identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03B587C9FFB2AF2BB09CED0576BE59D4.text	03B587C9FFB2AF2BB09CED0576BE59D4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acerocnema Becker 1894	<div><p>Genus Acerocnema Becker, 1894</p> <p>[Japanese name: Kuchiboso-funbae-zoku]</p> <p>Acerocnema Becker, 1894, Berl. ent. Z. 39: 154. Type-species: Cordylura breviseta Zetterstedt, 1846: 2022, by original designation [= Cordylura macrocera (Meigen, 1826)].</p> <p>The Palaearctic species of the genus Acerocnema were revised and defined by Ozerov (2006, 2014) and Iwasa (2014). The Nearctic fauna was recently revised by Chagnon &amp; Sinclair (2020).</p> <p>Diagnosis. Small, about 3.0-6.0 mm in body length; head rounded in profile; gena and postgena wide; male postpedicel developed and rectangular; arista bare to short-haired and not longer than maximum length of postpedicel measured on its dorsolateral surface in male; proboscis slender and sclerotized with reduced labella; palpus basally slender and dilated apically; chaetotaxy (yellow to black): 3-4 fr, 2-3 or, 1 oc, 1 vti, 1 vte, 1 poc. Thorax yellow to black and shiny or white to grey micotrichose, covered with yellowish hairs; chaetotaxy (yellow): 1-2 pprn, 2 npl, 1+2 sa, 2+3 dc, 2 pa, 1 b sc, 1 d sc, 1 ap sc,1 prepst, 1 prepm, 2-8 anepst; 1 kepst; legs slender, normally without black or stout setae; wings hyaline without any brown spots or black setae; R 1 with or without dorsal setulae on apical portion; vein A 1 not reaching wing margin. Male abdomen: enlarged apically and covered with long yellowish hairs; sternite V deeply bilobate posteriorly; surstylus articulated with epandrium and long, slender, but varied in shape; aedeagus basally chitinous and apically membranous, and sometimes whip-like; cercal plate developed and divided into complex lobes. Female abdomen: segments VI to VIII compressed; ovipositor short, strongly sclerotized and compressed posteriorly.</p></div> 	http://treatment.plazi.org/id/03B587C9FFB2AF2BB09CED0576BE59D4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Iwasa, Mitsuhiro	Iwasa, Mitsuhiro (2021): Three new species and six new records of Scathophagidae (Diptera) from Japan with a key to the Japanese genera. Zootaxa 4981 (3): 531-553, DOI: https://doi.org/10.11646/zootaxa.4981.3.6
03B587C9FFB2AF29B09CE83674985977.text	03B587C9FFB2AF29B09CE83674985977.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acerocnema incisa Iwasa 2021	<div><p>Acerocnema incisa sp. nov.</p> <p>(Figs 1–6)</p> <p>[Japanese name: Kiashi-kuchiboso-funbae]</p> <p>Type material examined. Holotype ♂, Japan: Inada, Obihiro City, Tokachi, Hokkaido, 18. v. 2014, leg. H. Torikura (OUAVM). Paratypes: Hokkaido, 3 ♂, 4 ♀, same locality as holotype, 15-25. v. 2014, leg. H. Torikura (OUAVM); 1 ♂, Kawanishi, Obihiro City, Tokachi, 11. v. 2014, leg. H. Torikura (OUAVM); 6 ♂, 34 ♀, Otofuke Town, Tokachi, 18-21. v. 2017, leg. H. Torikura (OUAVM). Kyushu, 2 ♀, south of Ito campus, Kyushu Univ. Fukuoka City, 4. iv. 2015, leg. D. Kato (OUAVM).</p> <p>Diagnosis. Acerocnema incisa sp. nov. can be recognized by thorax with grey microtrichose, dorsal setulae on apical portion of vein R 1 of wing, and male surstylus anterodistally dilated</p> <p>Description. Male. Head (Figs 1, 2): eyes reddish-brown; frons reddish; ocellar triangle black; face, parafacial, and gena yellow; occiput black and shining; postpedicel black and enlarged, approximately 2.5 times long as wide; arista basally pubescent and apically short-haired, approximately same length as postpedicel; palpi yellow and clavate; proboscis slender and sclerotized with reduced labella; vibrissal angle with a strong yellow vibrissa. Chaetotaxy (black): 3-4 fr, 3 or (1 proclinate, 1 lateroclinate, 1 latero-reclinate), 1 oc, 1 poc, 1 vti (strong), 1 vte. Thorax: wholly black; prescutum and scutum thinly grey microtrichose, with two longitudinally shiny stripes on scutum and with inconspicuous hairs (Fig. 1); scutellum shining; all thoracic pleura grey microtrichose; proepisternum anteriorly with hairs; anepisternum posteriorly hairy; katepisternum sparsely with long hairs; anepimeron, meron, and katatergite bare; metanotum shining in lower part. Chaetotaxy (yellow): 2 pprn, 2 npl, 1+1 ial, 1 sa, 2 pa, 1+3 dc, 1 bsc, 1 apsc,1 prepst, 1 prepm, 6-7 anepst, 1 kepst. Wings: hyaline, slightly tinged with brown; veins brown; R 1 with dorsal setulae on apical portion; veins R 4+5 and M 1 almost parallel; vein A 1 +CuA 2 disappear well before wing margin; calypter yellow, with hairs marginally; halteres yellow. Legs: entirely yellow, covered with hair-like setulae; fore femur with several pd setae distally; mid and hind femora with preapical setae; fore tibiae with 1 ad at two-fifths; mid tibia 1 ad at near half and subapical setae; hind tibia with 2 ad at one-third and half, and subapical setae; all tarsi yellow; fifth tarsomere somewhat compressed. Abdomen: tergites black and shining, covered with short hairs; sternites II to IV longitudinally rectangular, sparsely covered with hair-like setulae; sternite V (Fig. 3) deeply bilobate posteriorly; epandrium (Fig. 4) developed and sparsely with hairs but without setae; surstyli (Figs 4, 5) long, slender, curved and dilated antero-apically; aedeagus (Fig. 4) slender, apically swollen and membranous; epiphallus flattened, curved and pointed apically; parameres rod-shaped; hypandrium large, thick, and V-shaped; cerci stout and membranous, covered with hairs.</p> <p>Female. Basal and ventral portions of postpedicel, postpronotum and proepisternum paler than those of male; abdominal tergites II to VI centrally black and laterally yellow to light brown; sternites III to VI longitudinally elongate; segments (tergites and sternites) VII to VIII (Fig. 6) compressed and tergite VIII strongly sclerotized; cerci, epiproct, and hypoproct weakly pigmented. Other characteristics as those of male.</p> <p>Length (♂ ♀). Body, 4.0–4.2 mm; wings, 4.0–4.3 mm.</p> <p>Distribution. Japan (Hokkaido and Kyushu).</p> <p>Etymology. This new species is named after the scientific name of the flower species which has a close relationship with this fly.</p> <p>Remarks. This new species is similar to A. kishimotoae Iwasa in general appearance, but differs from it in having the following characteristics: frons reddish; scutum grey microtrichose with inconspicuous hairs; male surstyli anterodistally dilated; female postpedicel same length as that of male. In A. kishimotoae, frons black; scutum shining, sparsely covered with distinct hairs; male surstyli gradually dilated toward the tip, not anterodistally; female postpedicel smaller than that of male.</p> <p>Bionomics. Adults were collected from a flower community of Corydalis incisa (Thumb.) Per. in early spring. The larvae probably penetrate into the seed capsules of this flower.</p> </div>	http://treatment.plazi.org/id/03B587C9FFB2AF29B09CE83674985977	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Iwasa, Mitsuhiro	Iwasa, Mitsuhiro (2021): Three new species and six new records of Scathophagidae (Diptera) from Japan with a key to the Japanese genera. Zootaxa 4981 (3): 531-553, DOI: https://doi.org/10.11646/zootaxa.4981.3.6
