin; disc extremely finely and sparsely punctate, punctures becoming gradually larger laterally; sparse erect setae present near anterior angles and along lateral margin; anterior angles acute and strongly produced; posterior angles obtuse; sides of pronotum weakly converging anteriad in posterior two thirds, then strongly curved, strongly converging, and slightly sinuate in anterior third; basal marginal line interrupted in middle between level of elytral costal interval 2 on each side; all other marginal lines complete. Scutellum nearly semicircular, broader than long, moderately fine and sparsely punctate. Elytra regularly striate; three inner costal intervals (1, 2 and 3) slightly more convex than interstices; strial punctures distinct; subsutural interstice with secondary stria being irregularly doubled anteriorly and almost reaching posterior elytral margin; distinct but discontinuous secondary striae distinct on interstices 2 and 3; elytral surface with sparse micropunctation; humeral umbone and apical protuberance prominent; opaque area at apical curvature narrow; epipleuron broad near humerus, ending slightly posteriorly of elytral midlength; epipleuron with numerous soft short white setae near humerus and stout spiniform black setae beginning posteriorly of humerus and extending along lateral margin to apical curvature (becoming gradually larger apicad); two or three additional large spiniform setae present near apico-sutural angle. Propygidium with dense fringe of white setae along posterior margin covering approximately 1 / 2 of propygidial length in middle and 1 / 3 of length at sides. Pygidium strongly convex; apex broadly rounded; punctation transverse on disc, transforming into concentrically arranged dense striolation at sides and base; with two large spots of white setae near base; apex with several long, erect brownish setae. Ventral thoracic surface densely covered with soft, long, white setae. Meso-metaventral process short, compressed between mesocoxae, projecting slightly downward in lateral view, anteriorly vertical and straight; apex subcircular; bulbiform in ventral view. Abdominal ventrites with transverse band of dense, long, white setae in posterior half (broadly interrupted in middle) and irregular white setae on each side of anterior half of ventrites 2 – 4; ventrites 1 – 3 carinate laterally. Legs. Meso- and metafemur with two bands of long white setae, one along anterior margin, the second emerging from a transverse row of punctures parallel to posterior margin. Protibia bidentate, broadened, approximately 3.6 times longer than wide; proximal tooth short, situated close to the rather short, weakly curved apical tooth; inner spur short, articulated at level of proximal tooth. Metatibia strongly fusiform; protarsus slender; protarsomere 5 (without claws) slightly longer than tarsomeres 1 – 4 combined; inner protarsal claw approximately 3 / 4 as long as protarsomere 5, slightly widened and deeply incised apically, upper branch spiniform, lower branch broad, obliquely truncated; outer mesotarsal claw approximately as long as combined length of mesotarsomeres 1 – 4, deeply incised at apex, upper branch spiniform; metatarsal claws very unequal, outer claw approximately twice as thick and 1 / 3 longer than inner. Aedeagus as in Figs 5 B, H, N and 8 F – H. Female. Protibia slender, apical tooth of protibia long and spatulate; protarsus articulated slightly basally of level of proximal tooth; inner spur long, articulated between 1 / 2 and 2 / 3 of tibial length; protarsus very slender, protarsomere 5 (without claws) shorter than tarsomeres 1 – 4 combined; modified claws of pro- and mesotarsi shorter, two apical branches more equal than in males; antennal club as long as antennomeres 2 – 6 combined. Measurements. Total body length 10.2 – 11.4 mm (HT 10.2 mm), total body width 5.7 – 6.3 mm (HT 5.7 mm). Morphological variation. Elytral yellow spots vary slightly in shape and size. Number of elytral spiniform setae near apico-sutural angle varies between one and three. Shape of parameres very constant.	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA35C771CC141FBA6FEFFFA94.taxon	diagnosis	Differential diagnosis. Glenopopillia maculata is most similar to G. nagaii and G. forceps sp. nov. with which it shares the same elytral color pattern. It differs from G. forceps sp. nov. in having a distinctly broader protibia in males, the secondary stria on interstice 2 reaching the posterior quarter of elytra, and in having the yellow lateral spot of the elytron not divided. Glenopopillia maculata differs from G. nagaii in the punctation of the pronotum which is more distinct between the lateral impressions. The parameres of G. maculata are more or less symmetric (asymmetric in G. forceps sp. nov.), and the terminal and subapical lateral tooth of the parameres are more stout compared to G. nagaii (Figs 8 F – H).	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA35C771CC141FBA6FEFFFA94.taxon	distribution	Distribution. China (Guangxi Zhuang Autonomous Region) (LIN 1980). First country record from Vietnam (B ắc Giang Province).	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA35E771DC0A8FAA6FBB7FD14.taxon	description	(Figs 3 C, I; 4 C, I, O; 5 C, I, O)	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA35E771DC0A8FAA6FBB7FD14.taxon	materials_examined	Type locality. China, Yunnan, Jinghong, Naban River Watershed National Nature Reserve, Anmaxinzhai (forest), 22 ° 11.75 ʹN 100 ° 38.72 ʹE. Type material (13 spec.). HOLOTYPE: ♂ (IZAS), ‘ Anmaxinzhai V D | 16.05.2009 | leg. LZ. Meng | ffltñĸf ẓṅƶ [= donated by Naban River Watershed National Nature Reserve] || ± • ạėffltñĸfẓ [= Yunnan, Jinghong, Naban River Watershed National Nature Reserve] | Ẋ 麻kż (ƛff) [= Anmaxinzhai (forest)] | 2009. V. 16 772 m | 中fl ḣäb ffi [= IZAS] || 22.19577 ° N | 100.64532 ° E I ḛ [= flight interception traps] | JƗλ ： aƚł [= leg. Meng Lingzeng] | 中fl ḣäbffi [= IZAS] || IOZ (E) 1966471 [p] ’. PARATYPES: 2 ♂♂ (IZAS), same data as holotype, IOZ (E) 1966472, IOZ (E) 19664731; 1 ♂ (CZPC), same location data as holotype, 2009. V. 26, IOZ (E) 1966474; 1 ♀ (IZAS), ‘ Naban II / I D | 16.05.2009 | leg. LZ. Meng | ffltñĸf ẓṅƶ [= donated by Naban River Watershed National Nature Reserve] || ± • ạėffltñĸfẓ [= Yunnan, Jinghong, Naban River Watershed National Nature Reserve] | fflv ḬƑ (dƙff) [= Tea Factory of Naban (rubber forest)] | 2009. V. 16 732 m | 中fl ḣäbffi [= IZAS] || 22.15843 ° N | 100.66487 ° E I ḛ [= flight interception traps] | JƗλ ： aƚł [= leg. Meng Lingzeng] | 中fl ḣäbffi [= IZAS] || IOZ (E) 1966475 [p] ’; 1 ♀ (NMEC), ‘ CHINA: S-YUNNAN | (Xishuangbanna), | 23 km NW Jinghong, | vic [inity]. Na Ban (NNNR) [= Naban River Watershed National Nature Reserve] | N 22 ° 10.04, E 100 ° 39.52 | 28. VI. 2008, 730 m, forest | leg. A. Weigel MF 1 [= Malaise-Falle (Malaise trap)] ’; 2 ♂♂ 5 ♀♀ (PPCB), ‘ LAOS, 21 ° 09 ’ N 101 ° 19 ’ E, | Louangnamtha pr. | Namtha ~ MuangSing. | 5 – 31. v. 1997, 900 – | Vit Kubáň leg. – 1200 m, || Coll. P. Pacholátko | Brno | Merhautova 68 | Czech republic [p] ’.	