taxonID	type	description	language	source
B01A87E94B5CFFEDCE98B2B7FDFFB8E1.taxon	description	Figs 1 A – B, 2 A – C, 6 A, 7 A	en	Xi, Huacong, Wang, Younan, Yang, Xingke, Liu, Haoyu, Yang, Yuxia (2021): New species and taxonomic notes on Lycocerus hickeri species-group (Coleoptera Cantharidae). Zootaxa 4980 (3): 541-557, DOI: https://doi.org/10.11646/zootaxa.4980.3.5
B01A87E94B5CFFEDCE98B2B7FDFFB8E1.taxon	materials_examined	Type material examined. Holotype: ♀ (MNHN), [h] “ Yunnan / China ”, [h] “ type ”, [h] “ nigrobilineatus / Pic ”, [h] “ Athemus s. str. / nigrobilineatus / (Pic) / det. W. Wittmer ”, [h] “ 61 / 2 ”, [p] “ TYPE ”. Additional material examined. China, Yunnan: 1 ♀ (IZAS, IOZ (E) 1119866), Weixi, Pantiange, 2400 m, 25. vii. 1981, leg. S. Y. Wang; 1 ♀ (IZAS), Cangshan, 2850 m, 30. vi. 1981, leg. S. Y. Wang; 1 ♀ (IZAS, IOZ (E) 1435293), Weixi, Pantiange, 2500 m, 26. vii. 1981, leg. S. Y. Wang; 1 ♂ (NWAFU), Dali, Diancangshan, 2940 m, 19. – 20. vi. 1974, leg. Y. Zhou & F. Yuan; 1 ♂ (MHBU), Daxing, Laduhe, 26. vii. 2011, leg. H. Y. Liu; 1 ♂ (MHBU), “ Habashan mts. SE slope, 2000 – 3000 m, 27 ˚ 20 ’ N, 100 ˚ 11 ’ E, 10. – 13. vii. 1992, leg. Vít Kubáň ”.	en	Xi, Huacong, Wang, Younan, Yang, Xingke, Liu, Haoyu, Yang, Yuxia (2021): New species and taxonomic notes on Lycocerus hickeri species-group (Coleoptera Cantharidae). Zootaxa 4980 (3): 541-557, DOI: https://doi.org/10.11646/zootaxa.4980.3.5
B01A87E94B5CFFEDCE98B2B7FDFFB8E1.taxon	description	Redescription. Male (Fig. 1 B). Body black, mandibles brown, pronotum red, with two large dark brown markings on both sides which conjoint in center of disc, elytra red, darkened at the whole length of all elytral interstices. Body densely covered with short, decumbent brown pubescence. Head evenly narrowed behind eyes, surface densely and finely punctate; eyes moderately protruding, head width across eyes distinctly wider than anterior margin of pronotum; terminal maxillary palpomeres long-triangular, arcuate and sharp at apical half length of inner margins; antennae subfiliform, nearly extending to apical one-third length of elytra when reclined, antennomere II shortest, nearly as long as wide at apices, III ‒ VII feebly widened apically, III about three times as long as wide at apices, III ‒ XI each with an oblong to round smooth impression at apical part of inner margin, IV widest and slightly longer than III, VIII longest, VIII ‒ XI nearly parallel-sided, XI longer than X and pointed at apices. Pronotum 1.1 times longer than wide, widest at base, anterior margin arcuate, lateral margins moderately diverging posteriorly, posterior margin slightly arcuate and narrowly bordered, anterior angles rounded, posterior angles rectangular, disc moderately convex at postero-lateral parts, surface semillustrous, sparsely punctate than that on head. Elytra about 4.8 times longer than pronotum, 3.3 times longer than humeral width, which about 1.5 times wider than posterior margin of pronotum, lateral margins nearly parallel, apices rounded, elytra venations weakly developed, not costate, surface matt, wrinkled, sparsely and finely punctate. Legs slender, all claws simple. Aedeagus (Figs 2 A – C): ventral process of each paramere nearly straight both in ventral and lateral views, with apex feebly expanded and rounded; dorsal plate shorter than ventral process, longitudinally ridged laterally on inner surface, with inner margin hardly protuberant in middle, apical margin rounded and obviously emarginate at outer apical angle, which subrounded in lateral view; laterophyse feebly shorter than dorsal plate, with apex acutely hooked and bent dorsally, opposite to apical end of longitudinal ridge of dorsal plate. Female (Fig. 1 A). Similar to the male, but eyes less protruding, antennae shorter, extending to basal one-third length of elytra when reclined, III ‒ XI without impressions, pronotum wider, about 1.2 times wider than long, feebly convex on postero-lateral parts of disc, pro- and meso-outer tarsal claws each with a digitiform tooth at base. Abdominal sternite VIII (Fig. 6 A): roundly emarginate in middle and deeply and nearly triangularly emarginate on both sides of posterior margin, lateral emarginate parts much deeper than the middle one, the portion between lateral and middle emarginate parts obtuse-angled at apex, lateral emarginate parts membranous-lobed laterally, latero-apical angles subrounded at apices, not reaching apices of the portions between lateral and middle emarginate parts. Internal organ of reproductive system (Fig. 7 A): vagina stout and abruptly thinned at ventro-apical portion into a stout tube, where diverticulum and spermathecal duct arising separately; diverticulum moderately long, evenly thin and spiral; spermathecal duct obviously shorter and thicker than diverticulum; spermatheca provided with a spiral tube, longer than diverticulum, abruptly thinned at basal one-third, with the basal portion feebly thinned apically and much thicker than apical portion; accessory gland directly opening at base of spermatheca, thin and much longer than spermatheca. Body length: 7.5 – 9.0 mm; width: 1.8 ‒ 2.2 mm.	en	Xi, Huacong, Wang, Younan, Yang, Xingke, Liu, Haoyu, Yang, Yuxia (2021): New species and taxonomic notes on Lycocerus hickeri species-group (Coleoptera Cantharidae). Zootaxa 4980 (3): 541-557, DOI: https://doi.org/10.11646/zootaxa.4980.3.5