03B587C9FFB0AF29B09CEBD172815B18.text	03B587C9FFB0AF29B09CEBD172815B18.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acerocnema Becker 1894	<div><p>Key to the species of Japanese Acerocnema (male and females)</p> <p>1. R 1 of wing with dorsal setulae on apical portion............................................................. 2</p> <p>- R 1 of wing without dorsal setulae on apical portion........................................................... 3</p> <p>2. Prescutum, scutum and scutellum mostly grey microtrichose, with a pair of shiny stripes on scutum (Fig. 1); male surstylus anterodistally dilated (Fig. 4)............................................................... A. incisa sp. nov.</p> <p>- Prescutum, scutum and scutellum mostly shiny; male surstylus sinuate and tapering apically (see Iwasa (2014, figs 13, 14))........................................................................................ A. sinuata Iwasa</p> <p>3. Frons yellow; scutum longitudinally with a central grey microtrichose band; male surstylus filiform (see Iwasa (2014, figs 8, 9)).................................................................................. A. flavifrons Iwasa</p> <p>- Frons black; scutum wholly shining; surstylus clavate (see Iwasa (2014, figs 2, 3))................ A. kishimotoae Iwasa</p></div> 	http://treatment.plazi.org/id/03B587C9FFB0AF29B09CEBD172815B18	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Iwasa, Mitsuhiro	Iwasa, Mitsuhiro (2021): Three new species and six new records of Scathophagidae (Diptera) from Japan with a key to the Japanese genera. Zootaxa 4981 (3): 531-553, DOI: https://doi.org/10.11646/zootaxa.4981.3.6
03B587C9FFB0AF2EB09CE97373785D6B.text	03B587C9FFB0AF2EB09CE97373785D6B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chaetosa Coquillett 1898	<div><p>Genus Chaetosa Coquillett, 1898</p> <p>[Japanese name: Chibi-funbae-zoku]</p> <p>Chaetosa Coquillett, 1898: 163. Type-species: Cordylura punctipes Meigen, 1826: 239, by original designation.</p> <p>Diagnosis. Frons anteriorly yellow and posteriorly black; face, parafacial, and gena yellow; postpedicel yellow with acute apical corner, about 2.5 times as long as wide; arista pubescent; vibrissal angle with 1 vibrissa and 1 subvibrissa; palpus yellow without strong subapical seta; 2 fr, 3 or, 1 oc, 1 poc, 1 vte, 1 vti; thorax black; scutum and prescutum grey microtrichose: thoracic pleura greyish dusted; 1-2 pprn, 2 npl, 1+2 ial, 2+3 dc, 1+2 sa, 2 pa; 1 bsc, 1 apsc, 1 prepst, 1 prepm, 3 anepst, 2 kepst; vein A 1 +CuA 2 extending to wing margin; legs yellow; male sternite V bilobate posteriorly; male surstyli long and slender; male cerci with distinct long setae.</p></div> 	http://treatment.plazi.org/id/03B587C9FFB0AF2EB09CE97373785D6B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Iwasa, Mitsuhiro	Iwasa, Mitsuhiro (2021): Three new species and six new records of Scathophagidae (Diptera) from Japan with a key to the Japanese genera. Zootaxa 4981 (3): 531-553, DOI: https://doi.org/10.11646/zootaxa.4981.3.6
03B587C9FFB7AF2EB09CEFAC74125FAE.text	03B587C9FFB7AF2EB09CEFAC74125FAE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chaetosa punctipes (Meigen 1826)	<div><p>Chaetosa punctipes (Meigen, 1826)</p> <p>[Japanese name: Kita-chibi-funbae]</p> <p>(Figs 7–10)</p> <p>Cordylura punctipes Meigen, 1826: 239. Type-locality: not given.</p> <p>Material examined. Hokkaido, 2 ♂, 1 ♀, Tsurui Village, Kushiro, 14. vi. 1975, leg. H. Sasaki.</p> <p>Diagnosis. Chaetosa punctipes Meigen can be recognized by the following characteristics: postpedicel with acute upper corner (Fig. 7); thoracic pleura greyish dusted; legs wholly yellow; posterior lobes on sternite V with minute spinules on inner parts (Fig. 8); male surstyli slender and tapering apically (Fig. 9); male cercus with 3 long setae beyond surstyli (Figs 9, 10).</p> <p>Body length (♂). 4.3–4.9 mm.</p> <p>Distribution. Europe, Mongolia, North America, Russia (the European part, Siberia, the Far East) (Bagachanova et al., 2016), and Japan (Hokkaido). New to Japan.</p></div> 	http://treatment.plazi.org/id/03B587C9FFB7AF2EB09CEFAC74125FAE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Iwasa, Mitsuhiro	Iwasa, Mitsuhiro (2021): Three new species and six new records of Scathophagidae (Diptera) from Japan with a key to the Japanese genera. Zootaxa 4981 (3): 531-553, DOI: https://doi.org/10.11646/zootaxa.4981.3.6
03B587C9FFB7AF2EB09CEDEB760A5B70.text	03B587C9FFB7AF2EB09CEDEB760A5B70.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cleigastra Macquart 1835	<div><p>Genus Cleigastra Macquart, 1835</p> <p>[Japanese name: Hizaguro-funbae-zoku]</p> <p>Cleigastra Macquart, 1835: 384. Type-species: Cordylura apicalis Meigen, 1826: 236, by designation of Westwood, 1840.</p> <p>Cnemopogon Rondani, 1856: 100. Type-species: Cordylura apicalis Meigen, 1826: 236, by original designation.</p> <p>Nanna Strobl, 1894: 77 [as subgenus of Cordilura Fallén, 1810]. Type-species: Cordylura flavipes Fallén, 1819, by designation of Vockeroth, 1965.</p> <p>Amaurosoma Becker, 1894: 109. Type-species: Cordylura flavipes Fallén, 1819, by original designation.</p> <p>Spathephilus Becker, 1894: 121. Type-species: Cordylura breviventris Loew, 1873, by original designation.</p> <p>Pselaphephila Becker, 1894: 122. Type-species: Pselaphephila loewi Becker, 1894, by monotypy.</p> <p>Monochaeta Becker, 1894: 87, 186. Type-species: Cordylura breviventris Loew, 1873, by original designation.</p> <p>Orthacheta Becker, 1894: 101. Type-species: Cordylura pilosa Zetterstedt, 1838, by original designation.</p> <p>Gonatherus Rondani, 1856: 99. Type-species: Cordilura planiceps Fallén, 1826, by original designation.