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA35E771DC0A8FAA6FBB7FD14.taxon	description	Description of holotype (♂). Body shape. Elongate ovoid, weakly convex. Color. Ground color blackish-brown with strong green to red metallic luster; fore legs and major part of mesotibia dark orange with metallic sheen; mesofemur, apex of mesotibia, mesotarsus and posterior legs dark brown with green metallic luster; clypeus and antenna light brown; elytra predominantly medium brown with weak metallic sheen; lateral margin, epipleura, major part of sutural interval and humeral spot black; one vague yellow spot posteriorly of scutellum; small elongate black spots at costal intervals 1, 2 and 3 connected by dark brown curved zig-zag band reaching margin posteriorly of humeral umbone; three yellow spots just posteriorly to brown cingulum as follows: inner spanning costal interval 1 to 2, middle spot on costal interval 3 and outer spanning costal interval 4 to 5. Head. Clypeus subrectangular, disc very densely, transversely, confluently punctate; anterior corners rounded; anterior margin weakly reflexed; frons very shallowly impressed, laterally punctate like clypeus, separately punctate at middle; vertex sparsely punctate; ratio of interocular width / width of head approximately 0.66; antennal club longer than antennomeres 2 – 6 combined. Pronotum approximately 1.5 times wider than long, with two deep, oblique impressions on each side (posterior impression larger than anterior) and deep median longitudinal furrow; with steep decline along lateral margin; disc extremely finely and sparsely punctate, punctures gradually larger laterad; sparse erect setae present near anterior angles and along lateral margin; anterior angles acute and strongly produced; posterior angles obtuse; sides of pronotum weakly converging anteriad in posterior two thirds, then strongly curved, strongly converging, and slightly sinuate in anterior third; basal marginal line interrupted before scutellum between level of elytral costal interval 2 on each side; all other marginal lines complete. Scutellum nearly semicircular, broader than long, finely and sparsely punctate. Elytra regularly striate; three inner costal intervals (1, 2 and 3) slightly more convex than interstices; strial punctures distinct, subsutural interstice with secondary stria being irregularly doubled anteriorly and disapearing in posterior third; vague, widely discontinuous, and similarly disappearing secondary striae also present on interstice 2 and 3; elytral surface with sparse micropunctation; humeral umbone and apical protuberance very prominent; opaque area at apical curvature narrow; lateral margin with wide flat paramarginal extension between humerus and middle of elytra; epipleuron broad near humerus, ending slightly posteriorly of elytral midlength; epipleuron with numerous soft short white setae near humerus and stout spiniform black setae beginning posteriorly of humerus and extending along lateral margin to apical curvature (becoming gradually larger apicad); three additional large spiniform setae near apico-sutural angle; posterior margin evenly rounded. Propygidium with dense fringe of white setae along posterior margin covering approximately 1 / 3 of propygidial length in middle and 1 / 4 of length at sides. Pygidium strongly convex; apex broadly rounded; punctation rather sparse, transverse on disc (except across vague midline), transforming into concentrically arranged dense striolation at sides and base; with two spots of white setae near base and another two vague spots in a small depression at lateral margin; apex with several long, erect brownish setae. Ventral thoracic surface densely covered with soft, long, white setae. Meso-metaventral process short, compressed between mesocoxae, projecting slightly downward in lateral view, anteriorly vertical and straight; apex subrectangular and rounded; bulbiform in ventral view. Abdominal ventrites with transverse band of dense, long, white setae in posterior half (broadly interrupted in middle) and irregular white setae on each side of anterior half of ventrites 2 – 4; ventrites 1 – 2 and anterior half of ventrite 3 carinate laterally. Legs. Meso- and metafemur with two bands of long white setae, one along anterior margin, the second emerging from a transverse row of punctures parallel to posterior margin. Protibia bidentate, broadened, approximately 3.1 times longer than wide; proximal tooth short, situated close to the rather short, strongly curved apical tooth; inner spur short, articulated at level of proximal tooth. Metatibia fusiform; protarsus slender; protarsomere 5 (without claws) slightly longer than tarsomeres 1 – 4 combined; inner protarsal claw approximately 3 / 4 as long as protarsomere 5, deeply incised apically, upper branch spiniform, lower branch broad, obliquely truncated; outer mesotarsal claw approximately as long as combined length of mesotarsomeres 1 – 4, deeply incised at apex, upper branch spiniform; metatarsal claws very unequal, outer claw approximately twice as thick and 1 / 3 longer than inner. Aedeagus as in Figs 5 C, I, O. Female. Protibia slender, apical tooth of protibia long and spatulate; protarsus articulated slightly basally of level of proximal tooth; inner spur long, articulated between 1 / 2 and 2 / 3 of tibial length; protarsus very slender, protarsomere 5 (without claws) shorter than tarsomeres 1 – 4 combined; modified claws of pro- and mesotarsi shorter, two apical branches more equal than in males; antennal club as long as antennomeres 2 – 6 combined. Measurements. Total body length 10.0 – 12.0 mm (HT 10.8 mm), total body width 5.6 – 7.2 mm (HT 5.8 mm). Morphological variation. Elytral black-brown transversal band and yellow spots vary slightly in shape and size, sometimes with another vague yellow spot between scutellum and humerus. Number of elytral spiniform setae near apico-sutural angle varies between one and three. Shape of parameres very constant.	