identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03858793FFD0FFA3DA58FA7D7D68F85B.text	03858793FFD0FFA3DA58FA7D7D68F85B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Handaoia Seyrig 1952	<div><p>Genus Handaoia Seyrig, 1952</p> <p>Handaoia Seyrig, 1952: 26</p> <p>(type species: Handaoia spinosa Seyrig, 1952, by original designation).</p> <p>Diagnosis</p> <p>Handaoia can be recognized and separated from other Phygadeuontinae genera by the combination of the distally expanded and ventrally flattened antennal flagellum, complete posterior transverse carina of the mesosternum, isolated ʻpit’ (episternal scrobe) in the mesopleuron, and a single bulla in fore wing vein 2 m-cu. The following more complete diagnosis is modified from Townes (1970), who included Handaoia in a key to the genera of ‘Chiroticina’. Townes’ definition applies equally well to the Afrotropical species originally included as well as to the Neotropical and Palaearctic species: flagellum of female antenna fusiform (expanded beyond the middle and flattened ventrally) with a median white ring; genal carina reaching base of mandible; mandibular teeth subequal; maxillary palp long, reaching beyond centre of mesosternum; pronotum with two dorsal pits separated by weak longitudinal carina, usually flanked laterally by 3–4 longitudinal carinae; epomia absent; median lobe of mesoscutum without median longitudinal groove; scutellum without lateral carinae; mesopleural impression (episternal scrobe) ventral to speculum as an isolated pit, some distance from mesopleural furrow; posterior transverse carina of mesosternum complete; fore wing with areolet pentagonal, its outer side open (vein 3 rs-m missing), vein 2 m-cu with one wide bulla; area superomedia always separated from area petiolaris and usually confluent with area basalis; laterotergites of metasomal tergites 2 and 3 not separated or only weakly separated from tergite. Small species, body length 2–5 mm.</p> <p>Key to the New World species of Handaoia</p> <p>1. Apophysis of propodeum high, parallel sided or subtriangular, with blunt tip (Figs 2F, 4G, 7H, arrows).............................................................................................................................................. 2</p> <p>– Apophysis of propodeum as a low transverse rounded crest, or slightly pointed (Figs 3G, 5I, 8I, arrows).............................................................................................................................................. 4</p> <p>2. Area externa of propodeum confluent with area dentipara (Fig. 1A). Juxtacoxal carina absent (Fig. 2C). First and second flagellomeres about 5.6 and 6.5 × as long as wide respectively. Hind coxa tinged with white (Fig. 2A).............................................................. H. cuscoensis Bordera sp. nov.</p> <p>– Area externa of propodeum separated from area dentipara by strong carina (Fig. 1C, E). Juxtacoxal carina present, usually strong (Figs 4F, 7G). First and second flagellomeres about 4–5 and 3.2–5.4 × as long as wide, respectively. Hind coxa entirely dark brown (Figs 4A, 7A).................................. 3</p> <p>3. Fore wing with well-defined transverse dark brown band (Fig. 4A). Inner side of area externa 4.7 × as long as inner side of area dentipara (Fig. 1C). Juxtacoxal carina strong, irregular, joining submetapleural carina at its mid-length (Fig. 4F). Tergite I about 1.8× as long as its maximum width. First and second flagellomeres about 5 and 5.4× as long as wide respectively. Flagellum dark brown (except white band), with first and second flagellomeres light brown (Fig. 4A– B).................................................................................................... H. mercedensis Bordera sp. nov.</p> <p>– Fore wing evenly infuscate (Fig. 7A). Inner side of area externa 7.0× as long as inner side of area dentipara (Fig. 1E). Juxtacoxal carina weak but complete, joining anterior part of submetapleural carina (Fig. 7G). Tergite I about 2.4× as long as its maximum width. First and second flagellomeres 4.1 and 3.3× as long as wide respectively. Flagellum dark brown (except white band), with only first flagellomere light brown (Fig. 7A)................................................ H. ruizcancinoi Bordera sp. nov.</p> <p>4. Area externa of propodeum confluent with area dentipara (Fig. 1D). Juxtacoxal carina absent (Fig. 5H). Sternaulus very short and weak, reaching at most 0.3× the length of the mesopleuron (Fig. 5E). Head coarsely granulate (Fig. 5B–C)............................................. H. plaumanni sp. nov.