taxonID	type	description	language	source
03858793FFD0FFA3DA58FA7D7D68F85B.taxon	type_taxon	(type species: Handaoia spinosa Seyrig, 1952, by original designation).	en	Bordera, Santiago, Broad, Gavin R. (2021): Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World. European Journal of Taxonomy 757: 80-101, DOI: https://doi.org/10.5852/ejt.2021.757.1415, URL: http://dx.doi.org/10.5852/ejt.2021.757.1415
03858793FFD0FFA3DA58FA7D7D68F85B.taxon	diagnosis	Diagnosis Handaoia can be recognized and separated from other Phygadeuontinae genera by the combination of the distally expanded and ventrally flattened antennal flagellum, complete posterior transverse carina of the mesosternum, isolated ʻpit’ (episternal scrobe) in the mesopleuron, and a single bulla in fore wing vein 2 m-cu. The following more complete diagnosis is modified from Townes (1970), who included Handaoia in a key to the genera of ‘ Chiroticina’. Townes’ definition applies equally well to the Afrotropical species originally included as well as to the Neotropical and Palaearctic species: flagellum of female antenna fusiform (expanded beyond the middle and flattened ventrally) with a median white ring; genal carina reaching base of mandible; mandibular teeth subequal; maxillary palp long, reaching beyond centre of mesosternum; pronotum with two dorsal pits separated by weak longitudinal carina, usually flanked laterally by 3 – 4 longitudinal carinae; epomia absent; median lobe of mesoscutum without median longitudinal groove; scutellum without lateral carinae; mesopleural impression (episternal scrobe) ventral to speculum as an isolated pit, some distance from mesopleural furrow; posterior transverse carina of mesosternum complete; fore wing with areolet pentagonal, its outer side open (vein 3 rs-m missing), vein 2 m-cu with one wide bulla; area superomedia always separated from area petiolaris and usually confluent with area basalis; laterotergites of metasomal tergites 2 and 3 not separated or only weakly separated from tergite. Small species, body length 2 – 5 mm.	en	Bordera, Santiago, Broad, Gavin R. (2021): Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World. European Journal of Taxonomy 757: 80-101, DOI: https://doi.org/10.5852/ejt.2021.757.1415, URL: http://dx.doi.org/10.5852/ejt.2021.757.1415
03858793FFD0FFA3DA58FA7D7D68F85B.taxon	discussion	Key to the New World species of Handaoia	en	Bordera, Santiago, Broad, Gavin R. (2021): Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World. European Journal of Taxonomy 757: 80-101, DOI: https://doi.org/10.5852/ejt.2021.757.1415, URL: http://dx.doi.org/10.5852/ejt.2021.757.1415
03858793FFD2FFA4DA08FEB67926FE75.taxon	description	urn: lsid: zoobank. org: act: 49 A 19681 - 3 B 7 F- 4 E 6 F-B 95 B- 4 E 6847 D 1 C 5 AF Figs 1 A, 2, 9 B	en	Bordera, Santiago, Broad, Gavin R. (2021): Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World. European Journal of Taxonomy 757: 80-101, DOI: https://doi.org/10.5852/ejt.2021.757.1415, URL: http://dx.doi.org/10.5852/ejt.2021.757.1415
03858793FFD2FFA4DA08FEB67926FE75.taxon	diagnosis	Diagnosis Handaoia cuscoensis Bordera sp. nov. can be distinguished from all other New World species by the combination of the following characters: apophysis of propodeum high, subtriangular with blunt tip (Fig. 2 F, arrow); juxtacoxal carina absent (Fig. 2 C); area externa of propodeum confluent with area dentipara (Fig. 1 A); hind coxa tinged with white (Fig. 2 A).	en	Bordera, Santiago, Broad, Gavin R. (2021): Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World. European Journal of Taxonomy 757: 80-101, DOI: https://doi.org/10.5852/ejt.2021.757.1415, URL: http://dx.doi.org/10.5852/ejt.2021.757.1415
03858793FFD2FFA4DA08FEB67926FE75.taxon	etymology	Etymology The name of the species refers to the Department of Cusco (Peru), the type locality region.	en	Bordera, Santiago, Broad, Gavin R. (2021): Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World. European Journal of Taxonomy 757: 80-101, DOI: https://doi.org/10.5852/ejt.2021.757.1415, URL: http://dx.doi.org/10.5852/ejt.2021.757.1415
03858793FFD2FFA4DA08FEB67926FE75.taxon	materials_examined	Material examined Only known from the holotype female. Holotype PERU • ♀; Quincemil, 750 m nr. Marcapata; 10 – 15 Nov. 1962; Luis Peña leg.; AEIC.	en	Bordera, Santiago, Broad, Gavin R. (2021): Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World. European Journal of Taxonomy 757: 80-101, DOI: https://doi.org/10.5852/ejt.2021.757.1415, URL: http://dx.doi.org/10.5852/ejt.2021.757.1415
