identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03DF87B8D371FFFF5797F9EAFADEF97D.text	03DF87B8D371FFFF5797F9EAFADEF97D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pingasa rhadamaria subsp. signifrontaria (Mabille 1893)	<div><p>Pingasa rhadamaria signifrontaria (Mabille, 1893)</p> <p>(Fig. 1)</p> <p>Type locality. Comores: Mayotte.</p> <p>Material. ZSM: 1♀, Arussi ♀ (locality no. 1); Bale 1♀ (14); Keffa 1♂ (20); Gamo Gofa 1♂ (23).</p> <p>Genetic data. genetically well corresponding with P. rhadamaria from nine different countries in Africa.</p> </div>	http://treatment.plazi.org/id/03DF87B8D371FFFF5797F9EAFADEF97D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hausmann, Axel;Parisi, Francesco;Sciarretta, Andrea	Hausmann, Axel, Parisi, Francesco, Sciarretta, Andrea (2014): The geometrid moths of Ethiopia I: tribes Pseudoterpnini and Comibaenini (Lepidoptera: Geometridae, Geometrinae). Zootaxa 3768 (4): 460-468, DOI: http://dx.doi.org/10.11646/zootaxa.3768.4.4
03DF87B8D371FFFE5797F8CCFE6BFF77.text	03DF87B8D371FFFE5797F8CCFE6BFF77.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pingasa lahayei (Oberthur 1887)	<div><p>Pingasa lahayei (Oberthür, 1887)</p> <p>(Fig. 2)</p> <p>Type locality. Algeria: Oran, Aïn-Sefra, Lahaye.</p> <p>Material. ZSM: Gamo Gofa 1♂ (locality no. 3).</p> <p>Genetic data. genetically well fitting into the cluster of P. lahayei as represented by 18 specimens from eight different countries in Africa.</p> </div>	http://treatment.plazi.org/id/03DF87B8D371FFFE5797F8CCFE6BFF77	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hausmann, Axel;Parisi, Francesco;Sciarretta, Andrea	Hausmann, Axel, Parisi, Francesco, Sciarretta, Andrea (2014): The geometrid moths of Ethiopia I: tribes Pseudoterpnini and Comibaenini (Lepidoptera: Geometridae, Geometrinae). Zootaxa 3768 (4): 460-468, DOI: http://dx.doi.org/10.11646/zootaxa.3768.4.4
03DF87B8D370FFFE5797FD4DFE24FCDD.text	03DF87B8D370FFFE5797FD4DFE24FCDD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pingasa abyssinaria (Guenee 1858)	<div><p>Pingasa abyssinaria (Guenée, 1858)</p> <p>(Fig. 4)</p> <p>Type locality. Abyssinie (Ethiopia).</p> <p>Material. ZSM: Harerge 1♂ (locality no. 4); Keffa 3♂ (19); Keffa 2♂ (21); Keffa 10♂ 1♀ (18); DAEF: Shewa 2♂ (13); Bale 1♂ (14).</p> </div>	http://treatment.plazi.org/id/03DF87B8D370FFFE5797FD4DFE24FCDD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hausmann, Axel;Parisi, Francesco;Sciarretta, Andrea	Hausmann, Axel, Parisi, Francesco, Sciarretta, Andrea (2014): The geometrid moths of Ethiopia I: tribes Pseudoterpnini and Comibaenini (Lepidoptera: Geometridae, Geometrinae). Zootaxa 3768 (4): 460-468, DOI: http://dx.doi.org/10.11646/zootaxa.3768.4.4
03DF87B8D370FFFE5797FED9FE00FDFD.text	03DF87B8D370FFFE5797FED9FE00FDFD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pingasa distensaria (Walker 1860)	<div><p>Pingasa distensaria (Walker, 1860)</p> <p>(Fig. 3)</p> <p>Type locality. Not stated (BMNH).</p> <p>Material. ZSM: Keffa 1♂ (locality no. 7); Keffa 1♂ (10); Gamo Gofa 1♀ (18); Gamo Gofa 1♀ (3).</p> <p>Genetic data. Ethiopian populations genetically diverging from those of South Africa, Mozambique and Zimbabwe (subsp. respondens) by 3.3%, suggesting the potential need for raising the taxon respondens up to the rank of species. For verifying the identity (and the provenience) of nominotypical P. distensaria, the type specimen should be DNA barcoded.</p> </div>	http://treatment.plazi.org/id/03DF87B8D370FFFE5797FED9FE00FDFD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hausmann, Axel;Parisi, Francesco;Sciarretta, Andrea	Hausmann, Axel, Parisi, Francesco, Sciarretta, Andrea (2014): The geometrid moths of Ethiopia I: tribes Pseudoterpnini and Comibaenini (Lepidoptera: Geometridae, Geometrinae). Zootaxa 3768 (4): 460-468, DOI: http://dx.doi.org/10.11646/zootaxa.3768.4.4