</p> <p>Ozerov &amp; Krivosheina (2015) reviewed the genus Cleigastra Macquart and six related genera (Gonarcticus Becker, Gonatherus Rondani, Nana Strobl, Orthacheta Becker, and Spathephilus Becker) in Russia. Thereafter, Ozerov and Krivosheina (2016) proposed that Cleigastra comprises three subgenera Cleigastra sensu strico, Gonatherus, and Orthacheta. On the basis of these previous studies and the Japanese specimens, the characteristics of Cleigastra are defined as follows.</p> <p>Diagnosis. Frons black; face, parafacial and gena pale yellow or silvery microtrichose; 3-4 fr, 3 or, 1 oc, 1 poc,1 vti, 1 vte; postpedicel long approximately 4-4.5 times as long as wide, with acute upper corner; 1 vibrissa and 2 subvibrissae; thorax black, greyish dusted; 2 pprn, 2 npl, 1+2 ial, 2 sa, 2 pa, (2-3)+3 dc, 1 prepst, 1 prepm, 3 kpst, 1 dsc, 1 apsc; wings infuscate; vein A 1 +CuA 2 complete; legs black, brown to yellow; fore femur with rows of dorsal, posterodorsal, and sometimes with anteroventral strong setae; abdominal tergites black, greyish dusted, covered with hairs and setulae, and each segment often lateral and posterior strong setae; male sternite V distinctly bilobate posteriorly; male surstyli comparatively simple; male cerci fused to each other; female ovipositor long and normally cylindrical.</p></div> 	http://treatment.plazi.org/id/03B587C9FFB7AF2EB09CEDEB760A5B70	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Iwasa, Mitsuhiro	Iwasa, Mitsuhiro (2021): Three new species and six new records of Scathophagidae (Diptera) from Japan with a key to the Japanese genera. Zootaxa 4981 (3): 531-553, DOI: https://doi.org/10.11646/zootaxa.4981.3.6
03B587C9FFB6AF2FB09CEAB776525B0D.text	03B587C9FFB6AF2FB09CEAB776525B0D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cleigastra apicalis (Meigen 1826)	<div><p>Cleigastra apicalis (Meigen, 1826)</p> <p>[Japanese name: Hizaguro-funbae]</p> <p>(Figs 11–14)</p> <p>Cordylura apicalis Meigen, 1826: 236. Type-locality: not given.</p> <p>Material examined. Honshu, 2 ♂, 1 ♀, Mt. Hayachine, Iwate Pref., 17-18. vi. 1971, leg. S. Shinonaga; 1 ♂, Sawauchi Vill., Iwate Pref., 19. v. 1991, leg. T. Hasegawa; 1 ♂, 2 ♀, Mt. Shiritaka, Ishikawa Pref., 23. v. 1971, leg. S. Shinonaga; 1 ♂, Mt. Hakusan, Ishikawa Pref., 24. v. 1971, leg. S. Shinonaga.</p> <p>Diagnosis. This species can be characterized as follows: face, parafacial, and gena distinctly silvery shiny; fore coxae yellow; scutellum with apsc and dsc setae (Fig. 11); legs very bristled; fore femur yellow often with black stripe anteriorly with long black and pale hairs ventrally; female fore femur without pale hairs; mid and hind femora with black apical rings; posterior lobes on male sternite V with long hairs (Fig. 12); male surstyli apically broad in posterior view (Fig. 14); male cerci sclerotized and projected dorsally (Figs 13, 14).</p> <p>Body length (♂ ♀). 7.8–8.2 mm.</p> <p>Distribution. Europe (widespread), Iraq, Mongolia, Russia (widespread) (Ozerov &amp; Krivosheina, 2015), and Japan (Honshu).</p></div> 	http://treatment.plazi.org/id/03B587C9FFB6AF2FB09CEAB776525B0D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Iwasa, Mitsuhiro	Iwasa, Mitsuhiro (2021): Three new species and six new records of Scathophagidae (Diptera) from Japan with a key to the Japanese genera. Zootaxa 4981 (3): 531-553, DOI: https://doi.org/10.11646/zootaxa.4981.3.6
03B587C9FFB5AF22B09CE891749B5E33.text	03B587C9FFB5AF22B09CE891749B5E33.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cleigastra tibiella (Zetterstedt 1838)	<div><p>Cleigastra tibiella (Zetterstedt, 1838)</p> <p>[Japanese name: Togeashi-hizaguro-funbae]</p> <p>(Figs 15–20)</p> <p>Cordylura tibiella Zetterstedt, 1838: 731. Type-locality: “Juckasjervi Lapponiae Tornensis, ad Lycksele Lappon. Umensis, ut et ad Bjŏrkvik Nordlandiae… Dowre… Uplandia ad Nacka prope Stockhomiam… Smoland…” [Sweden, Norway].</p> <p>Cordylura nigripes Zetterstedt, 1846: 2026. Type-locality: “Smolandia… Jemtlandia boreali ad diversorium Skalstugan… Lapponia Lulensi” [Sweden].</p> <p>Amaurosoma nigrifrontata Becker, 1894: 120. Type-locality: “Sud-Tirol vom Luisier Pass” [now Passo di Lusia (Italy, Trentino)].</p> <p>Amaurosoma mutans Becker, 1894: 120. Type-locality: Legnica (formerly Liegnitz), Poland.</p> <p>Amaurosoma kamtschatkense Hendel, 1930: 9. Type-locality: “Klutchi” [Russia, Kamchatka Kray]</p> <p>Material examined. Hokkaido, 2 ♂, Kushiro, 16. v. 1988, leg. M. Iwasa (OUAVM).</p> <p>Diagnosis. Cleigastra tibiella (Zetterstedt) can be recognized by the following characteristics: frons black; face, parafacial, and genae pale yellow with silvery microtrichose; postpedicel black with acute upper corner (Fig. 15); thorax and abdominal tergites grey dusted (Fig. 15); katepisternum with 3 setae (Fig. 16); fore femora blackish with 7-10 distinct av setae (Fig. 17); male sternite V bilobate posteriorly (Fig. 18); male surstyli filiform, apically rounded, arched close to each other at the tips (Figs 19, 20).</p> <p>Body length (♂). 4.7–4.8 mm.</p> <p>Distribution. Europe, Japan (Hokkaido), Russia (Kamchatka Kray, Kranoyarsk Kray, Leningrad Oblast and St.- Petersburg, Magadan Oblast, Moscow Oblast, Sakhalin Oblast, Tumen’ Oblast) (Ozerov &amp; Krivosheina, 2015). New to Japan.</p> <p>Remarks. This species is very similar to C. puberula (Becker) in the structure of male fore femora and surstyli, but can be easily distinguished from it by the blackish fore femora.</p> </div>	http://treatment.plazi.org/id/03B587C9FFB5AF22B09CE891749B5E33	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Iwasa, Mitsuhiro	Iwasa, Mitsuhiro (2021): Three new species and six new records of Scathophagidae (Diptera) from Japan with a key to the Japanese genera. Zootaxa 4981 (3): 531-553, DOI: https://doi.org/10.11646/zootaxa.4981.3.6