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA35E771DC0A8FAA6FBB7FD14.taxon	diagnosis	Differential diagnosis. Glenopopillia mengi sp. nov. is distinguished from similar species by the combination of the following characters: unique color pattern of elytra – medium brown with a black / dark brown curved zig-zag band in the middle and additional vague yellow spots; punctures of subsutural interstice almost disappearing in the posterior third of elytra; epipleuron with numerous white short setae near humerus; protibia rather short and wide; white setae along posterior margin of propygidium rather sparse and short. The aedeagus is distinctive and cannot be confused with any other species of Glenopopillia. The parameres are similar to those of G. maculata, but are stouter, with the subapical lateral tooth very short. The apex of the ventral plate is equipped with a backward-pointed hook (Figs 5 C, I, O).	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA35E771DC0A8FAA6FBB7FD14.taxon	etymology	Etymology. Patronymic (noun in the genitive case), named in honour of Lingzeng Meng, the collector of most of the type material, who conducted an excellent field study in the Naban River Watershed National Nature Reserve.	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA35E771DC0A8FAA6FBB7FD14.taxon	discussion	Collecting circumstances. These specimens were collected in the Naban River Watershed National Nature Reserve with cross window traps in the tree canopy.	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA35E771DC0A8FAA6FBB7FD14.taxon	distribution	Distribution. China (Yunnan Province), Laos (Louang Namtha Province).	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA35F7703C3DDFD26FBDBF914.taxon	description	(Figs 9 A – I)	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA35F7703C3DDFD26FBDBF914.taxon	materials_examined	Type locality. ‘ North Vietnam, Mt Tamdao’. Type material studied (2 spec.). PARATYPES: 1 ♂ (MHNG) (Figs 9 A – I), ‘ North VIETNAM | Mt Tamdao, VI. 1993 | S. Nagai leg. [p] || Paratypus [p] ♂ [hw] | Callistethus nagai n. sp. | des. Sabatinelli 1997 [red] [p] ’ (Fig. 9 I); 1 ♀, (MHNG), ‘ North VIETNAM | Mt Tamdao, VI. 1993 | S. Nagai leg. [p] || Paratypus [p] ♀ [hw] | Callistethus nagai n. sp. | des. Sabatinelli 1997 [red] [p] ’. Additional material studied (49 spec.). VIETNAM: VĨNH PHÚC PROVINCE: 2 ♂♂ 1 ♀ (NMPC), Tam Dao, 3. – 11. vi. 1985, 900 – 1400 m, J. Jelínek lgt.; 1 ♂ (DKCP), Tam Dao, 5. vi. 1986, D. Král coll.; 4 ♂♂ 3 ♀♀ (MSPC, PPCB), Tamdao, 2. – 11. vi. 1985, Vít. Kubáň lgt.; 1 ♂ 3 ♀♀ (CZPC), Tam Dao, ca. 1000 m, 17. – 30. vi. 1999, A. Kallies lgt.; 1 ♂ (CZPC), Tam Dao, 8. – 22. v. 1990, Miloš Dudycha lgt.; 1 ♀ (CZPC), Tam Dao NP, 4. vi. 2012, 21 ° 27.577 ʹN 105 ° 38.489 ʹE, 1000 m, M. Pejcha lgt.; 1 ♀ (CZPC), Tam Dao, 930 m, vi. ‒ viii. 1997, native collector lgt.; 1 ♂ (PKLC), Tam Dao National Park, N 21.4603 ° E 105.64201 °, 1027 m, 1. – 4. vi. 2013, Nathan M. Schiff lgt; 9 ♂♂ 5 ♀♀ (MSPC), Tam Dao, 20. – 28. vi. 1990, Strnad Jan lgt.; 4 ♂♂ 6 ♀♀ (MSPC), Tam Dao, 20. – 27. vi. 1990, A. Olexa lgt.; 2 ♂♂ (MSPC), Tam Dao, 27. v. – 2. vi. 1986, 900 m, A. Olexa lgt.; 1 ♂ (MSPC), Tam Dao, 16. – 23. v. 1991, 900 m, Strnad Jan lgt. 2 ♂♂ (MSPC), Tam Dao, 6. – 25. v. 1990. LÀO CAI PROVINCE: 1 ♂ (CZPC), Mt. Fan-si-pan, W-Seite [western slope], Cha-pa (= Sapa), 2000 m, 22.15 ° N 103.45 °, E primär. Nebelwald [east, primary cloudy forest], 5. vii. 1994, Brechlin & Schintlmeister lgt. Doubtfully labeled material. CHINA: YUNNAN: 1 ♂ (CZPC), Baoshan city env., 2000 m a. s. l., 10. vii. 1998. A. Gorodinski lgt. (see Remarks below).	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA35F7703C3DDFD26FBDBF914.taxon	description	Redescription of paratype (♂). Body shape. Elongate ovoid, weakly convex. Color. Ground color blackish-brown with strong green to red metallic luster; legs dark orange except for meso- and metatarsus, meso- and metafemur, and apical part of metatibia being dark brown with green metallic luster; clypeus and antenna light brown; elytra including all margins, epipleura and suture blackish-brown with weak metallic sheen; a large oblique reniform yellow spot between humerus and costal interval 1 not reaching middle of elytron posteriorly, one small elongate yellow spot on anterior part of costal interval 5, two approximated small yellow spots slightly posteriorly of middle: one elongate in costal interval 3, the second transverse, spanning costal interval 4 to 5, and a subquadrate yellow spot in the apical 4 / 5 spanning costal interval 1 to 2. Head. Clypeus subsemicircular, disc very densely, partly transversely confluently punctate; anterior corners widely rounded; anterior margin weakly reflexed; frons very shallowly impressed, laterally punctate like clypeus, separately punctate at middle; vertex sparsely and finely punctate; ratio of interocular width / width of head approximately 0.65; antennal club longer than antennomeres 2 – 6 combined. Pronotum approximately 1.4 times wider than long, with two deep, oblique impressions on each side (posterior impression larger than anterior) and shallow median longitudinal furrow; with steep decline along lateral margin; disc extremely finely, shallowly and sparsely punctate, punctures becoming gradually larger laterally; with sparse erect setae present near anterior angles and along lateral margin; anterior angles acute and strongly produced; posterior angles obtuse; sides of pronotum distinctly converging anteriad in the posterior two thirds, then strongly curved, strongly converging, and slightly sinuate in anterior third; basal marginal line interrupted before scutellum between level of elytral interstice 2 on each side; all other marginal lines complete. Scutellum nearly semicircular, broader than long, moderately finely and sparsely punctate. Elytra