taxonID	type	description	language	source
E0399246B03058489FF01EE8B740D122.taxon	materials_examined	Examined material. Mesembrius arcuatus Jordaens, Goergen & De Meyer: Holotype, male, " HOLOTYPUS " " Entebbe, // Uganda. // 21.8.11. // C. C. Gowdey. // 1912 - 100. " " Mesembrius arcuatus // Det. K. Jordaens, 2019 " " NHMUK 010369965 " [NHMUK]. Paratypes: Uganda • 1 ♂; Entebbe; 11 Aug 1911; C. C. Gowdey leg.; NHMUK • 1 ♂; Entebbe; 14 Aug 1911; C. C. Gowdey leg.; NHMUK • 2 ♂♂; Entebbe; 14 Aug 1911; C. C. Gowdey leg.; NHMUK • 2 ♂♂, Entebbe; 16 Aug 1911; C. C. Gowdey leg.; NHMUK • 1 ♂; Entebbe; 17 Aug 1911; C. C. Gowdey leg.; NHMUK • 3 ♂♂; Entebbe; 21 Aug 1911; C. C. Gowdey leg.; NHMUK • 1 ♂; Entebbe; 3 - 4 Dec 1912; C. C. Gowdey leg.; NHMUK • 1 ♂; Entebbe; 13 Nov 1912; C. C. Gowdey leg.; NHMUK • 2 ♂♂; Entebbe; 16 Oct 1912; C. C. Gowdey leg.; NHMUK • 5 ♂♂; Entebbe; 1 - 11 Sep 1911; S. A. Neave leg.; NHMUK • 1 ♂; N. E. Side of Lake Albert; 1906; A. Hodges leg.; NHMUK • 1 ♂; near Entebbe; 5 Mar 1972; H. Falke leg.; CNC • 1 ♂; near Entebbe; 1 - 14 Feb 1973; H. Falke leg.; CNC • 1 ♂;? Kanue; 3 May 1911; collector unknown; NHMUK • 1 ♂; Mbarara; 29 May 1911; collector unknown; NHMUK; 1 ♂; Central Region, Wakisa District, Mabamba Swamps; 16 Dec 2018; X. Mengual; ZFMK • 2 ♂♂, Central Region, Wakisa District, Mabamba Swamps; 16 Dec 2018; M. Reemer leg.; RMNH • 1 ♂; Central Region, Wakisa District, Mabamba Swamps 1 ♂; 16 Dec 2018; K. Jordaens leg.; KMMA.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
E0399246B03058489FF01EE8B740D122.taxon	description	Description male (Fig. 3). Body length: 13.7 - 14.2 mm. Wing length: 9.8 - 10.4 mm. Head (Fig. 46). Eyes bare; holoptic, length of eye contiguity equal to approx. the length of ocellar triangle. Face dark with dark medial vitta; white pilose; white pollinose. Vertical triangle black pilose; white pollinose in dorsal half. Ocellar triangle dark; black pilose; white pollinose; distance between lateral ocellus and eye margin 1 / 2 width of ocellus. Occiput yellow; white pilose; yellow and white pollinose. Frontal triangle orange-brown; white pilose. Frontal prominence shiny black; black pilose. Antenna dark brown to black; postpedicel white pollinose; antennal arista reddish-brown. Thorax. Scutum black with a pair of dorsal, well-demarcated grey pollinose vittae; yellow and black pilose. Scutellum uniformly light yellow-brown; yellow pilose throughout, black pilose on posterior 2 / 3. Legs. All femora and tibiae with long, loose, yellow pile and, especially at distal end, loose, black pile. Proleg (Fig. 157): Femur dark brown to black; with a loose, apical pile brush which is black pilose dorsally, yellow pilose ventrally; with long, yellow pile posterodorsally and with shorter, black pile anterodorsally and anteroventrally. Tibia with long, black pile, except dorsally. Basitarsus orange-brown; with long, black pile posteriorly. Other tarsi orange-brown; yellow and black pilose dorsally; orange pilose ventrally. Mesoleg: Dark brown to black, except for tarsi which are reddish-brown; black pilose dorsally, orange pilose ventrally. Metaleg (Fig. 191): Femur dark brown to black; very slender; covered with long, thin yellow pile, except on ventral side which is almost bare; with shorter, black pile on posteroventral proximal 2 / 3; with a series of thick, black setulae at distal 1 / 3; with thick, black pile at distal end. Tibia curved; strongly dorsoventrally compressed in middle 1 / 3; with long, yellow and black pile, except in the flattened posterior section. Tarsi yellow and black pilose, except ventrally where orange pilose. Wing (Fig. 127). Entire wing uniformly microtrichose. Abdomen (Fig. 83). Tergite II with a pair of very large, yellow triangular to rounded maculae; yellow pilose; black markings hourglass-shaped; posterior black marking equal in size or somewhat narrower than anterior black marking, with a medial white pollinose area; posterior black marking with black pile that posterolaterally extends into the yellow maculae. Tergite III with yellow-orange fascia and a large, black and strongly white pollinose triangular marking; pile short, stiff and black, except on the lateral sides where it is longer, thinner and yellow. Tergite IV strongly white pollinose anteriorly, with a large posterior, rounded, white pollinose black marking; pile short, thick and black, except on the lateral sides where it is longer, thinner and yellow. Genitalia (Fig. 205). Epandrium: Dorsal lobe of surstylus short and stout, with a very long, sharp expansion on distal end; short black spinose on apex and long brown pilose on dorsal surface. Ventral lobe of surstylus strongly convex; bare.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
E0399246B03058489FF01EE8B740D122.taxon	distribution	Distribution. Uganda.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