B01A87E94B5CFFEDCE98B2B7FDFFB8E1.taxon	distribution	Distribution. China (Yunnan).	en	Xi, Huacong, Wang, Younan, Yang, Xingke, Liu, Haoyu, Yang, Yuxia (2021): New species and taxonomic notes on Lycocerus hickeri species-group (Coleoptera Cantharidae). Zootaxa 4980 (3): 541-557, DOI: https://doi.org/10.11646/zootaxa.4980.3.5
B01A87E94B5EFFEBCE98B290FDFFB8C5.taxon	description	Figs 1 C – D, 2 D – F, 6 B, 7 B	en	Xi, Huacong, Wang, Younan, Yang, Xingke, Liu, Haoyu, Yang, Yuxia (2021): New species and taxonomic notes on Lycocerus hickeri species-group (Coleoptera Cantharidae). Zootaxa 4980 (3): 541-557, DOI: https://doi.org/10.11646/zootaxa.4980.3.5
B01A87E94B5EFFEBCE98B290FDFFB8C5.taxon	materials_examined	Type material. Holotype: ♂ (MHBU), “ China, Yunnan, Hengduan mts-part MEILI, 3700 m, 28 ˚ 06 ’ N, 98 ˚ 52 ’ E, 10. – 13. vii. 1996, leg. Vít Kubáň ”. Paratypes: 1 ♀ (MHBU), same data as holotype; 2 ♀♀ (MHBU), “ China, Yunnan, Hengduan mts-part MEILI, 3300 m, 28 ˚ 06 ’ N, 98 ˚ 53 ’ E, 8. – 10. vii. 1996, leg. Vít Kubáň ”.	en	Xi, Huacong, Wang, Younan, Yang, Xingke, Liu, Haoyu, Yang, Yuxia (2021): New species and taxonomic notes on Lycocerus hickeri species-group (Coleoptera Cantharidae). Zootaxa 4980 (3): 541-557, DOI: https://doi.org/10.11646/zootaxa.4980.3.5
B01A87E94B5EFFEBCE98B290FDFFB8C5.taxon	description	Description. Male (Fig. 1 D). Body black, mandibles brown, pronotum red, with two large dark brown markings on both sides which conjoint in center of disc, elytra red, darkened at the whole length of all elytral interstices. Body densely covered with short, decumbent brown pubescence. Head evenly narrowed behind eyes, surface densely and finely punctate; eyes moderately protruding, head width across eyes wider than anterior margin of pronotum; terminal maxillary palpomeres long-triangular, arcuate and sharp at apical one-third length of inner margins; antennae subfiliform, extending to elytral mid-length when reclined, antennomere II shortest, nearly as long as wide at apices, III ‒ VIII slightly obliquely widened apically and long-triangular, III about 2.5 times as long as wide at apices, III ‒ XI each with an oblong to round smooth impression at apical part of inner margin, IV widest and slightly shorter than III, VII longest, IX ‒ XI nearly parallel-sided, XI longer than X and pointed at apices. Pronotum about 1.2 times wider than long, widest at base, anterior margin arcuate, lateral margins moderately diverging posteriorly, posterior margin slightly arcuate, anterior angles rounded, posterior angles rectangular, disc moderately convex at postero-lateral parts, surface semillustrous, sparsely punctate than that on head. Elytra about 5.1 times longer than pronotum, 3.1 times longer than humeral width, which about 1.4 times wider than posterior margin of pronotum, lateral margins moderately diverging posteriorly, elytra venations strongly developed and costate, surface semillustrous, wrinkled, densely and very finely punctate. Legs slender, all claws simple. Aedeagus (Figs 2 D – F): ventral process of each paramere nearly straight both in ventral and lateral views, with apex feebly expanded and rounded; dorsal plate nearly as long as ventral process, longitudinally ridged laterally on inner surface, with inner margin protuberant in middle, apical margin rounded and feebly emarginate at outer apical angle, which obtuse-angled in lateral view; laterophyse feebly shorter than dorsal plate, with apex acutely hooked and bent dorsally, opposite to apical end of longitudinal ridge of dorsal plate. Female (Fig. 1 C). Similar to the males, but eyes less protruding, antennae shorter, extending to basal one-third length of elytra when reclined, III ‒ XI without impressions, pronotum wider, 1.3 times as long as wide, feebly convex on postero-lateral parts of disc, elytra darkened only at the first and second elytral interstices, pro- and mesoouter tarsal claws each with a digitiform tooth at base. Abdominal sternite VIII (Fig. 6 B) roundly emarginate in middle and deeply and nearly triangularly emarginate on both sides of posterior margin, lateral emarginate parts deeper than the middle one, the portion between