</p> <p>– Area externa of propodeum separated from area dentipara by a strong carina (Fig. 1B, F). Juxtacoxal carina present (Figs 3F, 8H, arrows). Sternaulus deep, reaching at least 0.5 × the length of the mesopleuron (Figs 3D, 8G). Head finely granulate.......................................................................... 5</p> <p>5. Inner side of area externa 3.0–3.4 × as long as inner side of area dentipara (Fig. 1B); inner side of area dentipara (section of the lateromedian longitudinal carina) 0.4–0.5 × the length of the outer side (section of the lateral longitudinal carina) (Fig. 1B). Lateromedian longitudinal carina of first metasomal tergite weak, reaching at most 0.6× the length of the tergite, or sometimes inconspicuous (Fig. 1B). Malar space 1.15–1.25× as long as basal mandibular width (Fig. 3E). Tergite II smooth and shiny (Fig. 3B). Clypeus 2.1–2.2 × as wide as high (Fig. 3E)........................................................................................................................................................................................ H. fritzi sp. nov.</p> <p>– Inner side of area externa about 8.0× as long as inner side of area dentipara (Fig. 1F); inner side of area dentipara 0.25 × the length of the outer side (Fig. 1F). Lateromedian longitudinal carina of tergite I strong, reaching 0.8 × the length of the tergite (Fig. 1F). Malar space 1.4 × as long as mandible base (Fig. 8B). Tergite II finely granulate (Fig. 8D). Clypeus 1.6× as wide as high (Fig. 8B)................................................................................................................. H. urceus sp. nov.</p></div> 	http://treatment.plazi.org/id/03858793FFD0FFA3DA58FA7D7D68F85B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bordera, Santiago;Broad, Gavin R.	Bordera, Santiago, Broad, Gavin R. (2021): Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World. European Journal of Taxonomy 757: 80-101, DOI: https://doi.org/10.5852/ejt.2021.757.1415, URL: http://dx.doi.org/10.5852/ejt.2021.757.1415
03858793FFD2FFA4DA08FEB67926FE75.text	03858793FFD2FFA4DA08FEB67926FE75.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Handaoia cuscoensis Bordera 2021	<div><p>Handaoia cuscoensis Bordera sp. nov.</p> <p>urn:lsid:zoobank.org:act: 49A19681-3B7F-4E6F-B95B-4E6847D1C5AF</p> <p>Figs 1A, 2, 9B</p> <p>Diagnosis</p> <p>Handaoia cuscoensis Bordera sp. nov. can be distinguished from all other New World species by the combination of the following characters: apophysis of propodeum high, subtriangular with blunt tip (Fig. 2F, arrow); juxtacoxal carina absent (Fig. 2C); area externa of propodeum confluent with area dentipara (Fig. 1A); hind coxa tinged with white (Fig. 2A).</p> <p>Etymology</p> <p>The name of the species refers to the Department of Cusco (Peru), the type locality region.</p> <p>Material examined</p> <p>Only known from the holotype female.</p> <p>Holotype PERU • ♀; Quincemil, 750 m nr. Marcapata; 10–15 Nov. 1962; Luis Peña leg.; AEIC.</p> <p>Description</p> <p>Female</p> <p>MEASUREMENTS. Body length (without ovipositor) 3.8 mm. Fore wing 2.9 mm long.</p> <p>HEAD. Very finely and densely granulate, matt, distinctly narrowed behind eyes, occiput very concave centrally in dorsal view (Fig. 2E); gena about 0.15 × length of eye in dorsal view; posterior ocellus separated from eye by 1.43 × its own maximum diameter; distance between posterior ocelli 1.14 × maximum ocellar diameter (Fig. 2E); occipital carina gently V-shaped medially; malar space 1.33 × basal width of mandible; face moderately swollen medially (Fig. 2B); clypeus very weakly convex, almost flat, finely and slightly granulate with scattered punctures, its apex weakly rounded, about 2.3× as wide as high; mandible slightly tapered to apex, ventral tooth of mandible a little shorter than upper tooth; maxillary palp reaching at most ¾ length of mesosternum; flagellum widened in middle, tapered towards apex, flagellomeres 8–19 flattened ventrally (Fig. 1A); flagellum with 20 segments, lengththickness ratios: 1 st segment = 5.64, 2 nd = 6.55 and 10 th = 1.08.</p> <p>MESOSOMA. Very finely and densely granulate, matt (Fig. 2D); mesoscutum conspicuously convex, notauli absent (Fig. 2E); scutellum moderately convex, without lateral carinae; sternaulus very short and weak, as a wide V-shaped depression anteriorly, reaching at most 0.3× length of mesopleuron (Fig. 2D); juxtacoxal carina barely indicated anteriorly (Fig. 2C); propodeum (Fig. 1A) with mostly weak carinae; area superomedia confluent with area basalis and separated from area petiolaris; area externa and area dentipara confluent; area spiracularis confluent with area lateralis; area dentipara section of lateral longitudinal carina very weak, partly absent; posterior transverse carina strong, forming elevate apophysis joining lateral longitudinal carina, apophysis subtriangular, with blunt tip (Fig. 2D, F, arrow). Length of hind femur about 4.4 × its height. Hind wing with vein cu-a 0.73 as long as abscissa of CU between M and cu-a.