03B587C9FFBBAF22B09CEC947666591F.text	03B587C9FFBBAF22B09CEC947666591F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gimnomera Rondani 1866	<div><p>Genus Gimnomera Rondani, 1866</p> <p>[Japanese name: Shiributo-funbae-zoku]</p> <p>Gimnomera Rondani, 1866: 21 (1867: 135). Type-species: Cordylura tarsea Fallén, 1819, by original designation. Cochliarium Becker, 1894: 183. Type-species: Cordylura cuneiventris Zetterstedt, 1846, by original designation.</p> <p>Dasypleuron Malloch, 1919: 79. Type-species: Dasypleuron tibialis Malloch, 1919, by original designation.</p> <p>Paragymnomera Hendel, 1930: 80. Type-species: Gymnomera hirta Hendel, 1930, by original designation.</p> <p>Rhopochilus Enderlein, 1936: 136. Type-species: Cochliarum lasiostoma Becker, 1894, by monotypy.</p> <p>Langechristia Ozerov, 1999: 510. Type-species: Langechristia ziegleri Ozerov, 1999, by original designation.</p> <p>Diagnosis. Head variable from yellow to blackish; occiput covered with black setae in upper part and pale hairs in lower part; 2-5 fr, 2-3 or, 1 oc, 1 poc, 1 vte, 1 vti; postpedicel approximately 1.5-2 times as long as wide; arista pubescent or short-haired; palpi filiform, without long apical setae; 1 vibrissa and 1-2 subvibrissae; anterior margin of postpronotum with or without short erect spinules; 1-2 pprn, 0-2 npl, (0-1)+(0-1) ial; (0-1)+(1-2) sa; 1-2 pa, (0-2)+(1-3) dc, 1 apsc; 1 bsc or 1 dsc; 1 prepst; 0-1 prepm; 1-3 anepst; 1 ketpst; wings clear or slightly brownish; R 1 vein bare or setulose on apical half of dorsal surface; sternite V usually with median lobes, rarely with long triangular lateral lobes; male epandrium incompletely fused to surstyli; male surstyli often broadened apically; male cerci fused apically or medially forming cercal plate; female ovipositor short, compressed laterally, and tergite VIII strongly sclerotized.</p></div> 	http://treatment.plazi.org/id/03B587C9FFBBAF22B09CEC947666591F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Iwasa, Mitsuhiro	Iwasa, Mitsuhiro (2021): Three new species and six new records of Scathophagidae (Diptera) from Japan with a key to the Japanese genera. Zootaxa 4981 (3): 531-553, DOI: https://doi.org/10.11646/zootaxa.4981.3.6
03B587C9FFBBAF20B09CEB7977A258E7.text	03B587C9FFBBAF20B09CEB7977A258E7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gimnomera torikurai Iwasa 2021	<div><p>Gimnomera torikurai sp. nov.</p> <p>(Figs 21–27)</p> <p>[Japanese name: Torikura-shiributo-funbae]</p> <p>Type material examined. Holotype ♂, JAPAN: Kawanishi, Obihiro, Tokachi, Hokkaido, 29. iv. 2017, leg. H. Torikura (OUAVM). Paratypes: 1 ♂, same data as holotype (OUAVM); 1 ♂, same locality as holotype, 9. v. 2014, leg. H. Torikura (OUAVM); 2 ♂, 3 ♀, same locality, 1-7. v. 2015, leg. H. Torikura; 2 ♂, same locality, 30. iv. 2019, leg. H. Torikura (OUAVM); 1 ♂, Obihiro, Tokachi, 16. v. 2015, leg. H. Torikura (OUAVM); 1 ♀, Makubetsu, Tokachi, 27. iv. 2017, leg. H. Torikura; 4 ♂, Toyokoro, Tokachi, 4. v. 2013, leg. M. Iwasa (OUAVM); 1 ♂, 1 ♀, Obihiro, Tokachi, 16. v. 2000, leg. T. Watanabe (OUAVM); 1 ♀, same locality, 1. v. 2000, leg. T. Watanabe (OUAVM).</p> <p>Diagnosis. Gimnomera torikurai sp. nov. can be recognized by the black thoracic pleura with greyish dusting and the shape of male surstylus.</p> <p>Description. Male (Fig. 21). Head (Fig. 22): eyes reddish brown; frontal vitta reddish yellow; fronto-orbital plate and ocellar triangle black; face light brown with silvery microtrichose; parafacial yellow; genae narrow and yellow to grey with silvery microtrichose; postgenae and occiput dark grey microtrichose; postpedicel dark brown and approximately 2 times as long as wide; pedicel yellow; arista pubescent; palpi yellow and clavate, covered with short setulae; proboscis stout, with reduced labella; vibrissal angle with 1 strong and 2-3 subvibrissae. Chaetotaxy (black): 5 fr (medioclinate), 2 or (1 lateroclinate, 1 latero-reclinate), 1 oc, 1 poc, 1 vte, 1 vti. Thorax: wholly black with grey dusted; scutum with 4 longitudinal black stripes; postpronotum without erect black spinules on the anterior margin; proepisternum covered with hairs; anepisternum posteriorly with hairs; katepisternum sparsely with hairs; anepimeron, meron, and katatergite bare; metanotum bare and grey microtrichose. Chaetotaxy (black): 2 pprn, 2 npl, 1+2 ial; 2 sa, 2 pa, 2+3 dc, 1 dsc, 1 apsc; 1 prepm (yellow), 2-3 anepst, 1 kepst. Wings: hyaline, slightly tinged with brown; veins brown; vein R 1 with dorsal setulae on apical portion; veins R 4+5 and M 1 almost parallel; vein A 1 +CuA 2 abruptly end well before wing margin; calypter whitish yellow with hairs marginally; halteres yellow. Legs: coxae yellow to brown, femora and tibiae entirely yellow; fore femur ventrally covered with hairs; fore tibia with 1 apical d seta; mid tibia with apical d seta; hind tibia with 1 ad and 1 pd at middle, 1 pd basal one-third, 1 apical d, and 3-4 preapical setae; tarsi yellow, darkened apically. Abdomen: tergites I-V black and VI-VIII brown, covered with hairs; tergites I-II grey microtrichose and III-VIII rather shiny; sternite V (Fig. 23) with two broad median posterior lobes, covered with short spines; epandrium (Figs. 24, 25) sparsely with hairs but without setae; surstylus (Figs 24, 25) long and slender, tapered apically; aedeagus (Fig. 26) slender and apically membranous; epiphallus rod-shaped and slightly curved; paramere clavate, with setulae posteroapically; hypandrium strongly sclerotized and V-shaped; cercus (Figs 24, 25) somewhat slender, apically close to each other, sparsely covered with hairs and posteriorly with marginal setulae.</p> <p>Female. Abdominal sternites III to VI rounded square; segments VII to VIII sternites VII longitudinally elongate; segments (tergites and sternites) VII to VIII compressed and tegites VIII and sternite VIII strongly sclerotized and pigmented (Fig. 27); cercus, epiproct and hypoproct weakly sclerotized (Fig. 27). Other characteristics as those of male.</p> <p>Length (♂ ♀). Body, 6.0–6.5 mm; wings, 5.0 mm.</p> <p>Distribution. Japan (Hokkaido).</p> <p>Etymology. This new species is named after Mr. H. Torikura who first collected this species.</p> <p>Remarks. Ozerov (2019) synonymized Langechristia Ozerov with Gimnomera in his review of the Russian Gimnomera. Gimnomera torikurai sp. nov. belongs to the Langechristia group (G. amica (Ozerov), G. speciosa (Ozerov), and G. ziegleri (Ozerov)), which has no short erect black spinules on the anterior margin of the postpronotum. The present new species differs from other species of the Langechristia group in having the black thoracic pleura with greyish dusting and by the structure of male surstyli and cerci.</p> <p>Bionomics. This new species were collected from both flowers communities of Corydalis ambigua Cham. et Schltdl and C. incisa (Thunb.) Per. in early spring. The larvae possibly develop in the seed capsules of these flowers.</p> </div>	http://treatment.plazi.org/id/03B587C9FFBBAF20B09CEB7977A258E7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Iwasa, Mitsuhiro	Iwasa, Mitsuhiro (2021): Three new species and six new records of Scathophagidae (Diptera) from Japan with a key to the Japanese genera. Zootaxa 4981 (3): 531-553, DOI: https://doi.org/10.11646/zootaxa.4981.3.6