lateral and middle emarginate parts acute-angled at apex, middle emarginate part membranous-lobed basally and lateral emarginate parts membranous-lobed laterally, latero-apical angles subtruncate at apices, not reaching apices of the portions between lateral and middle emarginate parts. Internal organ of reproductive system (Fig. 7 B): vagina stout and abruptly thinned at ventro-apical portion into a stout tube, where diverticulum and spermathecal duct arising separately; diverticulum moderately long, evenly thin and spiral; spermathecal duct obviously shorter and thicker than diverticulum; spermatheca provided with a spiral tube, longer than diverticulum, abruptly thinned at basal one-third, with the basal portion even in diameter and obviously thicker than apical portion; basal portion of spermatheca extended into a short thick tube, where accessory gland opening, thin and much longer than spermatheca. Body length: 8.0 – 9.0 mm (8.5 mm in holotype); width: 2.1 ‒ 2.3 mm (2.2 mm in holotype).	en	Xi, Huacong, Wang, Younan, Yang, Xingke, Liu, Haoyu, Yang, Yuxia (2021): New species and taxonomic notes on Lycocerus hickeri species-group (Coleoptera Cantharidae). Zootaxa 4980 (3): 541-557, DOI: https://doi.org/10.11646/zootaxa.4980.3.5
B01A87E94B5EFFEBCE98B290FDFFB8C5.taxon	diagnosis	Diagnosis. The new species is similar to L. nigrobilineatus but can be distinguished from the latter by the following characters: male with the pronotum wider than long, elytra with strongly developed venations on disc, lateral margins diverging posteriorly, the surface semilustrous. The aedeagus has the dorsal plates of parameres nearly as long as the ventral processes and inner margins obviously protuberant in middle. Unlike the new species, L. nigrobilineatus has the pronotum longer than wide, elytra with weakly developed venations, lateral margins nearly parallel, surface matt, and dorsal plates of parameres is shorter than the ventral processes and inner margins feebly protuberant in middle. Female has the basal portion of spermatheca extended into a short tube, where accessory gland opening, the abdominal sternite VIII is emarginate in the middle part and membranous-lobed basally, the portion between lateral and middle emarginate part is acute-angled. For comparison, L. nigrobilineatus female has an accessory gland directly opening at spermatheca, abdominal sternite VIII emarginate in the middle and not membranous-lobed, the portion between lateral and middle emarginate parts is obtuse-angled apically.	en	Xi, Huacong, Wang, Younan, Yang, Xingke, Liu, Haoyu, Yang, Yuxia (2021): New species and taxonomic notes on Lycocerus hickeri species-group (Coleoptera Cantharidae). Zootaxa 4980 (3): 541-557, DOI: https://doi.org/10.11646/zootaxa.4980.3.5
B01A87E94B5EFFEBCE98B290FDFFB8C5.taxon	etymology	Etymology. The specific name is derived from the type locality, Mainri (in Tibetan, Meili in Chinese), Yunnan Province, China.	en	Xi, Huacong, Wang, Younan, Yang, Xingke, Liu, Haoyu, Yang, Yuxia (2021): New species and taxonomic notes on Lycocerus hickeri species-group (Coleoptera Cantharidae). Zootaxa 4980 (3): 541-557, DOI: https://doi.org/10.11646/zootaxa.4980.3.5
B01A87E94B5EFFEBCE98B290FDFFB8C5.taxon	distribution	Distribution. China (Yunnan).	en	Xi, Huacong, Wang, Younan, Yang, Xingke, Liu, Haoyu, Yang, Yuxia (2021): New species and taxonomic notes on Lycocerus hickeri species-group (Coleoptera Cantharidae). Zootaxa 4980 (3): 541-557, DOI: https://doi.org/10.11646/zootaxa.4980.3.5
B01A87E94B58FFE9CE98B2ACFDF2BA06.taxon	description	Figs 3 A – B, 5 A – C, 6 C, 7 C	en	Xi, Huacong, Wang, Younan, Yang, Xingke, Liu, Haoyu, Yang, Yuxia (2021): New species and taxonomic notes on Lycocerus hickeri species-group (Coleoptera Cantharidae). Zootaxa 4980 (3): 541-557, DOI: https://doi.org/10.11646/zootaxa.4980.3.5
B01A87E94B58FFE9CE98B2ACFDF2BA06.taxon	materials_examined	Type material. Holotype: ♂ (MHBU), China, Guangxi, Wuming, Damingshan, 1100 m, 27. v. 2011, leg. H. Y. Liu. Paratypes: China, Guangxi: 5 ♀♀ (MHBU), same data as holotype; 1 ♀ (MHBU), Wuming, Damingshan, 1230 – 1423 m, 20. v. 2011, leg. H. Y. Liu.	en	Xi, Huacong, Wang, Younan, Yang, Xingke, Liu, Haoyu, Yang, Yuxia (2021): New species and taxonomic notes on Lycocerus hickeri species-group (Coleoptera Cantharidae). Zootaxa 4980 (3): 541-557, DOI: https://doi.org/10.11646/zootaxa.4980.3.5