</p> <p>METASOMA. Tergite I (Figs 1A, 2G) finely granulate and very weakly strigose laterally, matt, 2.5× as long as its maximum width; lateromedian longitudinal carina absent; lateral longitudinal carina complete; tergite II (Fig. 2G) shallowly granulate, shiny, glabrous; remaining tergites (Fig. 2G) smooth and shiny with inconspicuous dense setiferous punctures. Ovipositor sheath about 0.4× as long as hind tibia.</p> <p>COLOUR. Head (Fig. 2A–B, D–E) light orange; antenna with scape and pedicel light brown, flagellum dark brown, distal part of flagellomere 4 to flagellomere 7 dorsally white (Fig. 2A). Mesosoma (Fig. 2D– E) light orange. Metasoma (Fig. 2G) mostly brown, tergites I–II and anterior part of IV light orange. Legs (Fig. 2A) mostly light orange, distal tarsomeres, dark brown, fore and mid coxae, trochanters and trochantelli white, hind coxa white proximally, dark brown distally. Wings (Fig. 2A) slightly evenly infuscate.</p> <p>Male</p> <p>Unknown.</p></div> 	http://treatment.plazi.org/id/03858793FFD2FFA4DA08FEB67926FE75	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bordera, Santiago;Broad, Gavin R.	Bordera, Santiago, Broad, Gavin R. (2021): Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World. European Journal of Taxonomy 757: 80-101, DOI: https://doi.org/10.5852/ejt.2021.757.1415, URL: http://dx.doi.org/10.5852/ejt.2021.757.1415
03858793FFD7FFA9DAB7FEB67926FB9E.text	03858793FFD7FFA9DAB7FEB67926FB9E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Handaoia fritzi Bordera & Broad 2021	<div><p>Handaoia fritzi sp. nov.</p> <p>urn:lsid:zoobank.org:act: 3FE16228-86F1-4A8C-8FA0-227999E65A4F</p> <p>Figs 1B, 3, 9B</p> <p>Diagnosis</p> <p>Handaoia fritzi sp. nov. can be distinguished from all other New World species by the combination of the following characters: apophysis of propodeum as a low transverse rounded or slightly pointed triangular crest (Fig. 3G, arrow); juxtacoxal carina strong, irregular, joining submetapleural carina at anterior 0.3–0.4 (Fig. 3F, arrow); area externa of propodeum separated from area dentipara by strong carina (Fig. 1B); inner side of area externa 3.0–3.4 × as long as inner side of area dentipara (Fig. 1B); inner side of area dentipara 0.46–0.5 × the length of outer side (Fig. 1B); tergite II smooth and shiny (Fig. 3B).</p> <p>Etymology</p> <p>This species is named in honour of Dr Fritz Plaumann, illustrious botanist and entomologist based in Brazil, who collected the type material. A noun in genitive case.</p> <p>Material examined</p> <p>Known from five females.</p> <p>Holotype BRAZIL • ♀; <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-52.383335&amp;materialsCitation.latitude=-27.183332" title="Search Plazi for locations around (long -52.383335/lat -27.183332)">Nova Teutonia</a>; 27º11′ S, 52º23′ W; 26 Aug. 1937; F. Plaumann leg.; B.M. 1937-748; B.M. Type Hym 3b.2871; NHMUK010636381.</p> <p>Paratypes BRAZIL • 1 ♀; same collection data as for holotype; Aug.1935;B.M. 1937-47; NHMUK010636388 • 1♀; same collection data as for preceding; 28 May 1938; B.M. 1938-458; NHMUK010636387 • 1 ♀; same collection data as for preceding; 11 Aug. 1938; B.M. 1938-682; NHMUK010636396 • 1 ♀; same collection data as for preceding; 16 Sep. 1938; B.M. 1938-682; NHMUK010636383.</p> <p>Description</p> <p>Female</p> <p>MEASUREMENTS. Body length (without ovipositor) 3.3–4.1 mm. Fore wing 2.7–3.3 mm long.</p> <p>HEAD. Finely granulate, matt, distinctly narrowed behind eyes, occiput very concave centrally in dorsal view (Fig. 3C); gena 0.2–0.25 × length of eye in dorsal view; posterior ocellus separated from eye by 0.75–1.25 × its own maximum diameter; distance between posterior ocelli 0.75–1.0 × maximum ocellar diameter (Fig. 3C); occipital carina gently V-shaped medially (Fig. 3B); malar space 1.15–1.25 × basal width of mandible; face slightly swollen medially; clypeus (Fig. 3E) weakly convex, faintly granulate dorsally, smooth and shiny ventrally, with scattered punctures, its apex almost straight, 2.1–2.2× as wide as high; mandible slightly tapered to apex, ventral tooth of mandible approximately same length as upper tooth; maxillary palp relatively short, reaching at most ⅔ length of mesosternum; flagellum widened in middle, tapered towards apex, flagellomeres 8–17 flattened ventrally; flagellum with 19–20 segments, length-thickness ratios: 1 st segment = 4.74–5.0, 2 nd = 5.0–5.28 and 10 th = 1.0.