regularly striate; three inner costal intervals (1, 2 and 3) slightly more convex than interstices; strial punctures distinct; subsutural interstice with secondary stria being irregularly doubled anteriorly and almost reaching posterior elytral margin; distinct but discontinuous secondary striae also present on interstice 2 and 3; elytral surface with sparse micropunctation; humeral umbone and apical protuberance very prominent; opaque area at apical curvature narrow; lateral margin with flat paramarginal extension in anterior half of elytra; epipleuron broad near humerus, ending slightly posteriorly of elytral midlength; epipleuron with several short white setae near humerus and stout, strongly spiniform black setae beginning at humerus and extending along lateral margin to apical curvature (becoming gradually larger apicad); 2 or 3 large spiniform setae present near apico-sutural angle; posterior margin evenly rounded. Propygidium with dense fringe of white setae along posterior margin covering approximately 1 / 2 of propygidial length in middle and 1 / 3 of length at sides. Pygidium strongly convex; apex broadly rounded; punctation transverse and rather sparse on disc (except across vague midline), transforming into concentrically arranged dense striolation at sides and base; with two spots of white setae near base and some white setae scattered at lateral margin; apex with several long, erect brownish setae. Ventral thoracic surface densely covered with soft, long, white setae. Meso-metaventral process short, compressed between mesocoxae, projecting slightly downward in lateral view, anteriorly vertical and straight; apex subrectangular, rounded; bulbiform in ventral view. Abdominal ventrites with transverse band of dense, long, white setae in posterior half (broadly interrupted in middle) and irregular white setae on each side of anterior half of ventrites 2 – 4; ventrites 1 – 2 and anterior half of ventrite 3 carinate laterally. Legs. Meso- and metafemur with two bands of long white setae, one along anterior margin, the second emerging from a transverse row of punctures parallel to posterior margin. Protibia bidentate, broadened, approximately 3.4 times longer than wide; proximal tooth short, situated close to the rather short, curved apical tooth; inner spur short, at level of proximal tooth. Metatibia strongly fusiform; protarsus slender; protarsomere 5 (without claws) slightly longer than tarsomeres 1 – 4 combined; inner protarsal claw approximately 3 / 4 as long as protarsomere 5, slightly widened and deeply incised apically, upper branch spiniform, lower branch broad, obliquely truncated; outer mesotarsal claw very long, approximately as long as combined length of mesotarsomeres 1 – 4, deeply incised at apex, upper branch spiniform; metatarsal claws very unequal, outer claw approximately twice as thick and 1 / 3 longer than inner. Aedeagus as in Figs 9 F – H. Female. Protibia slender, apical tooth of protibia long and spatulate; protarsus articulated slightly basally of level of proximal tooth; inner spur long, articulated between 1 / 2 and 2 / 3 of tibial length; protarsus very slender, protarsomere 5 (without claws) shorter than tarsomeres 1 – 4 combined; modified claws of pro- and mesotarsi shorter, two apical branches more equal than in males; antennal club as long as antennomeres 2 – 6 combined. Measurements. Total body length 10.6 – 11.5 mm, total body width 6.0 – 6.6 mm. Morphological variation. Elytral yellow spots vary very slightly in shape and size. Number of elytral spiniform setae near apico-sutural angle varies between two and four. Shape of parameres very constant.	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA35F7703C3DDFD26FBDBF914.taxon	diagnosis	Differential diagnosis. The elytra of Glenopopillia nagaii have the same basic color pattern as those of G. forceps sp. nov., G. klossi and G. maculata. While it is easily distinguishable from G. klossi by the presence of rigid setae at the apico-sutural angle and by several further characters (see ‘ Differential diagnosis’ of G. klossi), it is extremely similar to G. forceps and G. maculata. It differs from G. maculata in the following characters: punctation of pronotum extremely fine and shallow between the lateral impressions (distinct, extremely shallow and sparse only in very middle in G. maculata); end of each paramere with two laterally directed denticles, which are distinctly longer than those of G. maculata (Figs 9 F – H). Glenopopillia nagaii is distinguished from G. forceps by the following characters: scutellum more or less coarsely punctate; protibia in males slightly stouter; yellow lateral spot of elytra not divided; secondary stria on interstice 2 reaching the posterior quarter of elytra; aedeagus symmetrical (asymmetrical in G. forceps).	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA35F7703C3DDFD26FBDBF914.taxon	distribution	Distribution. Vietnam (Vĩnh Phúc and Lào Cai Provinces), so far known only from the Tam Đ ảo National Park and Mt. Phan Xi Păng (SABATINELLI 1997).	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA35F7703C3DDFD26FBDBF914.taxon	discussion	Remarks. This species was reported from Yunnan, China (ZORN 2006, ZORN & BEZDĚK 2016). We consider this record erroneous because the specimens on which it was based was presumably mislabeled.	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA3417701C33DF926FE5AFC34.taxon	description	(Figs 3 D, J; 4 D, J, P; 5 D, J, P; 8 K – R)	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA3417701C33DF926FE5AFC34.taxon	materials_examined	Type locality. ‘ Yunnan, Xishuangbanna, Meng’a, 1000 m a. s. l. ’. Type material studied (1 spec.). HOLOTYPE: ♂ (IZAS) ‘ ± • ıŋvfflŧ 啊 [= Yunnan, Xishuangbanna, Meng’a] [p] | 1000 [hw] » 尺 (m) | 中 ¤ fl $ ḣ [= Chinese Academy of Sciences] [p] || 1958. V. 25 [hw] JƗà Ŧħ ḟ [= leg. Wang Shuyong] || HOLOTYPE [red] [p] || Glenopopillia | rufipennis sp. nov. [hw] | ※ + à [p] ffl 1978 [= det. Lin Ping] [hw] || 421 [p] ’ (Fig. 8 R). Additional material studied (51 spec.). CHINA: YUNNAN PROVINCE: 1 ♀ (IZAS), Xishuangbanna, Meng’a, 1050 – 1080 m, 17. v. 1958, Wang Shuyong lgt., IOZ (E) 1966477; 1 ♀ (IZAS), Xishuangbanna, Meng’a, 1050 m, 17. v. 1958, Pu Fuji lgt., IOZ (E) 1966476; 1 ♀ (IZAS), Xishuangbanna, Meng’a, 1000 m, 25. v. 1958, Wang Shuyong lgt., IOZ (E) 1966478; 1 ♀ (IZAS), Xishuangbanna, Meng’a, 1000 m, 16. v. 1958, Pu Fuji lgt., IOZ (E) 1966479. LAOS: HOUAPHANH PROVINCE: 2 ♂♂ (NMPC), Ban Saluei → Phou Pane Mt., 20 ° 12 – 13.5 Nʹ 103 ° 59.5 – 104 ° 01 Eʹ, 1340 – 1870 m, 15. iv. – 15. v. 2008, Lao collectors lgt.; 22 ♂♂ 18 ♀♀ (CZPC, ZFKB), Ban Saleui, Phou Pan (Mt.) - 20 ° 12 ʹ N, 104 ° 01 ʹ E, 11. iv. ‒ 15. v. 2012, 1300 ‒ 1900 m, leg. C. Holzschuh ZFMK Ankauf 2012 / 13 (Figs 3 D, J; 4 D, J, P; 5 D, J, P); 1 ♂ (RBINS), Mt. Phu Phan, 2060 m, vi. 2015, S. Collard lgt.; 1 ♂ 3 ♀♀ (IZAS), Xam Neua, Mt. Pan, v. 2012.	