B01A87E94B58FFE9CE98B2ACFDF2BA06.taxon	description	Description. Male (Fig. 3 A). Body black, mandibles dark brown, pronotum red, with a large dark brown marking in center of disc, elytra red, darkened at the whole length of all elytral interstices. Body densely covered with short, decumbent black pubescence, mixed with brown pubescence on head, pronotum and elytra, of which slightly longer on elytra. Head evenly narrowed behind eyes, surface densely and finely punctate; eyes distinctly protruding, head width across eyes distinctly wider than anterior margin of pronotum; terminal maxillary palpomeres long-triangular, arcuate and sharp at apical half-length of inner margins; antennae almost reaching apices of elytra when reclined, antennomere II shortest, slightly longer than wide of apices, III ‒ XI dorso-ventrally flattened, III ‒ VIII obliquely widened apically, long-triangular, III about twice as long as wide at apices, IV widest and slightly longer than III, IV ‒ XI each with an oval to round smooth impression at apical or middle part of inner margin, VIII longest, IX ‒ XI nearly parallel-sided, XI slightly longer than X and pointed at apices. Pronotum about 1.2 times longer than wide, widest at base, anterior margin rounded, lateral margins slightly diverging posteriorly, posterior margin slightly arcuate and narrowly bordered, anterior angles rounded, posterior angles rectangular, disc moderately convex at postero-lateral parts, surface semillustrous, punctate like that of head. Elytra about 4.5 times longer than pronotum, 3.3 times longer than humeral width, which about 1.5 times wider than posterior margin of pronotum, lateral margins nearly parallel, apices rounded, elytra venations strongly developed and costate, disc matt, wrinkled, sparsely and very finely punctate. Legs slender, all tarsal claws simple. Aedeagus (Figs 5 A – C): ventral process of each paramere feebly bent inwards in ventral view and bent dorsally in lateral view, with apex rounded; dorsal plates shorter than ventral processes, with inner margins diverging posteriorly, apical margins straight, inner apical angles subrounded, outer apical angles rectangular in lateral view; laterophyse feebly shorter than dorsal plate, with apex acutely hooked, directing outwards and opposite to outer apical angle of dorsal plate. Female (Fig. 3 B). Similar to the males, but body larger; eyes less protruding, anternnae narrower and shorter, extending to elytral mid-length when reclined, IV ‒ XI without impressions; pronotum nearly as long as wide; pro- and meso-outer and inner tarsal claws each with a digitiform tooth at base. Abdominal sternite VIII (Fig. 6 C) shallowly and roundly emarginate on both sides and in middle of posterior margin, lateral emarginate parts as shallow as the middle one, between which the portion acute at apex, latero-apical angles acute at apex, which on the same horizontal line as the portion between lateral and middle emarginate parts. Internal organ of reproductive system (Fig. 7 C): vagina stout and abruptly thinned at ventroapical portion into a stout tube, where diverticulum and spermathecal duct arising separately; diverticulum moderately long, thinned apically and spiral; spermathecal duct shorter and much thicker than diverticulum; spermatheca provided with a spiral tube, gradually thinned apically, longer than diverticulum; basal portion of spermatheca extended into a short thick tube, where accessory gland opening; accessory gland thin and nearly as long as spermatheca. Body length: 9.0 – 11.0 mm (9.0 mm in holotype); width: 2.0 ‒ 2.4 mm (2.0 mm in holotype).	en	Xi, Huacong, Wang, Younan, Yang, Xingke, Liu, Haoyu, Yang, Yuxia (2021): New species and taxonomic notes on Lycocerus hickeri species-group (Coleoptera Cantharidae). Zootaxa 4980 (3): 541-557, DOI: https://doi.org/10.11646/zootaxa.4980.3.5
B01A87E94B58FFE9CE98B2ACFDF2BA06.taxon	etymology	Etymology. The specific name is derived from its type locality: Guangxi Zhuang Autonomous Region, China.	en	Xi, Huacong, Wang, Younan, Yang, Xingke, Liu, Haoyu, Yang, Yuxia (2021): New species and taxonomic notes on Lycocerus hickeri species-group (Coleoptera Cantharidae). Zootaxa 4980 (3): 541-557, DOI: https://doi.org/10.11646/zootaxa.4980.3.5
B01A87E94B58FFE9CE98B2ACFDF2BA06.taxon	diagnosis	Diagnosis. It is related to L. quadrilineatus Y. Yang et X. Yang, 2014 (located in Vietnam: Sa Pa), but differs in the longer male antennae that almost reach apices of elytra when reclined, the longer pronotum, about 1.2 times as long as wide, elytra tapered at apices in male. The aedeagus has the dorsal plates of parameres with inner margins diverging posteriorly at apical part. Unlike the new species, L. quadrilineatus has the antennae extending to apical one-fourth length of elytra, pronotum hardly longer than wide, elytra rounded at apices, and dorsal plates of parameres with nearly parallel inner margins. Female has the abdominal sternite VIII moderately deeply and roundly emarginate in middle of posterior margin, spermatheca is gradually thinned apically, diverticulum is thick at base and gradually thinned apically. For comparison, L. quadrilineatus has the abdominal sternite VIII hardly emarginate in middle, spermatheca is abruptly thinned apically, and diverticulum is uniformly thin in the whole length.	en	Xi, Huacong, Wang, Younan, Yang, Xingke, Liu, Haoyu, Yang, Yuxia (2021): New species and taxonomic notes on Lycocerus hickeri species-group (Coleoptera Cantharidae). Zootaxa 4980 (3): 541-557, DOI: https://doi.org/10.11646/zootaxa.4980.3.5
