taxonID	type	description	language	source
0391C576F26AFF8E6398FF07FD704700.taxon	description	(Figs 1 A – 1 H, 2 A – 2 V)	en	Kawase, Naoki (2021): Description of three new species of Psilotreta (Trichoptera: Odontoceridae) from Japan. Zootaxa 4980 (3): 589-598, DOI: https://doi.org/10.11646/zootaxa.4980.3.10
0391C576F26AFF8E6398FF07FD704700.taxon	diagnosis	Diagnosis. This species is easily distinguished from the previously known Japanese species by the shapes of the male genitalia. Although the male genitalia of this species are very similar to those of Psilotreta excavata Yuan et al. 2008 described from the Chinese mainland, Psilotreta clyssan Malicky 2014 from Taiwan, and Psilotreta moritai sp. nov., the male of P. voluta can be distinguished from latter three species by the shape of the ventral projection of the lateral processes and intermediate appendages of the male genitalia. The male of P. voluta is distinguishable from P. excavata by following structures: (1) the intermediate appendages of P. voluta are strongly curved, usually forming a C-shaped structure in lateral view, but those of P. excavata are more strongly curled, almost circular in shape; (2) ventral projections of the lateral processes are extended more anteriorly in P. voluta, but directed right ventrad in P. excavata; (3) the pair of parameres in the phallus are positioned near the center in P. voluta, but are positioned outside along the outer edges in P. excavata in ventral view. The male of P. voluta is distinguished from P. clyssan by following structures: (1) the median dorsal process of segment X is slightly clavate in P. voluta, but tapering to a weakly acuminate apex in P. clyssan; (2) the intermediate appendages of P. voluta are strongly curved, usually forming a C-shaped structure in lateral view, but those of P. c lyssan are more strongly curled, almost circular in shape; (3) with one or two spines along the outer edge of each intermediate appendage in P. voluta, but lacking spines in P. clyssan. Diagnosis of P. voluta and P. moritai is discussed in the description of the next following species, P. moritai. Adult Male and Female. (Figs 1 A, 1 B). General coloration brown for body, antennae, and wings. First and second segments of maxillary palps about same length. Head in dorsal view typical for genus (Parker & Wiggins 1987): anteromesal warts between bases of antennal sockets; pair of small oval anterior warts and larger oval posterior warts present; elongate, narrow posterolateral warts extending ventrad along posterior margins of eyes (Fig. 1 A). Length of each forewing: male 7.6 – 9.7 mm (mean = 8.9 ± 0.68, n = 7), female 9.2 – 10.3 mm (mean = 9.83 ± 0.36, n = 6). In venation (Fig. 1 B), discoidal cell present in each fore- and hind wing; male forewings each with apical forks I, II, and V (M 1 + 2, M 3 + 4 undivided); fork I rooted on discoidal cell about 1 / 3 its length, Cu 1 a fused with M 3 + 4; and Cu 1 a and Cu 1 b separate, Cu 2 apparently absent, row of setae parallel with posterior margin suggesting wing coupling mechanism; male hind wings each with apical forks I and II (M 1 + 2, M 3 + 4, and Cu 1 undivided), fork I rooted on discoidal cell about 2 / 3 its length, crossvein between M 3 + 4 and Cu 1 absent. Male Genitalia (Figs 1 C – 1 F, 2 A – 2 V). Tergum IX elongate, subtriangular in