taxonID	type	description	language	source
03F7A821D9543522FF67FDFC8B3C669D.taxon	materials_examined	Type species: Ckacus youcha n. sp. (Luoshui Dong cave, Xianheping, Anlong, Qianxinan, Guizhou) Generic characteristics. Medium-sized for cavernicolous trechine beetles, semi-aphaenopsian wholly pubescent, eyeless and depigmented; body rather thin, elongate, with moderate slender appendages. Head strongly elongate; genae nearly parallel-sided; two pairs of supra-orbital setiferous pores present; frontal furrows short and reduced posteriorly; vertex moderately convex; right mandibular tooth tridentate though median one reduced; labial suture completely disappeared; mentum bisetose, base largely and deeply concave; submentum with a row of 12 - setose; antennae moderately thin, not extending over apices of elytra. Propleura invisible from above; pronotum quadrate, slightly longer than wide, presence of two pairs of latero-marginal setae, both fore and hind angles sharpened. Elytra elongated ovate, with a distinct but rounded humeral angle on each side; slightly longer than fore body including mandibles; lateral margins ciliate throughout; disc strongly convex, punctation of striae noticeable, intervals convex, the 1 st intervals very narrowed in apical 1 / 3; two pairs of dorsal setiferous pores present on the 3 rd stria, preapical pores present; the 4 th pore of marginal umbilicate series far from the 3 rd, 5 th and 6 th pores close to each other. The 1 st and 2 nd protarsomeres modified in male, slightly widened and distinctly denticulate inwardly. Ventrite VII with one pair of apical setae in male. Male genitalia thin and elongate, moderately sclerotized.	en	Tian, Mingyi, Huang, Sunbin (2021): Discovery of a new cavernicolous trechine genus and species from a deep pit in Anlong, southwestern Guizhou Province (Coleoptera: Carabidae: Trechinae). Zootaxa 4985 (2): 194-202, DOI: https://doi.org/10.11646/zootaxa.4985.2.3
03F7A821D9543522FF67FDFC8B3C669D.taxon	discussion	Remarks. Ckacus looks like Bathytrechus Uéno, 2005, which was recorded from the cave Tiankeng Dong in Leye, northwestern most of Guangxi, in the medium and elongate body which is wholly pubescent, the tridentate right mandible, sharpened hind angles of pronotum, and entirely striate elytra with the first intervals extremely narrowed at the apical third of elytra. However, it differs from the latter genus in many characters, such as: (1) Mentum and submentum completely fused and submentum 12 - setose, versus labial suture traceable and submentum only 6 - setose in Bathytrechus; (2) Frontal furrows distinctly reduced posteriorly, versus entire in Bathytrechus; (3) 1 st and 2 nd protarsomeres modified in male, versus not modified in Bathytrechus; (4) the middle group of the marginal umbilicate pores close to each other, versus widely spaced in Bathytrechus; and (5) apical striole of elytra absent, versus present in Bathytrechus. In addition, male genitalia is much more elongate, with a sharp apical lobe in Ckacus, whereas shorter and apical lobe rod-like in Bathytrechus.	en	Tian, Mingyi, Huang, Sunbin (2021): Discovery of a new cavernicolous trechine genus and species from a deep pit in Anlong, southwestern Guizhou Province (Coleoptera: Carabidae: Trechinae). Zootaxa 4985 (2): 194-202, DOI: https://doi.org/10.11646/zootaxa.4985.2.3
03F7A821D9543522FF67FDFC8B3C669D.taxon	etymology	Etymology. The generic name is derived from the abbreviations of “ China Karst Cave Union ” (中国喀Dz特洞 Ë ḆDz). Gender masculine. Generic range. China (Guizhou).	en	Tian, Mingyi, Huang, Sunbin (2021): Discovery of a new cavernicolous trechine genus and species from a deep pit in Anlong, southwestern Guizhou Province (Coleoptera: Carabidae: Trechinae). Zootaxa 4985 (2): 194-202, DOI: https://doi.org/10.11646/zootaxa.4985.2.3
03F7A821D9543527FF67F95F8AD264E0.taxon	description	(Figs. 2 – 5)	en	Tian, Mingyi, Huang, Sunbin (2021): Discovery of a new cavernicolous trechine genus and species from a deep pit in Anlong, southwestern Guizhou Province (Coleoptera: Carabidae: Trechinae). Zootaxa 4985 (2): 194-202, DOI: https://doi.org/10.11646/zootaxa.4985.2.3