</p> <p>MESOSOMA. Finely granulate and matt (Fig. 3B–D); mesoscutum convex, notauli weak anteriorly, reaching about 0.3 distance to scuto-scutellar groove, sometimes inconspicuous (Fig. 3C); scutellum moderately convex, without lateral carinae; sternaulus moderately deep, reaching at least middle of mesopleuron (Fig. 3D); juxtacoxal carina strong, irregular, joining submetapleural carina at anterior 0.3–0.4 (Fig. 3F, arrow); propodeum (Fig. 1B) with strong and conspicuous carinae; area superomedia confluent with area basalis and separated from area petiolaris; inner side of area externa about 3.0– 4.0× as long as inner side of area dentipara; inner side of area dentipara 0.46–0.5× length of outer side; area spiracularis confluent with area lateralis; posterior transverse carina strong, forming a low and wide rounded or slightly subtriangular crest joining lateral longitudinal carina (Fig. 3G, arrow). Length of hind femur 4.0–4.7 × its height. Hind wing with vein cu-a 0.4–0.6 × as long as abscissa of CU between M and cu-a.</p> <p>METASOMA. Tergite I (Figs 1B, 3B), finely granulate, matt, 1.8–2.0 × as long as its maximum width; postpetiole laterally and posteriorly tending to fine strigosity; lateromedian longitudinal carina weak, reaching at least 0.6 × length of tergite; lateral longitudinal carina weak, complete or absent posterior to spiracles; tergite II (Fig. 3B) smooth and shiny, glabrous; remaining tergites (Fig. 3B) smooth and shiny with inconspicuous dense setiferous punctures. Ovipositor sheath 0.45–0.49 × as long as hind tibia.</p> <p>COLOUR. Head (Fig. 3A–E) dark brown, clypeus and mandibles reddish brown; palpi light brown; antenna with scape, pedicel and flagellomeres 1–4 light brown, darker dorsally, remaining flagellomeres dark brown, flagellomeres 4–7 dorsally white (Fig. 3A). Mesosoma (Fig. 3A–D) mostly dark orange; mesoscutum with wide dark brown longitudinal bands on central and lateral lobes; dorsal anterior part of propodeum dark brown. Metasoma (Fig. 3B) mostly brown, tergite I sometimes centrally tinged dark brown. Legs (Fig. 3A) light brown, distal tarsomeres dark brown, fore and mid coxae, part of hind coxa and all trochanters and trochantelli lighter. Wings (Fig. 3A) slightly infuscate.</p> <p>Male</p> <p>Unknown.</p></div> 	http://treatment.plazi.org/id/03858793FFD7FFA9DAB7FEB67926FB9E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bordera, Santiago;Broad, Gavin R.	Bordera, Santiago, Broad, Gavin R. (2021): Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World. European Journal of Taxonomy 757: 80-101, DOI: https://doi.org/10.5852/ejt.2021.757.1415, URL: http://dx.doi.org/10.5852/ejt.2021.757.1415
03858793FFD9FFABDA17FBE67926FA5B.text	03858793FFD9FFABDA17FBE67926FA5B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Handaoia mercedensis Bordera 2021	<div><p>Handaoia mercedensis Bordera sp. nov.</p> <p>urn:lsid:zoobank.org:act: 80A6A583-DB23-43D9-AF4A-4D16A6270918</p> <p>Figs 1C, 4, 9B</p> <p>Diagnosis</p> <p>Handaoia mercedensis Bordera sp. nov. can be distinguished from all other New World species by the combination of the following characters: apophysis of propodeum conspicuously elevated, parallelsided with blunt tip (Fig. 4G, arrow); fore wing with well-defined transverse dark brown band (Fig. 4A); juxtacoxal carina strong, irregular, joining submetapleural carina at mid length (Fig. 4F, arrow); flagellum dark brown (except white band), with first and second flagellomeres light brown (Fig. 4A–B).</p> <p>Etymology</p> <p>The name refers to La Merced, municipality of Chanchamayo in the Department of Junin (Peru), where the holotype was collected.</p> <p>Material examined</p> <p>Known only from the holotype female.</p> <p>Holotype PERU • ♀; La Merced, Fundo La Génova; Trampa Malaise 3; 3–17 Jan. 2009; AECID 013484/07; CEUA.</p> <p>Description</p> <p>Female</p> <p>MEASUREMENTS. Body length (without ovipositor) 4.3 mm. Fore wing 3.4 mm long.</p> <p>HEAD. Finely granulate, matt, strongly narrowed behind eyes, occiput very concave centrally in dorsal view (Fig. 4C); gena 0.18 × length of eye in dorsal view; posterior ocellus separated from eye by 1.3× its own maximum diameter; distance between posterior ocelli 0.9 × maximum ocellar diameter (Fig. 4C); occipital carina gently V-shaped medially; malar space about 1.4× basal width of mandible; face slightly swollen medially; clypeus (Fig. 4B) quite convex, gently granulate, with scattered coarse punctures in apical half, apex slightly rounded, about 2.0× as wide as high; mandible quite tapered to apex, ventral tooth of mandible a little shorter than upper tooth; maxillary palp reaching mid coxa; flagellum widened in middle, tapered towards apex, flagellomeres 8–20 flattened ventrally (Fig. 4A); flagellum with 22 segments, length-thickness ratios: 1 st segment = 5.0, 2 nd = 5.38 and 10 th = 1.0.