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA3417701C33DF926FE5AFC34.taxon	description	Redescription of holotype (♂). Body shape. Elongate ovoid, weakly convex. Color. Ground color blackish-brown with strong green to red metallic luster; legs dark orange except for mesoand metatarsus, meso- and metafemur, and apical part of metatibia being dark brown with green metallic luster; clypeus and antenna light brown; elytra predominantly light brown with weak metallic sheen; sutural margin narrowly blackened; some vague, pale yellow spots as follows: one posteriorly of scutellum; two near middle of outer margin: inner spot in costal interval 3, outer one slightly larger than inner and spanning costal interval 4 to 5; one subquadrate spot approximately between 2 / 3 and 3 / 4 of elytral length, from costal interval 1 to 2. Head. Clypeus subrectangular, disc very densely, transversely, confluently punctate; anterior corners rounded; anterior margin weakly reflexed; frons very shallowly impressed, punctate like clypeus, but punctures less confluent medially; vertex very sparsely and finely punctate; ratio of interocular width / width of head approximately 0.64; antennal club longer than antennomeres 2 – 6 combined. Pronotum approximately 1.4 times wider than long, with two deep, oblique impressions on each side (posterior impression larger than anterior) and a deep median longitudinal furrow; with steep decline along lateral margin; disc extremely finely and sparsely punctate, punctures becoming gradually larger laterally; sparse erect setae present near anterior angles and along lateral margin; anterior angles acute and strongly produced; posterior angles obtuse; sides of pronotum weakly converging anteriad in posterior two thirds, then strongly curved, strongly converging, and slightlysinuate in anterior third; basal marginal line interrupted before scutellum between level of elytral costal interval 3 on each side; all other marginal lines complete. Scutellum nearly semicircular, broader than long, finely and sparsely punctate. Elytra regularly striate; three inner costal intervals (1, 2 and 3) slightly more convex than interstices; strial punctures large, distinct; subsutural interstice with secondary stria being irregularly doubled anteriorly and almost reaching posterior elytral margin; distinct but discontinuous secondary striae also present on interstice 2 and 3; elytral surface with sparse micropunctation; humeral umbone and apical protuberance very prominent; opaque area at apical curvature broad, including interstice 4 laterally; lateral margin with wide flat paramarginal extension between humerus and middle of elytra; epipleuron broad near humerus, ending slightly posteriorly of elytral midlength; epipleuron with numerous soft short white setae near humerus and stout spiniform black setae beginning posteriorly of humerus and extending along lateral margin to apical curvature (becoming gradually larger apicad); 1 or 2 large spiniform setae present near apico-sutural angle; posterior margin evenly, separately rounded. Propygidium with dense fringe of white setae along posterior margin covering approximately 1 / 3 of propygidial length. Pygidium strongly convex; apex broadly rounded; punctation transverse and rather sparse on disc (except across vague midline), transforming into concentrically arranged dense striolation at sides and base; with two small spots of white setae near base and another two vague spots in a small depression at lateral margin; apex with several long, erect brownish setae. Ventral thoracic surface densely covered with soft, long, white setae. Meso-metaventral process short, compressed between mesocoxae, projecting slightly downward in lateral view, anteriorly vertical and straight; apex subrectangular, somewhat rounded; bulbiform in ventral view. Abdominal ventrites with transverse band of dense, long, white setae in posterior half (broadly interrupted in middle) and irregular white setae on each side of anterior half of ventrites 2 – 4; ventrites 1 – 2 and anterior half of ventrite 3 carinate laterally. Legs. Meso- and metafemur with two bands of long white setae, one along anterior margin, the second emerging from a transverse row of punctures parallel to posterior margin. Protibia bidentate, rather long, broadened, approximately 3.9 times longer than wide; proximal tooth short, situated close to the rather short, curved apical tooth; inner spur short, at level of proximal tooth. Metatibia fusiform; protarsus slender; protarsomere 5 (without claws) slightly longer than tarsomeres 1 – 4 combined; inner protarsal claw very long, slightly widened and deeply incised apically, upper branch spiniform; outer mesotarsal claw very long, curved, deeply incised at apex, upper branch spiniform; metatarsal claws very unequal, outer claw approximately twice as thick and 1 / 3 longer than inner. Aedeagus as in Figs 5 D, J, P and 8 O – Q. Female. Protibia slender, apical tooth of protibia long and spatulate; protarsus articulated slightly basally of level of proximal tooth; inner spur long, articulated between 1 / 2 and 2 / 3 of tibial length; protarsus very slender, protarsomere 5 (without claws) shorter than tarsomeres 1 – 4 combined; modified claws of pro- and mesotarsi shorter, two apical branches more equal than in males; antennal club as long as antennomeres 2 – 6 combined. Measurements. Total body length 10.7 – 13.3 mm (HT 10.7 mm), total body width 5.8 – 7.3 mm (HT 5.8 mm). Morphological variation. Vague yellow spots on elytra vary slightly in shape and size, sometimes with an additional vague yellow spot between scutellum and humerus, occasionally the inner of the two middle spots missing, sometimes all yellow spots very indistinct. Number of elytral spiniform setae near apico-sutural angle varies between one and three. Shape of parameres very constant.	