03858793FFD2FFA4DA08FEB67926FE75.taxon	description	Description Female MEASUREMENTS. Body length (without ovipositor) 3.8 mm. Fore wing 2.9 mm long. HEAD. Very finely and densely granulate, matt, distinctly narrowed behind eyes, occiput very concave centrally in dorsal view (Fig. 2 E); gena about 0.15 × length of eye in dorsal view; posterior ocellus separated from eye by 1.43 × its own maximum diameter; distance between posterior ocelli 1.14 × maximum ocellar diameter (Fig. 2 E); occipital carina gently V-shaped medially; malar space 1.33 × basal width of mandible; face moderately swollen medially (Fig. 2 B); clypeus very weakly convex, almost flat, finely and slightly granulate with scattered punctures, its apex weakly rounded, about 2.3 × as wide as high; mandible slightly tapered to apex, ventral tooth of mandible a little shorter than upper tooth; maxillary palp reaching at most ¾ length of mesosternum; flagellum widened in middle, tapered towards apex, flagellomeres 8 – 19 flattened ventrally (Fig. 1 A); flagellum with 20 segments, lengththickness ratios: 1 st segment = 5.64, 2 nd = 6.55 and 10 th = 1.08. MESOSOMA. Very finely and densely granulate, matt (Fig. 2 D); mesoscutum conspicuously convex, notauli absent (Fig. 2 E); scutellum moderately convex, without lateral carinae; sternaulus very short and weak, as a wide V-shaped depression anteriorly, reaching at most 0.3 × length of mesopleuron (Fig. 2 D); juxtacoxal carina barely indicated anteriorly (Fig. 2 C); propodeum (Fig. 1 A) with mostly weak carinae; area superomedia confluent with area basalis and separated from area petiolaris; area externa and area dentipara confluent; area spiracularis confluent with area lateralis; area dentipara section of lateral longitudinal carina very weak, partly absent; posterior transverse carina strong, forming elevate apophysis joining lateral longitudinal carina, apophysis subtriangular, with blunt tip (Fig. 2 D, F, arrow). Length of hind femur about 4.4 × its height. Hind wing with vein cu-a 0.73 as long as abscissa of CU between M and cu-a. METASOMA. Tergite I (Figs 1 A, 2 G) finely granulate and very weakly strigose laterally, matt, 2.5 × as long as its maximum width; lateromedian longitudinal carina absent; lateral longitudinal carina complete; tergite II (Fig. 2 G) shallowly granulate, shiny, glabrous; remaining tergites (Fig. 2 G) smooth and shiny with inconspicuous dense setiferous punctures. Ovipositor sheath about 0.4 × as long as hind tibia. COLOUR. Head (Fig. 2 A – B, D – E) light orange; antenna with scape and pedicel light brown, flagellum dark brown, distal part of flagellomere 4 to flagellomere 7 dorsally white (Fig. 2 A). Mesosoma (Fig. 2 D – E) light orange. Metasoma (Fig. 2 G) mostly brown, tergites I – II and anterior part of IV light orange. Legs (Fig. 2 A) mostly light orange, distal tarsomeres, dark brown, fore and mid coxae, trochanters and trochantelli white, hind coxa white proximally, dark brown distally. Wings (Fig. 2 A) slightly evenly infuscate. Male Unknown.	en	Bordera, Santiago, Broad, Gavin R. (2021): Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World. European Journal of Taxonomy 757: 80-101, DOI: https://doi.org/10.5852/ejt.2021.757.1415, URL: http://dx.doi.org/10.5852/ejt.2021.757.1415
03858793FFD7FFA9DAB7FEB67926FB9E.taxon	description	urn: lsid: zoobank. org: act: 3 FE 16228 - 86 F 1 - 4 A 8 C- 8 FA 0 - 227999 E 65 A 4 F Figs 1 B, 3, 9 B	en	Bordera, Santiago, Broad, Gavin R. (2021): Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World. European Journal of Taxonomy 757: 80-101, DOI: https://doi.org/10.5852/ejt.2021.757.1415, URL: http://dx.doi.org/10.5852/ejt.2021.757.1415
03858793FFD7FFA9DAB7FEB67926FB9E.taxon	diagnosis	Diagnosis Handaoia fritzi sp. nov. can be distinguished from all other New World species by the combination of the following characters: apophysis of propodeum as a low transverse rounded or slightly pointed triangular crest (Fig. 3 G, arrow); juxtacoxal carina strong, irregular, joining submetapleural carina at anterior 0.3 – 0.4 (Fig. 3 F, arrow); area externa of propodeum separated from area dentipara by strong carina (Fig. 1 B); inner side of area externa 3.0 – 3.4 × as long as inner side of area dentipara (Fig. 1 B); inner side of area dentipara 0.46 – 0.5 × the length of outer side (Fig. 1 B); tergite II smooth and shiny (Fig. 3 B).	en	Bordera, Santiago, Broad, Gavin R. (2021): Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World. European Journal of Taxonomy 757: 80-101, DOI: https://doi.org/10.5852/ejt.2021.757.1415, URL: http://dx.doi.org/10.5852/ejt.2021.757.1415
03858793FFD7FFA9DAB7FEB67926FB9E.taxon	etymology	Etymology This species is named in honour of Dr Fritz Plaumann, illustrious botanist and entomologist based in Brazil, who collected the type material. A noun in genitive case.	en	Bordera, Santiago, Broad, Gavin R. (2021): Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World. European Journal of Taxonomy 757: 80-101, DOI: https://doi.org/10.5852/ejt.2021.757.1415, URL: http://dx.doi.org/10.5852/ejt.2021.757.1415
