taxonID	type	description	language	source
52084A62FFB8FFB3FF236AFF980CFD00.taxon	description	Genus Chaetodellus Mašán & Halliday, 2013 Chaetodellus Mašán & Halliday, 2013. Type species Chaetodellus comatus Mašán & Halliday, 2013, by original designation. We described the new genus Chaetodellus to accommodate two species that have some character states that are not consistent with Pachylaelaps, namely (1) vertical setae j 1 reduced and minute; (2) genitiventral shield with three instead of two pairs of ventral setae; (3) tarsus II without setae modified into a spur; (4) sperm access system associated with coxae III; (5) genu I with 12 (2 - 5 / 3 - 2) instead of 11 or 13 setae (2 - 5 / 2 - 2 or 2 - 6 / 3 - 2); (6) dorsal setae heterogeneous in length, some with unusual positions on the dorsal shield, for example setae J 1 antiaxial to J 2; (7) strong reduction in the number of R setae inserted in the lateral soft integument.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FFB8FFB3FF236E449CF2F888.taxon	discussion	Berlese’s (1904) attempt to divide the genus Pachylaelaps into subgenera Pachylaelaps s. str. and Onchodellus was not generally accepted. Koroleva (1977 b) and Karg (1993) synonymised Onchodellus with Pachylaelaps. Later, the separate taxonomic status of Onchodellus was accepted by Evans & Till (1979) and Moraza & Johnston (1990), because the original Pachylaelaps exhibits two main morphological patterns previously noted by Koroleva (1977 b) and Evans & Till (1979) in their identification keys: (1) species with a slender hypostome and elongated corniculi, a sperm access system associated with coxae IV, sexual dimorphism of the palp tibia (male with palp tibial outgrowths), two spur-like distal setae on female tarsus II, and 13 setae on genu I; and (2) species with a wider and shorter hypostome and normal horn-like corniculi, a sperm access system associated with coxae III, sexual dimorphism of palp tibia not developed, one spur-like distal setae on female tarsus II, and 12 setae on genu I. The genus Onchodellus is now based especially on the following character states: (1) spermathecal tubes of sperm access system strongly elongated, fine, and associated with coxae III, (2) clunal setae J 5 needle-like and similar in size to the other dorsal shield setae, (3) gland opening gdZ 1 mostly not modified and circular (if slit-like, then antiaxial to alignment Z 1 – Z 2), (4) female tarsus II with one spur-like distal seta, and genu I with 12 pairs of setae. It is by far the largest genus of Pachylaelapidae, with up to 80 species distributed mainly in Europe, Asia and Africa.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FFB9FFB2FF236F0C9F0BFA61.taxon	discussion	Notes. We have provisionally placed this species in Onchodellus on the basis of the description of the male. Its correct placement must await the discovery of the female.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FFBCFFB7FF236F239F04FA50.taxon	discussion	Notes. We have provisionally placed this species in Onchodellus, based only on its phoresy on a scarabaeid beetle. We believe that phoresy does not occur in Pachylaelaps.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FFA1FFAAFF236BFD998EFCA0.taxon	diagnosis	Diagnosis and description. As given by Mašán (2007 a), pages 118 – 121, Figures 242 – 244, 248, 252, 257, 260, 269, 272, 273, Table 9. Specimens examined. Holotype female, Slovakia, Bodvianska Pahorkatina Wold, Hubovo Village, Ozvena Mt., 22 June 2005. Deposited in Institute of Zoology, Slovak Academy of Sciences, Bratislava. Paratypes as listed by Mašán (2007 a). Notes. Mašán (2007 a) did not designate a holotype for this species, so its name is not available from that date.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FFA7FFACFF236BAC9948FCE7.taxon	materials_examined	Type species Pachylaelaps sculptus Berlese, 1920, by original designation. Mašán (2007 a) erected the genus Pachydellus to accommodate species that share the following character states: (1) clunal setae J 5 minute, reduced to minute microsetae; (2) spermathecal tube of sperm access system strongly reduced to very short section mostly fully concealed inside the posterior margin of coxa III (Fig. 2); (3) slit-like opening of glands gdZ 1 placed in alignment with setae Z 1 - Z 2; (4) female tarsus II with one spur-like seta; (5) genu I with 12 setae; (6) male femur II always with axillary tubercle close to a large anterolateral distal projection; (7) male chelicera with spermatodactyl foliate, widened basally, curved medially and narrowed distally. Pachydellus is distributed exclusively throughout the Palaearctic region, except that Pachydellus hades may have been introduced into Australia by human activities (Halliday, 2001). It currently comprises 14 valid species from Europe and Asia, but we suspect that several of these species may include a complex of sibling species.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FFAAFFA1FF236CA19CD9F842.taxon	diagnosis	Diagnosis and description. As given by Mašán (2007 a), pages 194 – 195, Figures 472 – 477, Table 14. Specimens examined. Holotype female, Slovakia, Zlatno Village, Zlatnianske Skalky Forest, 20 June 2006. Deposited in Institute of Zoology, Slovak Academy of Sciences, Bratislava. Paratypes as listed by Mašán (2007 a). Notes. Mašán (2007 a) did not designate a holotype for this species, so its name is not available from that date.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FFAAFFA1FF236E929CC2F9A8.taxon	materials_examined	Type species Pachyglobolaelaps hallidayi Mašán, new species, here designated.