03DF87B8D370FFFE5797FC2CFEB4FA38.text	03DF87B8D370FFFE5797FC2CFEB4FA38.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pingasa pallidata de Joannis 1912	<div><p>Pingasa pallidata de Joannis, 1912</p> <p>(Fig. 5)</p> <p>Type locality. (Ethiopia): Eritrea.</p> <p>Material. ZSM: Shewa 1♂ (locality no. 2).</p> <p>Remarks: according to Prout (1932) possibly an individual aberration of P. abyssinaria, somewhat smaller and paler, postmedial line slightly less dentate. In the collection Herbulot, a specimen was found under this name, but attributed here as a synonym to P. distensaria. The dissection (♂) revealed this specimen as clearly different from both P. distensaria and P. abyssinaria, e.g. in the asymmetric valvae with a narrow spine at the centre of the left valva, a spinose process from the costa of the left valva, the absence of cornuti in the aedeagus a.s.o. (cf. Fig. 11). Therefore we accept and confirm the species status of this taxon as it was proposed also by Scoble (1999). The type specimen was not traced in the MNHN (Paris; Claude Tautel pers. comm.) and may be deposited in the NHM (London).</p> </div>	http://treatment.plazi.org/id/03DF87B8D370FFFE5797FC2CFEB4FA38	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hausmann, Axel;Parisi, Francesco;Sciarretta, Andrea	Hausmann, Axel, Parisi, Francesco, Sciarretta, Andrea (2014): The geometrid moths of Ethiopia I: tribes Pseudoterpnini and Comibaenini (Lepidoptera: Geometridae, Geometrinae). Zootaxa 3768 (4): 460-468, DOI: http://dx.doi.org/10.11646/zootaxa.3768.4.4
03DF87B8D370FFFE5797FA13FDBDF887.text	03DF87B8D370FFFE5797FA13FDBDF887.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pingasa ruginaria subsp. interrupta Warren 1901	<div><p>Pingasa ruginaria interrupta Warren, 1901</p> <p>(Fig. 6)</p> <p>Type locality. Uganda: Nandi Country, Rau.</p> <p>Material. ZSM: Bale 1♂ (locality no. 5).</p> <p>Genetic data. the single specimen of P. ruginaria from Ethiopia and populations from South-East Asia (examined from Thailand) diverge by 0.46% only, with very similar haplotypes occurring in most populations from western and eastern Africa. In West Africa, however, there is another cluster of haplotypes at a minimum distance of 4.1%, requiring further study.</p> </div>	http://treatment.plazi.org/id/03DF87B8D370FFFE5797FA13FDBDF887	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hausmann, Axel;Parisi, Francesco;Sciarretta, Andrea	Hausmann, Axel, Parisi, Francesco, Sciarretta, Andrea (2014): The geometrid moths of Ethiopia I: tribes Pseudoterpnini and Comibaenini (Lepidoptera: Geometridae, Geometrinae). Zootaxa 3768 (4): 460-468, DOI: http://dx.doi.org/10.11646/zootaxa.3768.4.4