03858793FFD7FFA9DAB7FEB67926FB9E.taxon	materials_examined	Material examined Known from five females. Holotype BRAZIL • ♀; Nova Teutonia; 27 º 11 ′ S, 52 º 23 ′ W; 26 Aug. 1937; F. Plaumann leg.; B. M. 1937 - 748; B. M. Type Hym 3 b. 2871; NHMUK 010636381. Paratypes BRAZIL • 1 ♀; same collection data as for holotype; Aug. 1935; B. M. 1937 - 47; NHMUK 010636388 • 1 ♀; same collection data as for preceding; 28 May 1938; B. M. 1938 - 458; NHMUK 010636387 • 1 ♀; same collection data as for preceding; 11 Aug. 1938; B. M. 1938 - 682; NHMUK 010636396 • 1 ♀; same collection data as for preceding; 16 Sep. 1938; B. M. 1938 - 682; NHMUK 010636383.	en	Bordera, Santiago, Broad, Gavin R. (2021): Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World. European Journal of Taxonomy 757: 80-101, DOI: https://doi.org/10.5852/ejt.2021.757.1415, URL: http://dx.doi.org/10.5852/ejt.2021.757.1415
03858793FFD7FFA9DAB7FEB67926FB9E.taxon	description	Description Female MEASUREMENTS. Body length (without ovipositor) 3.3 – 4.1 mm. Fore wing 2.7 – 3.3 mm long. HEAD. Finely granulate, matt, distinctly narrowed behind eyes, occiput very concave centrally in dorsal view (Fig. 3 C); gena 0.2 – 0.25 × length of eye in dorsal view; posterior ocellus separated from eye by 0.75 – 1.25 × its own maximum diameter; distance between posterior ocelli 0.75 – 1.0 × maximum ocellar diameter (Fig. 3 C); occipital carina gently V-shaped medially (Fig. 3 B); malar space 1.15 – 1.25 × basal width of mandible; face slightly swollen medially; clypeus (Fig. 3 E) weakly convex, faintly granulate dorsally, smooth and shiny ventrally, with scattered punctures, its apex almost straight, 2.1 – 2.2 × as wide as high; mandible slightly tapered to apex, ventral tooth of mandible approximately same length as upper tooth; maxillary palp relatively short, reaching at most ⅔ length of mesosternum; flagellum widened in middle, tapered towards apex, flagellomeres 8 – 17 flattened ventrally; flagellum with 19 – 20 segments, length-thickness ratios: 1 st segment = 4.74 – 5.0, 2 nd = 5.0 – 5.28 and 10 th = 1.0. MESOSOMA. Finely granulate and matt (Fig. 3 B – D); mesoscutum convex, notauli weak anteriorly, reaching about 0.3 distance to scuto-scutellar groove, sometimes inconspicuous (Fig. 3 C); scutellum moderately convex, without lateral carinae; sternaulus moderately deep, reaching at least middle of mesopleuron (Fig. 3 D); juxtacoxal carina strong, irregular, joining submetapleural carina at anterior 0.3 – 0.4 (Fig. 3 F, arrow); propodeum (Fig. 1 B) with strong and conspicuous carinae; area superomedia confluent with area basalis and separated from area petiolaris; inner side of area externa about 3.0 – 4.0 × as long as inner side of area dentipara; inner side of area dentipara 0.46 – 0.5 × length of outer side; area spiracularis confluent with area lateralis; posterior transverse carina strong, forming a low and wide rounded or slightly subtriangular crest joining lateral longitudinal carina (Fig. 3 G, arrow). Length of hind femur 4.0 – 4.7 × its height. Hind wing with vein cu-a 0.4 – 0.6 × as long as abscissa of CU between M and cu-a. METASOMA. Tergite I (Figs 1 B, 3 B), finely granulate, matt, 1.8 – 2.0 × as long as its maximum width; postpetiole laterally and posteriorly tending to fine strigosity; lateromedian longitudinal carina weak, reaching at least 0.6 × length of tergite; lateral longitudinal carina weak, complete or absent posterior to spiracles; tergite II (Fig. 3 B) smooth and shiny, glabrous; remaining tergites (Fig. 3 B) smooth and shiny with inconspicuous dense setiferous punctures. Ovipositor sheath 0.45 – 0.49 × as long as hind tibia. COLOUR. Head (Fig. 3 A – E) dark brown, clypeus and mandibles reddish brown; palpi light brown; antenna with scape, pedicel and flagellomeres 1 – 4 light brown, darker dorsally, remaining flagellomeres dark brown, flagellomeres 4 – 7 dorsally white (Fig. 3 A). Mesosoma (Fig. 3 A – D) mostly dark orange; mesoscutum with wide dark brown longitudinal bands on central and lateral lobes; dorsal anterior part of propodeum dark brown. Metasoma (Fig. 3 B) mostly brown, tergite I sometimes centrally tinged dark brown. Legs (Fig. 3 A) light brown, distal tarsomeres dark brown, fore and mid coxae, part of hind coxa and all trochanters and trochantelli lighter. Wings (Fig. 3 A) slightly infuscate. Male Unknown.	en	Bordera, Santiago, Broad, Gavin R. (2021): Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World. European Journal of Taxonomy 757: 80-101, DOI: https://doi.org/10.5852/ejt.2021.757.1415, URL: http://dx.doi.org/10.5852/ejt.2021.757.1415