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FFAAFFA1FF236E929CC2F9A8.taxon	diagnosis	Diagnosis and description. As given by Mašán (2007 a, pages 192 – 194). Pachyglobolaelaps is a highly specialised monotypic genus described by Mašán (2007 a). Perhaps as a result of its myrmecophilous mode of life, it differs from all the other pachylaelapid genera in having an extended and highly vaulted dorsal shield overlapping the lateral sides of the idiosoma and encroaching onto the ventral surface, a tubular sperm access system, minute microsetae J 5, and an unusual arrangement of dorsal setae, some of which are in ventral positions. The highly distorted and arched dorsal shield makes it extremely difficult to homologise these setae.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FFABFFA0FF236ABA9FB8FE7F.taxon	type_taxon	Type species Pachylaelapsoides longipedis Mašán, 2007, by original designation.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FFABFFA0FF236ABA9FB8FE7F.taxon	description	The monotypic genus Pachylaelapsoides may be distinguished from the other pachylaelapine genera by the following characters: (1) presence of an increased number of setae on some segments of leg IV: genu IV with eight setae and tibia IV with nine setae, instead of the usual seven setae on both segments; (2) at least legs I and IV longer than idiosoma; (3) enlarged slit-like gland pores gdZ 1, (4) well developed (not minute) needle-like clunal setae J 5; and (5) bipartite projection on male femur II (Mašán, 2007 a).	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FFABFFA0FF2368C8990CFB14.taxon	type_taxon	Type species Laelaps calcariger Berlese, 1902, by original designation.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FFABFFA0FF2368C8990CFB14.taxon	diagnosis	The diagnosis of the monotypic genus Pachysphaerolaelaps was based on the following character states: (1) dorsal shield and soft integument hypertrichous; (2) almost all leg segments with an increased number of setae; (3) epistome unipartite and elongate; (4) male femur II with two adjacent cuticular processes; (5) idiosoma subglobular; (6) legs longer than dorsal shield; (7) gland opening gdS 4 hypertrophied and circular; (8) anal shield subcircular; and (9) tarsus II uniformly setaceous and without spur-like distal setae. Pachysphaerolaelaps was proposed to accommodate the species Laelaps calcariger Berlese, 1902, which was a member of the genus Sphaerolaelaps as classified by Berlese (1904) and subsequent authors (Koroleva, 1977 b; Karg, 1993).	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FFABFFA3FF236C229996FEA8.taxon	materials_examined	Type species Laelaps holothyroides Leonardi, 1896, by original designation. The original spelling of the name of this genus was Spherolaelaps (Berlese, 1903 a: 243). Since then it has always been spelled as Sphaerolaelaps or Sphaerolaelaps, including by Berlese himself. That spelling is maintained here in the interest of stability. Berlese (1903 a) proposed the new genus Sphaerolaelaps with Laelaps holothyroides Leonardi, 1896 as type species, and placed it in the Laelapidae. Later, Berlese (1904) placed the genus together with Pachylaelaps into the Gamasidae. Hirschmann & Krauss (1965) and Karg (1971) placed Sphaerolaelaps holothyroides in Pachylaelaps, but their action has not gained general acceptance. The genus was regarded as valid by Berlese (1913 a), Vitzthum (1931), Koroleva (1977 b), Koroleva & Sklyar (1982), Moraza & Johnston (1990), Karg (1993) and Mašán (2007 a). This monotypic genus may be recognised by the presence of distinctive modified dorsal setae, which are elongated and thickened, with club-like tips, the atypical position of some dorsal setae, hypertrichous opisthogastric region, the unusual shape of the epistome, bifurcated processes on genu II and tibia II in the male, globular idiosoma, legs longer than the idiosoma, and uniformly setaceous tarsi II, without spur-like distal setae.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FFA8FFA3FF236ECD9965FA04.taxon	discussion	Notes. The holotype of Neopachylaelaps mancus is a female. That was not explicitly stated by Mašán (2007 a).	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FFA8FFA2FF236FC49EA1FEA8.taxon	type_taxon	Type species Pachylaelaps (Olopachys) scutatus Berlese, 1910, by original designation.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FFA8FFA2FF236FC49EA1FEA8.taxon	discussion	Berlese (1910 a) described Olopachys as a subgenus of Pachylaelaps, in which the females have the anal and genitiventral shields fused together. Koroleva (1977 b), Karg (1993) and Mašán (2007 a) considered this character state to be of generic importance because there is no other pachylaelapid genus with an entire genitiventrianal shield covering the posteroventral idiosoma. The genus was re-defined by Mašán (2007 a, 2007 b), and currently comprises 14 described species distributed mainly throughout the Caucasus and adjacent regions. In Olopachys, as in other genera of Pachylaelapinae, there are species with 1 – 2 spur-like distal setae on tarsus II, as well as species with dorsal setae J 5 either minute or normal. Mašán (2007 b) divided the genus into two subgenera, namely Olopachylaella and Olopachys s. str. The subgenus Olopachylaella may be reliably distinguished from Olopachys s. str. mainly by the presence of normal needle-like clunal setae J 5, and a shortened and foliate spermatodactyl. In general appearance, gnathosomal features and armature of legs II (not considering the fusion of the genitiventral shield and anal shield), Olopachylaella is similar to Pachylaelaps (Longipachylaelaps), which also has normally developed setae J 5. More significant similarities in external morphology between Olopachylaella and Longipachylaelaps than between Olopachylaella and Olopachys s. str. suggest convergent or parallel evolution of the two Olopachys subgenera as well as the secondary fusion of the genitiventral and anal shields in Olopachylaella.