E0399246B03058489FF01EE8B740D122.taxon	etymology	Etymology. The specific epithet Mesembrius arcuatus (Latin) means bent like a bow and was chosen with reference to the strongly curved metatibia. It is to be treated as an adjective (nominative singular masculine).	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
243A5D3ADB205ECF9714BB61D130610A.taxon	materials_examined	Examined material. Helophilus caffer Loew: Lectotype (hereby designated), male, " Helophilus // Helophilus caffer // ♂ " " 206 " " 207 " " Loan // 575 / 99 " " NHRS-BYWS // 000002618 " [NRMS]. Helophilus caffer Loew: Paralectotype, female, " Helophilus // Helophilus caffer // ♀ " " 206 " " Loan // 575 / 99 " " NHRS-BYWS // 000002617 " [NRMS]. Mesembrius ctenifera Hull: Holotype, male, " Mesembrius // Mesembrius ctenifera // Hull n. sp. " " TYPE 6596 // Mesembrius // Mesembrius ctenifera // F. M. Hull " " Oriental forest // Fanovana Dist. // Fianarantsoa // Madagascar " " 1 - V, 1937. // (C. Lamberton) " [ANSP]. Mesembrius ctenifera Hull: Allotype, female, " Allotype ♀ // Mesembrius // Mesembrius ctenifera // F. M. Hull " " Oriental forest // Fanovana, Dist. // Fianarantsoa // Madagascar " [ANSP].	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
243A5D3ADB205ECF9714BB61D130610A.taxon	materials_examined	Other material (nominal morph; see below). Burundi • 1 ♀; Bujumbura; 20 Feb 2017; G. Goergen leg.; IITA. Cameroon • 1 ♀; Maroua, Meskina; 23 May 2018; M. Azo'o Ela leg.; MAPC. Democratic Republic of the Congo • 1 ♂,? Adu; 8 Apr 1955; collector unknown; KMMA • 1 ♀; Coquilhatville [= Mbandaka], Bamania, Equateur; 21 Jul 1924; J. Bequaert leg.; KMMA • 1 ♂; Bambesa, Bas-Uele; Dec 1933; H. J Bredo leg.; KMMA • 1 ♀; Barumbu, Leopoldville [= Kinshasa]; 15 Oct 1910; Dr. Bequaert leg.; KMMA • 1 ♀; Boma; 15 Jul 1920; H. Schouteden leg.; KMMA • 1 ♀; Bondo, Bas-Uele; J. J. Rodhain leg.; KMMA • 1 ♂; Bukama, Haut-Lomami; 28 Mar 1911; J. Bequaert leg.; KMMA • 1 ♀; Bukama, Haut-Lomami; 24 May 1911; J. Bequaert leg.; KMMA • 1 ♀; Bunia, Ituri; 1938; P. Lefevre leg.; KMMA • 3 ♂♂ 3 ♀♀; Faradje, Haut-Uele; Nov 1912; Lang and Chapin leg.; KMMA • 4 ♂♂; Garamba, Haut-Uele; Jun-Jul 1912; Lang and Chapin leg.; KMMA • 2 ♀♀; Kalemie; 1 - 20 Jan 1919; R. Mayne leg.; KMMA • 1 ♂; Kasenyi, Lac Albert, Ituri; 15 May 1935; H. J. Bredo leg.; KMMA • 1 ♀; Kasongo, Maniema; date unknown; Dr. Pons leg.; KMMA • 1 ♂; Kasonsero, Ituri; 17 Jul 1914, J. Bequaert; KMMA • 1 ♂ 1 ♀; Stanleyville [= Kisangani], Tshopo; Mar 1915; Lang and Chapin leg.; KMMA • 1 ♂; Komi, Sankuru; May 1930; J. Ghesquiere leg.; KMMA • 1 ♂; N'Gwese, Lac Kivu; date unknown; Carlier leg.; KMMA • 1 ♂; Elisabethville [= Lubumbashi], Haut-Katanga; 3 Jun 1920; M. Bequaert leg.; KMMA • 1 ♀; Elisabethville [= Lubumbashi], Haut-Katanga; Dec 1925; Van Sackeghem leg.; KMMA; 1 ♂; Elisabethville [= Lubumbashi], Haut-Katanga; Jan 1933; M. Bequaert leg.; RMNH • 1 ♀; Elisabethville [= Lubumbashi], Haut-Katanga; 16 Nov 1921; M. Bequaert leg.; RMNH • 1 ♀; Elisabethville [= Lubumbashi], Haut-Katanga; 9 May 1920; M. Bequaert leg.; RMNH • 1 ♂; Faradje, Ituri; 11 Apr 1930; A. Collart leg.; KMMA • 1 ♀; Malima; 14 Oct 1910; J. Bequaert leg.; KMMA • 1 ♀; Mayumbe, Luki; 1924; L. Pieters leg.; KMMA • 1 ♀; Nyangwe, Maniema; 13 Dec 1910; Dr. Bequaert leg.; KMMA • 1 ♀; Nyangwe, Maniema; 19 Nov 1910; Dr. Bequaert leg.; KMMA • 1 ♀; Nyangwe, Maniema; 12 Nov 1910; J. Bequaert leg.; KMMA • 1 ♂ 1 ♀; Nyangwe, Maniema; Apr-May 1918; R. Mayne leg.; KMMA • 1 ♂ 1 ♀, Apr-May 1918, R. Mayne leg.; RMNH • 1 ♀; Ubundu, Tshopo; 21 Oct 1910; J. Bequaert leg.; KMMA • 1 ♂; Wombali, Mai-Ndombe; Jul 1913, P. Vanderijst leg.; KMMA • 1 ♀; unknown locality; 14 Dec 1951; H. De Saeger leg.; KMMA. • 1 ♂ 1 ♀; 4 Jan 1951; H. De Saeger leg.; KMMA • 1 ♀; 30 Oct 1951; H. De Saeger leg.; KMMA. Ethiopia • 1 ♀; Koka; 14 Jan 1968; J. W. Boyes leg.; CNC • 1 ♂, N. W. shore of lake Zwai; 3 Nov 1926; H. Scott leg.; NHMUK • 1 ♂; unknown locality; Nov 1911; R. J. Stordy leg.; NHMUK. Ghana • 1 ♂; Tamale; Nov 1916; J. J. Simpson leg.; NHMUK. Kenya • 2 ♂♂; Kabete; 24 May 1916; T. J. Anderson leg.; NHMUK • 4 ♂♂; Kanyamkago; 13 Jun 1911; J. Pugh leg.; NHMUK • 1 ♂; Marsabit District, Rendili Njoro; date unknown; C. A. Neave leg.; NHMUK • 1 ♂; Mbuyuni, Serengetti Plains; 25 May 1916; T. J. Anderson leg.; NHMUK • 1 ♂; Voi; 8 - 10 Feb 1912; S. A. Neave leg.; NHMUK • 1 ♀; Fisherman's Camp, Naivasha; 14 Mar 1993; M. De Meyer leg.; KMMA • 1 ♀; Chawia, near Wundanyi; 25 Jan 2017; A. Ssymank leg.; ASPC • 1 ♀; Ologassai; Apr 1986; J. Muhangani leg.; NMK • 1 ♀; Rift Valley; 27 - 29 May 2013; R. Copeland leg.; ICIPE • 1 ♂ 1 ♀; Taita Hills; 2017; A. Ssymank leg.; ASPC • 1 ♂; Turkana, Lothagam; Jul-Aug 1994; A. M. George leg.; NMK. Liberia • Bong 1 ♂; County, Suakoko; 6 Feb 1988; G. G. M. Schulten leg.; RMNH. Madagascar • 1 ♀; Andasibe, Perinet; 29 Oct 2010; A. Ssymank leg.; ASPC • 3 ♂♂; Antananarivo; 28 Feb 2016; G. Goergen leg.; IITA • 2 ♂♂ 1 ♀; Antananarivo; 28 Feb 2016; G. Goergen leg.; KMMA • 1 ♂; Antananarivo; 8 Mar 2016; G. Goergen leg.; KMMA • 1 ♂; Antananarivo, Tsimbazaza; 7 Feb 1968; J. W. Boyes leg.; CNC • 1 ♀; Antananarivo, Antananarivo, Tsimbazaza; 6 Nov 