03B587C9FFB9AF20B09CEB2073CF5A9D.text	03B587C9FFB9AF20B09CEB2073CF5A9D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Megaphthalma Becker 1894	<div><p>Genus Megaphthalma Becker, 1894</p> <p>[Japanese name: Kiiro-funbae-zoku]</p> <p>Megaphthalma Becker, 1894: 105. Type-species: Scatomyza pallida Fallén, 1819, by original designation.</p> <p>Diagnosis. Head yellowish except for eyes; 3 fr, 3 or (1 proclinate, 2 latero-reclinate), 1 oc, 1 poc, 1 vte, 1 vti, vibrissal angle with 1 vibrissa; postpedicel short and rounded apically; palpi slender and clavate; scutum yellow to dark brown; thoracic sclerites yellow; 2 pprn, 2 npl, 1+2 ial, 1+2 sa, 2 pa, 2 +3 dc, 1 dsc; apsc absent; 1 prepst and 1 prepm; 4 anepst, 1 kepst; proepimeron sparsely with yellowish hairs; abdominal tergites yellow; male sternite V prominently bilobate posteriorly; male surstyli long and slender; male cerci clothed with setulae.</p></div> 	http://treatment.plazi.org/id/03B587C9FFB9AF20B09CEB2073CF5A9D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Iwasa, Mitsuhiro	Iwasa, Mitsuhiro (2021): Three new species and six new records of Scathophagidae (Diptera) from Japan with a key to the Japanese genera. Zootaxa 4981 (3): 531-553, DOI: https://doi.org/10.11646/zootaxa.4981.3.6
03B587C9FFB9AF21B09CE8FF767B5DDF.text	03B587C9FFB9AF21B09CE8FF767B5DDF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Megaphthalma pallida (Fallen 1819)	<div><p>Megaphthalma pallida (Fallén, 1819)</p> <p>[Japanese name: Seobi-kiiro-funbae]</p> <p>(Figs 28–29)</p> <p>Cordylura pallida Fallén, 1819: 8. Type-locality: “Westrogothis… Scania ” [Sweden].</p> <p>Material examined. Hokkaido, 2 ♀, Iwanai Riv., Obihiro, Tokachi, 19. ix. 1995, leg. M. Iwasa. Diagnosis. This species is recognized by the following characteristics: frons, face, parafacial, and gena yellow; prescutum and scutum dark brown; postpronotum and thoracic pleura yellow; scutellum yellow with a pair of apical setae (Fig. 28); legs wholly yellow; abdominal tergites wholly yellow with narrow black line on each posterior margin (Fig. 29).</p> <p>Body length (♀). 5.5–6.0 mm.</p> <p>Distribution. Europe, Japan (Hokkaido), Russia (the European part, Siberia, the Far East) (Bagachanova et al., 2016), North America. New to Japan.</p> <p>Remarks. Available specimens of this species are females, but as a result of my examination of specimens of M. pallida preserved in the Museum National d’Histoire naturelle (Paris), the characteristics agreed with those of female M. pallida.</p> </div>	http://treatment.plazi.org/id/03B587C9FFB9AF21B09CE8FF767B5DDF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Iwasa, Mitsuhiro	Iwasa, Mitsuhiro (2021): Three new species and six new records of Scathophagidae (Diptera) from Japan with a key to the Japanese genera. Zootaxa 4981 (3): 531-553, DOI: https://doi.org/10.11646/zootaxa.4981.3.6
03B587C9FFB8AF21B09CEB6075C15AF9.text	03B587C9FFB8AF21B09CEB6075C15AF9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Megaphthalmoides Ringdahl 1936	<div><p>Genus Megaphthalmoides Ringdahl, 1936</p> <p>[Japanese name: Sesuji-funbae-zoku]</p> <p>Megaphthalmoides Ringdahl, 1936, 179. Type-species: Cordylura unilineata Zetterstedt, 1838: 727, by original designation.</p> <p>Diagnosis. Head yellowish except for eyes; 2-3 fr, 3 or (1proclinate, 2 latero-reclinate), 1 oc, 1 poc, 1 vte, 1 vti, vibrissal angle with 1 vibrissa and 1 subvibrissa (small); postpedicel short and rounded apically; palpi filiform and whitish; thorax yellow to light brown; scutum and prescutum often with dark longitudinal stripes; 2 pprn, 2 npl, 1+2 ial, 1+2 sa, 2 pa, 2+3 dc, 1 dsc, 1 apsc; 1 prepst and 1 prepm; proepiperon with yellowish hairs, 2-4 anepst, 1 kepst; abdomen basally flat and apically cylindrical; male sternite V prominently bilobate posteriorly; male surstyli long and slender; male cerci covered with strong setulae.</p></div> 	http://treatment.plazi.org/id/03B587C9FFB8AF21B09CEB6075C15AF9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Iwasa, Mitsuhiro	Iwasa, Mitsuhiro (2021): Three new species and six new records of Scathophagidae (Diptera) from Japan with a key to the Japanese genera. Zootaxa 4981 (3): 531-553, DOI: https://doi.org/10.11646/zootaxa.4981.3.6
03B587C9FFB8AF26B09CE953742C5E30.text	03B587C9FFB8AF26B09CE953742C5E30.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Megaphthalmoides japonicus Ozerov 2008	<div><p>Megaphthalmoides japonicus Ozerov, 2008</p> <p>[Japanese name: Sesuji-funbae]</p> <p>(Figs 30–33)</p> <p>Megaphthalmoides japonicus Ozerov, 2008: 420. Type-locality: ”Izumi Katswiagi”, Osaka, Japan.</p> <p>Material examined. Hokkaido, 7 ♂, 1 ♀, Shirogane-onsen, Biei, 26. viii. 1979, leg. M. Iwasa; 1 ♂, Mt. Ponyaoromappu, Hidaka range, 11. viii. 1998, leg. M. Iwasa; 1 ♀, Mt. Tsurugi, Hidaka range, 11. viii. 1998, leg. M. Iwasa. Honshu, 3 ♂, Mt. Hayachine, Iwate Pref., 17-18. vi. 1971, leg. S. Shinonaga; 1 ♀, Karuizawa, Nagano Pref., 11. viii. 1970, leg. R. Kano; 1 ♀, Mt. Houou, Yamanashi Pref., 6. vi. 1972, leg. S. Shinonaga. Kyushu, 1 ♂, Izumimachimoniki, Yatsushiro City (alt. 1350 m), Kumamoto Pref., 11. v. 2016, leg. D. Kato.</p> <p>Diagnosis. This species is a large species and characterized as follows:characteristics: head yellow except for eyes; postpedicel with long-haired arista (Fig. 30); thorax yellow with 2 longitudinal stripes on prescutum to scutum; abdominal tergites blackish, covered with hair-like setulae and strong lateral setae; male sternite V posterior lobes which (Fig. 31) are basally broadened; male surstyli (Figs 32, 33) long and slender, with rows of setulae posteriorly; male cerci clothed with strong setulae.</p> <p>Body length (♂ ♀). 7.8–10.0 mm.</p> <p>Distribution. Japan (Hokkaido, Honshu, and Kyushu).</p></div> 	http://treatment.plazi.org/id/03B587C9FFB8AF26B09CE953742C5E30	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Iwasa, Mitsuhiro	Iwasa, Mitsuhiro (2021): Three new species and six new records of Scathophagidae (Diptera) from Japan with a key to the Japanese genera. Zootaxa 4981 (3): 531-553, DOI: https://doi.org/10.11646/zootaxa.4981.3.6
