taxonID	type	description	language	source
03DF87B8D371FFFF5797F9EAFADEF97D.taxon	description	(Fig. 1)	en	Hausmann, Axel, Parisi, Francesco, Sciarretta, Andrea (2014): The geometrid moths of Ethiopia I: tribes Pseudoterpnini and Comibaenini (Lepidoptera: Geometridae, Geometrinae). Zootaxa 3768 (4): 460-468, DOI: http://dx.doi.org/10.11646/zootaxa.3768.4.4
03DF87B8D371FFFF5797F9EAFADEF97D.taxon	materials_examined	Type locality. Comores: Mayotte. Material. ZSM: 1 ♀, Arussi ♀ (locality no. 1); Bale 1 ♀ (14); Keffa 1 ♂ (20); Gamo Gofa 1 ♂ (23). Genetic data. genetically well corresponding with P. rhadamaria from nine different countries in Africa.	en	Hausmann, Axel, Parisi, Francesco, Sciarretta, Andrea (2014): The geometrid moths of Ethiopia I: tribes Pseudoterpnini and Comibaenini (Lepidoptera: Geometridae, Geometrinae). Zootaxa 3768 (4): 460-468, DOI: http://dx.doi.org/10.11646/zootaxa.3768.4.4
03DF87B8D371FFFE5797F8CCFE6BFF77.taxon	description	(Fig. 2)	en	Hausmann, Axel, Parisi, Francesco, Sciarretta, Andrea (2014): The geometrid moths of Ethiopia I: tribes Pseudoterpnini and Comibaenini (Lepidoptera: Geometridae, Geometrinae). Zootaxa 3768 (4): 460-468, DOI: http://dx.doi.org/10.11646/zootaxa.3768.4.4
03DF87B8D371FFFE5797F8CCFE6BFF77.taxon	materials_examined	Type locality. Algeria: Oran, Aïn-Sefra, Lahaye. Material. ZSM: Gamo Gofa 1 ♂ (locality no. 3). Genetic data. genetically well fitting into the cluster of P. lahayei as represented by 18 specimens from eight different countries in Africa.	en	Hausmann, Axel, Parisi, Francesco, Sciarretta, Andrea (2014): The geometrid moths of Ethiopia I: tribes Pseudoterpnini and Comibaenini (Lepidoptera: Geometridae, Geometrinae). Zootaxa 3768 (4): 460-468, DOI: http://dx.doi.org/10.11646/zootaxa.3768.4.4
03DF87B8D370FFFE5797FD4DFE24FCDD.taxon	description	(Fig. 4)	en	Hausmann, Axel, Parisi, Francesco, Sciarretta, Andrea (2014): The geometrid moths of Ethiopia I: tribes Pseudoterpnini and Comibaenini (Lepidoptera: Geometridae, Geometrinae). Zootaxa 3768 (4): 460-468, DOI: http://dx.doi.org/10.11646/zootaxa.3768.4.4
03DF87B8D370FFFE5797FD4DFE24FCDD.taxon	materials_examined	Type locality. Abyssinie (Ethiopia).	en	Hausmann, Axel, Parisi, Francesco, Sciarretta, Andrea (2014): The geometrid moths of Ethiopia I: tribes Pseudoterpnini and Comibaenini (Lepidoptera: Geometridae, Geometrinae). Zootaxa 3768 (4): 460-468, DOI: http://dx.doi.org/10.11646/zootaxa.3768.4.4
03DF87B8D370FFFE5797FD4DFE24FCDD.taxon	materials_examined	Material. ZSM: Harerge 1 ♂ (locality no. 4); Keffa 3 ♂ (19); Keffa 2 ♂ (21); Keffa 10 ♂ 1 ♀ (18); DAEF: Shewa 2 ♂ (13); Bale 1 ♂ (14).	en	Hausmann, Axel, Parisi, Francesco, Sciarretta, Andrea (2014): The geometrid moths of Ethiopia I: tribes Pseudoterpnini and Comibaenini (Lepidoptera: Geometridae, Geometrinae). Zootaxa 3768 (4): 460-468, DOI: http://dx.doi.org/10.11646/zootaxa.3768.4.4
03DF87B8D370FFFE5797FED9FE00FDFD.taxon	description	(Fig. 3)	en	Hausmann, Axel, Parisi, Francesco, Sciarretta, Andrea (2014): The geometrid moths of Ethiopia I: tribes Pseudoterpnini and Comibaenini (Lepidoptera: Geometridae, Geometrinae). Zootaxa 3768 (4): 460-468, DOI: http://dx.doi.org/10.11646/zootaxa.3768.4.4