1993; M. Hauser leg.; CAS • 1 ♂; Antananarivo, Antananarivo, Tsimbazaza; 9 Oct 1993; M. Hauser leg.; CAS • 1 ♂ 1 ♀; Antananarivo, Tsimbazaza; 16 - 22 Oct 1993; C. Kassebeer leg.; CNC • 1 ♂ 1 ♀; Antananarivo, Tsimbazaza; 16 - 22 Oct 1993; C. Kassebeer leg.; NMK • 1 ♀; Fianarantsoa, Mahabo Mananivo, Ampitavananima forest; 17 - 24 Mar 2007; M. Irwin, F. Parker and R. Harin'Hala leg.; CAS • 1 ♀; Tananarive; 28 - 30 Apr 1968; K. M. Guichard leg.; NHMUK • 1 ♂; Tananarive; 15 Oct 1957; F. Keiser leg.; NMB • 1 ♀; Tananarive; 20 Oct 1957; F. Keiser leg.; NMB • 1 ♂ 1 ♀; Fianarantsoa; Ambodimanga; 8 Aug 1958; F. Keiser leg.; NMB • 1 ♂; Ambongamaranitra; 20 Jun 1958; F. Keiser leg.; NMB • 2 ♂♂; Tananarive; 28 - 30 Apr 1968; K. M. Guichard leg.; NHMUK • 1 ♀; Tananarive; 9 Sep 1980; J. Stelleman leg.; RMNH. Malawi • 1 ♂; Blantyre; 25 Apr 1910; J. E. S. Old. leg.; NHMUK • 2 ♂♂; Chiromo; J. E. S. Old. leg.; NHMUK • 1 ♂; Cholo; R. C. Wood leg.; NHMUK • 1 ♂ 2 ♀♀; Mulanje Mountain Forest Reserve; 12 - 15 Nov 2016; K. Jordaens leg.; KMMA • 1 ♂ 6 ♀♀; Mulanje Mountain; Likhubula; 12 - 14 Nov 2016; K. Jordaens leg.; KMMA • 3 ♂♂; N. Malawi; 1916; N. M. Leys leg.; NHMUK • 3 ♂♂; Ruo; R. C. Wood leg.; NHMUK • 1 ♂; Chinteche; H. S. Stannus leg.; NHMUK. Mozambique • 3 ♂♂; Sofala, Gorongosa Park; 20 - 30 Apr 2015; M. Hauser and A. Runig leg.; CAS • 1 ♀; Luaba, lower Zambesi; Jun-Jul 1957; P. J. Usher and B. Stuckenberg leg.; CNC • 1 ♂; Luabo, lower Zambesi; Apr 1958; P. J. Usher leg.; CNC • 1 ♂; E of Mount Mulanje; 3 - 7 Oct 1913; S. A. Neave leg.; NHMUK. Nigeria • 1 ♀; Samaru; 15 - 22 Jun 1972; P. H. Ward leg.; NHMUK • 1 ♀; Ibadan; 28 May 1987; G. G. M. Schulten leg.; RMNH. Senegal • 1 ♂; Dakar; 14 Jan 1945; collector unknown; RMNH. South Africa • 1 ♀; Mariepskop, Mpumulanga; 20 - 22 Jan 2017; K. Jordaens leg.; KMMA • 1 ♂; St. Lucia Bay, Natal; 3 Nov 1959; D. J. Greathead leg.; NHMUK • 2 ♂♂; KwaZulu-Natal, Durban; 1 Jul 1903; G. Burn leg.; NMSA • 1 ♂; Rinkarla; date unkown; H. Junod leg.; NMSA. Tanzania • 2 ♂♂; Bondei; Jan 1986; C. W. Schmidt leg.; MNB • 1 ♂; Zanzibar; Jan-Feb 1925; H. J. Snell leg.; NHMUK • 5 ♂♂; Zanzibar, Kizimbani; 15 Jul 1985; G. G. M. Schulten leg.; RMNH • 5 ♂♂ 3 ♀♀; Kilanbeo; 1 May 1971; W. S. Bos leg.; RMNH. Togo • 1 ♂; Kloto Forest; Feb 2017; G. Goergen leg.; KMMA. Uganda • 2 ♂♂; Budongo, forest near Lake Albert; Apr 1972; E. B. Babyetagara leg.; CNC • 1 ♂; Entebbe; 3 - 4 Dec 1912; C. C. Gowdey leg.; NHMUK • 1 ♂; Entebbe; 12 - 20 Jan 1912; S. A. Neave leg.; NHMUK • 2 ♂♂; Ibanda; 23 - 28 Dec 1972; H. Falke leg.; CNC • 1 ♂; Kigezi, Kayonza Forest; Sep 1972; H. Falke leg.; CNC • 1 ♂; Kigezi, Kayonza Forest; Dec 1972; H. Falke leg.; CNC • 1 ♀; Kitende; 11 Sep 1927; J. Bequaert leg.; KMMA • 1 ♂; Masindi; 15 - 19 Dec 1911; S. A. Neave leg.; NHMUK • 1 ♂; locality unknown; Nov 1904; E. D. W. Greig leg.; NHMUK • 2 ♂♂ 4 ♀♀; Western Region, Kamwege District, Kibale Forest; 8 Dec 2018; K. Jordaens; leg. KMMA • 3 ♂♂ 1 ♀; Mabamba swamps, Nkima lodge and surroundings; 16 Dec 2018; K. Jordaens leg.; KMMA. Zambia • 1 ♂ 1 ♀; Lusaka Province 8.5 km NW Katondwe; 20 Apr 2016; M. Hauser leg.; CSCA. Zimbabwe • 1 ♀; Chishi Island, Bangweolo; 26 Jun 1908; S. A. Neave leg.; OXUM • 1 ♂; Chinsali District, mid-Chambezi Valley; 17 Apr 1908; S. A. Neave leg.; OXUM • 1 ♂; Mirongo; 3 Apr 1908; S. A. Neave leg.; OXUM • 1 ♂; upper Luangwa Valley; 29 Feb 1908; S. A. Neave leg.; OXUM • 1 ♂; upper Luangwa Valley; 17 Mar 1908; S. A. Neave leg.; OXUM • 1 ♂; upper Luangwa Valley; 20 Mar 1908; S. A. Neave leg.; OXUM • 1 ♂; upper Luangwa Valley; 5 - 7 Mar 1908; S. A. Neave leg.; OXUM • 1 ♂; upper Luangwa Valley; 23 - 24 Mar 1908; S. A. Neave leg.; OXUM. (Morphotype with conspicuous spine on metatibia; see Variation, comments and discussion). " ARABIA " • 1 ♂; Abu; date unknown; C. G. Nurse leg.; NHMUK. Benin • 3 ♂♂ 3 ♀♀; Calavi; 8 Dec 2013; K. Jordaens and G. Goergen leg.; KMMA • 2 ♂♂ 3 ♀♀; 9 Dec 2013; K. Jordaens and G. Goergen leg; KMMA • 1 ♀; Calavi; Jan 2014; G. Goergen leg.; KMMA • 1 M; Calavi; Mar 2000; G. Goergen leg.; IITA • 1 ♂; Calavi; 17 Jul 2015; G. Goergen leg.; IITA • 1 ♂; Calavi; Oct 2015; G. Goergen leg.; IITA • 1 ♂; Cotonou; Dec 2003; G. Goergen leg.; IITA • 1 ♂ 2 ♀♀; Cotonou; 14 Dec 2013; K. Jordaens and G. Goergen leg.; KMMA • 4 ♀♀; Cotonou; 28 Jan 2016; K. Jordaens and G. Goergen leg.; KMMA • 1 ♂; Cotonou; 1 May 1989; G. G. M. Schulten leg.; RMNH • 1 ♂; Lama Forest; 23 Jan 1995; G. Goergen leg.; IITA • 1 ♂; Lama Forest; 26 Jul 1995; G. Goergen leg.; IITA • 2 ♀♀; Lama Forest; 23 Jun 1995; G. Goergen leg.; IITA • 1 ♀; Niaouli; 2 Feb 2014; G. Goergen leg.; IITA • 1 ♂ 1 ♀; Pobe; 27 Jan 2016; G. Goergen leg.; IITA • 1 ♂ 1 ♀; Porto Novo; 27 Jan 2016; K. Jordaens and G. Goergen leg.; KMMA • 1 ♂; Seme; 7 Feb 2016; G. Goergen leg.; KMMA • 1 ♂; Tanougou Waterfalls; Nov 2016; G. Goergen leg.; KMMA. Cameroon • 20 ♂ 20 ♀; Maroua, Meskina; 23 May 2018; M. Azo'o Ela leg.; MAPC • 1 ♂; Kribi, Lobe Falls; 22 May 2006; A. Ssymank leg.; ASPC • 1 ♂; Victoria; 5 - 18 Nov 1975; W. Schacht