03858793FFD9FFABDA17FBE67926FA5B.taxon	description	urn: lsid: zoobank. org: act: 80 A 6 A 583 - DB 23 - 43 D 9 - AF 4 A- 4 D 16 A 6270918 Figs 1 C, 4, 9 B	en	Bordera, Santiago, Broad, Gavin R. (2021): Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World. European Journal of Taxonomy 757: 80-101, DOI: https://doi.org/10.5852/ejt.2021.757.1415, URL: http://dx.doi.org/10.5852/ejt.2021.757.1415
03858793FFD9FFABDA17FBE67926FA5B.taxon	diagnosis	Diagnosis Handaoia mercedensis Bordera sp. nov. can be distinguished from all other New World species by the combination of the following characters: apophysis of propodeum conspicuously elevated, parallelsided with blunt tip (Fig. 4 G, arrow); fore wing with well-defined transverse dark brown band (Fig. 4 A); juxtacoxal carina strong, irregular, joining submetapleural carina at mid length (Fig. 4 F, arrow); flagellum dark brown (except white band), with first and second flagellomeres light brown (Fig. 4 A – B).	en	Bordera, Santiago, Broad, Gavin R. (2021): Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World. European Journal of Taxonomy 757: 80-101, DOI: https://doi.org/10.5852/ejt.2021.757.1415, URL: http://dx.doi.org/10.5852/ejt.2021.757.1415
03858793FFD9FFABDA17FBE67926FA5B.taxon	etymology	Etymology The name refers to La Merced, municipality of Chanchamayo in the Department of Junin (Peru), where the holotype was collected.	en	Bordera, Santiago, Broad, Gavin R. (2021): Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World. European Journal of Taxonomy 757: 80-101, DOI: https://doi.org/10.5852/ejt.2021.757.1415, URL: http://dx.doi.org/10.5852/ejt.2021.757.1415
03858793FFD9FFABDA17FBE67926FA5B.taxon	materials_examined	Material examined Known only from the holotype female. Holotype PERU • ♀; La Merced, Fundo La Génova; Trampa Malaise 3; 3 – 17 Jan. 2009; AECID 013484 / 07; CEUA.	en	Bordera, Santiago, Broad, Gavin R. (2021): Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World. European Journal of Taxonomy 757: 80-101, DOI: https://doi.org/10.5852/ejt.2021.757.1415, URL: http://dx.doi.org/10.5852/ejt.2021.757.1415
03858793FFD9FFABDA17FBE67926FA5B.taxon	description	Description Female MEASUREMENTS. Body length (without ovipositor) 4.3 mm. Fore wing 3.4 mm long. HEAD. Finely granulate, matt, strongly narrowed behind eyes, occiput very concave centrally in dorsal view (Fig. 4 C); gena 0.18 × length of eye in dorsal view; posterior ocellus separated from eye by 1.3 × its own maximum diameter; distance between posterior ocelli 0.9 × maximum ocellar diameter (Fig. 4 C); occipital carina gently V-shaped medially; malar space about 1.4 × basal width of mandible; face slightly swollen medially; clypeus (Fig. 4 B) quite convex, gently granulate, with scattered coarse punctures in apical half, apex slightly rounded, about 2.0 × as wide as high; mandible quite tapered to apex, ventral tooth of mandible a little shorter than upper tooth; maxillary palp reaching mid coxa; flagellum widened in middle, tapered towards apex, flagellomeres 8 – 20 flattened ventrally (Fig. 4 A); flagellum with 22 segments, length-thickness ratios: 1 st segment = 5.0, 2 nd = 5.38 and 10 th = 1.0. MESOSOMA. Finely granulate and matt (Fig. 4 C – D); mesoscutum slightly convex, notauli absent (Fig. 4 C); scutellum moderately convex, without lateral carinae; sternaulus deep, exceeding half the length of mesopleuron (Fig. 4 D); juxtacoxal carina strong, irregular, joining submetapleural carina at mid length (Fig. 4 F, arrow); propodeum (Fig. 1 C) with strong and conspicuous carinae; area superomedia confluent with area basalis and separated from area petiolaris; inner side of area externa about 4.7 × as long as inner side of area dentipara; inner side of area dentipara about 0.3 × length of outer side; area spiracularis confluent with area lateralis; posterior transverse carina strong, forming an elevate apophysis joining lateral longitudinal carina, apophysis parallel-sided with blut tip (Fig. 4 G, arrow). Length of hind femur about 4.0 × its height. Hind wing with vein cu-a 0.5 as long as abscissa of CU between M and cu-a. METASOMA. Tergite I (Fig. 1 C) finely granulate, slightly shiny, 1.77 × as long as its maximum width; postpetiole laterally not finely strigose; lateromedian longitudinal carina moderately strong, reaching about 0.7 × length of tergite; lateral longitudinal carina complete; tergite II (Fig. 4 E) shiny, very slightly granulate anteriorly, glabrous, smooth posteriorly, with moderately dense short setae; remaining tergites (Fig. 4 E) smooth and shiny with inconspicuous dense setiferous punctures. Ovipositor sheath 0.53 × as long as hind tibia. COLOUR. Head (Fig. 4 A – D) orange brown, vertical orbits and interocellar area dark brown; palpi dark brown; mandibular teeth black; antenna with scape, pedicel and flagellomeres 1 – 2 light brown, remaining flagellomeres dark brown, flagellomeres 4 – 7 and base of 8 dorsally white (Fig. 4 A). Mesosoma (Fig. 4 A, C – D) entirely orange-brown. Metasoma (Fig. 4 E) except tergite I dark brown, tergite I light brown. Fore and mid legs (Fig. 4 A) light brown, tarsi fuscous; hind leg dark brown, trochanter and base of tibia white. Wings slightly infuscate, fore wing with well-defined transverse dark brown band (Fig. 4 A). Male Unknown.	