B01A87E94B58FFE9CE98B2ACFDF2BA06.taxon	distribution	Distribution. China (Guangxi).	en	Xi, Huacong, Wang, Younan, Yang, Xingke, Liu, Haoyu, Yang, Yuxia (2021): New species and taxonomic notes on Lycocerus hickeri species-group (Coleoptera Cantharidae). Zootaxa 4980 (3): 541-557, DOI: https://doi.org/10.11646/zootaxa.4980.3.5
B01A87E94B5AFFE7CE98B0F7FDFFBAB1.taxon	description	Figs 4 A – B, 5 D – F, 6 D, 7 D	en	Xi, Huacong, Wang, Younan, Yang, Xingke, Liu, Haoyu, Yang, Yuxia (2021): New species and taxonomic notes on Lycocerus hickeri species-group (Coleoptera Cantharidae). Zootaxa 4980 (3): 541-557, DOI: https://doi.org/10.11646/zootaxa.4980.3.5
B01A87E94B5AFFE7CE98B0F7FDFFBAB1.taxon	materials_examined	Type material. Holotype: ♂ (MHBU), China, Yunnan, Dali, Baiquesi, 2645 m, 25 ˚ 69 ’ N, 100 ˚ 11 ’ E, 25. vi. 2020, leg. L. L. Yuan. Paratypes: China, Yunnan: 2 ♂♂, 1 ♀ (MHBU), same data as holotype; 3 ♀ (IZAS, IOZ (E) 1436574, 1437044, 1436575), Yunlong, Zhibenshan, 2500 m, 22. vi. 1981, leg. X. Z. Zhang; 1 ♀ (IZAS, IOZ (E) 1436576), same locality as the proceeding one, 21. vi. 1981, leg. S. Y. Wang.	en	Xi, Huacong, Wang, Younan, Yang, Xingke, Liu, Haoyu, Yang, Yuxia (2021): New species and taxonomic notes on Lycocerus hickeri species-group (Coleoptera Cantharidae). Zootaxa 4980 (3): 541-557, DOI: https://doi.org/10.11646/zootaxa.4980.3.5
B01A87E94B5AFFE7CE98B0F7FDFFBAB1.taxon	description	Description. Male (Fig. 4 A). Body black, mandibles dark brown, pronotum red, with a large dark brown marking in center of disc, elytra red, darkened at basal halves of first elytral interstices. Body densely covered with short, decumbent black pubescence, mixed with brown pubescence on head, pronotum and elytra, of which slightly longer on elytra. Head evenly narrowed behind eyes, surface densely and finely punctate; eyes moderately protruding, head width across eyes wider than anterior margin of pronotum; terminal maxillary palpomeres long-triangular, arcuate and sharp at apical half-length of inner margins; antennae nearly extending to apical one-sixth length of elytra when reclined, antennomere II shortest, slightly longer than wide of apices, III ‒ XI dorso-ventrally flattened, III ‒ IX obliquely widened apically, long-triangular, III about twice as long as wide at apices, IV widest and about one-third longer than III, IV ‒ XI each with an oval to round smooth impression at apical or middle part of inner margin, IX longest, X ‒ XI nearly parallel-sided, XI longer than X and pointed at apex. Pronotum 1.1 times longer than wide, widest at base, anterior margin moderately rounded, lateral margins moderately diverging posteriorly, posterior margin slightly arcuate, anterior angles rounded, posterior angles rectangular, disc moderately convex at postero-lateral parts, surface semillustrous, slightly sparsely punctate than head. Elytra about 5.8 times longer than pronotum, 3.9 times longer than humeral width, which about 1.4 times wider than posterior margin of pronotum, lateral margins nearly parallel, apices tapered, elytra venations strongly developed and costate, surface matt, wrinkled, sparsely and very finely punctate. Legs slender, all tarsal claws simple. Aedeagus (Figs. 5 D – F): ventral process of each paramere slightly bent inwards in ventral view, nearly straight in lateral view, with apex rounded; dorsal plate shorter than ventral process, longitudinal-arcuately ridged laterally on inner surface, with inner margin widely emarginate at apical half, apical margin nearly straight, inner apical angle acute-angled, outer angle obtuse-angled; laterophyse shorter than dorsal plate, with apex acutely hooked and bent dorsally, opposite to the apical curved end of longitudinal ridge of dorsal plate. Female (Fig. 4 B). Similar to the males, but body larger; eyes less protruding, head width across eyes slightly wider than anterior margin of pronotum; antennae shorter, extending to basal one-third length when reclined, antennomeres III ‒ X obliquely widened apically, each slightly longer than wide at apex and nearly triangular, IV ‒ XI without impressions, XI widened in middle; pronotum 1.1 times wider than long, lateral margins nearly parallel, with a small dark brown marking in center of disc; elytra uniformly red, with lateral margins diverging posteriorly. Abdominal sternite VIII (Fig. 6 D) moderately deeply and roundly emarginate on both sides