leg.; RMNH. Ghana • 1 ♂; Japi; Nov 1915; J. J. Simpson leg.; NHMUK • 1 ♀; Tamale; Nov 1915; J. J. Simpson leg.; NHMUK • 9 ♂♂; Tamale; Nov 1916; J. J. Simpson leg.; NHMUK. Malawi • 1 ♀; Mulanje Mountain; 12 - 14 Dec 2016; K. Jordaens leg.; KMMA. Mali • 1 ♂ 2 ♀♀; Kogoni; Aug 1983; B. Sidibe leg.; NHMUK. Nigeria • 1 ♂; Samaru; 7 - 14 Jul 1970; P. H. Ward leg.; NHMUK. South Africa • 1 ♂; Dukuduku Forest Reserve; 16 - 17 Jul 1981; J. G. H. Londt and K. Craddock leg.; NMSA. Senegal • 3 ♀♀; Dassilame Serere; 30 Nov 2011; S. Cavailles leg.; SCPC • 1 ♀; Dindefelo; 5 Nov 2016; S. Cavailles and R. Bou leg.; SCPC. Togo • 1 ♂; Marais d'Asrama; 8 Apr 2008; A. Ssymank leg.; MZH • 1 ♀; Kloto Forest; Nov 2016; G. Goergen leg.; KMMA. Zambia • 1 ♂; Chilanga; 8 Sep 1913; R. C. Wood leg.; NHMUK • 1 ♂; Chilanga; 2 Jan 1914; R. C. Wood leg.; NHMUK. (Morphotype with low spine on metatibia; see Variation, comments and discussion). Ghana • 9 ♂♂; Tamale; Nov 1916; J. J. Simpson leg.; NHMUK. Nigeria • 1 ♂; Samaru; 7 - 14 Jul 1970; P. H. Ward leg.; NHMUK. Zambia • 1 ♂; Chilanga; 2 Jan 1914; R. C. Wood leg.; NHMUK.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
243A5D3ADB205ECF9714BB61D130610A.taxon	description	Re-description of male (Figs 4 - 6). Body length: 12.1 - 13.8 mm. Wing length: 8.2 - 10.0 mm. Head (Figs 47 - 49). Eyes bare; slightly dichoptic, distance between eyes approx. the width of ocellus. Face white with dark medial vitta; white pollinose; white pilose. Frontal triangle white; white pilose; white pollinose. Vertical triangle black pilose in ventral half and at ocellar triangle, yellow pilose on vertex; yellow pollinose until just before anterior ocellus; distance between lateral ocellus and eye margin less than 1 / 2 width of ocellus. Frontal prominence shiny black. Occiput yellow; yellow and white pollinose; yellow pilose. Antenna black, antennal arista reddish-brown. Thorax. Scutum black with only vague grey pollinose pair of vittae; yellow to rufous pilose. Scutellum uniformly yellow-brown; yellow pilose. Legs. Proleg: Femur without apical pile brush; black; pile long, yellow, ventrally of equal size over entire length; without black pile. Tibia yellow in proximal half, black in distal half; with long, yellow pile. Tarsi dark brown; black pilose dorsally; orange pilose ventrally with some thick black pile, especially posteroventrally. Mesoleg (Figs 178, 182): Femur dark brown to black; long yellow pilose, but short black pilose on dorsal distal 1 / 3. Tibia yellow, distally darkened; long yellow pilose, but with black pile interspersed in distal half. Tarsi dark brown; black pilose, with some yellow pile interspersed on basitarsus. Metaleg (Figs 195, 196): Femur dark brown to black; bare in anteroventral and ventral proximal 1 / 4, posteroventral proximal 1 / 4 with thicker pile varying from entirely yellow (as in Fig. 195: red arrow) to entirely black (as in Fig. 196: red arrow); otherwise with long and thin yellow pile interspersed with shorter black pile ventrally. Tibia black; long yellow and black pilose; distal end variable: either with (as in Fig. 196; referred to as spined morph) or without (as in Fig. 195; referred to as nominal morph) a tooth-like projection (" spine ") at the ventral distal end; unmodified, but with two shallow depressions on posterior side; long yellow pilose on anterior and dorsal side, black pilose on posterior and ventral side. Basitarsus long, as long as tarsomeres 2 + 3; dark brown; black pilose. Other tarsi dark brown; black pilose. Wing (Fig. 128). Entire wing uniformly dense microtrichose. Abdomen (Figs 84 - 86). Tergite II with a pair of very large, yellow rounded maculae; black marking hourglass-shaped; posterior black marking sometimes nearly absent (as in Fig. 85), if present, then narrower than anterior black marking; with short, stiff black setulae which do not extend to the lateral margins. Tergite III and IV with yellow fascia of variable size, often occupying almost the entire tergite, but sometimes strongly reduced; Tergite V strongly white pollinose, except for a black medial zone; with short, black stiff pile posteriorly which does not reach the lateral tergal sides. Genitalia (Figs 206 - 208). Epandrium: Dorsal lobe of surstylus somewhat elongated, broadly rounded; with short, black spines on almost entire surface; dorsally long yellow pilose. Ventral lobe of surstylus straight without conspicuous pilosity.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