03DF87B8D377FFF95797F932FACEF81E.text	03DF87B8D377FFF95797F932FACEF81E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microbaena pulchra subsp. minor Hausmann 1996	<div><p>Microbaena pulchra minor Hausmann, 1996</p> <p>(Fig. 7)</p> <p>Type locality. Sudan: Kassala Province, Ercowit, 1000–1300m.</p> <p>Material. ZSM: Keffa 1♂ (locality no. 8); Keffa 1♂ (1); Sidamo 1♂ (11); Sidamo 1♀ (22); Sidamo 1♀ (16); Sidamo 1♀ (17).</p> <p>Genetic data. populations of M. pulchra split up into six different genetic lineages, awaiting further study.</p> </div>	http://treatment.plazi.org/id/03DF87B8D377FFF95797F932FACEF81E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hausmann, Axel;Parisi, Francesco;Sciarretta, Andrea	Hausmann, Axel, Parisi, Francesco, Sciarretta, Andrea (2014): The geometrid moths of Ethiopia I: tribes Pseudoterpnini and Comibaenini (Lepidoptera: Geometridae, Geometrinae). Zootaxa 3768 (4): 460-468, DOI: http://dx.doi.org/10.11646/zootaxa.3768.4.4
03DF87B8D376FFFA5797FF69FAF3FB96.text	03DF87B8D376FFFA5797FF69FAF3FB96.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Comibaena theodori Hausmann & Parisi & Sciarretta 2014	<div><p>Comibaena theodori Hausmann &amp; Parisi sp. nov.</p> <p>(Figs 8, 9)</p> <p>Type material. Holotype: ♂, Ethiopia, Prov. Kaffa, 10 km n <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.416668&amp;materialsCitation.latitude=7.35" title="Search Plazi for locations around (long 36.416668/lat 7.35)">Bonga</a>, 1550m 7°21’N 36°25’E, 29.IV.2008, leg. S. Naumann, H. Schnitzler, coll. ZSM (locality no. 10); gen.prp 19376. DNA Barcode specimen ID BC ZSM Lep 14842. Paratypes: 1♀, Ethiopie, Choa, Debre-Zeit, 1800m, 21.III.1986, leg. F. Thiaucourt (locality no. 2), coll. Herbulot / ZSM; 1♀, Äthiopien, Äthiopisches Hochland, Prov. Sidamo / Oromia, Yabello, 17 km Wildlife Sanctuary OSO, 1580m, (Dornbuschgebiet), 04°51.010”N, 38°14.783’E, 16. V.2012, leg. Dietl M. + S. &amp; R. Beck, coll. ZSM (locality no. 17); 1♀, Äthiopien, Ostafrikanischer Graben, Prov. Gamu Goffa, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-37.58255&amp;materialsCitation.latitude=6.1385164" title="Search Plazi for locations around (long -37.58255/lat 6.1385164)">Arba Minch</a>, 12 km NNO 1620m, 06°08.311’N, 37°34.953’O, 23. V.2012, leg. Dietl M. + S. &amp; R. Beck, coll. ZSM (locality no. 18); 1♀, E. Ethiopia, Hararge, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=41.14&amp;materialsCitation.latitude=9.24" title="Search Plazi for locations around (long 41.14/lat 9.24)">Kara Jara</a> [9.24° N 41.14° E], 2180m, 20.IV.2006, leg. R. Beck &amp; Tamrat (locality no. 4). Other material: DAEF: 1♀, Ethiopia, near Abelti, Ghibe river, 1096m, 30.I.2012, coord. UTM 37P 338350E, 913109N, Sciarretta A., Parisi F. leg, gen.prp. R; 1♂, Ethiopia, Wellega zone, Didessa river, 1280 m, 24.I.2012, coord. UTM 37P 215814E, 961832N, Sciarretta A., Parisi F. leg, gen.prp. N, not included in the type series because of the poor condition of these specimens (worn, almost without scales) and because of the unusually long process of sternum A 8 in the male (see below under ‘male genitalia’).</p> <p>Description. Adult (Figs 8–9). Wingspan: ♂ ♀ 23–29 mm. Ground colour leaf green, ante- and postmedial lines fine, whitish. Medial area dilated towards forewing costa. Discal dots red brown, fine and sharp on all wings. Terminal line uninterrupted, red brown, slightly dilated at forewing tornus and hindwing apex. Fringe chequered pale yellow and red brown, the latter in triangles. Underside shiny pale green with postmedial fascia darker green, discal dots fine on all wings, fringe chequered as on upper side. Frons pale green, vertex darker green. ♂ palpi with mixed brown and white scales, length 1.5 mm, last segment smoothly scaled. ♂ antennae long bipectinate, length of branches up to 1.3 mm, approx. 