03F7A821D9543527FF67F95F8AD264E0.taxon	materials_examined	Type material. Holotype male, Luoshui Dong cave, Xianheping, Anlong Xian (County), Qianxinan Miao & Buyi Zizhizhou (Autonomous Prefecture), southwest Guizhou, 105 ° 35 ’ 31.42 ’’ E, 24 ° 59 ’ 16.11 ’’ N, 1163 m, 2020 - XII- 06, leg. Huanghe Zhou (“ You Cha ”), in SCAU.	en	Tian, Mingyi, Huang, Sunbin (2021): Discovery of a new cavernicolous trechine genus and species from a deep pit in Anlong, southwestern Guizhou Province (Coleoptera: Carabidae: Trechinae). Zootaxa 4985 (2): 194-202, DOI: https://doi.org/10.11646/zootaxa.4985.2.3
03F7A821D9543527FF67F95F8AD264E0.taxon	diagnosis	Diagnosis. A medium-sized semi-aphaenopsian trechine species, wholly pubescent, with moderately elongated body and thin appendages.	en	Tian, Mingyi, Huang, Sunbin (2021): Discovery of a new cavernicolous trechine genus and species from a deep pit in Anlong, southwestern Guizhou Province (Coleoptera: Carabidae: Trechinae). Zootaxa 4985 (2): 194-202, DOI: https://doi.org/10.11646/zootaxa.4985.2.3
03F7A821D9543527FF67F95F8AD264E0.taxon	description	Description. Length: 7.0 mm, width: 2.0 mm. Habitus as in Fig. 2. Body yellowish-brown, but palps and tarsi yellow; wholly covered with long setae, moderately shining, underside of head with a few setae, thorax glabrous ventrally but prosternum with several setae, abdominal ventrites shortly and densely pubescent. Macrosculpture: transversely meshed on head, densely and transversely striate on pronotum and elytra. Head much longer than wide, HLm / HW = 2.1, HLl / HW = 1.6; widest at about middle of head excluding mandibles, neck constriction well-marked; frons and vertex moderately convex; frontal furrows shallow and short, ended at about middle of head; anterior and posterior supraorbital pores located at middle and basal 1 / 4 of head respectively; clypeus 6 - setose; labrum transverse, frontal margin straight, 6 - setose; mentum bisetose on either side of tooth base, tooth moderately long, unified at tip; ligula thick and short, 8 - setose at apex, inner two much longer than other; palps thin and elongate, all glabrous but bisetose on inner margin of 2 nd labial palpomere and two additional setae at outer margin medially and subapically, penultimate palpomere longer than the apical one in labium and maxillaries (cf. 1.2 times); suborbital pores intermedially sited; antennae slender, 1 st antennomere with several cilia then densely pubescent from the 2 nd antennomere, 1 st stouter than others, as long as 2 nd, 3 rd the longest, relative length of each antennomere as follows: the 1 st (1.0), 2 nd (1.0), 3 rd (1.9), 4 th (1.8), 5 th (1.7), 6 th (1.7), 7 th (1.5), 8 th (1.5), 9 th (1.3, 10 th (1.1) and 11 th (1.5). Propleura invisible from above. Pronotum slightly longer than wide, PnL / PnW = 1.1, shorter than head excluding mandibles, PL / HLl = 0.8; lateral margins finely bordered throughout; widest at about 1 / 3 from front, gently contracted forwards and backwards, contracted and nearly straight near hind angles which are acute; fore angles slightly protruded, nearly acutangular; base faintly bisinuate, front nearly straight, both unbordered, front slightly wider than base, PfW / PbW = 1.2; anterior and posterior latero-marginal setae at 5 / 6 from base and just before hind angles respectively; disc moderately convex, mid-line distinct. Scutellum small and narrow. Elytra slightly longer than fore body including mandibles, much longer than wide, EL / EW = 1.7, twice as wide as pronotum; base unbordered, lateral margins finely ciliate throughout, nearly obliquely truncate at humeral portions, humeral angles broad, widest a little behind middle of elytra; disc strongly convex; basal pore present at sides of scutellum, anterior and posterior dorsal pores on the 3 rd stria at basal 1 / 4 and middle of elytra respectively, preapical pore at exactly the anastomotic point between the 2 nd and 3 rd striae, closer to elytral suture than to apical margin; locations of umbilicate marginal pores as in Fig. 3 A. Legs densely pubescent; 1 st and 2 nd protarsomeres slightly widened in male, shortly and bluntly denticulate at inner apex; 1 st tarsomere much shorter