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FFA8FFA3FF23689F9FC4FB0A.taxon	description	Genus Neopachylaelaps Mašán, 2007 Neopachylaelaps Mašán, 2007 a: 102. Type species Neopachylaelaps mancus Mašán, 2007, by original designation. Mašán (2007 a) based his diagnosis of the monotypic European genus Neopachylaelaps on the following character states: (1) deficient chaetotaxy of some leg segments, specifically genu I lacking ventral seta av 2 and dorsal seta pd 3, femur I and tibia I lacking ventral seta av 2, and genu III lacking ventral seta av 1; (2) deficient chaetotaxy of the dorsal shield, with setae S 1 and R absent; (3) moderate reduction of the anterior section of the peritremes, with tips not reaching the anterior dorsal surface; (4) proximal insertion of the spermatodactyl on the movable digit in the male; (5) reduction of palp tibial projections in the male, with only a simple conical tubercle present; and (6) epistome produced into long, lanceolate and apically bifurcate projection.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FFAFFFA4FF236B929ECBF98F.taxon	materials_examined	Type species Gamasus pectinifer G. Canestrini, 1881, by subsequent designation (Berlese, 1904), but see below. The type species of Pachylaelaps is always said to be Gamasus pectinifer G. Canestrini, 1881, but that is impossible. The genus name Pachylaelaps was first used by Berlese (1888 b, 30 November, date from Poggi, 2008), with two included species, Pachylaelaps haeros Berlese, 1888 and Pachylaelaps athleticus Berlese, 1888. The type species of the genus must therefore be either P. haeros or P. athleticus. Gamasus pectinifer cannot be the type species of the genus because it was not one of the species originally included when the genus was described (International Code of Zoological Nomenclature, Article 67.2). Only ten days later Berlese (1888 a, 10 December) redescribed the genus and added a third species, " Gamase Savignii ", apparently referring to Acarus savignyi Audouin, 1826. Berlese (1892 c) then moved P. haeros and P. athleticus into his new subgenus Pachylaelaps (Megalolaelaps), and placed G. pectinifer in Pachylaelaps (Pachylaelaps). This arrangement would make Pachylaelaps (Megalolaelaps) an objective junior synonym of Pachylaelaps (Pachylaelaps). Berlese (1904) then re-described Pachylaelaps yet again, and stated that its type species was P. pectinifer. That invalid type species designation has been followed ever since. Pachylaelaps haeros and P. athleticus are both now placed in the genus Megalolaelaps, which is the type genus of the family Megalolaelapidae. The use of either of these species as the type species of Pachylaelaps would seriously disrupt the taxonomic concept of the genus, which has been stable for over 100 years. We therefore maintain the current usage of P. pectinifer as the type species of Pachylaelaps in the interest of stability. Further re-descriptions of Pachylaelaps were provided by Evans & Hyatt (1956), Costa (1971) and Koroleva (1977 b). Mašán (2007 a) clarified the concept of the genus Pachylaelaps by removing some species that obviously belong in other genera (e. g. Onchodellus, Pachydellus), and described the new subgenus Longipachylaelaps, which may be easily separated from Pachylaelaps s. str. by the presence of normal needle-like clunal setae J 5 (minute microsetae in Pachylaelaps s. str.) and only one pair of hypertrophied slit-like gland openings gdS 4 on the posterolateral margin of the dorsal shield (two pairs gdZ 1 and gdS 4 in Pachylaelaps s. str.). Pachylaelaps can be now easily identified by the following characters: (1) sperm access system of females associated with coxae IV (Fig. 4); (2) female tarsus II with two spur-like distal setae, pl 1 and pl 2; (3) genu I with 13 setae; and (4) tibial projections on male palp present (except for species of Pachylaelaps pectinifer group). It currently comprises up to 60 known species found mostly in the Palaearctic Region, and only rarely in Africa, North America, Australia and South Pacific Islands.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FF97FF9CFF23692C9FAEFA7C.taxon	materials_examined	Type species Pachylaelaps (Elaphrolaelaps) fenestratus Berlese, 1910, by original designation.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FF97FF9CFF23692C9FAEFA7C.taxon	description	The type species E. fenestratus has never been described or re-described in detail. As a result, it is difficult to construct a reliable diagnosis for the genus. The most valuable information on the genus was presented by Costa (1974, 1975) and Elsen (1974 a, 1974 b). Elsen (1974 a) divided the genus into two subgenera, Elaphrolaelaps (Incisosternum) and Elaphrolaelaps s. str. His new subgenus was based primarily on the presence of a deep incision on the posterior margin of the sternal shield, the reduced number of setae in the soft integument, and the greatly elongated setae inserted around the margins of the dorsal shield.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FF97FF9CFF236ABA9D48FD51.taxon	materials_examined	Type genus Paralaelaps Trägårdh, 1908.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FF97FF9CFF236ABA9D48FD51.taxon	description	Description. Dorsal idiosoma with holodorsal shield, the shield entire but not covering whole dorsal surface, bearing 30 – 38 pairs of setae. Dorsal shield setae simple, smooth and needle-like. Presternal plates absent. In females, sternal shield fused with endopodal and metasternal platelets to form a sternal-metasternal complex, bearing four pairs of setae and three pairs of lyrifissures; epigynal shield fused with ventral shield to form a genitiventral shield, this shield not strongly expanded towards the anal shield, suboval to tongue-shaped, normally rounded on posterior margin, bearing a pair of genital setae and usually one pair of ventral setae (in Paralaelaps pietersburgensis, the genitiventral shield is polytrichous). Peritrematal shields fused with exopodal and parapodal platelets, free from lateral margins of genitiventral shield, and only moderately produced posteriorly. Anal shield free, subtriangular to suboval, bearing three circumanal setae and small anus. Males with holoventral shield; peritrematal shields connected to ventral scutal complex. Legs shorter than idiosoma, tarsi I – IV with minute claws and well formed empodium (claws may absent), tarsi II with no distal setae modified into a spur. Sperm access system unknown.