03B587C9FFBFAF27B09CE8AA76275E33.text	03B587C9FFBFAF27B09CE8AA76275E33.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Megaphthalmoides unilineatus (Zetterstedt 1838)	<div><p>Megaphthalmoides unilineatus (Zetterstedt, 1838)</p> <p>[Japanese name: Kiiro-sesuji-funbae]</p> <p>(Figs 34–37)</p> <p>Cordylura unilineata Zetterstedt, 1838: 727. Type-locality: “Nordlandia…Bjorkvik” [Norway].</p> <p>Material examined. Hokkaido, 3 ♂, 2 ♀, Mt. Kurodake (1900 m), Kamikawa, 24.vii. 1996, leg. M. Iwasa; 1♂, Kogen-onsen (1300 m), Mt. Daisetsu, Kamikawa, 21. vii. 2016, leg. M. Iwasa; 3 ♀, same locality, 6 viii. 2016, leg. M. Iwasa; 1 ♂, same locality, 2. viii. 2012, leg. S. Shinonaga; 1 ♀, Ginsendai (1500 m), Mt. Daisetsu, Kamikawa, 5. viii. 1998, leg. M. Iwasa; 1 ♀, Nukabira, 28. vii. 1987, Tokachi, leg. M. Iwasa; 1 ♀, Mt. Tsurugi, Hidaka range, Tokachi, 25. vii. 1978, leg. M. Iwasa.</p> <p>Diagnosis. This species is medium-sized species and characterized as follows: head yellow except for eyes; postpedicel black in apical two-thirds and yellow in basal one-third (Fig. 34); arista short-haired; thorax yellow with a longitudinal stripe on prescutum to scutum; abdominal tergites yellow and each tergite with a horizontal black line on posterior margin; male sternite V bilobate posteriorly (Fig. 35); male surstyli long, slender, and tapered apically (Figs 36, 37); cerci clothed with strong setulae.</p> <p>Distribution. Europe, Russia (the European part, Siberia, the Far East), North America (Ozerov and Krivosheina 2014), Japan (Hokkaido). New to Japan.</p> <p>Body length (♂ ♀). 5.5–6.0 mm.</p> <p>Remarks. This species is easily distinguished from M. japonicus Ozerov by the characteristics mentioned in the key below.</p> </div>	http://treatment.plazi.org/id/03B587C9FFBFAF27B09CE8AA76275E33	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Iwasa, Mitsuhiro	Iwasa, Mitsuhiro (2021): Three new species and six new records of Scathophagidae (Diptera) from Japan with a key to the Japanese genera. Zootaxa 4981 (3): 531-553, DOI: https://doi.org/10.11646/zootaxa.4981.3.6
03B587C9FFBDAF24B09CE82D72815B09.text	03B587C9FFBDAF24B09CE82D72815B09.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Megaphthalmoides Ringdahl 1936	<div><p>Key to the species of Japanese Megaphthalmoides (male and females)</p> <p>1. Large-sized, 7.8–10.0 mm in body length; postpedicel wholly yellow; arista plumose (Fig. 30)........ M. japonicus Ozerov</p> <p>- Medium-sized, 5.0–6.0 mm in body length; postpedicel wholly or partly black; arista short-haired..................... 2</p> <p>2. Frontal vitta with a distinct black pattern (Fig. 39); postpedicel wholly black (Fig. 40)................ M. vittatus sp. nov.</p> <p>- Frontal vitta wholly yellow without a black pattern; postpedicel black in distal two-thirds and yellow in basal one-third (Fig. 34)............................................................................ M. unilineatus Zetterstedt</p></div> 	http://treatment.plazi.org/id/03B587C9FFBDAF24B09CE82D72815B09	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Iwasa, Mitsuhiro	Iwasa, Mitsuhiro (2021): Three new species and six new records of Scathophagidae (Diptera) from Japan with a key to the Japanese genera. Zootaxa 4981 (3): 531-553, DOI: https://doi.org/10.11646/zootaxa.4981.3.6
03B587C9FFBDAF24B09CEE8D75B959EB.text	03B587C9FFBDAF24B09CEE8D75B959EB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Megaphthalmoides vittatus Iwasa 2021	<div><p>Megaphthalmoides vittatus sp. nov.</p> <p>[Japanese name: Kurotsuno-sesuji-funbae]</p> <p>(Figs 38–43)</p> <p>Type material examined. Holotype ♂, JAPAN: Kamikochi, Azumimura, Nagano Pref., Honshu, 3-4. x. 1988, leg. S. Shinonaga (OUAVM). Paratypes: Honshu, 1 ♂, Tôshichi Spa, Hachimantai City, Iwate Pref., 3. viii. 2013, leg. D. Kato (OUAVM); 1 ♂, Hatomati-toge, Oze, Gumma Pref. 7. vii. 1951, leg. H. Hasegawa (IAES); 1 ♂, Gumma Pref., 23. ix. 1950, leg. N. Fukuhara (IAES).</p> <p>Diagnosis. Megaphthalmoides vittatus sp. nov. can be recognized by frontal vitta with a distinct black pattern and wholly black postpedicel.</p> <p>Description. Male (Fig. 38). Head (Fig. 39): eyes reddish brown; frontal vitta with a distinct black pattern; fronto-orbital plate yellow; ocellar triangle dark; face yellow and shiny; parafacial very narrow; gena and postgena yellow slightly with silvery microtrichose; occiput yellow and shiny; postpedicel (Fig. 40) black and approximately 2 times as long as wide; pedicel yellow; arista short-haired; palpus yellow and filiform, sparsely with short setulae anteroventrally; proboscis yellow to brown; vibrissal angle with 1 strong seta and 1 subvibrissae. Chaetotaxy (black): 2-3 fr (medioclinate), 2 or (1 proclinate, 1 latero-reclinate), 1 oc, 1 vti, 1 vte, 1 poc. Thorax: wholly yellow and shiny; scutum and prescutum with weak longitudinal stripe; proepisternum bare; anepisternum, katepisternum and anepimeron sparsely covered with short hair-like setulae; meron and katatergite bare; metanotum bare and slightly silvery microtrichose. Chaetotaxy (black): 2 pprn, 2 npl, 1+2 ial; 2 sa, 2 pa, 2+3 dc, 1 bsc, 1 apsc; 1 prepm, 2-1 (2 long, 1 short) anepst, 1 kepst. Wings: hyaline, slightly tinged with brown; veins brown; veins R 4+5 and M 1 almost parallel; vein A 1 +CuA 2 reach wing margin; calypter yellow with hairs marginally; halter yellow. Legs: coxae yellow to light brown; femora and tibiae yellow; fore femur yellow, ventrally and posterodorsally covered with hairs and setulae; mid femur apically black, with several a setulae; hind femur apically black with a row of ad setae; fore tibia with 1 p seta at basal two-fifths, 1 d seta near half and 1 preapical pd and 1 preapical d; mid tibia with 1 ad and 1 av setae at distal one-third and 1 apical av and 1 apical pv; hind tibia with 3 ad and 3 pd setae at regular intervals and 1 apical av; tarsi dark brown. Abdomen: tergites brown to dark brown and shiny, covered with hair-like setulae and distinct setae at posterior margin of each segment; sternite V (Fig. 41) bilobate posteriorly, covered with long and short setae on lobes; epandrium (Figs 42, 43) sparsely covered with short setae; surstylus (Figs 42, 43) long and slender, tapered apically; aedeagus stout and sclerotized; epiphallus rod-shaped and tapering apically; paramere comparatively thick with 1 short setula posteroventrally; hypandrium sclerotized and rounded, rhombus-shaped; cercus (Figs. 42, 43) somewhat elongated, covered with hair-like setulae.</p> <p>Length (♂). Body, 5.0–5.2 mm; wings, 5.8 mm.</p> <p>Female. Unknown.</p> <p>Distribution. Japan (Honshu).</p> <p>Etymology. The scientific name derives from the Latin vitta meaning band, referring the presence of a distinct black band on the frons of the head.</p> <p>Remarks. This species is closely similar to M. unilineatus (Zetterstedt), but can be distinguished from it by characteristics mentioned in the following key.</p> </div>	http://treatment.plazi.org/id/03B587C9FFBDAF24B09CEE8D75B959EB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Iwasa, Mitsuhiro	Iwasa, Mitsuhiro (2021): Three new species and six new records of Scathophagidae (Diptera) from Japan with a key to the Japanese genera. Zootaxa 4981 (3): 531-553, DOI: https://doi.org/10.11646/zootaxa.4981.3.6