than 2 nd – 4 th combined in fore legs, whereas as long as in middle and hind legs; tibiae without longitudinal sulci. Each abdominal ventrites V-VI bisetose paramedially, VII bisetose in male. Male genitalia (Fig. 3 B, C): The median lobe of aedeagus medium-sized, quite elongate, gently arcuate towards apex which is sharp; basal opening quite small, with a large and almost rounded sagittal aileron; inner sac armed with a long copulatory piece, which is about 1 / 4 as long as aedeagus; in dorsal view, apical lobe thin, much longer than wide, broadly rounded at apex; parameres well-developed, but much shorter than median lobe, each paramere with four long setae at apex.	en	Tian, Mingyi, Huang, Sunbin (2021): Discovery of a new cavernicolous trechine genus and species from a deep pit in Anlong, southwestern Guizhou Province (Coleoptera: Carabidae: Trechinae). Zootaxa 4985 (2): 194-202, DOI: https://doi.org/10.11646/zootaxa.4985.2.3
03F7A821D9543527FF67F95F8AD264E0.taxon	etymology	Etymology. Dedicated to “ You Cha ”, a nickname of Ms. Huanghe Zhou, an active member of Guilin Cave Exploration Team, Guilin.	en	Tian, Mingyi, Huang, Sunbin (2021): Discovery of a new cavernicolous trechine genus and species from a deep pit in Anlong, southwestern Guizhou Province (Coleoptera: Carabidae: Trechinae). Zootaxa 4985 (2): 194-202, DOI: https://doi.org/10.11646/zootaxa.4985.2.3
03F7A821D9543527FF67F95F8AD264E0.taxon	distribution	Distribution. China (Guizhou). Known only from the limestone cave Luoshui Dong, sympatric with Satotrechus rieae Uéno, 2006. Luoshui Dong is located within Xianheping Forest Park. This vertical pothole is 515 m deep, one of the deepest caves in China (Figs. 4, 5 A, B). There is a lot of debris in the muddy terrace of about 400 m below the entrance. The specimens sampled, as well as many other Trechini, observed wandering on the ground and walls between - 330 m to - 450 m (Fig. 4). Apart from beetle species, other animals living inside the cave were cricket Tachycines (Gymnaeta) sp., millipedes of the genera Glyphiulus and Paracortina, harvestmen, bats, frog Oreolalax rhodostigmatus Hu & Fei, 1979 and crab Diyutamon cereum Huang et al., 2017 (Fig. 5 C-E).	en	Tian, Mingyi, Huang, Sunbin (2021): Discovery of a new cavernicolous trechine genus and species from a deep pit in Anlong, southwestern Guizhou Province (Coleoptera: Carabidae: Trechinae). Zootaxa 4985 (2): 194-202, DOI: https://doi.org/10.11646/zootaxa.4985.2.3
03F7A821D9513527FF67FA808D5966DF.taxon	description	Tian & Huang, 2018: 246	en	Tian, Mingyi, Huang, Sunbin (2021): Discovery of a new cavernicolous trechine genus and species from a deep pit in Anlong, southwestern Guizhou Province (Coleoptera: Carabidae: Trechinae). Zootaxa 4985 (2): 194-202, DOI: https://doi.org/10.11646/zootaxa.4985.2.3
03F7A821D9513527FF67FA808D5966DF.taxon	materials_examined	Material examined. 3 females, from deep area of - 330 to - 450 m inside the cave Luoshui Dong, Xianheping, Anlong Xian (County), Qianxinan Miao & Buyi Zizhizhou (Autonomous Prefecture), southwest Guizhou, 105 ° 35 ’ 31.42 ’’ E / 24 ° 59 ’ 16.11 ’’ N, 1163 m, 2020 - XII- 06, leg. Huanghe Zhou (You Cha), in SCAU; 1 male and 1 female, Cave Da Dong, Yangfan, Rongdu, Ceheng, Qianxinan Miao & Buyi Zizhizhou, southwest Guizhou, 105 ° 39 ’ 40 ” E / 25 ° 2 ’ 31 ” N, 1190 m, 2015 - VI- 20, Mingyi Tian, Weixin Liu, Xinhui Wang and Mingruo Tang leg.	en	Tian, Mingyi, Huang, Sunbin (2021): Discovery of a new cavernicolous trechine genus and species from a deep pit in Anlong, southwestern Guizhou Province (Coleoptera: Carabidae: Trechinae). Zootaxa 4985 (2): 194-202, DOI: https://doi.org/10.11646/zootaxa.4985.2.3
03F7A821D9513527FF67FA808D5966DF.taxon	distribution	Distribution: China (Guizhou). Known from four caves in Anlong: Xiniu Dong, Xianren Dong, Rongnei Dong (Uéno, 2006) and Luoshui Dong and Da Dong in Ceheng (Tian & Huang, 2018).	en	Tian, Mingyi, Huang, Sunbin (2021): Discovery of a new cavernicolous trechine genus and species from a deep pit in Anlong, southwestern Guizhou Province (Coleoptera: Carabidae: Trechinae). Zootaxa 4985 (2): 194-202, DOI: https://doi.org/10.11646/zootaxa.4985.2.3