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FF94FF9FFF236FD19FD9F865.taxon	materials_examined	Type species Pachylaelaps (Paralaelaps) kibonotensis Trägårdh, 1908, by monotypy.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FF95FF9EFF236E13986CF8C1.taxon	materials_examined	Type species Mirabulbus qinbaensis Liu & Ma, 2001, by original designation. Liu & Ma (2001) erected Mirabulbus to accommodate two species from China, the type species Mirabulbus qinbaensis Liu & Ma, 2001, and Mirabulbus yadongensis (Ma & Wang, 1997). Unfortunately, their generic diagnosis contains only some very vague and general features which do not allow precise characterisation and classification of the new genus. In addition, they incorrectly placed Mirabulbus in the family Bulbogamasidae Gu, Wang & Duan, 1991, which may be a synonym of Neoparasitidae Oudemans, 1939 (see remarks for the genus Neoparasitus). Three more free-living species have been described on the basis of specimens collected from soil substrates, and which appear similar to Mirabulbus, viz. Neoparasitus scleoides Ishikawa, 1969 and Neoparasitus punctatus Ishikawa, 1987 from Japan, and Pachyseius malimingi Bei, Chen & Wu, 2010 from China. Based on the illustrations accompanying the two original species of Mirabulbu s, together with more precise descriptions of the other species from China and Japan, Mirabulbus appears to be closely related to Pachyseius. These two genera share the following character states: (1) dorsal shield with 30 pairs of setae; (2) sternal shield with three pairs of setae; (3) a pair of metasternal platelets present, each with a seta and associated lyrifissure; (4) separate epigynal shield with a pair of setae and minute suboval post-epigynal sclerites close to its posterior margin; (5) ventri-anal shield bearing at least two pairs of ventral setae; (6) epistome with anterior margin not produced into a narrowly necked projection; (7) palp apotele three-tined; (8) chelicera with one setal brush at the base of the movable digit; (9) male with spermatodactyl long and projecting posterodorsally; (10) male with separate sternitigenital and ventri-anal shields, and with free posterior ends of peritrematal shields. For the main distinguishing characters between Mirabulbus and Pachyseius refer to the key to genera.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FF9AFF91FF236EAB9F87F83A.taxon	materials_examined	Type species Pachyseius humeralis Berlese, 1910, by original designation. The genus Pachyseius was proposed by Berlese (1910 a), but has sometimes been considered as a synonym of Neoparasitus Oudemans, 1901 in the family Neoparasitidae Oudemans, 1939, for example by Vitzthum (1942) and Baker & Wharton (1952). Evans (1957 a) placed Pachyseius as a separate genus in a very broad and heterogeneous family Neoparasitidae. Later Evans & Till (1979) placed this genus in the family Pachylaelapidae, but Karg (1971, 1993) placed Pachyseius in the family Macrochelidae rather than the Pachylaelapidae. Karg (1971) proposed a separate tribe Pachyseiini Karg, 1971 for the genus Pachyseius, primarily because of the simple form of the epistome, short dorsal setae, straight peritremes, and fused ventral and anal shields. Although most other authors have placed Pachyseius in the family Pachylaelapidae, the systematic position of the genus in the family seems to be fragile and provisional. Mašán (2007 a) placed it in the subfamily Pachyseiinae, and that usage is followed here. The relationship of Pachyseius to the other genera of Pachylaelapid is unclear. On the basis of the recently described species (Mašán & Mihál, 2007), it appears that the general evolutionary trend in Pachyseius is towards increased sclerotisation, including fusion of some ventral shield components to form a contiguous exopodalperitrematal complex, a sternal-metasternal complex, and abnormally large metapodal platelets widely abutting the parapodal platelets and ventri-anal shield.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FF98FF92FF236D6A996DFE1F.taxon	type_taxon	Type species Pachyseiulus hispanicus Moraza & Johnston, 1990 (= Pachylaelaps singularis Schweizer, 1961), by original designation.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FF98FF92FF236D6A996DFE1F.taxon	description	The monotypic genus Pachyseiulus was placed in the family Pachylaelapidae by Moraza & Johnston (1990), and their description needs no amendment. The genus is based mainly on the following character states: (1) dorsal shield with 37 pairs of setae, including a full complement of J, Z and S opisthonotal setae; (2) tarsus II in female with distal setae al 1 and pl 1 spur-like; (3) palp apotele 2 - tined; (4) epistome nearly truncate, without anterior projection; (5) genitiventral shield bearing four pairs of setae; (6) male chelicera with fixed digit distally truncate. The type species of the genus, Pachyseiulus hispanicus, described from Northern Spain, was synonymised with Pachylaelaps singularis Schweizer, 1961 by Mašán (2007 a), who examined Schweizer’s original type material.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FF99FF92FF23689F9D2AF945.taxon	materials_examined	Type species Pseudoparasitus (Pseudopachys) parasitizans Berlese, 1916, by original designation.