243A5D3ADB205ECF9714BB61D130610A.taxon	description	Re-description of female (Figs 27 - 29). Body length: 12.5 - 15.0 mm. Wing length: 10.1 - 10.3 mm. Head. Eyes bare; dichoptic. Face white with dark medial vitta; white pilose, white pollinose. Frons black on dorsal 2 / 5, yellow-white on ventral 3 / 5; black pilose on ocellar triangle and just ventrally of ocellar triangle, otherwise white pilose; pollinosity variable, but mostly strongly white pollinose on ventral 3 / 5. Distance between lateral ocellus and eye margin approx. width of ocellus. Occiput yellow-white; yellow-white pilose; yellow-white pollinose. Frontal prominence shiny black; antenna dark brown to black; antennal arista reddish-brown. Thorax. Scutum dark brown with one pair of dorsolateral yellow pollinose vittae which are connected posteriorly and with lateral, yellow pollinose vitta; yellow pilose. Scutellum yellow-orange; yellow pilose. Legs. Proleg: Femur black, distal end orange-brown; yellow pilose, short, black pilose on dorsal distal end. Tibia orange-brown in proximal 2 / 3, dark brown to black in distal 1 / 3; yellow pilose with thicker black pile interspersed on ventral side. Tarsi orange-brown; black pilose dorsally, yellow pilose ventrally. Mesoleg: Femur black; yellow-white pilose. Tibia orange-brown in proximal half, darkened in distal half; yellow-white pilose, with some shorter and thicker black pile ventrally, especially in distal half. Tarsi black; black pilose ventrally and dorsally, yellow pilose on posterior and anterior side. Metaleg: Femur black; yellow-white pilose with shorter and thicker black pile on ventral distal half. Tibia black; yellow-white pilose with some black pile interspersed ventrally. Tarsi black; short black pilose dorsally; densely yellow-orange pilose ventrally. In some specimens from Benin, the metafemur and metatibia is brown (but not as light as in M. senegalensis) and without the interspersed black pile ventrally. Wing. Entire wing uniformly microtrichose. Abdomen (Figs 107 - 109). Tergite II with a pair of very large, rounded yellow-orange maculae (Figs 107, 109; nominal morph) or with a yellow-orange fascia (Fig. 108; spined morph); yellow pilose on maculae, yellow and black pilose on hourglass-shaped black marking. Tergite III with broad (approx. 3 / 5 of tergal length in medial section), yellow-orange fascia, with posterior black marking; yellow pilose on fascia, predominantly black pilose on black marking and just anterior of black marking. Tergite IV with narrower yellow-orange fascia (approx. 1 / 3 of tergal length in medial section); yellow pilose on fascia, yellow and black pilose on black marking. Tergite V black; yellow pilose.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
243A5D3ADB205ECF9714BB61D130610A.taxon	distribution	Distribution. ' Arabia', Benin, Burundi, Cameroon, Democratic Republic of the Congo, Ethiopia, Ghana, Kenya, Liberia, Madagascar, Malawi, Mali, Mozambique, Nigeria, Senegal, South Africa, Tanzania, Togo, Uganda, Zambia and Zimbabwe.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
0A6DC797C79055499B7ED02757AC502D.taxon	materials_examined	Examined material. Helophilus capensis Macquart: Lectotype (hereby designated), male, " SYNTYPE " " MNHN, Paris // ED 6791 " " 1 ♂ Helophilus // Helophilus capensis Macq // C. F. Kassebeer 1999 " [MNHN]. Helophilus capensis Macquart: Paralectotype, female, " SYNTYPE " " MNHN, Paris // ED 6790 " " SYNTYPES // Vockeroth ' 69 " " afrique // Delalande " [MNHN]. Helophilus lagopus Loew: Holotype, female, " Helophilus // Mesembrius lagopus " " 208 " " Cap. B. // Spei. " " Victo- // rin. " " LECTOTYPUS // Helophilus lagopus // Loew, 1858 " " design. Kassebeer 1993 " " NHRS-BYWS // 000002620 " [NRMS].	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
0A6DC797C79055499B7ED02757AC502D.taxon	materials_examined	Other material. Angola • 1 ♂ 1 ♀; 30 km NE of Duque de Braganza; Nov-Dec. 1957; G. H. Heinrich leg.; NHMUK • 1 ♀; Duque de Braganza; Nov-Dec 1957; G. H. Heinrich leg.; NHMUK. Botswana • 1 ♂; Gabarone; 12 Nov 1988; W. H. O. Ernst leg.; RMNH. Democratic Republic of the Congo • 1 ♀; Sakania; Sep 1931; N. P. Cockerell leg.; NHMUK • 1 ♂ 1 ♀; Lualaba; Bunkeya; Oct 1907; S. A. Neave leg.; NHMUK • 1 ♂; Katanga; Kafubu Mission; Nov 1931; J. Ogilvie leg.; CNC • 1 ♀; South Kivu; Kalambelembe-Baraka; Jul 1918; R. Mayne leg.; KMMA • 1 ♀; Kapanga; Lulua; Nov 1928; Walker leg.; KMMA • 1 ♂ 1 ♀ Leopoldville [= Kinshasa]; 27 May 1915; Lang and Chapin leg.; KMMA • 1 ♂; Elisabethville [= Lubumbashi]; 11 - 17 Sep 1931; A. Mackie leg.; NHMUK • 1 ♀; Elisabethville [= Lubumbashi]; 23 May 1920; M. Bequaert leg.; KMMA • 1 ♂ 1 ♀; Elisabethville [= Lubumbashi]; 1927; M. Bequaert leg.; KMMA • 1 ♀; Elisabethville [= Lubumbashi]; 12 Nov 1928; M. Bequaert leg.; KMMA • 1 ♂ 1 ♀; Elisabethville [= Lubumbashi]; 1 Dec 1929; M. Bequaert leg.; KMMA • 1 ♂; Elisabethville [= Lubumbashi]; 11 - 17 Nov 1931; T. D. A. Cockerell leg.; CNC • 1 ♂ 1 ♀; Elisabethville [= Lubumbashi]; 9 Sep 1932; De Loose leg. • KMMA • 2 ♀♀; Elisabethville [= Lubumbashi]; 1 Dec 1929; M. Bequaert leg.; RMNH • 1 ♂; Elisabethville [= Lubumbashi]; 15 Mar 1928; M. Bequaert leg.; RMNH • 1 ♂; Elisabethville [= Lubumbashi]; 23 May 1920; M. Bequaert leg.; RMNH • 1 ♂; Sakania; Nov 1931; V. P. Cockerell leg.; CNC • 1 ♀; Tumbwe; 16 Nov 1921; M. Bequaert leg.; KMMA • 1 ♀; locality unknown; 8 Nov 1951; H. De Saeger leg.; KMMA. Eswatini; 1 ♀; Manzini; 7 May 1991; J. A. W. Lucas leg.; RMNH. Ethiopia • 1 ♂ 1 ♀; Errer River; date unknown; G. Kristensen