03B587C9FFA3AF3AB09CEE8D740C5E00.text	03B587C9FFA3AF3AB09CEE8D740C5E00.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microprosopa Becker 1894	<div><p>Genus Microprosopa Becker, 1894</p> <p>[Japanese name: Hoso-funbae-zoku]</p> <p>Microprosopa Becker, 1894, Berl. Ent. Z., 39:147. Type-species: Cordylura haemorrhoidalis Meigen, 1826: Syst. Beschr., 5: 237, by original designation.</p> <p>Diagnosis. Head blackish, only gena partly whitish; 2 fr, 3 or (2 proclinate, 1 latero-reclinate), 1 oc, 1 poc, 1 vte, 1 vti, vibrissal angle with 1 vibrissa and 1 subvibrissa; postpedicel short and rounded apically; palpi spatulate and whitish; proepimeron with yellowish hairs; 2 pprn, 2 npl, 1+2 ial, 1+2 sa, 2 pa, (1-2) +3 dc, 1 bsc, 1 apsc; 1 prepst (yellow) and 1 prepm (yellow); 3 anepst, 1 kepst; abdomen slender and elongate; male sternite V prominently bilobate posteriorly; male surstyli stout and tapered apically.</p></div> 	http://treatment.plazi.org/id/03B587C9FFA3AF3AB09CEE8D740C5E00	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Iwasa, Mitsuhiro	Iwasa, Mitsuhiro (2021): Three new species and six new records of Scathophagidae (Diptera) from Japan with a key to the Japanese genera. Zootaxa 4981 (3): 531-553, DOI: https://doi.org/10.11646/zootaxa.4981.3.6
03B587C9FFA3AF3BB09CEC7A77C85E6F.text	03B587C9FFA3AF3BB09CEC7A77C85E6F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microprosopa haemorrhoidalis (Meigen 1826)	<div><p>Microprosopa haemorrhoidalis (Meigen, 1826)</p> <p>[Japanese name: Miyama-hoso-funbae]</p> <p>(Figs 44–48)</p> <p>Cordylura haemorrhoidalis Meigen, 1826: 237. Type-locality: “ Schweden ”.</p> <p>Cordylura pallipes (Zetterstedt, 1838): 732. Type-localities: “Lapponia; Scania ” [Sweden].</p> <p>Cordylura lividepes Zetterstedt, 1848: 2042. Type-locality: “Lapponia Lulensi ad Qvickjock” [Sweden].</p> <p>Material examined. Hokkaido, 3♂, Mt. Kaundake-Tenninkyo, Mt. Daisetsu-zan, 28. vii. 1983, leg. M. Iwasa.</p> <p>Diagnosis. This species is comparatively smaller-sized species and characterized as follows: frons black in anterior and posterior parts and yellow in central part; postpedicel black; arista pubescent; thorax black with grey dusting; katepisternum (Fig. 44) with a single seta; fore tibia ventrally with a row of short black spines (Fig. 45); abdominal tergites black and subshiny; legs yellow; posterior lobes on male sternite V tapered apically and curved inward (Fig. 46); male surstyli broad, tapering apically in lateral view (Figs. 47); male cerci covered with setulae, apically close to each other (Fig. 48).</p> <p>Body length (♂ ♀). 5.5–5.6 mm.</p> <p>Distribution. Europe, Russia (The European part, Siberia, the Far East), North America (Bagachanova et al., 2016), Japan (Hokkaido). New to Japan.</p> <p>Remarks. This species can be easily distinguished from other known species of the Microprosopa by having posterior lobes on sternite V which are tapered apically and curved inward, and broad surstyli in posterior view in male.</p> </div>	http://treatment.plazi.org/id/03B587C9FFA3AF3BB09CEC7A77C85E6F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Iwasa, Mitsuhiro	Iwasa, Mitsuhiro (2021): Three new species and six new records of Scathophagidae (Diptera) from Japan with a key to the Japanese genera. Zootaxa 4981 (3): 531-553, DOI: https://doi.org/10.11646/zootaxa.4981.3.6
03B587C9FFA2AF38B09CECA974485D4F.text	03B587C9FFA2AF38B09CECA974485D4F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microprosopa pallidicauda (Zetterstedt 1838)	<div><p>Microprosopa pallidicauda (Zetterstedt, 1838)</p> <p>[Japanese name: Haiiro-hoso-funbae]</p> <p>(Figs 49–51)</p> <p>Cordylura pallidicauda Zetterstedt, 1838: 733. Type-locality: “Lycksele Lapponiae Umensis … Juckasjervi Lapponiae Tornensis… (Lapponia Suecica et sylvatica)” [Sweden].</p> <p>Cordylura melanura Zetterstedt, 1838: 731. Type-locality: “Kengis Lapponiae Tornensis… Stensele, Umenaes, Wilhelmina et Asele, Lapponia Umensis” [Sweden].</p> <p>Material examined. Honshu, 9 ♂, Mt. Norikura, Gifu Pref., 4. viii. 1952, leg. I. Hattori; 2 ♀, same locality and date, leg. S. Kato.</p> <p>Diagnosis. This species can be recognized by the following characteristics: frontal vitta reddish yellow, face and parafacial black to pale yellow; gena and postgena pale yellow; palpi spatulate and white; postpedicel rounded apically, about 2 times as long as wide; thorax black with grey dusting; legs yellow, only mid and hind coxae black; fore femur with a row of posterodorsal setae, with numerous hairs anteriorly and ventrally; abdominal tergites black and greyish dusted; posterior lobes on male sternite V basally wide, with one row of spinules on inside margin (Fig. 49); male surstyli simple and slender (Figs 50, 51).</p> <p>Body length (♂ ♀). 6.0–6.2 mm.</p> <p>Distribution. Europe, Russia (European territory, West and East Siberia, Far East) (Ozerov, 2017) and Japan (Honshu).</p> <p>Remarks. This species is easily distinguished from M. haemorrhoidalis Meigen by having posterior lobes on sternite V which are basally stout, and by the shape of surstyli.</p> </div>	http://treatment.plazi.org/id/03B587C9FFA2AF38B09CECA974485D4F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Iwasa, Mitsuhiro	Iwasa, Mitsuhiro (2021): Three new species and six new records of Scathophagidae (Diptera) from Japan with a key to the Japanese genera. Zootaxa 4981 (3): 531-553, DOI: https://doi.org/10.11646/zootaxa.4981.3.6