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FF99FF92FF23689F9D2AF945.taxon	materials_examined	Type species Pseudopachyseiulus ignacii Moraza & Johnston, 1993, by original designation. The genus Pseudopachys was proposed and briefly described by Berlese (1916 b), as a subgenus of Pseudoparasitus Oudemans, 1902, which is now classified in the Laelapidae. There are two slides of Pseudopachys parasitizans in the Berlese Acaroteca (and one vial, 39 ° / 1917, which was not examined by us). One of the slides (181 / 43) is labelled Pseudopachys parasitizans, " tipico ", from the European mole, Talpa europaea, Italy (Padova). This slide contains one male and one female, of which only the female is in relatively good condition for examination. Another slide (206 / 13) in the collection is labelled Pseudoparasitus (Pseudopachys) parasitizans, from moss, Italy (Tiarno-Trentino), and it bears one barely observable female. In addition, we have examined specimens of this species from south-eastern France (French Prealps, Vercors, Petit Veymont, Saint- Michel-les-Portes, 1,385 m a. s. l., August 26, 2007, leaf litter of beech forest, coll. Peter Fenďa). Examination of the female specimen on slide 181 / 43 confirmed the conspecificity of Pseudoparasitus (Pseudopachys) parasitizans with Pseudopachyseiulus ignacii, which was originally found in Spain, and described in detail by Moraza & Johnston (1993). These two nominal species share following characters: (1) tarsus II with two hypertrophied distal setae al 1 and pl 1, located in characteristic opposed anterolateral and posterolateral positions; (2) holodorsal shield with more than 30 pairs of short, needle-like setae; (3) anterior margin of sternum with a pair of presternal plates; (4) sternal shield with convex posterior margin and three pairs of setae; (5) genitiventral shield constricted at the level of genital setae st 5, bearing two pairs of setae (st 5 and Jv 1), and with posterior margin closely adjacent to anterior margin of subtriangular anal shield; (6) endopodal and exopodal platelets not fused, clearly separated between coxae II – IV; (7) parapodal platelets well developed behind coxae IV, widely rounded on posterior margin; (8) femur II with a small ventral tubercle; (9) separate small suboval metapodal platelets. Because Pseudoparasitus (Pseudopachys) parasitizans was designated as the type species of the genus Pseudopachys by Berlese (1916 b) and Pseudopachyseiulus ignacii as the type species of the genus Pseudopachyseiulus by Moraza & Johnston (1993), Pseudopachyseiulus becomes a junior synonym of Pseudopachys.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FF99FF92FF236D39986FF863.taxon	description	Pseudoparasitus parasitizans. — Hallan, 2005.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FF9DFF96FF236AFC9F0AFD80.taxon	discussion	Notes. Franz (1975, Table 227) referred to a species as " Pachylaelaps parvulus Willm. ". Schmölzer (1995) used the same name, in the form " Onchodellus parvulus Willmann (nom. nud.) ", which he considered to be a junior synonym of Pachylaelaps strigifer Berlese, 1892. We have not been able to find any description of Pachylaelaps parvulus Willmann or Onchodellus parvulus Willmann. However, in the Willmann collection in Munich, there is a slide labelled Pachylaelaps (Onchodellus) parvulus (shelf No. 46 / slide No. 14). This slide is in poor condition and only legs II and the gnathosoma of a male mite are visible (Mašán, personal observations). The comments by Franz (1975) and Schmölzer (1995) may refer to the same slide.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FF9DFF96FF2368489E76FCA0.taxon	discussion	Notes. Castagnoli & Pegazzano (1985) referred to a specimen in the Berlese Acaroteca as Pachylaelaps brachiosus Berlese, but this species was apparently never described by Berlese. Pachylaelaps brachiosus Hirschmann & Krauss, 1965 could be based on the male specimen deposited in the Berlese collection but that cannot be confirmed on the basis of the available information.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FF9DFF96FF2369A89882FBA4.taxon	discussion	Notes. A species with this name is listed in the catalogue of the Oudemans collection by Buitendijk (1945), based on specimens from Indonesia. We have not been able to find a description of this species in any Oudemans publication, and it is not listed in the index to Oudemans' works by Van Eyndhoven (1943).	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FF9DFF96FF236EA49882FAB8.taxon	discussion	Notes. A species with this name is listed in the catalogue of the Oudemans collection by Buitendijk (1945), based on specimens from Indonesia. We have not been able to find a description of this species in any Oudemans publication, and it is not listed in the index to Oudemans' works by Van Eyndhoven (1943).	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FF9DFF96FF236FA09F07F990.taxon	discussion	Notes. Castagnoli & Pegazzano (1985) referred to a specimen in the Berlese Acaroteca as Pachylaelaps ovatus Berlese, but this species was apparently never described.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FF9DFF96FF236C789D82F820.taxon	discussion	Notes. Franz (1975, Table 227) referred to a species as " Pachylaelaps pannonicus Willm. ". Schmölzer (1995) used the same name, in the form " Pachylaelaps pannonicus Willmann (nom. nud.) ", which he considered to be a junior synonym of Pachylaelaps pectinifer (G. & R. Canestrini). We have not been able to find any published description of Pachylaelaps pannonicus Willmann. However, in the Willmann collection in Munich, there is a slide labelled Pachylaelaps pannonicus n. sp. (det. C. Willmann) (shelf No. 45 / slide No. 12). The specimen on this slide is represented only by legs II and the gnathosoma of a male mite. It appears to be a specimen of Pachylaelaps littoralis Halbert, 1915 (Mašán, personal observations). The comments by Franz (1975) and Schmölzer (1995) may refer to the same slide.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FF82FF89FF236E1C985FFAE7.taxon	discussion	Notes. Actinoseius, as a subgenus of the genus Epicriopsis Berlese, 1916, is included in the list of genera of Pachylaelapidae compiled by Hallan (2005). The original description of the genus and its type species Actinoseius terrificans are useless for species and genus recognition. We see no reason to include this genus in the Pachylaelapidae, since Epicriopsis is now classified in the family Ameroseiidae. Actinoseius terrificans is not included in the Catalogue of the Berlese Acaroteca (Castagnoli & Pegazzano, 1985).	