and in middle of posterior margin, lateral emarginate parts nearly as deep as the middle one, between which the portion acute at apex, latero-apical angles subtruncate at apex, exceeding over the portion between lateral and middle emarginate parts. Internal organ of reproductive system (Fig. 7 D): vagina stout and abruptly thinned at ventro-apical portion into a stout tube, where diverticulum and spermathecal duct arising separately; diverticulum short, thinned apically and spiral; spermathecal duct hardly shorter and obviously thicker than diverticulum; spermatheca provided with a spiral tube, gradually thinned apically, obviously longer than diverticulum; basal portion of spermatheca extended into a short tube, where accessory gland opening; accessory gland thin and longer than spermatheca. Body length: 9.2 ‒ 11.5 mm (10.5 mm in holotype); width: 1.7 ‒ 2.8 mm (2.2 mm in holotype).	en	Xi, Huacong, Wang, Younan, Yang, Xingke, Liu, Haoyu, Yang, Yuxia (2021): New species and taxonomic notes on Lycocerus hickeri species-group (Coleoptera Cantharidae). Zootaxa 4980 (3): 541-557, DOI: https://doi.org/10.11646/zootaxa.4980.3.5
B01A87E94B5AFFE7CE98B0F7FDFFBAB1.taxon	etymology	Etymology. The specific name is derived from Latin rufus (red) + pennis (wing), referring to its elytra uniformly red, without black interstices.	en	Xi, Huacong, Wang, Younan, Yang, Xingke, Liu, Haoyu, Yang, Yuxia (2021): New species and taxonomic notes on Lycocerus hickeri species-group (Coleoptera Cantharidae). Zootaxa 4980 (3): 541-557, DOI: https://doi.org/10.11646/zootaxa.4980.3.5
B01A87E94B5AFFE7CE98B0F7FDFFBAB1.taxon	diagnosis	Diagnosis. This new species is similar to L. acutiapices sp. nov., but can be distinguished by the male with moderately protruding eyes, longer antennomere IV, which is about one-third longer than III, pronotum with anterior margin moderately rounded, elytra mostly red, which are only black at basal halves of first elytra interstices. The aedeagus has the ventral process of each paramere nearly straight in lateral view, dorsal plate with inner margin roundly emarginate at apical half, inner apical angles acute and outer apical angle rectangular. Unlike in the new species, L. acutiapices sp. nov. has the eyes strongly protruding, antennomere VI feebly longer than III, pronotum with anterior margin strongly rounded, elytra darkened at all elytral interstices, and ventral process of each paramere is feebly bent dorsally in lateral view, dorsal plate bears nearly straight inner margin, subrounded inner apical angle and slightly emarginate outer apical angle.	en	Xi, Huacong, Wang, Younan, Yang, Xingke, Liu, Haoyu, Yang, Yuxia (2021): New species and taxonomic notes on Lycocerus hickeri species-group (Coleoptera Cantharidae). Zootaxa 4980 (3): 541-557, DOI: https://doi.org/10.11646/zootaxa.4980.3.5
B01A87E94B5AFFE7CE98B0F7FDFFBAB1.taxon	distribution	Distribution. China (Yunnan).	en	Xi, Huacong, Wang, Younan, Yang, Xingke, Liu, Haoyu, Yang, Yuxia (2021): New species and taxonomic notes on Lycocerus hickeri species-group (Coleoptera Cantharidae). Zootaxa 4980 (3): 541-557, DOI: https://doi.org/10.11646/zootaxa.4980.3.5
B01A87E94B54FFE4CE98B338FC44BC97.taxon	description	Figs 4 C, 5 G – I	en	Xi, Huacong, Wang, Younan, Yang, Xingke, Liu, Haoyu, Yang, Yuxia (2021): New species and taxonomic notes on Lycocerus hickeri species-group (Coleoptera Cantharidae). Zootaxa 4980 (3): 541-557, DOI: https://doi.org/10.11646/zootaxa.4980.3.5
B01A87E94B54FFE4CE98B338FC44BC97.taxon	materials_examined	Type material. Holotype: ♂ (MHBU), China, Yunnan, Dali, Baiquesi, 2645 m, 25 ˚ 69 ’ N, 100 ˚ 11 ’ E, 25. vi. 2020, leg. L. L. Yuan & Y. N. Wang. Paratypes: 2 ♂♂ (MHBU), same data as holotype.	en	Xi, Huacong, Wang, Younan, Yang, Xingke, Liu, Haoyu, Yang, Yuxia (2021): New species and taxonomic notes on Lycocerus hickeri species-group (Coleoptera Cantharidae). Zootaxa 4980 (3): 541-557, DOI: https://doi.org/10.11646/zootaxa.4980.3.5
B01A87E94B54FFE4CE98B338FC44BC97.taxon	description	Description. Male (Fig. 4 C). Body black, mandibles dark brown, pronotum red, with a large dark brown marking in center of disc, elytra red, darkened at the whole length of all elytral interstices. Body densely covered with short, decumbent black pubescence, mixed with brown pubescence on head, pronotum and elytra, of which slightly longer on elytra. Head evenly narrowed behind eyes, surface densely and finely punctate; eyes strongly protruding, head width across eyes distinctly wider than anterior margin of pronotum; terminal maxillary palpomeres long-triangular, arcuate and sharp at apical half-length of inner margins; antennae nearly extending to apical one-fifth length of elytra when reclined, antennomere II shortest, slightly longer than wide at apices, III ‒ XI dorso-ventrally flattened, III ‒ X obliquely widened apically, long-triangular, III about twice as long as wide at apices, IV widest and slightly longer than III, IX longest, IV ‒ XI each with an oval smooth impression at apical part or middle of inner margin, XI longer than X, nearly parallel-sided and pointed at apex. Pronotum as long as wide, widest at base, anterior margin strongly rounded, lateral margins moderately diverging posteriorly, posterior margin slightly arcuate, anterior angles rounded, posterior angles rectangular, disc moderately convex at postero-lateral parts, surface semillustrous, slightly sparsely punctate than head. Elytra about 5.4 times longer than pronotum, 3.8 times longer than humeral width, which about 1.5 times wider than posterior margin of pronotum, lateral margins nearly parallel, apices tapered, elytra venations strongly developed and costate, surface matt, wrinkled, sparsely and very finely punctate. Legs slender, all tarsal claws simple. Aedeagus (Figs. 5 G – I): ventral process of each paramere straight in ventral view and feebly bent dorsally in lateral view, with apex feebly expanded and rounded; dorsal plate shorter than ventral process, longitudinally ridged laterally on inner surface, with inner margin nearly straight, apical margin slightly rounded, inner apical angle subrounded, outer apical angle slightly emarginate in lateral view; laterophyse shorter than dorsal plate, with apex acutely hooked, bent dorsally and directing outwards, opposite to apical end of longitudinal ridge of dorsal plate. Female. Unknown. Body length: 10.0 ‒ 11.0 mm (10.5 mm in holotype); width: 2.1 ‒ 2.4 mm (2.2 mm in holotype).	en	Xi, Huacong, Wang, Younan, Yang, Xingke, Liu, Haoyu, Yang, Yuxia (2021): New species and taxonomic notes on Lycocerus hickeri species-group (Coleoptera Cantharidae). Zootaxa 4980 (3): 541-557, DOI: https://doi.org/10.11646/zootaxa.4980.3.5
B01A87E94B54FFE4CE98B338FC44BC97.taxon	etymology	Etymology. The specific name is derived from Latin acutus (sharp) and apices (plural form of apex, tips), referring to the acute apices of elytra.	en	Xi, Huacong, Wang, Younan, Yang, Xingke, Liu, Haoyu, Yang, Yuxia (2021): New species and taxonomic notes on Lycocerus hickeri species-group (Coleoptera Cantharidae). Zootaxa 4980 (3): 541-557, DOI: https://doi.org/10.11646/zootaxa.4980.3.5
B01A87E94B54FFE4CE98B338FC44BC97.taxon	diagnosis	Diagnosis. This species is most related to L. parahickeri nom. nov., but differs in the following characters: male with the longer antennae, elytra tapered at apices, all tarsal claws simple in male. The aedeagus has the dorsal plate of each paramere with nearly straight inner margin, subrounded inner apical angle and slightly emarginate outer apical angle in lateral view. For comparison, L. parahickeri nom. nov. has the antennae only extending to apical one-third length, elytra subrounded at apices, pro- and meso-tarsal claws each bears a digitiform tooth at base of outer claw in male, and the dorsal plate of each paramere is equipped with bisinuate inner margin, acute inner apical angle and rectangular outer apical angle in lateral view.	en	Xi, Huacong, Wang, Younan, Yang, Xingke, Liu, Haoyu, Yang, Yuxia (2021): New species and taxonomic notes on Lycocerus hickeri species-group (Coleoptera Cantharidae). Zootaxa 4980 (3): 541-557, DOI: https://doi.org/10.11646/zootaxa.4980.3.5
B01A87E94B54FFE4CE98B338FC44BC97.taxon	distribution	Distribution. China (Yunnan). IGURE 5. Aedeagus (A, D, G. ventral view; B, E, H. dorsal view; C, F, I. lateral view): A – C. Lycocerus guangxiensis sp. nov.; D – F. L. rufipennis sp. nov.; G – I. L. acutiapicis sp. nov. Scale bars: 1.0 mm.	en	Xi, Huacong, Wang, Younan, Yang, Xingke, Liu, Haoyu, Yang, Yuxia (2021): New species and taxonomic notes on Lycocerus hickeri species-group (Coleoptera Cantharidae). Zootaxa 4980 (3): 541-557, DOI: https://doi.org/10.11646/zootaxa.4980.3.5
B01A87E94B56FFE5CE98B406FDF8BCA9.taxon	materials_examined	Additional material examined. China: Sichuan: 1 ♂ (MHBU), Anzihe, Shaoyaogou, 10. viii. 2016, leg. W. Y. Zhou; 1 ♂ (MHBU), “ 70 km West Chengdu, Qingcheng Hou Shan mts, 1500 m, 30 ˚ 44 ’ N, 103 ˚ 08 ’ E, 17. vi. 2005, leg. S. Murzin ”; 1 ♂ (MHBU), “ 70 km West Chengdu, Qingcheng Hou Shan mts, 1500 m, 30 ˚ 44 ’ N, 103 ˚ 08 ’ E, 8. – 14. vi. 2005, leg. S. Murzin ”; 1 ♀ (MHBU), “ Mts ling Shan, E. Mianning, 2500 – 3000 m, 14. – 18. vii. 2009, leg. S. Murzin ”.	en	Xi, Huacong, Wang, Younan, Yang, Xingke, Liu, Haoyu, Yang, Yuxia (2021): New species and taxonomic notes on Lycocerus hickeri species-group (Coleoptera Cantharidae). Zootaxa 4980 (3): 541-557, DOI: https://doi.org/10.11646/zootaxa.4980.3.5