leg.; NHMUK. Ghana • 1 ♂; Kumasi; 28 Oct 1946; J. Bowden leg.; NMSA. Kenya • 2 ♂♂ 3 ♀♀; Nairobi Prov., Kasarani; 10 - 14 Dec 2016; K. Jordaens and R. Copeland leg.; NMK • 1 ♀; Nairobi Prov., Kasarani; IV. 2014; R. Copeland leg.; ICIPE • 1 ♀; Central Prov., Kinnyaga, Mucagara Farm; 16 - 18 Dec 2016; K. Jordaens and R. Copeland leg.; NMK • 2 ♂♂; Nyeri; X. 1948; van Someren leg.; NHMUK • 2 ♂♂ 2 ♀♀; Sola District, Lonje Valley, Laikipia escarpment; 12 Sep 1919; T. J. Anderson leg.; NHMUK. Madagascar • 1 ♀; Antananarivo; 28 Feb 2016; G. Goergen leg.; IITA. Malawi • 1 ♀; Ruo; date unknown; R. C. Wood leg.; NHMUK. Mozambique • 1 ♂; Luabo; Lower Zambezi River, Port East Africa; 1 Apr 1958; P. Usher leg.; NMSA • 1 ♂; Rikatla; date unknown; H. Junod leg.; NMSA. Rwanda • 1 ♀; Nduga; Mar 1953; P. Basilewsky leg.; KMMA. South Africa • 1 ♀; Barberton, Aloe Bridge Farm; 22 Oct 2016; A. Vujic et al.; leg.; UNS • 2 ♂♂; KwaZulu-Natal, Bisley Valley Nature Reserve; 22 Dec 1993; J. G. H. Londt and Craddock leg.; NMSA • 1 ♂ 1 ♀; Bontebok National Park; 4 Dec 2016; A. Vujic; S. Radenkovic; N. Velickovic and Z. Petanidou leg.; UNS • 1 ♂ 1 ♀; Capland; date unknown; S. Krebs leg.; MNB; 1 ♂; Western Cape, Cederberg; 17 - 21 Oct 2017; K. Jordaens leg.; KMMA • 1 ♂; KwaZulu-Natal, Doonybrook; 10 Oct 2015; Vujic et al. leg.; UNS • 1 ♀; Eastern Cape, East London; 9 Apr 1922; H. K. Munro leg.; RMNH • 1 ♂; Eastern Cape, East London; 9 Apr 1922; H. K. Munro leg.; NMSA • 1 ♂; Hottentot, Holland; 5 Dec 2016; A. Vujic; S. Radenkovic; N. Velickovic and Z. Petanidou leg.; UNS • 1 ♂; KwaZulu-Natal, Ashburton; 8 Nov 1982; D. A. Barraclough leg.; NMSA • 1 ♂; KwaZulu-Natal, Barlett Estate, Cato Ridge; 9 Oct 2018; G. Theron leg.; NMSA • 1 ♂; KwaZulu-Natal, Bishopstown, near Pietermaritzburg; 11 Dec 1982; A. Seymour leg.; NMSA • 2 ♂♂; KwaZulu-Natal, Bisley Valley Reserve; 22 Dec 1993; J. G. H. Londt leg.; NMSA • 1 ♂; KwaZulu-Natal, Congelia; 26 Oct 1906; G. F. Leigh leg.; NMSA • 2 ♂♂; KwaZulu-Natal; Ferncliff; 16 Nov 2018; K. Jordaens leg.; NMSA • 1 ♂; KwaZulu-Natal, Hudley; date unknown; E. Pinhey leg.; NMSA • 1 ♂; KwaZulu-Natal, Illovo; 14 Jun 1919; collector unknown; NMSA • 1 ♂; KwaZulu-Natal, Ingwavuma; 21 Feb 1979; J. G. H. Londt leg.; NMSA • 1 ♂; KwaZulu-Natal, Kosi Bay Nature Reserve; 30 Nov 1982; B. R. Stuckenberg leg.; NMSA • 1 ♀; KwaZulu-Natal, Kosi Lake; 22 - 27 Jan 1967; D. Gilissen leg.; RMNH • 1 ♂; KwaZulu-Natal, Mkuzi Game Reserve, Nsumu Pan Area; 12 Jan 1994; Natal Museum Staff leg.; NMSA • 1 ♂; KwaZulu-Natal, Mtunzini; 7 Feb 1965; T. Schofield leg.; NMSA • 1 ♂; KwaZulu-Natal, Port Shepstone, Uvongo; 23 Sep 2005; A. Wilson leg.; NMSA • 1 ♂; KwaZulu-Natal, Salt Rock; 5 Oct 1991; J. G. H. Londt leg.; NMSA • 2 ♂♂; KwaZulu-Natal; Zululand, Ndumu Game Reserve; 26 Oct 1972; M. E. Irwin leg.; NMSA • 3 ♂♂ 4 ♀♀; KwaZulu-Natal, Howick; 18 Oct 2015; A. Vujic et al. leg.; UNS • 1 ♀; KwaZulu-Natal, Howick, near Curry's Post; 14 Feb 2016; A. Vujic; S. Radenkovic leg.; UNS • 1 ♂; Western Cape, Keniworth Racecourse Cons. Area; 5 Nov 2014; A. Vujic et al. leg.; UNS • 1 ♀; Mpumulanga, Mariepskop National Park; 23 - 24 Jan 2017; K. Jordaens leg.; KMMA • 1 ♂ 1 ♀; Mpumulanga, Molele Farm; 28 Jan 2017; K. Jordaens leg.; KMMA • 1 ♂; Muizenberg, False Bay; 3 Jan 1972; Southern African Expedition leg.; NHMUK • 1 ♀; KwaZulu-Natal, N. from Pietermaritzburg along Otto's Bluff; 19 Oct 2015; X. Mengual leg.; ZFMK • 4 ♂♂ 8 ♀♀; KwaZulu-Natal, near Howick; 18 Oct 2015; X. Mengual leg.; ZFMK • 1 ♂; Cape Province, Port Elizabeth; 22 - 27 Dec 1985; J. G. H. Londt leg.; NMSA • 1 ♂; Stellenbosch, Delheim winery; 10 Feb 2009; E. M. & L. Laasonen leg.; MZH • 1 ♂; Western Cape, Cederberg NP; 20 Nov 2011; A. Vujic leg.; MZH • 1 ♂ 1 ♀; KwaZulu-Natal, Winterton; 1 Oct 2015; X. Mengual leg.; ZFMK • 1 ♂; KwaZulu-Natal, Winterton; 27 Sep 2015; A. Vujic et al. leg.; UNS • 1 ♂ 2 ♀♀; Zastron; 16 Dec 2016; A. Vujic; S. Radenkovic; N. Velickovic and Z. Petanidou leg.; UNS • 1 ♂ 1 ♀; locality and date unknown; S. Krebs leg.; MNB • 1 ♂; Limpopo Province, Moorddrift; date and collector unknown; NMSA • 1 ♂; Limpopo Province, Plat River; 1 Jan 1903; V. Judzncka leg.; NMSA • 1 ♂; Mongosi; May 1916; W. E. Jones leg.; RMNH • 2 ♂♂; Mpumalanga, Barberton, De Kaap; 18 and 29 Apr 1929; H. K. Munro leg.; NMSA • 2 ♂♂; Mpumalanga, Lomati River; 7 Nov 1970; B. R. Stuckenberg leg.; NMSA • 1 ♀; Piet Retief; 15 Mar 1918; Dr. Brauns leg.; RMNH • 1 ♀; Port Elizabeth; 24 Feb 1922; H. K. Munro leg.; RMNH • 1 ♂; Western Cape, Knysna; 1 Jan 1910; H. Brauns leg.; NMSA. Togo • 1 ♀; Kloto Forest; Feb 2005; G. Goergen leg.; IITA. Uganda • 1 ♂; Entebbe; 23 - 31 Jan 1973; H. Falke leg.; CNC • 1 ♀; Entebbe; 1 - 15 Apr 1983; G. G. M. Schulten leg.; RMNH • 1 ♂; Ibanda; 23 - 28 Dec 1972; H. Falke leg.; CNC. Zambia • 1 ♂; Lake Bangweulu, Mbawala Island; Nov-Dec 1946; collector unknown; NHMUK • 1 ♂; N. of lake Bangweulu, near Milambo; 20 Oct 1946; collector unknown; NHMUK • 1 ♂; N. of Lake Bangweulu; N'Sombo; 11 Dec 1946; collector unknown; NHMUK • 1 ♂; Chilanga; 2 Jan 1914; R. C. Wood leg.; NHMK. Zimbabwe • 1 ♂; upper Kalungwsisi valley; 10 Sep 1908; S. A. Neave leg.; OXUM • 1 ♂; Salisbury [= Harare]; date unknown; G. A. K. Marshall leg.; NHMUK • 1 ♀; Umtali District; 2 Jan 1931; P. A. Sheppard leg.; RMNH • 1 ♂; N. Vumba; 6 Oct 1963; D. Cookson leg.; NMSA.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