dorsal view, with finely granular surface and steep sides above basal setal warts. Basal segment of each inferior appendage shorter than preanal appendages, cylindrical and slightly tapered near apex, covered with setae; apical segment about 0.4 X as long as basal segment, cylindrical with many small brown teeth on apical half. Preanal appendages elongate, compressed (flattened, about 4 times as long as thick in dorsal view), parallel-sided in lateral and dorsal views, extending beyond posterior margins of lateral processes in lateral view. Tergum X forming median dorsal process slightly clavate apically, posterior margin usually rounded but sometimes shallowly notched (Figs 1 D, 1 E); lateral processes each forming subtriangular or oval plate with ventral projection heavily sclerotized and variously shaped (even at same localities), usually curved anteroventrad and acutely pointed at apex (Figs 2 A – 2 H); intermediate appendages dark brown, heavily sclerotized, strongly curved, usually forming C-shaped structure in lateral view, each with 1 or 2 spines along outer edge, ending in acute apex directed ventrad (or rarely posteroventrad). Phallus with phallotheca long, cylindrical; endotheca ventrally with pair of parameres variously prolonged among localities: in Shikoku (Kochi, Ehime, and Tokushima), its length about 4 to 5 times its basal width; in Honshu (Okayama and Shimane) and northern Kyushu (Nagasaki), the length about 5 to 6 times; and in southern Kyushu (Kagoshima), the length more than 6 times as long; aedeagus with ventral plate slightly sclerotized, membranous apex rounded and slightly enlarged, internal phallotremal sclerite V-shaped in ventral view, strongly curved dorsad in lateral view (Figs 2 S – 2 V). Female Genitalia (Figs 1 G – 1 H). Length of segment IX about as long as wide in ventral view. Vaginal chamber within segment IX; anterior vaginal sclerite weakly sclerotized; posterior vaginal sclerites heavily sclerotized distally. Larva, Pupa. Unknown.	en	Kawase, Naoki (2021): Description of three new species of Psilotreta (Trichoptera: Odontoceridae) from Japan. Zootaxa 4980 (3): 589-598, DOI: https://doi.org/10.11646/zootaxa.4980.3.10
0391C576F26AFF8E6398FF07FD704700.taxon	materials_examined	Holotype. male (in alcohol), Japan, Shikoku, Kochi, Ino-cho, Teragawa (Yosakoi pass), 33 ° 45 ' N, 133 ° 11 ' E, alt. 1350 m, 3. viii. 2001, I. Yamashita [Izumi Yamashita] (LBM 141001205). Paratypes. Japan, Shikoku, Kochi: 1 male, 1 female, same locality as holotype, 7. vii. 2001, M. Takai [Mikio Takai] (LBM 141001206 – 141001207); 1 male, same locality as holotype, 16. vii. 2000, I. Yamashita (LBM 141001208); 1 female, same locality as holotype, 18. vii. 2005, M. Takai (LBM 141001209); 1 male, same locality as holotype, 14. vii. 2006, M. Takai (MKNC 2021001); 1 female, same locality as holotype, 14. vii. 2001, I. Yamashita (MKNC 2021002); 1 female, same locality as holotype, 9. vii. 2004, M. Takai (MKNC 2021003); 1 female, same locality as holotype, 17. vii. 2004, M. Takai (MKNC 2021004); 1 male and 1 female, Kami-shi, Monobe-son, Kubokage, 33 ° 48 ' N, 133 ° 59 ' E, 25. vi. 2006, M. Takai (CBM 0180204 – 0180205); 1 male, Kami-shi, Monobe-son, Sasa, 33 ° 47 ' N, 133 ° 53 ' E, 3. vii. 2004, M. Takai (CBN 0180206). Ehime: 1 male (pinned), Saijyo-shi, Komatsu-cho, Ishizuchi, Ishizuchi