en	Bordera, Santiago, Broad, Gavin R. (2021): Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World. European Journal of Taxonomy 757: 80-101, DOI: https://doi.org/10.5852/ejt.2021.757.1415, URL: http://dx.doi.org/10.5852/ejt.2021.757.1415
03858793FFDBFFAFDA59F9A37A13FB8F.taxon	description	urn: lsid: zoobank. org: act: 7 A 2 E 9032 - B 90 C- 4 EB 7 - 9 D 7 F- 139 AA 2 D 3 D 4 B 9 Figs 1 D, 5 – 6, 9 B	en	Bordera, Santiago, Broad, Gavin R. (2021): Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World. European Journal of Taxonomy 757: 80-101, DOI: https://doi.org/10.5852/ejt.2021.757.1415, URL: http://dx.doi.org/10.5852/ejt.2021.757.1415
03858793FFDBFFAFDA59F9A37A13FB8F.taxon	diagnosis	Diagnosis Handaoia plaumanni sp. nov. can be distinguished from all other New World species by the combination of the following characters (both male and female): apophysis of propodeum as a low transverse crest (Fig. 5 I, arrow); sternaulus very short and weak, reaching at most 0.3 × the length of the mesopleuron (Figs 5 E, 6 D); juxtacoxal carina absent (Figs 5 H, 6 E); area externa of propodeum confluent with area dentipara (Figs 1 D, 6 F).	en	Bordera, Santiago, Broad, Gavin R. (2021): Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World. European Journal of Taxonomy 757: 80-101, DOI: https://doi.org/10.5852/ejt.2021.757.1415, URL: http://dx.doi.org/10.5852/ejt.2021.757.1415
03858793FFDBFFAFDA59F9A37A13FB8F.taxon	etymology	Etymology This species is named in honour of Dr Fritz Plaumann, illustrious botanist and entomologist based in Brazil, who collected the type series. A noun in the genitive case.	en	Bordera, Santiago, Broad, Gavin R. (2021): Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World. European Journal of Taxonomy 757: 80-101, DOI: https://doi.org/10.5852/ejt.2021.757.1415, URL: http://dx.doi.org/10.5852/ejt.2021.757.1415
03858793FFDBFFAFDA59F9A37A13FB8F.taxon	materials_examined	Material examined Known from three females and two males. Holotype BRAZIL • ♀; Nova Teutonia; 27 º 11 ′ S, 52 º 23 ′ W; 24 May 1938; F Plaumann leg.; B. M. 1938 - 452; B. M. Type Hym 3 b. 2872; NHMUK 010636386. Paratypes BRAZIL • 1 ♀; same collection data as for holotype; 13 Apr. 1938; B. M. 1938 - 682; NHMUK 010636382 • 1 ♂; same collection data as for preceding; 10 May 1938; B. M. 1938 - 682; NHMUK 010636384 • 1 ♀; same collection data as for preceding; 13 May 1938; B. M. 1938 - 682; NHMUK 010636395 • 1 ♂; same collection data as for preceding; 7 Jun. 1938; B. M. 1938 - 458; NHMUK 010636385.	en	Bordera, Santiago, Broad, Gavin R. (2021): Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World. European Journal of Taxonomy 757: 80-101, DOI: https://doi.org/10.5852/ejt.2021.757.1415, URL: http://dx.doi.org/10.5852/ejt.2021.757.1415
03858793FFDBFFAFDA59F9A37A13FB8F.taxon	description	Description Female MEASUREMENTS. Body length (without ovipositor) 2.6 – 2.9 mm. Fore wing 2.3 – 2.4 mm long. HEAD. Moderately coarsely granulate, matt, distinctly narrowed behind eyes, occiput very concave centrally in dorsal view (Fig. 5 B – E); gena 0.16 – 0.25 × length of eye in dorsal view; posterior ocellus separated from eye by 0.9 – 1.0 × its maximum diameter; distance between posterior ocelli 0.75 – 1.3 × maximum ocellar diameter (Fig. 5 D); occipital carina gently V-shaped medially (Fig. 5 D); malar space 1.17 – 1.33 × basal width of mandible; face weakly swollen medially (Fig. 5 C); clypeus (Fig. 5 C) weakly convex, smooth and shiny with scattered punctures, apex almost straight, about 2.0 × as wide as high; mandible slightly tapered to apex, ventral tooth approximately same length as upper tooth; maxillary palp relatively short, reaching at most ⅔ length of mesosternum; flagellum widened in middle, tapered towards apex, flagellomeres 8 – 17 flattened ventrally (Fig. 5 A); flagellum with 18 segments, lengththickness ratios: 1 st segment = 4.67 – 5.0, 2 nd = 5.0 – 5.33 and 10 th = 1.0. MESOSOMA. Moderately coarsely granulate, matt (Fig. 5 D – E); mesoscutum slightly convex, flat posteriorly, notauli very weak anteriorly or absent (Fig. 5 B, D), when present reaching at most 