03DF87B8D370FFFE5797FED9FE00FDFD.taxon	materials_examined	Type locality. Not stated (BMNH). Material. ZSM: Keffa 1 ♂ (locality no. 7); Keffa 1 ♂ (10); Gamo Gofa 1 ♀ (18); Gamo Gofa 1 ♀ (3). Genetic data. Ethiopian populations genetically diverging from those of South Africa, Mozambique and Zimbabwe (subsp. respondens) by 3.3 %, suggesting the potential need for raising the taxon respondens up to the rank of species. For verifying the identity (and the provenience) of nominotypical P. distensaria, the type specimen should be DNA barcoded.	en	Hausmann, Axel, Parisi, Francesco, Sciarretta, Andrea (2014): The geometrid moths of Ethiopia I: tribes Pseudoterpnini and Comibaenini (Lepidoptera: Geometridae, Geometrinae). Zootaxa 3768 (4): 460-468, DOI: http://dx.doi.org/10.11646/zootaxa.3768.4.4
03DF87B8D370FFFE5797FC2CFEB4FA38.taxon	description	(Fig. 5)	en	Hausmann, Axel, Parisi, Francesco, Sciarretta, Andrea (2014): The geometrid moths of Ethiopia I: tribes Pseudoterpnini and Comibaenini (Lepidoptera: Geometridae, Geometrinae). Zootaxa 3768 (4): 460-468, DOI: http://dx.doi.org/10.11646/zootaxa.3768.4.4
03DF87B8D370FFFE5797FC2CFEB4FA38.taxon	materials_examined	Type locality. (Ethiopia): Eritrea. Material. ZSM: Shewa 1 ♂ (locality no. 2). Remarks: according to Prout (1932) possibly an individual aberration of P. abyssinaria, somewhat smaller and paler, postmedial line slightly less dentate. In the collection Herbulot, a specimen was found under this name, but attributed here as a synonym to P. distensaria. The dissection (♂) revealed this specimen as clearly different from both P. distensaria and P. abyssinaria, e. g. in the asymmetric valvae with a narrow spine at the centre of the left valva, a spinose process from the costa of the left valva, the absence of cornuti in the aedeagus a. s. o. (cf. Fig. 11). Therefore we accept and confirm the species status of this taxon as it was proposed also by Scoble (1999). The type specimen was not traced in the MNHN (Paris; Claude Tautel pers. comm.) and may be deposited in the NHM (London).	en	Hausmann, Axel, Parisi, Francesco, Sciarretta, Andrea (2014): The geometrid moths of Ethiopia I: tribes Pseudoterpnini and Comibaenini (Lepidoptera: Geometridae, Geometrinae). Zootaxa 3768 (4): 460-468, DOI: http://dx.doi.org/10.11646/zootaxa.3768.4.4
03DF87B8D370FFFE5797FA13FDBDF887.taxon	description	(Fig. 6)	en	Hausmann, Axel, Parisi, Francesco, Sciarretta, Andrea (2014): The geometrid moths of Ethiopia I: tribes Pseudoterpnini and Comibaenini (Lepidoptera: Geometridae, Geometrinae). Zootaxa 3768 (4): 460-468, DOI: http://dx.doi.org/10.11646/zootaxa.3768.4.4
03DF87B8D370FFFE5797FA13FDBDF887.taxon	materials_examined	Type locality. Uganda: Nandi Country, Rau. Material. ZSM: Bale 1 ♂ (locality no. 5). Genetic data. the single specimen of P. ruginaria from Ethiopia and populations from South-East Asia (examined from Thailand) diverge by 0.46 % only, with very similar haplotypes occurring in most populations from western and eastern Africa. In West Africa, however, there is another cluster of haplotypes at a minimum distance of 4.1 %, requiring further study.	en	Hausmann, Axel, Parisi, Francesco, Sciarretta, Andrea (2014): The geometrid moths of Ethiopia I: tribes Pseudoterpnini and Comibaenini (Lepidoptera: Geometridae, Geometrinae). Zootaxa 3768 (4): 460-468, DOI: http://dx.doi.org/10.11646/zootaxa.3768.4.4
03DF87B8D377FFF95797F932FACEF81E.taxon	description	(Fig. 7)	en	Hausmann, Axel, Parisi, Francesco, Sciarretta, Andrea (2014): The geometrid moths of Ethiopia I: tribes Pseudoterpnini and Comibaenini (Lepidoptera: Geometridae, Geometrinae). Zootaxa 3768 (4): 460-468, DOI: http://dx.doi.org/10.11646/zootaxa.3768.4.4
