identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
231E878BFFD5FFF0D5DA6255EACDF8D4.text	231E878BFFD5FFF0D5DA6255EACDF8D4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thinobius Kiesenwetter 1844	<div><p>Genus Thinobius Kiesenwetter, 1844</p> <p>Thinobius Kiesenwetter, 1844: 355; Herman, 1970: 395; 2001: 1735. Yosiityphlus Sawada, 1971: 327.</p> <p>See Herman (2001) for more synonymies and references.</p> <p>Diagnosis. Maxillary palpomere 4 aciculate; protrochantin exposed; epipleural ridge of elytra absent; longitudinal row of tibial spines absent; tarsal formula 2-2-2; basolateral ridges on tergites II–VII present (Herman 1970).</p> <p>Habitats. They occur on banks of rivers, streams, ponds and other freshwater, but a few are found in intertidal mudflats or fine gravels (Frank &amp; Ahn 2011).</p> <p>Key to the species of Korean coastal Thinobius</p> <p>1. Antennomeres 4–6 transverse (Fig. 2A); metaventral carinae present (Fig. 2F); paramere short, lateral part of aedeagus without serrated pattern (Fig. 3B)............................................................ Thinobius jejuensis sp. n</p> <p>- Antennomeres 4–6 elongate (Figs. 4A and 6A); metaventral carinae absent (Figs. 4E and 6E); paramere long, lateral part of aedeagus with serrated pattern (Figs. 5B and 7B)............................................................ 2</p> <p>2. Eye present (Fig. 4B); hind wings not reduced; anterior part of metaventrite with setae.......... Thinobius koreanus sp. n.</p> <p>- Eye absent (Fig. 6B); hind wings extremely reduced; anterior part of metaventrite without setae....... Thinobius kuroshio</p></div> 	http://treatment.plazi.org/id/231E878BFFD5FFF0D5DA6255EACDF8D4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lee, Jae-Seok;Yoo, In-Seong;Ahn, Kee-Jeong	Lee, Jae-Seok, Yoo, In-Seong, Ahn, Kee-Jeong (2021): Taxonomy of the coastal Thinobius Kiesenwetter (Coleoptera: Staphylinidae Oxytelinae) in Korea. Zootaxa 4985 (2): 261-268, DOI: https://doi.org/10.11646/zootaxa.4985.2.9
231E878BFFD7FFF2D5DA65FDEA37FE06.text	231E878BFFD7FFF2D5DA65FDEA37FE06.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thinobius jejuensis Lee & Yoo & Ahn 2021	<div><p>Thinobius jejuensis Lee &amp; Ahn sp. n.</p> <p>(Figs. 1A, 2–3)</p> <p>Description. Length 1.04–1.30 mm. Body flattened, surface pubescent; dark brown. Head. Almost as long as wide and slightly narrower than pronotum; widest across eyes; clypeus trapezoidal; eye moderate in size and rounded in lateral aspect, with large eye facets and interfacetal setae; temple well developed, about 0.8 times as long as eye length in dorsal view; gular suture fused and broadly divergent in anterior and posterior part; neck well developed, distinctly transversely reticulate sculptures with some short setae (Figs. 2B–C); antenna slender, moderately elongate, and pubescent, antennomere 1 longest, 4 and 6 shortest, 1–2 and 7–11 elongate, 3 and 5 slightly elongate, 4 and 6 transverse (Fig. 2A). Mouthparts. Labrum transverse, semicircula; mandible triangle, apex bifid, with one blunt subapical tooth, prostheca and molar tooth well developed; maxillary palpus with four palpomeres, palpomere 1 transverse and small, 2 slightly dilated to apex, 3 largest, 2–3 with dense setae; apical margin of 3 about 3.0 times as wide as basal margin of 4; labial palpus with three elongate palpus, apex of palpomere 1 with two long setae and one pore, apex of 2 with one pore, 3 with few short sensilla at apex; mentum quadrate with one pair of long macrosetae and many setae. Thorax. Pronotum transverse, width 0.24–0.27 mm, widest about anterior third, surface pubescent; prosternum with few setae (Fig. 2D); scutellum an inverted triangle, anterior margin of scutellum emarginate, posterior part reticulate with many setae (Fig. 2E); elytron elongate and pubescent, posterior margin with