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FF82FF89FF236FE69976F945.taxon	discussion	Notes. Oudemans (1929) described the genus Beaurieuia with Neopodocinum nederveeni as its type species. Krantz (1965) placed B. nederveeni in the genus Neopodocinum (Macrochelidae), but did not make any reference to Beaurieuia. Bregetova (1977 a) confirmed the synonymy of Beaurieuia with Neopodocinum, but the genus name Beaurieuia can be still found in some inventories of the family Pachylaelapidae (Hallan, 2005; Walter, 2006).	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FF82FF89FF236AB59846FEF2.taxon	discussion	Notes. A species with this name is mentioned in the abstract of the paper published by Moraza & Peña (2005 c), as a newly described species. Pachylaelaps subkarawaiewi was never described in that paper (or in other publications by these authors), but it may be the same species as Pachylaelaps minutus, which is not listed among the seven species introduced as new in the abstract summary given by Moraza & Peña (2005 c).	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FF82FF89FF236B959F07FDAA.taxon	discussion	Notes. Castagnoli & Pegazzano (1985) referred to a specimen in the Berlese Acaroteca as Pachylaelaps terebrifer Berlese, but this species was apparently never described.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FF82FF89FF2368AD9F07FC82.taxon	discussion	Notes. Castagnoli & Pegazzano (1985) referred to a specimen in the Berlese Acaroteca as Pachyseius spectandus Berlese, but this species was apparently never described.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FF82FF88FF236D079F2EFE3A.taxon	discussion	Notes. There is only one slide bearing the type species Pachylaelaps (Brachylaelaps) rotundus in the Berlese Acaroteca, on slide number 107 / 47, labelled " tipico ". It contains a poor quality male specimen from Java, Indonesia, which can be characterised as having some leg segments strongly armed with spurs (femur II, genu II, tarsus II, trochanters III−IV, femur IV), a long and slim spermatodactyl originating at the middle of the movable digit, and an unusual arrangement of ventral shields, with separate sternitigenital, ventri-anal and peritrematal shields. Its classification among the Pachylaelapidae cannot be established with any confidence until more material is collected. It may be better placed in the Neoparasitidae, perhaps as a synonym of Neoparasitus Oudemans, 1901, but that cannot be confirmed with the available information.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FF83FF88FF2368229D27FD74.taxon	discussion	Notes. We consider this genus to be a synonym of Neoparasitus Oudemans, 1901. See below for notes on the genus Neoparasitus.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FF83FF8BFF23690F990FFD3F.taxon	discussion	Notes. When Berlese (1892 c) described Megalolaelaps, as a subgenus of Pachylaelaps, he included two species, Pachylaelaps heros Berlese, 1888 (sic, should be haeros) and Pachylaelaps athleticus Berlese, 1888, but did not designate a type species. He also doubtfully included Pachylaelaps tetragonoides (Dugès), P. gigas (Dugès) and P. savignyi (Audouin) in this subgenus. Vitzthum (1942, page 761) designated M. haeros as the type species. The original descriptions of M. haeros and Megalolaelaps are of limited value for species and genus recognition. Fonseca (1946) provided a more useful re-description of Megalolaelaps immanis Berlese, 1910, and also proposed the new monotypic family Megalolaelapidae (as Megalolaelaptidae) to accommodate this genus. We consider this action to be reasonable because M. immanis exhibits a combination of characters not observed in Pachylaelapidae. These species are 3 – 4 mm long and up to 2 mm wide, so they are among the biggest known Mesostigmata. More valuable information about Megalolaelaps can be found in Gorirossi-Bourdeau (1956) and Hennessey & Farrier (1988). Fonseca (1946), Gorirossi-Bourdeau (1956) and Hennessey & Farrier (1988) redescribed and illustrated two species of Megalolaelaps, from which we derived the brief differential comparison between Megalolaelapidae and Pachylaelapidae (see below). Fonseca (1946) described the female and male of Megalolaelaps immanis, and Gorirossi-Bourdeau (1956) and Hennessey & Farrier (1988) redescribed the female of Megalolaelaps ornatus (Keegan, 1946) (Gorirossi-Bourdeau only gnathosomal structures of the species). Females of these species can be easily separated based especially on the position of the metasternal lyrifissures, the form of the epigynal shield, and the form and size of the metapodal platelets. In M. ornatus, (1) the metasternal gland openings are unusually situated on the sternal shield; (2) the epigynal shield is pyriform, constricted between coxae IV and widened posteriorly, with the genital setae almost medially positioned on the shield; (3) the metapodal platelets are bacillate and inconspicuous in size. In M. immanis, (1) the metasternal gland openings are not placed on the sternal shield and the shield has the normal two pairs of lyrifissures; (2) the epigynal shield is irregular, subpentagonal, with the anterior portion more expanded than the narrow posterior part, concave on its posterior margin, with genital setae inserted closer to the lateral margins; (3) metapodal platelets are suboval and conspicuously enlarged. Both species possess the following specific character states that are not found in Pachylaelapidae, and which could be considered as the main diagnostic characters for the family Megalolaelapidae: (1) ventral shield absent (in Pachylaelapidae, the ventral shield bears at least one pair of