03DF87B8D377FFF95797F932FACEF81E.taxon	materials_examined	Type locality. Sudan: Kassala Province, Ercowit, 1000 – 1300 m. Material. ZSM: Keffa 1 ♂ (locality no. 8); Keffa 1 ♂ (1); Sidamo 1 ♂ (11); Sidamo 1 ♀ (22); Sidamo 1 ♀ (16); Sidamo 1 ♀ (17). Genetic data. populations of M. pulchra split up into six different genetic lineages, awaiting further study.	en	Hausmann, Axel, Parisi, Francesco, Sciarretta, Andrea (2014): The geometrid moths of Ethiopia I: tribes Pseudoterpnini and Comibaenini (Lepidoptera: Geometridae, Geometrinae). Zootaxa 3768 (4): 460-468, DOI: http://dx.doi.org/10.11646/zootaxa.3768.4.4
03DF87B8D376FFFA5797FF69FAF3FB96.taxon	description	(Figs 8, 9)	en	Hausmann, Axel, Parisi, Francesco, Sciarretta, Andrea (2014): The geometrid moths of Ethiopia I: tribes Pseudoterpnini and Comibaenini (Lepidoptera: Geometridae, Geometrinae). Zootaxa 3768 (4): 460-468, DOI: http://dx.doi.org/10.11646/zootaxa.3768.4.4
03DF87B8D376FFFA5797FF69FAF3FB96.taxon	materials_examined	Type material. Holotype: ♂, Ethiopia, Prov. Kaffa, 10 km n Bonga, 1550 m 7 ° 21 ’ N 36 ° 25 ’ E, 29. IV. 2008, leg. S. Naumann, H. Schnitzler, coll. ZSM (locality no. 10); gen. prp 19376. DNA Barcode specimen ID BC ZSM Lep 14842. Paratypes: 1 ♀, Ethiopie, Choa, Debre-Zeit, 1800 m, 21. III. 1986, leg. F. Thiaucourt (locality no. 2), coll. Herbulot / ZSM; 1 ♀, Äthiopien, Äthiopisches Hochland, Prov. Sidamo / Oromia, Yabello, 17 km Wildlife Sanctuary OSO, 1580 m, (Dornbuschgebiet), 04 ° 51.010 ” N, 38 ° 14.783 ’ E, 16. V. 2012, leg. Dietl M. + S. & R. Beck, coll. ZSM (locality no. 17); 1 ♀, Äthiopien, Ostafrikanischer Graben, Prov. Gamu Goffa, Arba Minch, 12 km NNO 1620 m, 06 ° 08.311 ’ N, 37 ° 34.953 ’ O, 23. V. 2012, leg. Dietl M. + S. & R. Beck, coll. ZSM (locality no. 18); 1 ♀, E. Ethiopia, Hararge, Kara Jara [9.24 ° N 41.14 ° E], 2180 m, 20. IV. 2006, leg. R. Beck & Tamrat (locality no. 4). Other material: DAEF: 1 ♀, Ethiopia, near Abelti, Ghibe river, 1096 m, 30. I. 2012, coord. UTM 37 P 338350 E, 913109 N, Sciarretta A., Parisi F. leg, gen. prp. R; 1 ♂, Ethiopia, Wellega zone, Didessa river, 1280 m, 24. I. 2012, coord. UTM 37 P 215814 E, 961832 N, Sciarretta A., Parisi F. leg, gen. prp. N, not included in the type series because of the poor condition of these specimens (worn, almost without scales) and because of the unusually long process of sternum A 8 in the male (see below under ‘ male genitalia’).	en	Hausmann, Axel, Parisi, Francesco, Sciarretta, Andrea (2014): The geometrid moths of Ethiopia I: tribes Pseudoterpnini and Comibaenini (Lepidoptera: Geometridae, Geometrinae). Zootaxa 3768 (4): 460-468, DOI: http://dx.doi.org/10.11646/zootaxa.3768.4.4
03DF87B8D376FFFA5797FF69FAF3FB96.taxon	description	Description. Adult (Figs 8 – 9). Wingspan: ♂ ♀ 23 – 29 mm. Ground colour leaf green, ante- and postmedial lines fine, whitish. Medial area dilated towards forewing costa. Discal dots red brown, fine and sharp on all wings. Terminal line uninterrupted, red brown, slightly dilated at forewing tornus and hindwing apex. Fringe chequered pale yellow and red brown, the latter in triangles. Underside shiny pale green with postmedial fascia darker green, discal dots fine on all wings, fringe chequered as on upper side. Frons pale green, vertex darker green. ♂ palpi with mixed brown and white scales, length 1.5 mm, last segment smoothly scaled. ♂ antennae long bipectinate, length of branches up to 1.3 mm, approx. 