0A6DC797C79055499B7ED02757AC502D.taxon	description	Re-description male (Fig. 7). Body length: 10.6 - 14.5 mm. Wing length: 8.1 - 10.4 mm. Head (Fig. 50). Eyes bare; slightly dichoptic, distance between eyes approx. the width of ocellus. Face white with dark medial vitta; white pollinose; white pilose. Vertical triangle with black pile in lower half and at ocellar triangle, yellow pile on vertex; yellow pollinose until just before anterior ocellus; distance between lateral ocellus and eye margin 1 / 2 width of ocellus. Frontal triangle white; white pilose; white pollinose. Frontal prominence shiny black. Occiput yellow; yellow pilose; yellow and white pollinose. Antenna black, antennal arista reddish-brown. Thorax. Scutum black with, dorsally, pair of well-demarcated yellow vittae and a faint yellow medial line, which are both connected at anterior and posterior parts of scutum; with lateral, yellow vitta; pile rufous. Scutellum uniformly yellow-brown; yellow pilose. Legs. Proleg (Fig. 160): femur dark brown to black, without apical pile brush; yellow pilose, pile shorter on dorsal side; with a small patch of black pile at posteroventral proximal end. Tibia yellow to orange-brown, long yellow pilose with a few thick black pile at distal end; tarsi orange-brown, black pilose dorsally, yellow-orange pilose ventrally. Mesoleg (Fig. 179): femur as in proleg, but without the posteroventral black pile at proximal end; tibia yellow-orange; with black pile on anterodorsal side; otherwise long yellow pilose. Basitarsus and second tarsal segment orange-brown; black pilose dorsally, long yellow pilose posteriorly. Other tarsi brown; black pilose, with long dark brown pile posteriorly. Metaleg: femur dark brown to black; yellow pilose, with long black pile interspersed at posteroventral proximal half and with some short and thicker black pile at distal end. Tibia dark brown; unmodified; long yellow pilose with sorter, black pile on ventrally and posteriorly, ventral side also with some long, black pile. Tarsi dark brown to black; black pilose dorsally, dark brown and orange pilose ventrally. Wing (Fig. 129). Entire wing uniformly dense microtrichose. Abdomen (Fig. 87). Tergite II with a pair of large, yellow triangular maculae; black marking hourglass-shaped; posterior black marking, though sometimes very vague (as in Fig. 87), equal in size or somewhat narrower than anterior black marking. Posterior black part with short, stiff black setulae which to not extend to the lateral margins. Tergite III and IV with yellow fascia of variable size, often occupying almost the entire tergite, but always with black triangular marking posteriorly. Tergite V strongly white pollinose, except for a black medial zone; with short, black stiff pile posteriorly which do not reach the lateral tergal sides, this pile being absent in specimens where the posterior black marking is strongly reduced. Genitalia (Fig. 209). Epandrium: Dorsal lobe of surstylus elongated, more or less rectangular with upwardly curved apex; with short, black spines in distal half; dorsolaterally with a few longer, black setulae; dorsally and laterally with long, yellow pile. Ventral lobe of surstylus with large expansion ventrally with, on ventral side of the expansion, very long and dense black setulae.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
0A6DC797C79055499B7ED02757AC502D.taxon	description	Re-description female (Fig. 30). Body length: 9.6 - 14.4 mm. Wing length: 8.1 - 10.2 mm. Head (Fig. 70). Eyes bare; dichoptic. Face yellow-white with dark medial vitta; white pilose, white pollinose. Frons yellow-white; predominantly black pilose on ocellar triangle and just ventrally of ocellar triangle, otherwise white pilose; strongly white pollinose on ventral 3 / 5. Distance between lateral ocellus and eye margin approx. 11 / 2 x width of ocellus. Occiput yellow-white; yellow-white pilose; yellow-white pollinose. Frontal prominence shiny black, orange-brown at distal end; antenna black; antennal arista reddish-brown. Thorax. Scutum dark brown with one pair of dorsolateral lighter, yellow pollinose vittae which are connected posteriorly; with lateral, yellow pollinose vitta; yellow pilose. Scutellum yellow-orange; yellow pilose. Legs. Proleg: Femur black, distal end yellow-orange; yellow pilose; white pollinose. Tibia orange, darkened on dorsal distal 1 / 3; yellow pilose. Tarsi orange-brown to black; black pilose dorsally, yellow pilose ventrally with some thick, black pile ventrally. Mesoleg: Femur black; yellow-white pilose, with some short black spines at distal end ventrally. Tibia orange; yellow pilose, with some