B01A87E94B56FFE5CE98B406FDF8BCA9.taxon	etymology	Etymology. This specific name is derived from Latin mimus (imitate), referring to its similarity to L. hickeri.	en	Xi, Huacong, Wang, Younan, Yang, Xingke, Liu, Haoyu, Yang, Yuxia (2021): New species and taxonomic notes on Lycocerus hickeri species-group (Coleoptera Cantharidae). Zootaxa 4980 (3): 541-557, DOI: https://doi.org/10.11646/zootaxa.4980.3.5
B01A87E94B56FFE5CE98B406FDF8BCA9.taxon	discussion	Remarks. Lycocerus sichuanus Y. Yang et X. Yang, 2014, is preoccupied by L. sichuanus (Wittmer, 1995), originally described in Athemus. When two or more names are homonyms, only the senior name, as determined by the Principle of Priority, may be used as a valid (ICZN, 4 th ed., article 52.2), so L. sichuanus (Wittmer, 1995) remains valid. A junior homonym must be rejected and replaced either by an available and potentially valid synonym or, for lack of such a name, by a new substitute name (ICZN, 4 th ed., article 60.1), therefore, L. hickerimimus nom. nov. is proposed here to replace L. sichuanus Y. Yang et X. Yang, 2014.	en	Xi, Huacong, Wang, Younan, Yang, Xingke, Liu, Haoyu, Yang, Yuxia (2021): New species and taxonomic notes on Lycocerus hickeri species-group (Coleoptera Cantharidae). Zootaxa 4980 (3): 541-557, DOI: https://doi.org/10.11646/zootaxa.4980.3.5
B01A87E94B56FFE5CE98B406FDF8BCA9.taxon	distribution	Distribution. China (Sichuan).	en	Xi, Huacong, Wang, Younan, Yang, Xingke, Liu, Haoyu, Yang, Yuxia (2021): New species and taxonomic notes on Lycocerus hickeri species-group (Coleoptera Cantharidae). Zootaxa 4980 (3): 541-557, DOI: https://doi.org/10.11646/zootaxa.4980.3.5
B01A87E94B50FFE3CE98B1E8FBFCBF6E.taxon	materials_examined	Additional material examined. China, Hubei: 8 ♀♀, 6 ♂♂ (MHBU), Luoyanghe, 31. v. 2019, leg. P. Wang; 1 ♀ (MHBU), Shennongjia, Dongxi, 819 m, 31 ˚ 34 ’ 15 ” N, 110 ˚ 08 ’ 47 ” E, 7. – 10. vii. 2014, leg. Y. B. Ba & S. Y. Tang; 1 ♂ (MHBU), Shennongjia, 27. v. 2018, leg. P. Wang; 1 ♂ (MHBU), Yichang, Longmenhe, 25. vi. 2010, leg. Y. Li; 1 ♂ (MHBU), Dalaoling Forest Farm, 10. vii. 2014, leg. Y. P. Lin; 1 ♂ (MHBU), Dalaoling Forest Farm, 13. vii. 2014, leg. L. F. Xiang. Shaanxi: 2 ♀♀ (MHBU), Zhenan, Cigou, 1100 m, 33 ˚ 30 ’ 0 ” N, 109 ˚ 1 ’ 14 ” E, 21. vi. 2014, leg. C. X. Yuan & Y. Tian; 1 ♀ (MHBU), Danfeng, Caichuan, Mangshan, 1417 m, 33 ˚ 56 ’ 20 ” N, 110 ˚ 19 ’ 34 ” E, 2. vii. 2014, leg. C. X. Yuan & Y. Tian; 1 ♂ (MHBU), Zhashui, Zhonghe, 900 m, 33 ˚ 34 ’ 7 ” N, 109 ˚ 24 ’ 2 ” E, 25. vi. 2014, leg. C. X. Yuan & Y. Tian.	en	Xi, Huacong, Wang, Younan, Yang, Xingke, Liu, Haoyu, Yang, Yuxia (2021): New species and taxonomic notes on Lycocerus hickeri species-group (Coleoptera Cantharidae). Zootaxa 4980 (3): 541-557, DOI: https://doi.org/10.11646/zootaxa.4980.3.5
B01A87E94B50FFE3CE98B1E8FBFCBF6E.taxon	etymology	Etymology. This specific name is derived from Greek para - (near), referring to its close relationship to L. hickeri.	en	Xi, Huacong, Wang, Younan, Yang, Xingke, Liu, Haoyu, Yang, Yuxia (2021): New species and taxonomic notes on Lycocerus hickeri species-group (Coleoptera Cantharidae). Zootaxa 4980 (3): 541-557, DOI: https://doi.org/10.11646/zootaxa.4980.3.5
B01A87E94B50FFE3CE98B1E8FBFCBF6E.taxon	discussion	Remarks. Lycocerus hubeiensis Y. Yang et X. Yang, 2014, is preoccupied by L. hubeiensis (Švihla, 2004), originally described in Athemus. When two or more names are homonyms, only the senior, as determined by the Principle of Priority, may be used as a valid name (ICZN, 4 th ed., article 52.2), so L. hubeiensis (Švihla, 2004) remains valid. A junior homonym must be rejected and replaced either by an available and potentially valid synonym or, for lack of such a name, by a new substitute name (ICZN, 4 th ed., article 60.1), therefore, L. parahickeri nom. nov. is proposed here to replace L. hubeiensis Y. Yang et X. Yang, 2014. Additionally, in the original description (Yang et al. 2014), the tarsal claws of female were described having basal teeth at pro- and meso-outer tarsal claws. What’s noted here, not only outer claws but also inner claws of pro- and meso-legs are equipped with digitiform teeth in female, based on the examination of a large series of additional material available for the present study.	en	Xi, Huacong, Wang, Younan, Yang, Xingke, Liu, Haoyu, Yang, Yuxia (2021): New species and taxonomic notes on Lycocerus hickeri species-group (Coleoptera Cantharidae). Zootaxa 4980 (3): 541-557, DOI: https://doi.org/10.11646/zootaxa.4980.3.5
B01A87E94B50FFE3CE98B1E8FBFCBF6E.taxon	distribution	Distribution. China (Hubei, Shaanxi). Newly recorded to Shaanxi Province.	en	Xi, Huacong, Wang, Younan, Yang, Xingke, Liu, Haoyu, Yang, Yuxia (2021): New species and taxonomic notes on Lycocerus hickeri species-group (Coleoptera Cantharidae). Zootaxa 4980 (3): 541-557, DOI: https://doi.org/10.11646/zootaxa.4980.3.5