Shrine Jojusha, 33 ° 47 ' N, 133 ° 07 ' E, 26. vii. 1952, “ J. I. & T. I. ” (EUM). Tokushima: 1 male, Mima-shi, Minokoshi, 33 ° 52 ’ N, 134 ° 05 ’ E, 4. viii. 2003, Kaori Nio (MKNC 2021005). Honshu, Okayama: 2 males, Kagamino-cho, Hadenishitani, 35 ° 13 ' N, 133 ° 49 ' E, 12. vii. 2013, Koichi Nojima (MKNC 2021006 – 2021007). Shimane: 1 female, Okuizumo-cho, takezaki (Mt. Sentsu), 35 ° 10 ' N 133 ° 10 ' E, 26. vi – 5. vii. 2007, Masakazu Hayashi (MKNC 2021008); 2 females, ibid., 5 – 22. vii. 2007, Masakazu Hayashi (MKNC 2021009 – 2021010); 1 male, ibid., 6 – 7. vii. 2019, N. Kawase & H. Morita [Hisayuki Morita] (MKNC 2021011). Kyushu, Nagasaki: 1 male, Goto-shi, Tomie-machi, Shigejiki, 32 ° 39 ' N, 128 ° 45 ' E, larva collected on 28. viii. 2018, adult emerged on iii. 2019 by Noriyoshi Shimura (MKNC 2021012); 2 females, ibid., larvae collected on 28. viii. 2018, adult emerged on 16. v. 2020 by Noriyoshi Shimura (MKNC 2021013 – 2021014). Kagoshima: 2 males, Yakushima-cho, Kurio, 30 ° 18 ‘ N, 130 ° 24 ‘ E, 10 – 11. v. 2006, Tomiko Ito (MKNC 2021015 – 2021016).	en	Kawase, Naoki (2021): Description of three new species of Psilotreta (Trichoptera: Odontoceridae) from Japan. Zootaxa 4980 (3): 589-598, DOI: https://doi.org/10.11646/zootaxa.4980.3.10
0391C576F26AFF8E6398FF07FD704700.taxon	etymology	Etymology. The Latin specific epithet voluta, “ volute ” in English, is derived from the sharply curved intermediate appendages in the male genitalia.	en	Kawase, Naoki (2021): Description of three new species of Psilotreta (Trichoptera: Odontoceridae) from Japan. Zootaxa 4980 (3): 589-598, DOI: https://doi.org/10.11646/zootaxa.4980.3.10
0391C576F26AFF8E6398FF07FD704700.taxon	distribution	Distribution. Western Honshu (Okayama, Shimane), Shikoku (Ehime, Kochi, Tokushima), and Kyushu (Kagoshima-Yakushima, Nagasaki-Fukue) Islands. Habitat and biology. Adults collected near small mountain streams by net sweeping or light traps in late June to August. Japanese name. Uzu-kiso-tobikera.	en	Kawase, Naoki (2021): Description of three new species of Psilotreta (Trichoptera: Odontoceridae) from Japan. Zootaxa 4980 (3): 589-598, DOI: https://doi.org/10.11646/zootaxa.4980.3.10
0391C576F26AFF8E6398FF07FD704700.taxon	discussion	Remarks. Kuwada (1965) and Tanida (2008) recorded P. kisoensis from Shikoku based on a male specimen. I examined the specimen deposited in Ehime University Museum and determined it as P. voluta sp. nov. There is no reliable record of P. kisoensis from Shikoku at present.	en	Kawase, Naoki (2021): Description of three new species of Psilotreta (Trichoptera: Odontoceridae) from Japan. Zootaxa 4980 (3): 589-598, DOI: https://doi.org/10.11646/zootaxa.4980.3.10
0391C576F26FFF8C6398FA7EFDF04660.taxon	description	(Figs 3 A – 3 L)	en	Kawase, Naoki (2021): Description of three new species of Psilotreta (Trichoptera: Odontoceridae) from Japan. Zootaxa 4980 (3): 589-598, DOI: https://doi.org/10.11646/zootaxa.4980.3.10