0.3 distance to scuto-scutellar groove; scutellum moderately convex, without lateral carinae; sternaulus very short and weak, as a wide V-shaped depression anteriorly, reaching at most 0.3 × the length of mesopleuron (Fig. 5 E); juxtacoxal carina absent (Fig. 5 H); propodeum (Figs 1 D, 5 I) with moderately strong carinae; area superomedia confluent with area basalis and separated from area petiolaris; area externa and area dentipara confluent; area spiracularis confluent with area lateralis; posterior transverse carina forming a very low transverse crest joining lateral longitudinal carina (Fig. 5 I, arrow). Length of hind femur 4.0 – 4.26 × its height. Hind wing with vein cu-a 0.4 – 0.6 × as long as abscissa of CU between M and cu-a. METASOMA. Tergite I (Figs 1 D, 5 J) finely granulate, matt, 1.6 – 1.8 × as long as its maximum width; lateromedian longitudinal carina absent; lateral longitudinal carina weak, absent posterior to spiracles; tergite II (Fig. 5 F – G) smooth and shiny, glabrous; remaining tergites (Fig. 5 F – G) smooth and shiny with inconspicuous dense setiferous punctures. Ovipositor sheath about 0.5 – 0.6 × as long as hind tibia. COLOUR. Head (Fig. 5 B – E) dark brown; antenna with scape and pedicel light brown and flagellomeres 1 – 4 light brown sometimes infuscate, remaining flagellomeres dark brown, flagellomeres 4 – 7 dorsally white (Fig. 5 A). Mesosoma (Fig. 5 B, D – E) mostly orange-brown; mesoscutum tending to entirely dark brown, sometimes anterior part of propodeum infuscate. Metasoma (Fig. 5 F – G) brown, tergites I – II and sometimes anterior part of III light brown, remaining tergites dark brown. Legs (Fig. 5 A) light brown, distal tarsomeres dark brown, sometimes fore and mid coxae, part of hind coxa and all trochanters and trochantelli lighter. Wings (Fig. 5 A) slightly infuscate. Male MEASUREMENTS. Body length 2.3 – 2.9 mm. Fore wing 2.0 – 2.6 mm long. HEAD. Less narrowed behind eyes and less concave centrally in dorsal view than female (Fig. 6 C); gena 0.31 – 0.42 × length of eye in dorsal view; posterior ocellus separated from eye by 1.25 – 1.33 × its maximum diameter; distance between posterior ocelli 0.9 – 1.3 × maximum ocellar diameter (Fig. 6 C); malar space 1.33 × basal width of mandible; clypeus about 1.7 × as wide as high; flagellum filiform (Fig. 6 A – B) with 19 segments, length-thickness ratios: 1 st = segment 5.0, 2 nd = 5.0 and 10 th = 3.4 – 4.0 MESOSOMA. Notauli absent (Fig. 6 C – D); length of hind femur about 4.5 – 4.6 × its heigth. METASOMA. Tergite I finely granulate, matt, 2.1 × as long as its maximum width. COLOUR. As in female (Fig. 6 A – F) but antenna entirely dark brown. In one paratype (NHMUK 010636385), mesosoma and metasoma mostly dark brown (Fig. 6 A, D). Otherwise similar to female.	en	Bordera, Santiago, Broad, Gavin R. (2021): Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World. European Journal of Taxonomy 757: 80-101, DOI: https://doi.org/10.5852/ejt.2021.757.1415, URL: http://dx.doi.org/10.5852/ejt.2021.757.1415
03858793FFDFFFB1DA15FBF77926FA5B.taxon	description	urn: lsid: zoobank. org: act: 151 C 284 F- 8024 - 4 A 2 A- 97 FD-D 0 BBDE 6 AF 048 Figs 1 E, 7, 9 A	en	Bordera, Santiago, Broad, Gavin R. (2021): Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World. European Journal of Taxonomy 757: 80-101, DOI: https://doi.org/10.5852/ejt.2021.757.1415, URL: http://dx.doi.org/10.5852/ejt.2021.757.1415
03858793FFDFFFB1DA15FBF77926FA5B.taxon	diagnosis	Diagnosis Handaoia ruizcancinoi Bordera sp. nov. can be distinguished from all other New World species by the combination of the following characters: apophysis of propodeum conspicuously high, subtriangular with blunt tip (Fig. 7 H, arrow); juxtacoxal carina weak but complete, joining submetapleural carina at its anterior part (Fig. 7 G, arrow); inner side of area externa about 7.0 × as long as inner side of area dentipara (Fig. 1 E).	en	Bordera, Santiago, Broad, Gavin R. (2021): Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World. European Journal of Taxonomy 757: 80-101, DOI: https://doi.org/10.5852/ejt.2021.757.1415, URL: http://dx.doi.org/10.5852/ejt.2021.757.1415
03858793FFDFFFB1DA15FBF77926FA5B.taxon	etymology	Etymology The species is named in honour of Dr Enrique Ruiz Cancino from Universidad Autónoma de Tamaulipas (Mexico), in recognition of his contribution to the study of the Ichneumonidae of Mexico. A noun in the genitive case.	en	Bordera, Santiago, Broad, Gavin R. (2021): Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World. European Journal of Taxonomy 757: 80-101, DOI: https://doi.org/10.5852/ejt.2021.757.1415, URL: http://dx.doi.org/10.5852/ejt.2021.757.1415