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA3417701C33DF926FE5AFC34.taxon	diagnosis	Differential diagnosis. Only two other species share the entirely light brown elytra as seen in G. rufipennis rufipennis: G. fossulata and G. skalei sp. nov. Of these, G skalei sp. nov. is easily distinguished by the partly blackened costal interval 1, the longer propygidial white setae and the wider protibia in males. Glenopopillia rufipennis rufipennis is virtually identical externally to G. fossulata, except that the setae at the abdominal ventrites form two transverse bands in G. rufipennis rufipennis; whereas the anterior areas are nearly glabrous in the syntypes of G. fossulata. Moreover, the setation of the metafemur is more distinct. There are, however, distinct differences in the shape of the parameres, which are strongly asymmetric and subapically dentate in G. rufipennis, but strongly curved and not dentate in G. fossulata (Figs 5 D, J, P; 8 O – Q). Both, G. rufipennis rufipennis and G. fossulata, are larger than all other species, the spiniform setae on the elytral margin are especially stout, and the series begins right posteriorly of the humeral umbone. For the separation from G. rufipennis nigropicta subsp. nov., see ‘ Diagnosis’ under that subspecies.	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA3417701C33DF926FE5AFC34.taxon	distribution	Distribution. China (Yunnan) (LIN 1980). First country record from Laos (Houaphanh).	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA3437706C08CFC46FB41FEB4.taxon	description	(Figs 3 E, K; 4 E, K, Q; 5 E, K, Q)	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA3437706C08CFC46FB41FEB4.taxon	materials_examined	Type locality. Laos, Attapeau Province, Dong Ampham National Biodiversity Conservation Area, Nong Fa (crater lake) enviroment, 15 ° 05.9 ʹN 107 ° 25.6 ʹE, 1160 m a. s. l. Type material (2 spec.). HOLOTYPE: ♂ (IRSB), ‘ Coll. I. R. Sc. N. B. | LAOS, Attapeau Prov. | Annam Highlands Mts. Dong | Amphan NBCA, ca 1160 m NONG | FA (crater lake) env. | 15 ° 05.9 ’ N, 107 ° 25.6 ’ E | 30 - IV- 6 - V- 2010 | Lgt. St Jakl / I. G. 31.970 [p] ’. PARATYPE: 1 ♀ (IRSB), ‘ Coll. I. R. Sc. N. B. | LAOS, Attapeau Prov. | Annam Highlands Mts. Dong | Amphan NBCA, ca 1160 m NONG | FA (crater lake) env. | 15 ° 05.9 ’ N, 107 ° 25.6 ’ E | 30 - IV 6 - V- 2010 | Lgt. St Jakl / I. G. 31.970 [p] ’.	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA3437706C08CFC46FB41FEB4.taxon	description	Description of holotype (♂). Body shape. Elongate ovoid, weakly convex. Color. Ground color blackish-brown with strong green to red metallic luster; legs dark orange except for mesoand metatarsus, meso- and metafemur, and apical part of metatibia being dark brown with green metallic luster; clypeus and antenna light brown; elytra light brown with weak metallic sheen in anterior half to two thirds; a humeral spot, anterior margin, inner half of sutural interval, lateral margin and posterior third of elytra black; black color enclosing a transverse yellow / light brown spot near lateral margin as well as a subapical subquadrate yellow / light brown spot spanning costal interval 1 to 2; another yellow elongate spot in subsutural interstice posteriorly of scutellum. Head. Clypeus subrectangular, disc very densely, transversely, confluently punctate; anterior corners rounded; anterior margin weakly reflexed; frons very shallowly impressed, punctate like clypeus, with punctures less confluent at middle; vertex very sparsely and finely punctate; ratio of interocular width / width of head approximately 0.65; antennal club longer than antennomeres 2 – 6 combined. Pronotum approximately 1.4 times wider than long, with two deep, oblique impressions on each side (posterior impression larger than anterior) and deep median longitudinal furrow; with steep decline along lateral margin; disc extremely finely and sparsely punctate, punctures becoming gradually larger laterally; sparse erect setae present near anterior angles and along lateral margin; anterior angles acute and strongly produced; posterior angles obtuse; sides of pronotum weakly converging anteriad in posterior two thirds, then strongly curved, strongly converging, and slightly sinuate in anterior third; basal marginal line interrupted before scutellum between third costal interval on each side; all other marginal lines complete. Scutellum nearly semicircular, broader than long, finely and sparsely punctate. Elytra regularly striate; three inner costal intervals (1, 2 and 3) slightly more convex than interstices; subsutural interstice with secondary stria being irregularly doubled anteriorly and almost reaching posterior elytral margin; distinct but discontinuous secondary striae also present on interstices 2 and 3; elytral surface with sparse micropunctation; humeral umbone and apical protuberance very prominent; opaque area at apical curvature broad, including interstice 4 laterally; lateral margin with wide flat paramarginal extension between humerus and middle of elytra; epipleuron broad near humerus, ending slightly posteriorly of elytral midlength; epipleuron with numerous soft short white setae near humerus and stout spiniform black setae beginning posteriorly of humerus and extending along lateral margin to apical curvature (becoming gradually larger apicad); one or two large spiniform setae present near apico-sutural angle; posterior margin evenly, separately rounded. Propygidium with dense fringe of white setae along posterior margin covering approximately 1 / 3 of propygidial length. Pygidium strongly convex; apex broadly rounded; punctation transverse and rather sparse on disc (except across vague midline near apex), transforming into concentrically arranged dense striolation at sides and base; with scattered white setae near base and in an indistinct depression at lateral margin; apex with several long, erect brownish setae. Ventral thoracic surface densely covered with soft, long, white setae. Meso-metaventral process short, compressed between mesocoxae, projecting slightly downward in lateral view, anteriorly vertical and straight; apex