</p> <p>MESOSOMA. Finely granulate and matt (Fig. 4C–D);mesoscutum slightly convex, notauli absent (Fig. 4C); scutellum moderately convex, without lateral carinae; sternaulus deep, exceeding half the length of mesopleuron (Fig. 4D); juxtacoxal carina strong, irregular, joining submetapleural carina at mid length (Fig. 4F, arrow); propodeum (Fig. 1C) with strong and conspicuous carinae; area superomedia confluent with area basalis and separated from area petiolaris; inner side of area externa about 4.7× as long as inner side of area dentipara; inner side of area dentipara about 0.3× length of outer side; area spiracularis confluent with area lateralis; posterior transverse carina strong, forming an elevate apophysis joining lateral longitudinal carina, apophysis parallel-sided with blut tip (Fig. 4G, arrow). Length of hind femur about 4.0× its height. Hind wing with vein cu-a 0.5 as long as abscissa of CU between M and cu-a.</p> <p>METASOMA. Tergite I (Fig. 1C) finely granulate, slightly shiny, 1.77 × as long as its maximum width; postpetiole laterally not finely strigose; lateromedian longitudinal carina moderately strong, reaching about 0.7× length of tergite; lateral longitudinal carina complete; tergite II (Fig. 4E) shiny, very slightly granulate anteriorly, glabrous, smooth posteriorly, with moderately dense short setae; remaining tergites (Fig. 4E) smooth and shiny with inconspicuous dense setiferous punctures. Ovipositor sheath 0.53 × as long as hind tibia.</p> <p>COLOUR. Head (Fig. 4A–D) orange brown, vertical orbits and interocellar area dark brown; palpi dark brown; mandibular teeth black; antenna with scape, pedicel and flagellomeres 1–2 light brown, remaining flagellomeres dark brown, flagellomeres 4–7 and base of 8 dorsally white (Fig. 4A). Mesosoma (Fig. 4A, C–D) entirely orange-brown. Metasoma (Fig. 4E) except tergite I dark brown, tergite I light brown. Fore and mid legs (Fig. 4A) light brown, tarsi fuscous; hind leg dark brown, trochanter and base of tibia white. Wings slightly infuscate, fore wing with well-defined transverse dark brown band (Fig. 4A).</p> <p>Male</p> <p>Unknown.</p></div> 	http://treatment.plazi.org/id/03858793FFD9FFABDA17FBE67926FA5B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bordera, Santiago;Broad, Gavin R.	Bordera, Santiago, Broad, Gavin R. (2021): Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World. European Journal of Taxonomy 757: 80-101, DOI: https://doi.org/10.5852/ejt.2021.757.1415, URL: http://dx.doi.org/10.5852/ejt.2021.757.1415
03858793FFDBFFAFDA59F9A37A13FB8F.text	03858793FFDBFFAFDA59F9A37A13FB8F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Handaoia plaumanni Bordera & Broad 2021	<div><p>Handaoia plaumanni sp. nov.</p> <p>urn:lsid:zoobank.org:act: 7A2E9032-B90C-4EB7-9D7F-139AA2D3D4B9</p> <p>Figs 1D, 5–6, 9B</p> <p>Diagnosis</p> <p>Handaoia plaumanni sp.nov. can be distinguished from all other New World species by the combination of the following characters (both male and female): apophysis of propodeum as a low transverse crest (Fig. 5I, arrow); sternaulus very short and weak, reaching at most 0.3× the length of the mesopleuron (Figs 5E, 6D); juxtacoxal carina absent (Figs 5H, 6E); area externa of propodeum confluent with area dentipara (Figs 1D, 6F).</p> <p>Etymology</p> <p>This species is named in honour of Dr Fritz Plaumann, illustrious botanist and entomologist based in Brazil, who collected the type series. A noun in the genitive case.</p> <p>Material examined</p> <p>Known from three females and two males.</p> <p>Holotype BRAZIL • ♀; <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-52.383335&amp;materialsCitation.latitude=-27.183332" title="Search Plazi for locations around (long -52.383335/lat -27.183332)">Nova Teutonia</a>; 27º11′ S, 52º23′ W; 24 May 1938; F Plaumann leg.; B.M. 1938-452; B.M. Type Hym 3b.2872; NHMUK010636386.</p> <p>Paratypes BRAZIL • 1 ♀; same collection data as for holotype; 13 Apr. 1938; B.M. 1938-682; NHMUK010636382 • 1 ♂; same collection data as for preceding; 10 May 1938; B.M. 1938-682; NHMUK010636384 • 1 ♀; same collection data as for preceding; 13 May 1938; B.M. 1938-682; NHMUK010636395 • 1 ♂; same collection data as for preceding; 7 Jun. 1938; B.M. 1938-458; NHMUK010636385.</p> <p>Description</p> <p>Female</p> <p>MEASUREMENTS. Body length (without ovipositor) 2.6–2.9 mm. Fore wing 2.3–2.4 mm long.