03858793FFDFFFB1DA15FBF77926FA5B.taxon	materials_examined	Material examined Known only from the holotype female. Holotype MEXICO • ♀; Tamaulipas, Ocampo, J. de Manantiales; T. Amarillas; 13 May 1995; J. Coronado, S. Nino and C. Hernández leg.; CEUA.	en	Bordera, Santiago, Broad, Gavin R. (2021): Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World. European Journal of Taxonomy 757: 80-101, DOI: https://doi.org/10.5852/ejt.2021.757.1415, URL: http://dx.doi.org/10.5852/ejt.2021.757.1415
03858793FFDFFFB1DA15FBF77926FA5B.taxon	description	Description Female MEASUREMENTS. Body length (without ovipositor) 4.0 mm. Fore wing about 3.0 mm long. HEAD. Finely granulate, matt, strongly narrowed behind eyes, occiput very concave centrally in dorsal view (Fig. 7 B – D); gena 0.14 × length of eye in dorsal view; posterior ocellus separated from eye by 1.55 × its maximum diameter; distance between posterior ocelli 1.27 × maximum ocellar diameter; occipital carina gently V-shaped medially (Fig. 7 D); malar space 1.3 × basal width of mandible; face conspicuously swollen medially (Fig. 7 B); clypeus (Fig. 7 B) quite convex, gently granulate with scattered coarse punctures, apex slightly rounded, about 1.8 × as wide as high; mandible quite tapered to apex, ventral tooth of mandible a little shorter than upper tooth; flagellum widened in middle, tapered towards apex, flagellomeres 8 – 18 flattened ventrally; flagellum with 20 segments, length-thickness ratios: 1 st segment = 4.1, 2 nd = 3.27 and 10 th = 0.72. MESOSOMA. Finely granulate and matt (Fig. 7 A, C – D); mesoscutum slightly convex, almost flat posteriorly; notauli moderately deep over 0.7 × length of mesoscutum (Fig. 7 D); scutellum moderately convex, without lateral carinae; sternaulus as a wide V-shaped depression anteriorly, reaching about 0.3 × the length of mesopleuron (Fig. 7 C); juxtacoxal carina weak but complete, joining submetapleural carina at its anterior part (Fig. 7 G, arrow); propodeum (Figs 1 E, 7 C, H) with strong and conspicuous carinae; area superomedia confluent with area basalis and separated from area petiolaris; inner side of area externa about 7.0 × as long as inner side of area dentipara; inner side of area dentipara about 0.3 × length of outer side; area spiracularis confluent with area lateralis; posterior transverse carina strong, forming an elevated apophysis joining lateral longitudinal carina, apophysis subtriangular with blunt tip (Fig. 7 C, H, arrow). Length of hind femur 3.9 × its height. Hind wing with vein cu-a 0.77 × as long as abscissa of CU between M and cu-a. METASOMA. Tergite I (Fig. 7 E) finely granulate, shiny, strigose laterally, about 2.4 × as long as its maximum width; lateromedian longitudinal carina moderately strong, reaching posterior 0.8 × length of tergite; lateral longitudinal carina present from spiracle to posterior margin of postpetiole; tergite II (Fig. 7 F) shiny, smooth and almost glabrous; remaining tergites (Fig. 7 F) smooth and shiny, with short and moderately dense setiferous punctures. Ovipositor sheath about 0.5 × as long as hind tibia. COLOUR. Head (Fig. 7 B – D) brown, darker dorsally, lighter ventrally, mandibular teeth and labrum dark brown; antenna with scape, pedicel and first flagellomere light brown, remaining flagellomeres dark brown, flagellomeres 5 – 7 dorsally white (Fig. 7 A). Mesosoma (Fig. 7 A, C) brown, more infuscate ventrally. Metasoma (Fig. 7 E – F) dark brown. Legs (Fig. 7 A) mostly dark brown; fore and mid coxae and trochanters, and hind trochanter, white. Wings (Fig. 7 A) evenly infuscate. Male Unknown.	en	Bordera, Santiago, Broad, Gavin R. (2021): Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World. European Journal of Taxonomy 757: 80-101, DOI: https://doi.org/10.5852/ejt.2021.757.1415, URL: http://dx.doi.org/10.5852/ejt.2021.757.1415
03858793FFC1FFB2DA43F9A37926FEC2.taxon	description	urn: lsid: zoobank. org: act: 5413 C 7 C 8 - D 785 - 4583 - 803 F- 0 C 1 E 75 B 7 EA 9 E Figs 1 F, 8, 9 B	en	Bordera, Santiago, Broad, Gavin R. (2021): Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World. European Journal of Taxonomy 757: 80-101, DOI: https://doi.org/10.5852/ejt.2021.757.1415, URL: http://dx.doi.org/10.5852/ejt.2021.757.1415
03858793FFC1FFB2DA43F9A37926FEC2.taxon	diagnosis	Diagnosis Handaoia urceus sp. nov. can be distinguished from all other New World species by the combination of the following characters: apophysis of propodeum as a low transverse rounded crest (Fig. 8 I, arrow); tergite II finely granulate (Fig. 8 D); juxtacoxal carina strong anteriorly, weak posteriorly, joining submetapleural carina at its half length (Fig. 8 H, arrow); area externa of propodeum separated from area dentipara by strong carina (Fig. 1 F); inner side of area externa about 8.0 × as long as inner side of area dentipara (Fig. 1 F).	en	Bordera, Santiago, Broad, Gavin R. (2021): Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World. European Journal of Taxonomy 757: 80-101, DOI: https://doi.org/10.5852/ejt.2021.757.1415, URL: http://dx.doi.org/10.5852/ejt.2021.757.1415