ventral setae and is always developed, either as a part of the genitiventral or ventri-anal shield in females, or as a part of the holoventral or ventri-anal shield of males); (2) exopodal platelets are absent, so the inner margin of the peritremes is adjacent to the soft integument of the circum-coxal surface (in Pachylaelapidae, exopodals are present, usually well developed and covering the soft integument of the intercoxal areas, closely abutting or fused to the peritrematal shield to form a well developed endopodal-exopodal-peritrematal shield complex); (3) the anal shield in the male is free (in Pachylaelapidae, the anal shield of males is always fused and incorporated into the holoventral or ventri-anal shield); (4) the spermatodactyl is long, tightly coiled, and inserted medially in the outer lateral surface of the movable digit (in Pachylaelapidae, the spermatodactyl is never tightly coiled); (5) the size of the three cheliceral segments exhibits sexual dimorphism, in which the cheliceral segments of the male are larger and more robust than those of the female (in Pachylaelapidae, the chelicerae of both sexes are similar in size); (6) the palp trochanter has a large, horn-like projection on the anteroventral surface, which does not occur in Pachylaelapidae. In addition, there may be other characters in Megalolaelapidae which are considered as unusual or rarely developed in Pachylaelapidae, for example: (1) metasternal setae inserted on soft integument, (2) hypertrichy of lateral and ventral soft integument, (3) epistome with anterior apex bifid, (4) palp apotele 2 - tined. Unfortunately no information on the sperm access system or dorsal shield and leg setation in Megalolaelaps is available in the published literature. It is not clear why some authors (Vitzthum, 1931; Baker & Wharton, 1952; Krantz, 1965; Hennessey & Farrier, 1988) classified Megalolaelaps in the Pachylaelapidae, except spur-like setae pl 1 and al 1 on the tarsus II (in female of M. ornatus), which can also be found in some Macrochelidae (Krantz, 1965).	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FF80FF8BFF2368DE9E0BFB75.taxon	discussion	Notes. Turk (1948) described Meliponapachys pallidus on the basis of a single male specimen from the nest of the bee Melipona fasciata trinitatis Cockerell, 1919 in Trinidad. Because of the brevity of his original description and the lack of further collections, the genus has remained essentially unknown. The described specimen displays some unusual features that are not typical of the Pachylaelapidae, e. g. a pair of presternal plates each bearing a sternal seta st 1, and minute free peritrematal shields. In the same paper and from the same nest of the host bee, Turk (1948) described a female mite as Hypoaspis favosus. This species also has a pair of presternal plates each bearing a sternal seta st 1, which is unknown in Hypoaspis Canestrini, 1884. In addition, both species possess similar reticulate ornamentation of the ventral shield. It is very likely that these two species are different adult stages of the same species. This hypothesis should be confirmed by further comparative study of mites collected in similar habitats in the Neotropical Region.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FF80FF8AFF236F179F88FDF7.taxon	discussion	Notes. Neoparasitus, as type genus of the family Neoparasitidae Oudemans, 1939, has been classified in the same family by several later authors including Vitzthum (1941) and Baker & Wharton (1952). The family was based especially on the presence of a 3 - tined palp tarsal apotele and a holodorsal shield. It originally included various genera, most of which have since been transferred to other families, especially Ameroseiidae, Laelapidae, Macrochelidae and Ologamasidae, and the family now includes only the type genus Neoparasitus. Present knowledge of the type genus Neoparasitus was recently summarised in detail by Moraza & Johnston (2002). They considered Neoparasitus and Bulbogamasus as synonyms, and discussed problems with the classification of the genus. Moraza & Johnston left the systematic position of Neoparasitus open, stating that " although the relationships between Neoparasitus and other genera of the family Pachylaelapidae appear strong enough to consider them as members of the same family, this genus has characteristics shared with other eviphidioid families ". It should be noted that the concept of Neoparasitus used by Moraza & Johnston (2002) is based on N. orientalis and N. sinicus, not the type species N. oudemansi Oudemans, 1901. We here retain the monotypic family Neoparasitidae in the superfamily Eviphidoidea, rather than placing Neoparasitus in either the Neoparasitinae or Pachyseiinae within the Pachylaelapidae, for the following reasons. Neoparasitus has in the female (1) a dorsal shield with more than 30 pairs of setae; (2) salivary styli well sclerotised; (3) corniculi weakly sclerotised, adjacent each to other and with hyaline fringe laterally, not horn-like; (4) cheliceral digits robust, each with a coarsely striate major tooth; (5) palp genu with seta al 2 spatulate; (6) palp tarsus with elongate distal chemoreceptor; (8) exopodal platelets absent; parapodal platelets free and each well separated from peritrematal shield; (9) tarsi II – IV with minute claws and well developed empodium; (10) trochanters II−IV usually with apical dorsal spur; (11) genital setae sometimes off the epigynal shield; and in the male, (7) spermatodactyl strongly elongate and coiled. In addition, species of Neoparasitus are mostly insecticolous species associated with coprid beetles, unlike the free-living edaphic species classified in the Pachyseiinae. The question of the correct classification and systematic placement of Neoparasitus will not be resolved until more detailed morphological analysis of these mites is carried out, including study of their developmental stages and a complete re-description of the type species Neoparasitus oudemansi Oudemans, 1901 (= Pachyseius quartus Vitzthum, 1926).	