membranous lobe; hind wing present; mesoventral process slightly extended and blunted at apex; metaventral carinae incomplete; metaventrite with two pairs of long macrosetae and dense setae (Fig. 2F); all tibiae with one long macroseta in middle. Abdomen. Posterior margin of tergite VII with fringe of setae; postero-lateral margin of tergite VIII slightly convex. Secondary sexual characteristics. Male sternite VIII emarginate, female with linear form; male tergite IX with ventral structs, struts as long as remainder, female tergite IX without ventral struts. Male sternite IX elongate, female sternite IX pentagon. Genitalia. Aedeagus oval (Figs. 3A–B); paramere short; spermatheca as in Fig. 3C.</p> <p>Type material. Holotype, 1 ♂, labeled as follows: KOREA: Jeju Prov., Seogwipo-si, Haye-dong, N33°13'54.45" E126°22'19.81" 2 m, 19 X 2014, KJ Ahn, IS Yoo, JS Lee, 10-20 cm deep under/in gravels on seashore. ‘ Holotype, Thinobius jejuensis Lee and Ahn, Desig. Jae-Seok Lee and Kee-Jeong Ahn 2021 ’ ‘ Deposited in the Chungnam National University Insect Collection, Korea’. Paratypes, 3 exx., labeled as same as Holotype (1 on slide; 1 on CNUIC voucher); 30 exx., same data as former except for ‘ N33°13'54.20" E126°22'18.49" 2 m, 05 VIII 2020, JS Lee, JY Park, YJ Kim, flotation’. 24 exx., same data as former except for ‘Yeraehaean-Ro, N33.232254, E126.371442, 19 X 2020, KJ Ahn, Under pebbles/gravels by flotation’.</p> <p>Distribution. Korea (Jeju island).</p> <p>Remarks. This species is similar to T. marinus, but can distinguished by the length of temple and the shape of antennomeres. Temple is shorter than the diameter of eye in T. jejuensis Lee &amp; Ahn, sp. n. (Fig. 2B), but longer than the diameter of eye in T. marinus (Cameron, 1917). In addition, antennomeres 5 and 7–8 are elongate in the new species (Fig. 2A), but 4–8 are transverse in T. marinus. All specimens were collected on Jeju-do island, Korea</p> </div>	http://treatment.plazi.org/id/231E878BFFD7FFF2D5DA65FDEA37FE06	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lee, Jae-Seok;Yoo, In-Seong;Ahn, Kee-Jeong	Lee, Jae-Seok, Yoo, In-Seong, Ahn, Kee-Jeong (2021): Taxonomy of the coastal Thinobius Kiesenwetter (Coleoptera: Staphylinidae Oxytelinae) in Korea. Zootaxa 4985 (2): 261-268, DOI: https://doi.org/10.11646/zootaxa.4985.2.9
231E878BFFD6FFF5D5DA607FEE35F8F9.text	231E878BFFD6FFF5D5DA607FEE35F8F9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thinobius koreanus Lee & Yoo & Ahn 2021	<div><p>Thinobius koreanus Lee &amp; Yoo sp. n.</p> <p>(Figs. 1B, 4–5)</p> <p>Description. Length 1.60–1.80 mm. Body flattened, surface pubescent; light brown to dark brown, head and abdomen dark brown, pronotum, elytra and antenna brown, mouthparts and legs light brown. Head. Almost as long as wide and slightly narrower than pronotum; widest across eyes; clypeus trapezoidal; eye moderate in size and rounded in lateral aspect, with large eye facet and interfacetal setae; temple well developed, diameter of eye slightly longer than temple; gular suture fused and broadly divergent at posterior part; neck well developed, transversely reticulate sculptures distinct, seta absent (Fig. 4B); antenna slender, moderately elongate, and pubescent; antennomere 1 longest, 3 shortest, 1–11 elongate (Fig. 4A). Mouthparts. Labrum transverse, lateral margin rounded, anterior margin slightly emarginate; mandible triangular, apex bifid with subapical tooth, prostheca and molar tooth well developed; maxillary palpus with four palpomeres, palpomere 1 transverse and small, 2 slightly dilated to apex, 3 largest, 2–3 with many setae, 4 short and slender; apical margin of 3 more than 2.0 times as wide as basal margin of 4; labial palpus with three elongate palpi, palpomere 1 with one long, one short seta and two pores, 2 with one long seta and one pore, 3 with few short sensilla at