03858793FFC1FFB2DA43F9A37926FEC2.taxon	etymology	Etymology From the Latin for ‘ pitcher’, named after the flask- or pitcher-shaped combined area basalis + superomedia. A noun in apposition.	en	Bordera, Santiago, Broad, Gavin R. (2021): Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World. European Journal of Taxonomy 757: 80-101, DOI: https://doi.org/10.5852/ejt.2021.757.1415, URL: http://dx.doi.org/10.5852/ejt.2021.757.1415
03858793FFC1FFB2DA43F9A37926FEC2.taxon	materials_examined	Material examined Known only from the holotype female. Holotype BRAZIL • ♀; Nova Teutonia; 27 º 11 ′ S, 52 º 23 ′ W; 8 Feb. 1939; F. Plaumann leg.; B. M. 1939 - 181; B. M. Type Hym 3 b. 2873; NHMUK 010636380.	en	Bordera, Santiago, Broad, Gavin R. (2021): Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World. European Journal of Taxonomy 757: 80-101, DOI: https://doi.org/10.5852/ejt.2021.757.1415, URL: http://dx.doi.org/10.5852/ejt.2021.757.1415
03858793FFC1FFB2DA43F9A37926FEC2.taxon	description	Description Female MEASUREMENTS. Body length (without ovipositor) 4.3 mm. Fore wing 3.7 mm long. HEAD. Finely granulate, matt, distinctly narrowed behind eyes, occiput very concave centrally in dorsal view (Fig. 8 B – C); gena 0.22 × length of eye in dorsal view; posterior ocellus separated from eye by 1.0 × its maximum diameter; distance between posterior ocelli 1.0 × maximum ocellar diameter (Fig. 8 C); occipital carina gently V-shaped medially (Fig. 8 E); malar space 1.44 × basal width of mandible; face moderately swollen medially (Fig. 8 B); clypeus (Fig. 8 B) weakly convex, gently granulate with scattered punctures, apex slightly rounded, about 1.6 × as wide as high; mandible slightly tapered to apex, ventral tooth approximately same length as upper tooth; maxillary palp long, almost reaching middle coxa; flagellum widened in middle, tapered towards apex (Fig. 8 A), flagellomeres 8 – 18 flattened ventrally; flagellum with 22 segments, length-thickness ratios: 1 st segment = 4.9, 2 nd = 6.1 and 10 th = 1.13. MESOSOMA. Finely granulate and matt (Fig. 8 E, G); mesoscutum (Fig. 8 E, G) convex, notauli moderately deep anteriorly, reaching about 0.3 × distance to scuto-scutellar groove; scutellum moderately convex, without lateral carinae; sternaulus moderately deep, reaching about mid-point of mesopleuron (Fig. 8 G); juxtacoxal carina strong anteriorly, weak posteriorly, joining submetapleural carina at mid length (Fig. 8 H, arrow); propodeum (Figs 1 F, 8 I) with strong and conspicuous carinae; area superomedia confluent with area basalis and separated from area petiolaris; inner side of area externa about 8.0 × as long as inner side of area dentipara; inner side of area dentipara 0.25 × length of outer side; area spiracularis confluent with area lateralis; posterior transverse carina strongly elevated, forming a low transverse crest joining lateral longitudinal carina (Fig. 8 I, arrow). Length of hind femur 4.2 × its height. Hind wing with vein cu-a about 1.0 × as long as abscissa of CU between M and cu-a. METASOMA. Tergite I (Figs 1 F, 8 D) finely granulate, matt, about 1.70 × as long as its maximum width; postpetiole laterally and posteriorly tending to fine strigosity; lateromedian and lateral longitudinal carinae strong, reaching at least 0.8 × length of tergite; tergite II (Fig. 8 D, F) finely granulate, shiny; remaining tergites (Fig. 8 D, F) smooth and shiny with inconspicuous setiferous punctures. Ovipositor sheath about 0.54 × as long as hind tibia. COLOUR. Head (Fig. 8 B – C) dark brown tending to be more reddish on clypeus and mandibles; palpi light brown; antenna with scape, pedicel and flagellomeres I – II (III) light brown, remaining flagellomeres dark brown, distal part of flagellomere 4 to flagellomere 7 dorsally white (Fig. 8 A). Mesosoma (Fig. 8 E – G) entirely orange. Metasoma (Fig. 8 D – F) with a gradient from orange at anterior of tergite I to dark brown on posterior tergites, posterior rim of tergites I – III yellowish. Legs (Fig. 8 A) light brown, distal tarsomeres dark brown, fore and mid coxae, part of hind coxa and all trochanters and trochantelli lighter. Wings (Fig. 8 A) slightly infuscate. Male Unknown.	en	Bordera, Santiago, Broad, Gavin R. (2021): Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World. European Journal of Taxonomy 757: 80-101, DOI: https://doi.org/10.5852/ejt.2021.757.1415, URL: http://dx.doi.org/10.5852/ejt.2021.757.1415