10 times width of flagellum. ♂ hindtibia with four spurs and a terminal process, length of the latter 0.75 mm. Male genitalia (Figs 10a–10c): Uncus with lateral projections much longer than central, 0.5–0.6 mm deepest incision. Valva comparatively broad at centre. Saccus projections comparatively short, length 0.5–0.6 mm from central excavation. Aedeagus straight, length 2.25 mm. Sternum A8 with moderately long projection, length 0.3 mm in the holotype (but 0.95 mm in the specimen from Didessa river).</p> <p>Differential diagnosis. African members of genus Comibaena often show interspecific differential features in coloration and wing pattern, though the differences in male genitalia are often small, e.g. in the rufitornus / leucospilata / esmeralda species-group. The specific differential traits in the habitus, however, are well correlated with COI-haplotypes at interspecific genetic distances of usually 5 to 8%. Anticipating a comprehensive revision of the African species of the genus Comibaena (probably raising the species number from actually 12 to more than 20) we describe a new Ethiopian species here, which is well differentiated from all other African species in male genitalia and DNA barcodes. The new species differs from C. rufitornus Prout, 1916 (dissections examined from Kenya, Cape Verde), C. leucospilata (Walker, 1863) (South Africa, Natal) and C. esmeralda (Warren, 1898) (Nigeria, Guinea) by a shorter projection of sternum A8 (0.3 mm rather than 0.5–0.9 mm) by the long lateral projections of uncus (0.55 mm rather than 0.35–0.50 mm) which are clearly exceeding the length of the central projections and by the comparatively short saccus projections. C. leucospilata from South Africa (Natal) very similar in habitus, on average with narrower medial area, length of ♂ antennal branches 1.1–1.2 mm, length of ♂ hindtibial process 0.5 mm. Genetically the nearest neighbour (6.2%) is an undescribed species from Kenya and Tanzania which strongly differs in habitus (transverse lines absent, discal spots weak, triangle pattern of terminal line and fringe absent) and genitalia (aedeagus much longer, with termino-lateral lobe, saccus projections and posterior spine on sternum A8 much longer). C. rufitornus (barcoded from Uganda, Yemen and Cape Verde, the last tentatively considered conspecific), C. leucospilata (South Africa, Zimbabwe, Zambia, Tanzania) and C. esmeralda (Ghana, Cameroon, Sierra Leone) diverge from C. theodori sp.nov. by 6.8%, 7.5% and 6.5% respectively.</p> <p>Etymology. The new species is dedicated to Theodor Beck, the father of the collector of two paratypes.</p></div> 	http://treatment.plazi.org/id/03DF87B8D376FFFA5797FF69FAF3FB96	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hausmann, Axel;Parisi, Francesco;Sciarretta, Andrea	Hausmann, Axel, Parisi, Francesco, Sciarretta, Andrea (2014): The geometrid moths of Ethiopia I: tribes Pseudoterpnini and Comibaenini (Lepidoptera: Geometridae, Geometrinae). Zootaxa 3768 (4): 460-468, DOI: http://dx.doi.org/10.11646/zootaxa.3768.4.4