10 times width of flagellum. ♂ hindtibia with four spurs and a terminal process, length of the latter 0.75 mm. Male genitalia (Figs 10 a – 10 c): Uncus with lateral projections much longer than central, 0.5 – 0.6 mm deepest incision. Valva comparatively broad at centre. Saccus projections comparatively short, length 0.5 – 0.6 mm from central excavation. Aedeagus straight, length 2.25 mm. Sternum A 8 with moderately long projection, length 0.3 mm in the holotype (but 0.95 mm in the specimen from Didessa river).	en	Hausmann, Axel, Parisi, Francesco, Sciarretta, Andrea (2014): The geometrid moths of Ethiopia I: tribes Pseudoterpnini and Comibaenini (Lepidoptera: Geometridae, Geometrinae). Zootaxa 3768 (4): 460-468, DOI: http://dx.doi.org/10.11646/zootaxa.3768.4.4
03DF87B8D376FFFA5797FF69FAF3FB96.taxon	diagnosis	Differential diagnosis. African members of genus Comibaena often show interspecific differential features in coloration and wing pattern, though the differences in male genitalia are often small, e. g. in the rufitornus / leucospilata / esmeralda species-group. The specific differential traits in the habitus, however, are well correlated with COI-haplotypes at interspecific genetic distances of usually 5 to 8 %. Anticipating a comprehensive revision of the African species of the genus Comibaena (probably raising the species number from actually 12 to more than 20) we describe a new Ethiopian species here, which is well differentiated from all other African species in male genitalia and DNA barcodes. The new species differs from C. rufitornus Prout, 1916 (dissections examined from Kenya, Cape Verde), C. leucospilata (Walker, 1863) (South Africa, Natal) and C. esmeralda (Warren, 1898) (Nigeria, Guinea) by a shorter projection of sternum A 8 (0.3 mm rather than 0.5 – 0.9 mm) by the long lateral projections of uncus (0.55 mm rather than 0.35 – 0.50 mm) which are clearly exceeding the length of the central projections and by the comparatively short saccus projections. C. leucospilata from South Africa (Natal) very similar in habitus, on average with narrower medial area, length of ♂ antennal branches 1.1 – 1.2 mm, length of ♂ hindtibial process 0.5 mm. Genetically the nearest neighbour (6.2 %) is an undescribed species from Kenya and Tanzania which strongly differs in habitus (transverse lines absent, discal spots weak, triangle pattern of terminal line and fringe absent) and genitalia (aedeagus much longer, with termino-lateral lobe, saccus projections and posterior spine on sternum A 8 much longer). C. rufitornus (barcoded from Uganda, Yemen and Cape Verde, the last tentatively considered conspecific), C. leucospilata (South Africa, Zimbabwe, Zambia, Tanzania) and C. esmeralda (Ghana, Cameroon, Sierra Leone) diverge from C. theodori sp. nov. by 6.8 %, 7.5 % and 6.5 % respectively.	en	Hausmann, Axel, Parisi, Francesco, Sciarretta, Andrea (2014): The geometrid moths of Ethiopia I: tribes Pseudoterpnini and Comibaenini (Lepidoptera: Geometridae, Geometrinae). Zootaxa 3768 (4): 460-468, DOI: http://dx.doi.org/10.11646/zootaxa.3768.4.4
03DF87B8D376FFFA5797FF69FAF3FB96.taxon	etymology	Etymology. The new species is dedicated to Theodor Beck, the father of the collector of two paratypes.	en	Hausmann, Axel, Parisi, Francesco, Sciarretta, Andrea (2014): The geometrid moths of Ethiopia I: tribes Pseudoterpnini and Comibaenini (Lepidoptera: Geometridae, Geometrinae). Zootaxa 3768 (4): 460-468, DOI: http://dx.doi.org/10.11646/zootaxa.3768.4.4