subrectangular, somewhat rounded; bulbiform in ventral view. Abdominal ventrites with a transverse band of dense, long, white setae on posterior half (broadly interrupted in middle) and irregular white setae on each side of anterior half of ventrites 2 – 4; ventrites 1 – 2 and anterior half of ventrite 3 carinate laterally. Legs. Meso- and metafemur with two bands of long white setae, one along anterior margin, the second emerging from a transverse row of punctures parallel to posterior margin. Protibia bidentate, broadened, approximately 3.75 times longer than wide; proximal tooth short, situated close to the rather short, curved apical tooth; inner spur short, at level of proximal tooth. Metatibia fusiform; protarsus slender; protarsomere 5 (without claws) slightly longer than tarsomeres 1 – 4 combined; inner protarsal claw approximately 3 / 4 as long as protarsomere 5, deeply incised apically, upper branch spiniform, lower branch broad, obliquely truncated; outer mesotarsal claw approximately as long as combined length of mesotarsomeres 1 – 4, curved, deeply incised at apex, upper branch spiniform; metatarsal claws very unequal, outer claw approximately twice as thick and 1 / 3 longer than inner. Aedeagus as in Figs 5 E, K, Q. Female. Protibia slender, apical tooth of protibia long and spatulate; protarsus articulated slightly basally of level of proximal tooth; inner spur long, articulated between 1 / 2 and 2 / 3 of tibial length; protarsus very slender, protarsomere 5 (without claws) shorter than tarsomeres 1 – 4 combined; modified claws of pro- and mesotarsi shorter, two apical branches more equal than in males; antennal club as long as antennomeres 2 – 6 combined. Measurements. Total body length 10.3 – 10.8 mm (HT 10.3 mm), total body width 6.0 – 6.2 mm (HT 6.0 mm). Morphological variation. The extent of the black and yellow elytral markings does not differ significantly between the two known specimens.	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA3437706C08CFC46FB41FEB4.taxon	diagnosis	Differential diagnosis. Glenopopillia rufipennis nigropicta subsp. nov. differs from the nominotypical subspecies in the following characters: large areas of elytra black laterally and posteriorly (light brown in G. r. rufipennis), body size smaller than majority of specimens of the nominotypical form, setae at lateral margin of elytra less stout and less numerous, starting near the middle of elytra (starting near humerus in G. r. rufipennis), opaque area on apical curvature of elytra weakly expressed, indistinct at apicosutural angle and laterally extending only to interstice 5 (broad in G. r. rufipennis, including interstice 4 laterally, including apico-sutural angle).	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA3437706C08CFC46FB41FEB4.taxon	etymology	Etymology. The specific epithet (adjective in the nominative singular) translates as ‘ black-colored’ and alludes to the black coloration of the elytra.	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA3437706C08CFC46FB41FEB4.taxon	distribution	Distribution. Laos (Attapeu).	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA3437706C08CFC46FB41FEB4.taxon	discussion	Remarks. We decided to give this taxon subspecies status because there are substantial external morphological differences between the two available specimens from southern Laos and all examined specimens of the nominotypical subspecies (n = 52) from China and northern Laos, but there are virtually no differences in the shape and structure of the male genitalia (Figs 5 E, K, Q). We found that all other species in this genus have a very low degree of variation in the external morphological characters, especially in color pattern (except for G. albopilosa sp. nov.). Therefore, we consider the observed external morphological differences between these two subspecies to be stable and realiable, even if only two specimens of the subspecies G. f. nigropicta subsp. nov. were available to us. Thus, these are allopatric, phenetically distinct populations that show no differences in the morphology of the male genitalia. These findings comply best with a relationship at subspecies level (DE QUEIROZ 2007). To what extent there is reproductive isolation between the two populations, which are separated by approximately 500 km distance, remains uncertain. Further material is needed to verify the taxonomical status.	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA3447704C3C6FEC6FDC8F7B4.taxon	description	(Figs 3 F, L; 4 F, L, R; 5 F, L, R)	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA3447704C3C6FEC6FDC8F7B4.taxon	materials_examined	Type locality. N Vietnam, Cao Bang Province, Vin Den, Nui Pla Oac Nature Reserve, 22 ° 33 ʹ 53 ″ N, 105 ° 52 ʹ 53 ″ E, 900 – 1300 m a. s. l. Type material (6 spec.). HOLOTYPE: ♂ (NMEC), ‘ N-VIETNAM Cao Bang Prov., vic. | Vin Den, Nui Pia Oac Nature Res. | 10. – 15. V. 2014, 22 ° 33 ’ 53 ’’ N, | 105 ° 52 ’ 53 ’’ E 900 – 1300 m, A. Skale [p] ’. PARATYPES: 1 ♂ 2 ♀♀ (ASPC, CZPC), same data as holotype; 1 ♂ 1 ♀ (CZPC, NMEC), ‘ N-VIETNAM Cao Bang Pr. | vic. Vin Den, Nui Pia Oac | Nature Res., 6. – 10. V. 2013 | 22 ° 33 ’ 53 ’’ N, 105 ° 52 ’ 53 ’’ E | 900 – 1300 m leg. A. Weigel || collection | NATURKUNDE – | MUSEUM ERFURT [yellow] [p] ’.	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA3447704C3C6FEC6FDC8F7B4.taxon	description	Description of holotype (♂). Body shape. Elongate ovoid, weakly convex. Color. Ground color blackish-brown with strong green to red metallic luster; legs dark orange except for mesoand metatarsus, meso- and metafemur, and apical part of metatibia being dark brown with green metallic luster; clypeus and antenna light brown; elytra light brown with weak metallic sheen; anterior margin before humeral umbone and mid-lateral margin dark brown; suture and inner 1 / 3 to 1 / 2 of costal interval 1 blackish-brown; some vague, pale yellow spots present as follows: one posteriorly of scutellum; two near middle of lateral margin; one between 2 / 3 and 3 / 4 of elytral length spanning costal interval 1 to 2. Head. Clypeus subrectangular, disc very densely, transversely, confluently punctate; anterior corners rounded; anterior margin weakly reflexed; frons very shallowly impressed, punctate like clypeus; vertex very sparsely and finely punctate; ratio