0391C576F26FFF8C6398FA7EFDF04660.taxon	diagnosis	Diagnosis. The male of this species is similar to that of P. voluta sp. nov., but can be distinguished from the latter by the following male genital structures: (1) the basal segment of each inferior appendage is thick and clearly dilated at the middle in lateral view (slender and almost cylindrical in P. voluta); (2) the preanal appendages are broadly oval, 2.5 times as long as wide in lateral view (more slender, about 4 times as long as wide in P. voluta); (3) each lateral process has a short, almost straight, sword-like ventral projection directed anterad with a slightly rounded or truncate apex (in P. voluta, the ventral projection is curved anteroventrad with an acutely pointed apex); (4) each intermediate appendage lacks spines on the outer edge (in P. voluta, each intermediate appendages has one (or two) spine (s) along the outer edge); and (5) the aedeagus has a membranous protrusion dorsally on the surface in lateral view (in P. voluta, with no protrusions dorsally on the surface). Adult Male. General coloration reddish to pale brown for body, antennae, and wings. Maxillary palps approximately same as for P. voluta sp. nov. Head in dorsal view similar to that of P. voluta sp., but oval anterior and posterior warts smaller (Fig. 3 A). Length of each male forewing 7.0 – 7.3 mm (mean = 7.18 ± 0.11, n = 4). Venation similar to that of P. voluta sp. nov., but discoidal cell of male hind wings open in two of five specimens examined in this study (Fig. 3 B). Male Genitalia (Figs 3 C – 3 L). Tergum IX subtriangular in dorsal view, with steep sides below basal setal warts. Basal segment of each inferior appendage thick, dilated at middle in lateral view and covered with setae, extending to posterior margin of preanal appendages; apical segment nearly 2 / 3 as long as basal segment in lateral view, cylindrical, with many brown teeth on apical half. Preanal appendages broadly oval in lateral view, compressed. Segment X with median dorsal process apically clavate, bearing several setae around apex; lateral processes each with ventral projection heavily sclerotized, stout and directed straight anterad, with its base derived from very posterior end of lateral process or just at base of its intermediate appendage (Figs 3 D – 3 G); each intermediate appendage heavily sclerotized, curved posterad and ventrad in lateral view, ending in acute apex directed ventrad (or rarely posterad). Phallus with phallotheca long, cylindrical; endotheca with pair of parameres, each only about 4 times as long as its basal width in ventral view; aedeagus with membranous protrusion dorsally, ventral plate slightly sclerotized, membranous apex slightly truncate and dilated, internal phallotremal sclerite strongly curved toward apex, L-shaped in lateral view, moderately broadened apically in ventral view (Figs 3 K, 3 L). Female, Larva, Pupa. Unknown.	en	Kawase, Naoki (2021): Description of three new species of Psilotreta (Trichoptera: Odontoceridae) from Japan. Zootaxa 4980 (3): 589-598, DOI: https://doi.org/10.11646/zootaxa.4980.3.10
0391C576F26FFF8C6398FA7EFDF04660.taxon	materials_examined	Holotype. male (in alcohol), Japan, Honshu, Mie, Kumano-shi, Hobo-cho (Tategasaki), 33 ° 56 ' N, 136 ° 11 ' E, alt. 50 m, 31. v. 2006, N. Kawase (LBM 141001210). Paratypes. Japan, Honshu, Mie: 1 male, same data as holotype (LBM 141001211); 2 males, same locality as holotype, 1. vi. 2002, H. Morita (MKNC 2021017 – 18); 1 male, Owase-shi, Mikisato-cho, 33 ° 59 ' N, 136 ° 10 ' E, 16. vii. 2016, H. Morita (MKNC 2021019); 1 male, Kihoku-cho, Aiga, 34 ° 07 ' N, 136 ° 10 ' E, 2. vii. 2016, H. Morita (CBM 0180207).	en	Kawase, Naoki (2021): Description of three new species of Psilotreta (Trichoptera: Odontoceridae) from Japan. Zootaxa 4980 (3): 589-598, DOI: https://doi.org/10.11646/zootaxa.4980.3.10