</p> <p>HEAD. Moderately coarsely granulate, matt, distinctly narrowed behind eyes, occiput very concave centrally in dorsal view (Fig. 5B–E); gena 0.16–0.25× length of eye in dorsal view; posterior ocellus separated from eye by 0.9–1.0× its maximum diameter; distance between posterior ocelli 0.75–1.3 × maximum ocellar diameter (Fig. 5D); occipital carina gently V-shaped medially (Fig. 5D); malar space 1.17–1.33 × basal width of mandible; face weakly swollen medially (Fig. 5C); clypeus (Fig. 5C) weakly convex, smooth and shiny with scattered punctures, apex almost straight, about 2.0× as wide as high; mandible slightly tapered to apex, ventral tooth approximately same length as upper tooth; maxillary palp relatively short, reaching at most ⅔ length of mesosternum; flagellum widened in middle, tapered towards apex, flagellomeres 8–17 flattened ventrally (Fig. 5A); flagellum with 18 segments, lengththickness ratios: 1 st segment = 4.67–5.0, 2 nd = 5.0–5.33 and 10 th = 1.0.</p> <p>MESOSOMA. Moderately coarsely granulate, matt (Fig. 5D–E); mesoscutum slightly convex, flat posteriorly, notauli very weak anteriorly or absent (Fig. 5B, D), when present reaching at most 0.3 distance to scuto-scutellar groove; scutellum moderately convex, without lateral carinae; sternaulus very short and weak, as a wide V-shaped depression anteriorly, reaching at most 0.3 × the length of mesopleuron (Fig. 5E); juxtacoxal carina absent (Fig. 5H); propodeum (Figs 1D, 5I) with moderately strong carinae; area superomedia confluent with area basalis and separated from area petiolaris; area externa and area dentipara confluent; area spiracularis confluent with area lateralis; posterior transverse carina forming a very low transverse crest joining lateral longitudinal carina (Fig. 5I, arrow). Length of hind femur 4.0–4.26 × its height. Hind wing with vein cu-a 0.4– 0.6× as long as abscissa of CU between M and cu-a.</p> <p>METASOMA. Tergite I (Figs 1D, 5J) finely granulate, matt, 1.6–1.8 × as long as its maximum width; lateromedian longitudinal carina absent; lateral longitudinal carina weak, absent posterior to spiracles; tergite II (Fig. 5F–G) smooth and shiny, glabrous; remaining tergites (Fig. 5F–G) smooth and shiny with inconspicuous dense setiferous punctures. Ovipositor sheath about 0.5–0.6 × as long as hind tibia.</p> <p>COLOUR. Head (Fig. 5B–E) dark brown; antenna with scape and pedicel light brown and flagellomeres 1–4 light brown sometimes infuscate, remaining flagellomeres dark brown, flagellomeres 4–7 dorsally white (Fig. 5A). Mesosoma (Fig. 5B, D–E) mostly orange-brown; mesoscutum tending to entirely dark brown, sometimes anterior part of propodeum infuscate. Metasoma (Fig. 5F–G) brown, tergites I–II and sometimes anterior part of III light brown, remaining tergites dark brown. Legs (Fig. 5A) light brown, distal tarsomeres dark brown, sometimes fore and mid coxae, part of hind coxa and all trochanters and trochantelli lighter. Wings (Fig. 5A) slightly infuscate.</p> <p>Male</p> <p>MEASUREMENTS. Body length 2.3–2.9 mm. Fore wing 2.0– 2.6 mm long.</p> <p>HEAD. Less narrowed behind eyes and less concave centrally in dorsal view than female (Fig. 6C); gena 0.31–0.42 × length of eye in dorsal view; posterior ocellus separated from eye by 1.25–1.33 × its maximum diameter; distance between posterior ocelli 0.9–1.3 × maximum ocellar diameter (Fig. 6C); malar space 1.33 × basal width of mandible; clypeus about 1.7 × as wide as high; flagellum filiform (Fig. 6A–B) with 19 segments, length-thickness ratios: 1 st = segment 5.0, 2 nd = 5.0 and 10 th = 3.4–4.0</p> <p>MESOSOMA. Notauli absent (Fig. 6C–D); length of hind femur about 4.5–4.6× its heigth.</p> <p>METASOMA. Tergite I finely granulate, matt, 2.1× as long as its maximum width.</p> <p>COLOUR.As in female (Fig. 6A–F) but antenna entirely dark brown.In one paratype (NHMUK010636385), mesosoma and metasoma mostly dark brown (Fig. 6A, D).</p> <p>Otherwise similar to female.</p></div> 	http://treatment.plazi.org/id/03858793FFDBFFAFDA59F9A37A13FB8F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bordera, Santiago;Broad, Gavin R.	Bordera, Santiago, Broad, Gavin R. (2021): Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World. European Journal of Taxonomy 757: 80-101, DOI: https://doi.org/10.5852/ejt.2021.757.1415, URL: http://dx.doi.org/10.5852/ejt.2021.757.1415