of interocular width / width of head approximately 0.67; antennal club longer than antennomeres 2 – 6 combined. Pronotum approximately 1.5 times wider than long, with two deep, oblique impressions on each side (posterior impression larger than anterior) and shallow median longitudinal furrow; with steep decline along lateral margin; disc extremely finely and sparsely punctate, punctures becoming gradually larger laterally; sparse erect setae present near anterior angles and along lateral margin; anterior angles acute and strongly produced; posterior angles obtuse; sides of pronotum distinctly converging anteriad in posterior two thirds, then strongly curved, strongly converging, and slightly sinuate in anterior third; basal marginal line interrupted before scutellum between level of elytral interstice 2 on each side; all other marginal lines complete. Scutellum nearly semicircular, broader than long, moderately finely and sparsely punctate. Elytra regularly striate; three inner costal intervals (1, 2 and 3) slightly more convex than interstices; strial punctures large, distinct; subsutural interstice with a secondary stria that is irregularly doubled anteriorly and almost reaches posterior elytral margin; distinct but discontinuous secondary stria also present on interstice 2, secondary stria on interstice 3 only present in anterior half; elytral surface with sparse micropunctation; humeral umbone and apical protuberance very prominent; opaque area at apical curvature narrow; lateral margin with flat paramarginal extension in anterior half of elytron; epipleuron broad near humerus, ending slightly posteriorly of elytral midlength; epipleuron with several short white setae near humerus and stout strongly spiniform black setae beginning at humerus and extending in a row along lateral margin to apical curvature (becoming gradually larger apicad); three large spiniform setae present near apico-sutural angle; posterior margin evenly, separately rounded. Propygidium with dense fringe of white setae along posterior margin covering approximately 1 / 3 of propygidial length in middle and 1 / 4 of length at sides. Pygidium strongly convex; apex broadly rounded; punctation transverse and rather sparse on disc (except across vague midline), transforming into concentrically arranged dense striolation at sides and base; with two big spots of white setae near base and some white setae scattered at lateral margin; apex with several long, erect brownish setae. Ventral thoracic surface densely covered with soft, long, white setae. Meso-metaventral process short, compressed between mesocoxae, projecting slightly downward in lateral view, anteriorly vertical and straight; apex subrectangular, somewhat rounded; bulbiform in ventral view. Abdominal ventrites with transverse band of dense, long, white setae in posterior half (broadly interrupted in middle) and irregular white setae on each side of anterior half of ventrites 2 – 4; ventrites 1, 2 and anterior half of ventrite 3 carinate laterally. Legs. Meso- and metafemur with two bands of long white setae, one along anterior margin, the second emerging from a transverse row of punctures parallel to posterior margin. Protibia bidentate, broadened, approximately 3.3 times longer than wide; proximal tooth short, situated close to the rather short, curved apical tooth; inner spur very short, at level of proximal tooth. Metatibia fusiform; protarsus slender; protarsomere 5 (without claws) slightly longer than tarsomeres 1 – 4 combined; inner protarsal claw approximately 3 / 4 as long as protarsomere 5, deeply incised apically, upper branch spiniform, lower branch broad, obliquely truncated; outer mesotarsal claw approximately as long as combined length of mesotarsomeres 1 – 4, deeply incised at apex, upper branch spiniform; metatarsal claws very unequal, outer claw approximately twice as thick and 1 / 3 longer than the inner one. Aedeagus as in Figs 5 F, L, R. Female. Protibia slender, apical tooth of protibia long and spatulate; protarsus articulated slightly basally of level of proximal tooth; inner spur long, articulated between 1 / 2 and 2 / 3 of tibial length; protarsus very slender, protarsomere 5 (without claws) shorter than tarsomeres 1 – 4 combined; modified claws of pro- and mesotarsi shorter, two apical branches more equal than in males; antennal club as long as antennomeres 2 – 6 combined. Measurements. Total body length 10.6 – 11.0 mm (HT 11.0 mm), total body width 5.6 – 6.5 mm (HT 6.3 mm). Morphological variation. Number of elytral spiniform setae near apico-sutural angle varies between 2 and 4. Secondary stria on interstice 3 may or may not be present in posterior half of elytra. Shape of parameres very constant.	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA3447704C3C6FEC6FDC8F7B4.taxon	diagnosis	Differential diagnosis. Glenopopillia skalei sp. nov. is very similar to G. rufipennis rufipennis and G. fossulata in having the same light brown elytra. In contrast to those two taxa, the inner 1 / 3 to 1 / 2 of costal interval 1 is blackened (only suture blackened in G. r. rufipennis and G. fossulata), the protibia is wider in G. skalei sp. nov., and the fringe of white setae of the propygidium is half as long as the length of the propygidium in the middle (one third as long in G. rufipennis and G. fossulata). The parameres are symmetrical and similar to those of G. maculata, but are broader in dorsal view, with the subapical tooth closer to the apex and bent upwards in lateral view. Moreover, the basal piece of the ventral plate is more strongly developed (Figs 5 F, L, R).	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA3447704C3C6FEC6FDC8F7B4.taxon	etymology	Etymology. Patronymic (noun in the genitive case), named in honor of André Skale who collected most of the type material including the holotype.	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
03E6755EA3447704C3C6FEC6FDC8F7B4.taxon	distribution	Distribution. Vietnam (Cao B ằng Province).	en	Lu, Yuan Yuan, Zorn, Carsten, Král, David, Bai, Ming, Yang, Xing Ke (2018): Taxonomic revision of the genus Glenopopillia (Coleoptera: Scarabaeidae: Rutelinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2): 297-320, DOI: 10.2478/aemnp-2018-0026, URL: http://dx.doi.org/10.2478/aemnp-2018-0026