03B587C9FFA1AF38B09CEF88760A5F5C.text	03B587C9FFA1AF38B09CEF88760A5F5C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spaziphora Rondani 1856	<div><p>Genus Spaziphora Rondani, 1856</p> <p>[Japanese name: Migiwa-funbae-zoku]</p> <p>Spaziphora Rondani, 1856: Dipt. Ital. Prodromus, 1: 99. Type-species: Cordylura hydromyzina Fallén, 1819: 7, by original designation.</p> <p>Diagnosis. Head whitish to black; 3-4 fr, 3 or (2 proclinate, 1 latero-reclinate), 1 oc, 1 poc, 1 vte, 1 vti, vibrissal angle with 1 vibrissa and 1-2 subvibrissae; postpedicel with acute or slightly angular upper corner; palpi distinctly spatulate and whitish; proepisternum and proepimeron with yellowish hairs; anepimeron covered with hairs in posterior half and row of black setae along posterior margin; thorax grey dusted; 2 pprn, 2 npl, 1+2 ial, 1+1 sa, 2 pa, 2+3 dc, 1 bsc, 1 apsc; 1 kepst; prepst and prepm absent; abdomen slender, elongate, and grey dusted; male sternite V with three pairs of tubercles; male surstyli short and stout, clothed with strong setae; female ovipositor short and cylindrical.</p></div> 	http://treatment.plazi.org/id/03B587C9FFA1AF38B09CEF88760A5F5C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Iwasa, Mitsuhiro	Iwasa, Mitsuhiro (2021): Three new species and six new records of Scathophagidae (Diptera) from Japan with a key to the Japanese genera. Zootaxa 4981 (3): 531-553, DOI: https://doi.org/10.11646/zootaxa.4981.3.6
03B587C9FFA1AF38B09CEDB976C859F7.text	03B587C9FFA1AF38B09CEDB976C859F7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spaziphora tomkovichi Ozerov. The 2011	<div><p>Spaziphora tomkovichi Ozerov, 2011</p> <p>[Japanese name: Hirata-migiwa-funbae]</p> <p>(Figs 52–56)</p> <p>Spaziphora tomkovichi Ozerov, 2011. Type-locality: Chukotka, Anadyr’Distr., Meynipyl’gino, lake-morene [Russia].</p> <p>Material examined. Hokkaido, 2 ♂, 1 ♀, Kushiro Marsh, Kushiro, 4. viii. 1993, leg. M. Iwasa.</p> <p>Diagnosis. Spaziphora tomkovichi Ozerov (Fig. 52) can be recognized by the following characteristics: anterior part of frons, face, parafacial, and gena whitish; postpedicel yellowish, with slightly angular upper corner (Fig. 53); palpi spatulate and white (Fig. 54); thorax black with grey dusting; legs yellow; abdomen slender, grey dusted, and subshiny; male sternite V ventrally with two distinct spur-like tubercles (Figs 55, 56); male surstyli stout and apically pointed, closing each other (Fig. 56).</p> <p>Body length (♂ ♀). 7.3–8.0 mm.</p> <p>Distribution. Russia (the Far East) (Ozerov &amp; Krivosheina, 2014) and Japan (Hokkaido). New to Japan.</p> <p>Remarks. Currently, the genus Spaziphora comprises three species in the world (Ozerov, 2011). Spaziphora tomkovichi is easily distinguished from other known congeneric species by having two distinct ventral spure-like tubercles on male sternite V.</p> </div>	http://treatment.plazi.org/id/03B587C9FFA1AF38B09CEDB976C859F7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Iwasa, Mitsuhiro	Iwasa, Mitsuhiro (2021): Three new species and six new records of Scathophagidae (Diptera) from Japan with a key to the Japanese genera. Zootaxa 4981 (3): 531-553, DOI: https://doi.org/10.11646/zootaxa.4981.3.6
03B587C9FFA7AF3EB09CEE8D72815FE2.text	03B587C9FFA7AF3EB09CEE8D72815FE2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scathophagidae	<div><p>Key to the genera of Japanese Scathophagidae (males and females)</p> <p>1. Fore femur and tibia ventrally with remarkable rows of spine-like setae, longer than diameter of femur and tibia (Hironaga &amp; Suwa 2005)........................................................................ Norellisoma Wahlgren</p> <p>- Fore femur and tibia without ventral rows of long spine-like setae............................................... 2</p> <p>2. Palpus short and small, less than length of proboscis (Hering 1955)............................. Parallelomma Becker</p> <p>- Palpus long and large, longer than length of proboscis........................................................ 3</p> <p>3. Palpus with strong subapical setae about as long as or longer than palpus (see Iwasa (2020, fig. 7))....... Cordilura Fallén</p> <p>- Palpus without very strong subapical setae (Fig. 2)........................................................... 4</p> <p>4. Katepisternum with two or three setae..................................................................... 5</p> <p>- Katepisternum with one seta (Fig. 44)..................................................................... 6</p> <p>5. Katepisternum with two setae........................................................... Chaetosa Coquillett</p> <p>- Katepisternum with three setae (Fig. 16)................................................... Cleigastra Macquart</p> <p>6. Fore tibia ventrally without rows of long spine-like setae but with 1 or more rows of short black spines (Fig. 45).......... 7</p> <p>- Fore tibia ventrally without rows of both long spine-like setae and short black spines................................ 8</p> <p>7. Apex of fore tibia with a stout ventral seta (Suwa 1986)..................................... Acanthocnema Becker</p> <p>- Apex of fore tibia without stout ventral seta............................................... Microprosopa Becker</p> <p>8. Anepisternum, katepisternum, and anepimeron wholly covered with dense, long erect fine hairs....................... 9</p> <p>- Anepisternum, katepisternum, and anepimeron covered with sparse fine hairs or bare.............................. 10</p> <p>9. Palpus distinctly spatulate (Fig. 54)....................................................... Spaziphora Rondani</p> <p>- Palpus filiform or clavate.............................................................. Scathophaga Meigen</p> <p>10. Arista plumose (Fig. 30) or short-haired (Figs 34, 40); body shiny yellow........................................ 11</p> <p>- Arista bare (Fig. 53) or pubescent (Fig. 7); body shiny black, grey microtrichose or dusted.......................... 12</p> <p>11. Scutellum with pair of strong setae (Fig. 28).............................................. Megaphthalma Becker</p> <p>- Scutellum with two pairs of strong setae............................................ Megaphthalmoides Ringdahl</p> <p>12. Arista relatively short, thick and slightly longer than postpedicel; postpedicel enlarged in males (Fig. 2)..................................................................................................... Acerocnema Becker</p> <p>- Arista rather thin, and distinctly longer than postpedicel; postpedicel not more enlarged in both sex (Fig. 22)................................................................................................ Gimnomera Rondani</p></div> 	http://treatment.plazi.org/id/03B587C9FFA7AF3EB09CEE8D72815FE2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Iwasa, Mitsuhiro	Iwasa, Mitsuhiro (2021): Three new species and six new records of Scathophagidae (Diptera) from Japan with a key to the Japanese genera. Zootaxa 4981 (3): 531-553, DOI: https://doi.org/10.11646/zootaxa.4981.3.6