03858793FFDFFFB1DA15FBF77926FA5B.text	03858793FFDFFFB1DA15FBF77926FA5B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Handaoia ruizcancinoi Bordera 2021	<div><p>Handaoia ruizcancinoi Bordera sp. nov.</p> <p>urn:lsid:zoobank.org:act: 151C284F-8024-4A2A-97FD-D0BBDE6AF048</p> <p>Figs 1E, 7, 9A</p> <p>Diagnosis</p> <p>Handaoia ruizcancinoi Bordera sp. nov. can be distinguished from all other New World species by the combination of the following characters: apophysis of propodeum conspicuously high, subtriangular with blunt tip (Fig. 7H, arrow); juxtacoxal carina weak but complete, joining submetapleural carina at its anterior part (Fig. 7G, arrow); inner side of area externa about 7.0× as long as inner side of area dentipara (Fig. 1E).</p> <p>Etymology</p> <p>The species is named in honour of Dr Enrique Ruiz Cancino from Universidad Autónoma de Tamaulipas (Mexico), in recognition of his contribution to the study of the Ichneumonidae of Mexico. A noun in the genitive case.</p> <p>Material examined</p> <p>Known only from the holotype female.</p> <p>Holotype MEXICO • ♀; Tamaulipas, Ocampo, J. de Manantiales; T. Amarillas; 13 May 1995; J. Coronado, S. Nino and C. Hernández leg.; CEUA.</p> <p>Description</p> <p>Female</p> <p>MEASUREMENTS. Body length (without ovipositor) 4.0 mm. Fore wing about 3.0 mm long.</p> <p>HEAD. Finely granulate, matt, strongly narrowed behind eyes, occiput very concave centrally in dorsal view (Fig. 7B–D); gena 0.14 × length of eye in dorsal view; posterior ocellus separated from eye by 1.55 × its maximum diameter; distance between posterior ocelli 1.27 × maximum ocellar diameter; occipital carina gently V-shaped medially (Fig. 7D); malar space 1.3× basal width of mandible; face conspicuously swollen medially (Fig. 7B); clypeus (Fig. 7B) quite convex, gently granulate with scattered coarse punctures, apex slightly rounded, about 1.8 × as wide as high; mandible quite tapered to apex, ventral tooth of mandible a little shorter than upper tooth; flagellum widened in middle, tapered towards apex, flagellomeres 8–18 flattened ventrally; flagellum with 20 segments, length-thickness ratios: 1 st segment = 4.1, 2 nd = 3.27 and 10 th = 0.72.</p> <p>MESOSOMA. Finely granulate and matt (Fig. 7A, C–D); mesoscutum slightly convex, almost flat posteriorly; notauli moderately deep over 0.7× length of mesoscutum (Fig. 7D); scutellum moderately convex, without lateral carinae; sternaulus as a wide V-shaped depression anteriorly, reaching about 0.3× the length of mesopleuron (Fig. 7C); juxtacoxal carina weak but complete, joining submetapleural carina at its anterior part (Fig. 7G, arrow); propodeum (Figs 1E, 7C, H) with strong and conspicuous carinae; area superomedia confluent with area basalis and separated from area petiolaris; inner side of area externa about 7.0× as long as inner side of area dentipara; inner side of area dentipara about 0.3× length of outer side; area spiracularis confluent with area lateralis; posterior transverse carina strong, forming an elevated apophysis joining lateral longitudinal carina, apophysis subtriangular with blunt tip (Fig. 7C, H, arrow). Length of hind femur 3.9× its height. Hind wing with vein cu-a 0.77 × as long as abscissa of CU between M and cu-a.</p> <p>METASOMA. Tergite I (Fig. 7E) finely granulate, shiny, strigose laterally, about 2.4× as long as its maximum width; lateromedian longitudinal carina moderately strong, reaching posterior 0.8× length of tergite; lateral longitudinal carina present from spiracle to posterior margin of postpetiole; tergite II (Fig. 7F) shiny, smooth and almost glabrous; remaining tergites (Fig. 7F) smooth and shiny, with short and moderately dense setiferous punctures. Ovipositor sheath about 0.5× as long as hind tibia.</p> <p>COLOUR. Head (Fig. 7B–D) brown, darker dorsally, lighter ventrally, mandibular teeth and labrum dark brown; antenna with scape, pedicel and first flagellomere light brown, remaining flagellomeres dark brown, flagellomeres 5–7 dorsally white (Fig. 7A). Mesosoma (Fig. 7A, C) brown, more infuscate ventrally. Metasoma (Fig. 7E–F) dark brown. Legs (Fig. 7A) mostly dark brown; fore and mid coxae and trochanters, and hind trochanter, white. Wings (Fig. 7A) evenly infuscate.</p> <p>Male</p> <p>Unknown.</p></div> 	http://treatment.plazi.org/id/03858793FFDFFFB1DA15FBF77926FA5B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bordera, Santiago;Broad, Gavin R.	Bordera, Santiago, Broad, Gavin R. (2021): Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World. European Journal of Taxonomy 757: 80-101, DOI: https://doi.org/10.5852/ejt.2021.757.1415, URL: http://dx.doi.org/10.5852/ejt.2021.757.1415