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FF81FF8AFF2368969EA2FA91.taxon	discussion	Notes. The genus Pachylaella was proposed by Berlese (1916 b) with Pachylaella robustissima as type species, and placed in the family Pachylaelapidae. There are four slides and seven adult specimens (three females, four males) of this species in the Berlese Acaroteca, viz. 182 / 9 (type) and 182 / 6 – 8, all from the dung beetle Atheucus sp. in La Plata, Argentina, but only one female specimen is in suitable condition for examination (on slide 182 / 6). Pachylaella robustissima does not appear to be a member of the family Pachylaelapidae. The sternal shield has three pairs of setae; there is a pair of small circular metasternal platelets separate from the sternal shield; the epigynal shield is tongue-shaped, not enlarged posteriorly, and bears only one pair of genital setae; the anal shield is subcircular, enlarged anteriorly, almost adjacent to the epigynal shield, and appears to have some ventral setae on its surface. Pachylaella could be classified in the family Macrochelidae rather than the Pachylaelapidae. The ventral idiosoma of P. robustissima resembles some species of Holostaspella Berlese, 1903. In addition, the peritremes of P. robustissima are looped posteriorly, and join the stigma posteriorly, which is a specific character typical of most of the known macrochelid genera. Unfortunately, some important characters on the chelicerae were obscure and not visible in the available material.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FF81FF8AFF236F7B98E4F85D.taxon	discussion	Notes. No type specimens of the monotypic genus Platylaelaps are available in the Berlese Acaroteca in Florence. The material is missing and must be considered lost (Castagnoli & Pegazzano, 1985). The original description in Berlese (1905) is too brief and insufficient to enable the genus and species to be recognised. A schematic illustration of the ventral side of a female is available in Berlese 1905 (Pl. XVI, Fig. 29). From that illustration we can derive some useful information: (1) tarsus II has one spur-like seta, legs II are enlarged, (2) a continuous sternal-metasternal-endopodal-exopodal-peritrematal shield complex is present, (3) the peritrematal shields are not strongly expanded behind coxae IV; (4) the parapodal platelets are very narrow and not connected to the epigynal or ventral sections of the geniti-ventri-anal shield; (5) the anal and epigynal shields are fused with the ventral shields to form a spindle-shaped geniti-ventri-anal shield. These features may be found in other pachylaelapid genera, but a complete geniti-ventri-anal shield can be found only in Olopachys. In contrast to Platylaelaps, Olopachys has the peritrematal shields strongly expanded behind coxae IV and connected to the lateral margins of the geniti-ventri-anal shield. The genus Platylaelaps appears to be a member of the Pachylaelapidae or Neoparasitidae, but we cannot be more specific until more material is collected.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FF86FF8DFF2368809CF2FBD7.taxon	discussion	Notes. The genus Zygoseius was revised by Halliday (1997) and Karg (1998), and is now well documented. It has been provisionally placed in the families Ascidae (Evans, 1958; Sheals 1962; Hyatt 1964), Pachylaelapidae (Lindquist & Evans 1965; Krantz & Ainscough, 1990; Halliday, 1997; Lindquist et al., 2009) and Halolaelapidae (Karg, 1988), but none of these proposals is convincing. The presence of a strongly sclerotised sternal shield fused with the endopodal platelets is not consistent with the Halolaelapidae as defined by Halliday (2008) and Lindquist et al. (2009). It is excluded from the Pachylaelapidae as defined here on the basis of its leg chaetotaxy and the presence of two dorsal shields in the deutonymph. At the moment we are unable to place Zygoseius in any known family.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FF86FF8DFF236FD89970F948.taxon	discussion	Notes. This species may belong in Neoparasitus Oudemans, 1901, based on examination of the type specimen (by PM). If this is true, it would make Brachylaelaps Berlese, 1910 a synonym of Neoparasitus Oudemans, 1901.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FF86FF8DFF236ABA9989FDD8.taxon	discussion	Notes. Different authors have classified Pseudolaelaps in different places within the cohort Gamasina. In their classification of the family Dermanyssidae, Evans & Till (1966) erected a separate subfamily Pseudolaelapinae for this genus, to accommodate a species with marked hypotrichy of the idiosoma and appendages, and a trispinate epistome. Bregetova (1977 b) considered the genus as a member of the Laelapidae. Karg (1993) later moved the genus to the Eviphidoidea and elevated it to family rank. In Lindquist et al. (2009), Pseudolaelaps is included provisionally as the sole representative of the pachylaelapid subfamily Pseudolaelapinae, despite the fact that the female displays characteristics not typical of the Pachylaelapidae. We provisionally agree with Karg (1993), in recognising the family Pseudolaelapidae, but further study is needed to determine its correct systematic placement.	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
52084A62FF87FF8CFF236AB5988DFE48.taxon	discussion	Notes. At the head of his account of this species (Oudemans, 1936, page 363), the genus name was spelled Dinocalaeno. This appears to be a lapsus, since the same name is spelled Dinocelaeno on pages xv and 2922 of the same work. We place this species in the family Celaenopsidae following Oudemans (1936).	en	MAŠÁN, PETER, HALLIDAY, BRUCE (2014): Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776 (1): 1-66, DOI: http://dx.doi.org/10.11646/zootaxa.3776.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3776.1.1