apex; mentum quadrate with two pairs of long macrosetae and many setae. Thorax. Pronotum transverse, width 0.29–0.30 mm, widest near half of length, surface pubescent with seven pairs of macrosetae and many setae; prosternum with one pair of long setae and a few short setae, apex of prosternal process sharp and elevated (Figs. 4C–D); scutellum an inverted triangle, anterior margin of scutellum emarginate, posterior part reticulate with a few setae and inverted triangle impression, impression about 1.6 times as wide as long, elytron elongate and pubescent; posterior margin with membranous lobe; hind wing present; mesoventral process slightly extended and pointed at apex; metaventral carina absent; metaventrite distinctly reticulate with two pairs of long macrosetae and many setae (Fig. 4E); all tibiae with one long macroseta in middle. Abdomen. Posterior margin of tergite VII with fringe of setae; posterior margin of tergite VIII emarginate; posterior margin of sternite VIII convex. Secondary sexual characteristics. Male tergite IX with ventral structs, struts shorter than other part, female gonocoxites present; female with one ring structure (Fig. 4F). Genitalia. Aedeagus oval (Fig. 5A) with serrated pattern in lateral aspect (Fig. 5B); paramere long and slender; spermatheca as in Fig. 5C.</p> <p>Type material. Holotype, 1 ♂, labeled as follows: ‘ KOREA: Gyeongbuk Prov., Pohang-si, Buk-gu, Cheongha-myeon, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=129.39365&amp;materialsCitation.latitude=36.177425" title="Search Plazi for locations around (long 129.39365/lat 36.177425)">Cheongjin-ri</a>, N36°10'38.73" E129°23'37.15" 17 m, 12 VII 2018, IS Yoo, JS Lee, JG Jung, in coarse sand/gravels on seashore, flotation’ ‘ Holotype, Thinobius koreanus Lee and Yoo, Desig. Jae-Seok Lee and In-Seong Yoo, 2021 ’ ‘ Deposited in the Chungnam National University Insect Collection, Korea’. Paratypes, 7 exx., same data as holotype, 5 exx., same data as holotype except for ‘ N36°10ʹ37.8 ʺE129°23ʹ38.0ʺ 9m, 27 VIII 2010, KJ Ahn, TK Kim, YH Kim, IS Yoo, JH Song, SG Lee, JH Jeon, under stones on seashore.’</p> <p>Distribution. Korea.</p> <p>Remarks. This species is similar to T. marinus, but can be distinguished by the shape of antennomeres. Antennomeres 1–11 are elongate in Thinobius koreanus sp. n. (Fig. 4A), but 4–8 transverse in T. marinus. The new species is also similar to T. frizzelli from Canada and USA but can be distinguished by the shape of antennomeres. Antennomeres 4–10 are quadrate and 4–6 are almost same length in the new species (Fig. 4A), but 4–10 rounded form and 5 larger than 4 or 6 in T. frizzelli (Kincaid, 1961). All specimens were collected in the southern part of the Korean peninsula.</p> </div>	http://treatment.plazi.org/id/231E878BFFD6FFF5D5DA607FEE35F8F9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lee, Jae-Seok;Yoo, In-Seong;Ahn, Kee-Jeong	Lee, Jae-Seok, Yoo, In-Seong, Ahn, Kee-Jeong (2021): Taxonomy of the coastal Thinobius Kiesenwetter (Coleoptera: Staphylinidae Oxytelinae) in Korea. Zootaxa 4985 (2): 261-268, DOI: https://doi.org/10.11646/zootaxa.4985.2.9
231E878BFFD0FFF6D5DA6166ED50FE96.text	231E878BFFD0FFF6D5DA6166ED50FE96.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thinobius kuroshio (Sawada 1971)	<div><p>Thinobius kuroshio (Sawada, 1971)</p> <p>(Figs. 1C, 6–7)</p> <p>Yosiityphlus kuroshio Sawada, 1971: 327. Thinobius kuroshio: Gusarov &amp; Makranczy, 2004: 3; Frank &amp; Ahn, 2011: 32.