03858793FFC1FFB2DA43F9A37926FEC2.text	03858793FFC1FFB2DA43F9A37926FEC2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Handaoia urceus Bordera & Broad 2021	<div><p>Handaoia urceus sp. nov.</p> <p>urn:lsid:zoobank.org:act: 5413C7C8-D785-4583-803F-0C1E75B7EA9E</p> <p>Figs 1F, 8, 9B</p> <p>Diagnosis</p> <p>Handaoia urceus sp. nov. can be distinguished from all other New World species by the combination of the following characters: apophysis of propodeum as a low transverse rounded crest (Fig. 8I, arrow); tergite II finely granulate (Fig. 8D); juxtacoxal carina strong anteriorly, weak posteriorly, joining submetapleural carina at its half length (Fig. 8H, arrow); area externa of propodeum separated from area dentipara by strong carina (Fig. 1F); inner side of area externa about 8.0× as long as inner side of area dentipara (Fig. 1F).</p> <p>Etymology</p> <p>From the Latin for ‘pitcher’, named after the flask- or pitcher-shaped combined area basalis + superomedia. A noun in apposition.</p> <p>Material examined</p> <p>Known only from the holotype female.</p> <p>Holotype BRAZIL • ♀; <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-52.383335&amp;materialsCitation.latitude=-27.183332" title="Search Plazi for locations around (long -52.383335/lat -27.183332)">Nova Teutonia</a>; 27º11′ S, 52º23′ W; 8 Feb. 1939; F. Plaumann leg.; B.M. 1939-181; B.M. Type Hym 3b.2873; NHMUK010636380.</p> <p>Description</p> <p>Female</p> <p>MEASUREMENTS. Body length (without ovipositor) 4.3 mm. Fore wing 3.7 mm long.</p> <p>HEAD. Finely granulate, matt, distinctly narrowed behind eyes, occiput very concave centrally in dorsal view (Fig. 8B–C); gena 0.22 × length of eye in dorsal view; posterior ocellus separated from eye by 1.0 × its maximum diameter; distance between posterior ocelli 1.0× maximum ocellar diameter (Fig. 8C); occipital carina gently V-shaped medially (Fig. 8E); malar space 1.44 × basal width of mandible; face moderately swollen medially (Fig. 8B); clypeus (Fig. 8B) weakly convex, gently granulate with scattered punctures, apex slightly rounded, about 1.6× as wide as high; mandible slightly tapered to apex, ventral tooth approximately same length as upper tooth; maxillary palp long, almost reaching middle coxa; flagellum widened in middle, tapered towards apex (Fig. 8A), flagellomeres 8–18 flattened ventrally; flagellum with 22 segments, length-thickness ratios: 1 st segment = 4.9, 2 nd = 6.1 and 10 th = 1.13.</p> <p>MESOSOMA. Finely granulate and matt (Fig. 8E, G); mesoscutum (Fig. 8E, G) convex, notauli moderately deep anteriorly, reaching about 0.3 × distance to scuto-scutellar groove; scutellum moderately convex, without lateral carinae; sternaulus moderately deep, reaching about mid-point of mesopleuron (Fig. 8G); juxtacoxal carina strong anteriorly, weak posteriorly, joining submetapleural carina at mid length (Fig. 8H, arrow); propodeum (Figs 1F, 8I) with strong and conspicuous carinae; area superomedia confluent with area basalis and separated from area petiolaris; inner side of area externa about 8.0 × as long as inner side of area dentipara; inner side of area dentipara 0.25 × length of outer side; area spiracularis confluent with area lateralis; posterior transverse carina strongly elevated, forming a low transverse crest joining lateral longitudinal carina (Fig. 8I, arrow). Length of hind femur 4.2 × its height. Hind wing with vein cu-a about 1.0 × as long as abscissa of CU between M and cu-a.</p> <p>METASOMA. Tergite I (Figs 1F, 8D) finely granulate, matt, about 1.70 × as long as its maximum width; postpetiole laterally and posteriorly tending to fine strigosity; lateromedian and lateral longitudinal carinae strong, reaching at least 0.8× length of tergite; tergite II (Fig. 8D, F) finely granulate, shiny; remaining tergites (Fig. 8D, F) smooth and shiny with inconspicuous setiferous punctures. Ovipositor sheath about 0.54× as long as hind tibia.</p> <p>COLOUR. Head (Fig. 8B–C) dark brown tending to be more reddish on clypeus and mandibles; palpi light brown; antenna with scape, pedicel and flagellomeres I–II (III) light brown, remaining flagellomeres dark brown, distal part of flagellomere 4 to flagellomere 7 dorsally white (Fig. 8A). Mesosoma (Fig. 8E– G) entirely orange. Metasoma (Fig. 8D–F) with a gradient from orange at anterior of tergite I to dark brown on posterior tergites, posterior rim of tergites I–III yellowish. Legs (Fig. 8A) light brown, distal tarsomeres dark brown, fore and mid coxae, part of hind coxa and all trochanters and trochantelli lighter. Wings (Fig. 8A) slightly infuscate.</p> <p>Male</p> <p>Unknown.</p></div> 	http://treatment.plazi.org/id/03858793FFC1FFB2DA43F9A37926FEC2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bordera, Santiago;Broad, Gavin R.	Bordera, Santiago, Broad, Gavin R. (2021): Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World. European Journal of Taxonomy 757: 80-101, DOI: https://doi.org/10.5852/ejt.2021.757.1415, URL: http://dx.doi.org/10.5852/ejt.2021.757.1415