0391C576F26FFF8C6398FA7EFDF04660.taxon	etymology	Etymology. The Latin species epithet moritai is derived from Hisayuki Morita who first collected specimens of this species.	en	Kawase, Naoki (2021): Description of three new species of Psilotreta (Trichoptera: Odontoceridae) from Japan. Zootaxa 4980 (3): 589-598, DOI: https://doi.org/10.11646/zootaxa.4980.3.10
0391C576F26FFF8C6398FA7EFDF04660.taxon	distribution	Distribution. Central Honshu (Kii Peninsula). Habitat and Biology. Adults were collected around mountainous streams at light from May to July. Japanese name. Morita-kiso-tobikera.	en	Kawase, Naoki (2021): Description of three new species of Psilotreta (Trichoptera: Odontoceridae) from Japan. Zootaxa 4980 (3): 589-598, DOI: https://doi.org/10.11646/zootaxa.4980.3.10
0391C576F26DFF826398FB5EFDEB41B8.taxon	description	(Figs 4 A – 4 K)	en	Kawase, Naoki (2021): Description of three new species of Psilotreta (Trichoptera: Odontoceridae) from Japan. Zootaxa 4980 (3): 589-598, DOI: https://doi.org/10.11646/zootaxa.4980.3.10
0391C576F26DFF826398FB5EFDEB41B8.taxon	diagnosis	Diagnosis. This new species is somewhat similar to P. voluta sp. nov. and P. moritai sp. nov. described in this study but can be distinguished from them by a character of the male genitalia. In P. flavida, each lateral process of segment X has a ventral projection directed posterad instead of anteroventrad as in P. voluta and P. moritai. Adult Male (Figs 4 A – 4 C). General coloration yellowish to pale brown in body, antennae and wings. Maxillary palps approximately same as for P. voluta sp. nov. Head in dorsal view similar to that of P. voluta, but notch between anteromesal warts deeper than for P. voluta (Fig. 4 A). Length of each forewing: male 6.6 – 7.3 mm (mean = 7.1 ± 0.19, n = 22), female 6.8 – 8.1 mm (mean = 7.5 ± 0.45, n = 7). Venation similar to P. voluta sp. nov. and P. moritai sp. nov., but cross-vein between M 3 + 4 and Cu 1 present in male hind wings (Fig. 4 C). Male Genitalia (Figs 4 D – 4 I). Tergum IX subtriangular in dorsal view, with steep sides above rounded basal setal warts. Basal segment of each inferior appendage long, extending beyond posterior margin of lateral process, cylindrical and covered with setae, slightly tapered; apical segment about 1 / 4 length of basal segment, cylindrical, with many brown teeth on apical half. Preanal appendages broad at basal 3 / 4 and abruptly tapered distally in lateral view, not extending beyond posterior apex of median dorsal process. Segment X with median dorsal process narrow subapically, slightly clavate, sparsely setose apically; lateral processes each with acute ventral projection apically directed caudad and upcurved; each intermediate appendage heavily sclerotized, sharply curved posteroventrad, usually forming C shape in lateral view, positioned on lower 2 / 3 of lateral process with its top never reaching dorsal margin of lateral process. Phallus with phallotheca short, cylindrical; endotheca with pair of short and broad parameres located lateroventrally, about 2 times as long as its basal width; aedeagus slightly depressed dorsally in lateral view, with ventral plate weakly sclerotized, membranous apex nearly rounded with posterior margin slightly notched in ventral view, internal phallotremal sclerite distinct, V-shaped in ventral view, strongly C-shaped in lateral view. Female Genitalia (Figs. 4 J – 4 K). Vaginal chamber almost 2 times as long as sternum IX in ventral view; posterior vaginal sclerites heavily sclerotized apically. Larva, Pupa. Unknown.	