</p> <p>Redescription. Length 1.02–1.32 mm. Body flattened, surface pubescent; light brown. Head. Slightly elongate and slightly narrower than pronotum; widest at about half of head; arrow shaped; clypeus slightly trapezoidal, anterolateral margin rounded; eye absent; temple well developed, gular suture fused and broadly divergent at anterior and posterior part; neck well developed, seta absent and reticulate sculptures distinct (Figs. 6B–C); antenna slender, moderately elongate and pubescent, antennomere 1 longest, 1–2 and 9–11 elongate, 3–8 slightly elongate (Fig. 6A). Mouthparts. Labrum transverse, lateral margin rounded, anterior margin slightly emarginate; epipharynx slightly developed; mandible triangle, apex bifid, with one blunt subapical tooth, prostheca and molar tooth well developed; maxillary palpus with four palpomeres, palpomere 1 transverse and small, 2 slightly dilated to apex; 3 largest, 2–3 with many setae; apical margin of 3 more than 2.0 times as wide as basal part of palpomere 4; labial palpus with three elongate palpomeres, palpomere 1 with one long and one short setae and two pores, 2 with one long seta and one pore, 3 with few short sensilla at apex; antero-middle part of hypopharynx with two blunted macro spines; mentum subquadrate with one pairs of long macrosetae and many setae. Thorax. Pronotum transverse, width 0.19–0.23 mm, surface pubescent with seven pairs of long macro setae; apex of prosternal process sharp and elevated; prosternum with one pair of long setae and few setae (Fig. 6D); scutellum an inverted triangle, middle of anterior margin emarginate, posterior part weakly reticulate with few setae and diamond shaped impression, impression about 1.7 times as wide as long; elytra elongate and pubescent, posterior margin with membranous lobe; hind wing extremely reduced; mesoventrite with one pair of long setae; mesoventral process slightly extended and pointed at apex; metaventral carina absent; metaventrite reticulate, anterior part without setae (Fig. 6E); all tibiae with one long macroseta in middle. Abdomen. Posterior margin of tergite VII with fringe of setae; posterior margin of sternite VIII convex. Secondary sexual characteristics. Male tergite IX with ventral struts, struts shorter than remainder, female tergite IX without ventral strut; male sternite IX present and female genital appendages present; female with one ring structure (Fig. 6F). Genitalia. Aedeagus oval (Fig. 7A) with serrated pattern in lateral aspect (Fig. 7B); paramere long and slender; spermatheca as in Fig. 7C.</p> <p>Materials examined. KOREA: Gyeongbuk Prov.: 3 exx., Gyeongju-si, Gampo-eup, Najeong-beach, N35°46'56" E129°29'27", 21 VI 2018, KJ Ahn, pebble beach in mid-tide, flotation; 16 exx., same data as former except for ‘ Najeong-ri, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=129.49084&amp;materialsCitation.latitude=35.782223" title="Search Plazi for locations around (long 129.49084/lat 35.782223)">Najeong-beach</a>, N35°46′56.79″ E129°29′30.95″ 0 m, 13 VII 2018, IS Yoo, JS Lee, JG Jung, 10–80 cm deep under gravels in mid-high tide, flotation’; 6 exx., Jeju Prov.: Seogwipo-si, Daejeong-eup, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=129.49193&amp;materialsCitation.latitude=35.78244" title="Search Plazi for locations around (long 129.49193/lat 35.78244)">Sangmori</a>, N33°11′55.6″ E126°16′30.7″ 2 m, 20 IX 2018, IS Yoo, JS Lee, WJ Jeong, 30–50 cm deep under fine gravels on gravel beach, flotation; 3 exx., same data as former except for ‘ N33°12'22.21" E126°17'25.76", 21 IX 2018, IS Yoo, JS Lee, WJ <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=126.29049&amp;materialsCitation.latitude=33.20617" title="Search Plazi for locations around (long 126.29049/lat 33.20617)">Jeong</a>, in gravels covered with seaweed, flotation’</p> <p>Distribution. Korea, Japan.</p></div> 	http://treatment.plazi.org/id/231E878BFFD0FFF6D5DA6166ED50FE96	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lee, Jae-Seok;Yoo, In-Seong;Ahn, Kee-Jeong	Lee, Jae-Seok, Yoo, In-Seong, Ahn, Kee-Jeong (2021): Taxonomy of the coastal Thinobius Kiesenwetter (Coleoptera: Staphylinidae Oxytelinae) in Korea. Zootaxa 4985 (2): 261-268, DOI: https://doi.org/10.11646/zootaxa.4985.2.9