en	Kawase, Naoki (2021): Description of three new species of Psilotreta (Trichoptera: Odontoceridae) from Japan. Zootaxa 4980 (3): 589-598, DOI: https://doi.org/10.11646/zootaxa.4980.3.10
0391C576F26DFF826398FB5EFDEB41B8.taxon	materials_examined	Holotype. male, Japan, Shikoku, Ehime, Uchiko-cho, Odamiyama (Namakusa valley), 33 ° 32 ’ N, 132 ° 50 ’ E, alt. 1200 m, 26. vii. 1998, E. Yamamoto [Eiji Yamamoto] (LBM 141001212). Paratypes. Japan, Shikoku, Ehime: 18 males and 5 females, same locality as holotype, 8. viii. 2000, E. Yamamoto. (LBM 141001213 – 141001235); 1 male, same locality as holotype, 28. vii. 2000, E. Yamamoto (CBM 0180208); 2 males, same locality as holotype, 9. viii. 2016, E. Yamamoto (CBM 0180209 – 0180210); 1 male, Oozu-shi, Hijikawa-cho, Yamatosaka, 33 ° 28 ’ N, 132 ° 43 ’ E, 7 – 9. viii. 2000, Tsukaguchi (MKNC 2021020); 1 male, Kumakogen-cho, Yurano, 33 ° 38 ’ N, 132 ° 53 E, 10. vii. 2007, E. Yamamoto (MKNC 2021021); 1 male, ibid, 20. vii. 2007, E. Yamamoto (MKNC 2021022); 1 male, ibid, 31. vii. 2007, E. Yamamoto (MKNC 2021023). Kochi: 4 males, Niyodogawa-cho, Ooue (Mt. Torigata), 33 ° 27 ’ N, 133 ° 04 ’ E, 25. vii. 2001, N. Kawase (MKNC 2021024 – 2021027); 2 males, Ino-cho, Teragawa (Yosakoi pass), 33 ° 45 ' N, 133 ° 11 ' E, 3. viii. 2001, I. Yamashita (MKNC 2021028 – 2021029); 1 male, Kamishi, Monobe-cho, Kubokage (Nishikuma valley), 33 ° 47 ' N, 133 ° 58 ' E, 5. viii. 2006, I. Yamashita (MKNC 2021030); 1 male, ibid, 26. viii. 2006, I. Yamashita (MKNC 2021031); 1 male, Kochi-shi, Tosayama-Takakawa (Mt. Kuishi), 33 ° 40 ’ N, 133 ° 30 ’ E, 8. vi. 2005, M. Takai (MKNC 2021032); 2 males, Tosashimizu-shi, Kaikake, 32 ° 51 ’ N, 132 ° 54 ’ E, 28. vi. 2003, M. Takai (MKNC 2021033 – 2021034); 3 males and 2 females, ibid, 26. vi. 2004 (CBM 0180211 – 0180215); 1 male, Sukumo-shi, Yamana-cho, Ittyubara, 33 ° 00 ’ N, 132 ° 48 ’ E, 15. vi. 2001, M. Takai (MKNC 2021035).	en	Kawase, Naoki (2021): Description of three new species of Psilotreta (Trichoptera: Odontoceridae) from Japan. Zootaxa 4980 (3): 589-598, DOI: https://doi.org/10.11646/zootaxa.4980.3.10
0391C576F26DFF826398FB5EFDEB41B8.taxon	etymology	Etymology. The Latin specific epithet flavida, “ yellow ” in English, refers to the yellowish general coloration of this new species.	en	Kawase, Naoki (2021): Description of three new species of Psilotreta (Trichoptera: Odontoceridae) from Japan. Zootaxa 4980 (3): 589-598, DOI: https://doi.org/10.11646/zootaxa.4980.3.10
0391C576F26DFF826398FB5EFDEB41B8.taxon	distribution	Distribution. Shikoku. Habitat and Biology. Adults were collected near streams in mountain forests from June (alt. about 100 m on Sukumo-shi and Tosashimizu-shi) to August (alt. about 1000 m in Uchiko-cho) by a light trap or Malaise trap. Japanese Name. Kiiro-kiso-tobikera	en	Kawase, Naoki (2021): Description of three new species of Psilotreta (Trichoptera: Odontoceridae) from Japan. Zootaxa 4980 (3): 589-598, DOI: https://doi.org/10.11646/zootaxa.4980.3.10