long, thick black pile at ventral distal end. Basitarsus orange; yellow pilose, with some long, thick black pile on dorsal distal end and ventrally. Other tarsi black; dorsally short black pilose, ventrally yellow pilose with some large, thick black pile. Metaleg: Femur black; yellow-white pilose with shorter and thicker black pile on ventral distal half; white pollinose. Tibia chocolate-brown; yellow-white pilose with some black pile interspersed ventrally, pile longer on distal ventral end. Tarsi black; short black and yellow pilose dorsally; densely yellow-orange pilose ventrally. Wing. Entire wing uniformly microtrichose. Abdomen (Fig. 110). Tergite II with a broad orange fascia, with a narrow black anterior marking and a larger, posterior black marking which, in the medial part, is approx. 1 / 2 the tergal length; yellow pilose, with some very short and thicker black pile posteriorly; especially the posterior part of the posterior black marking white pollinose. Tergite III with yellow-orange fascia which in the medial part is approx. 2 / 5 of tergal length; yellow pilose, with some very short and thicker black pile posteriorly; especially in the medial part strongly white pollinose. Tergite IV similar, but entire orange fascia strongly white pollinose and black pile in black marking almost absent. Tergite V black; yellow-white pilose; strongly white pollinose on anterior half, especially on the lateral sides.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
0A6DC797C79055499B7ED02757AC502D.taxon	distribution	Distribution. Angola, Botswana, Democratic Republic of the Congo, Eswatini, Ethiopia, Ghana, Kenya, Madagascar, Malawi, Mozambique, Rwanda, South Africa, Togo, Uganda, Zambia and Zimbabwe.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
7220A207FAF45ACBA7C83B7B13D2F214.taxon	materials_examined	Examined material. Mesembrius chapini Curran: Holotype, male, " Mesembrius // Mesembrius chapini // ♂ // Holotype // Curran " " Lukolela // left bank // Congo R. 1 ° 5 ' S / 7. I. 1931 " " J. P. Chapin // Ac. 31300 " [AMNH]. Type studied from picture on the website.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
7220A207FAF45ACBA7C83B7B13D2F214.taxon	materials_examined	Other material Benin • 2 ♂♂; Ahozon; date unknown; G. Goergen leg.; IITA • 3 ♂♂ 2 ♀♀; Calavi; Apr 2014; G. Goergen leg.; IITA • 1 ♀; Cotonou; 16 - 18 Jan 2016; G. Goergen leg.; IITA • 1 ♂ 1 ♀; Cotonou; 1 Nov 2013; G. Goergen and K. Jordaens leg.; KMMA • 1 ♂; Cotonou; 28 Jan 2016; K. Jordaens leg.; KMMA • 1 ♂; Dangbo; 13 Jun 2015; G. Goergen leg.; KMMA • 2 ♂♂; Lokossa; Nov 2005; G. Goergen leg.; IITA • 1 ♂; Niaouli; 15 Jan 1998; G. Goergen leg.; IITA • 2 ♂♂ 1 ♀; Pahou; 11 Jan 2014; G. Goergen leg.; IITA • 2 ♀♀; Porto Novo; 28 Nov 2002; G. Goergen leg.; IITA • 1 ♂; Porto Novo; Oct 2004; G. Goergen leg.; IITA • 1 ♂; Porto Novo; 27 Jan 2016; G. Goergen leg.; IITA • 1 ♂; Porto Novo; Jan 2016; K. Jordaens and G. Goergen leg.; KMMA • 2 ♂♂; 5 ♀♀; Porto Novo; 20 Jan 2018; G. Goergen leg.; IITA • 1 ♂ 12 ♀♀; Porto Novo; 27 Jan 2016; K. Jordaens and G. Goergen leg.; KMMA • 1 ♂ 1 ♀; Porto Novo; 27 Jan 2016; K. Jordaens and G. Goergen leg.; IITA • 5 ♂♂ 8 ♀♀; Porto Novo; 7 Mar 2018; G. Goergen & K. Jordaens leg.; KMMA • 4 ♂♂; Porto Novo; date unknown; K. Jordaens leg.; KMMA. Democratic Republic of the Congo • 1 ♀; Tshopo, Basoko; Oct 1948; P. L. G. Benoit leg.; KMMA • 1 ♂ 1 ♀; Equateur, Eala; Jul 1936; J. Ghesquiere leg.; KMMA; 1 ♂; Equateur, Eala; Mar 1936; J. Ghesquiere leg.; RMNH • 1 ♂; Equateur, Eala; 5 Apr 1935; J. Ghesquiere leg.; RMNH • 1 ♂; Equateur, Eala; Sep 1935; J. Ghesquiere leg.; RMNH • 1 ♀; Equateur, Eala; Jan 1936; J. Ghesquiere leg.; RMNH • 2 ♂♂ 2 ♀♀; Equateur, Eala; Feb 1936; J. Ghesquiere leg.; RMNH • 5 ♂♂ 1 ♀; Equateur, Eala; Mar 1936; J. Ghesquiere leg.; RMNH • 1 ♀; Equateur, Eala; Sep 1936; J. Ghesquiere leg.; RMNH • 1 ♀; Equateur, Eala; 28 Sep 1936; J. Ghesquiere leg.; RMNH • 1 ♂ 2 ♀♀; Equateur, Eala; Dec 1936; J. Ghesquiere leg.; RMNH • 1 ♂; Equateur, Eala; 25 Jul 1935; J. Ghesquiere leg.; KBIN • 2 ♂♂ 1 ♀; Equateur, Eala; Aug 1935; J. Ghesquiere leg.; KBIN • 1 ♂; Equateur, Eala; Sep 1935; J. Ghesquiere leg.; KBIN • 1 ♀; Equateur, Eala; Dec 1935; J. Ghesquiere leg.; KBIN • 1 ♀; Leopoldville [= Kinshasa]; 1 Jun 1915; Lang and Chapin leg.; KMMA. Nigeria • 1 ♂; Ibadan, IITA Station; 18 Nov 2004; G. Goergen leg.; IITA.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
7220A207FAF45ACBA7C83B7B13D2F214.taxon	description	Re-description male (Fig. 8). Body length: 14.0 - 17.2 mm. Wing length: 10.2 - 11.3 mm. Head (Fig. 51). Eyes bare; holoptic, eye contiguity almost as long as length of ocellar triangle. Face white with dark medial vitta; white pilose; white pollinose. Vertical triangle black; black pilose; yellow pollinose on ventral half. Distance between lateral ocellus and eye margin 1 / 2 width of ocellus. Frontal triangle short; yellow-white; with some long, black pile; yellow pollinose. Frontal prominence shiny black with orange-brown apex. Occiput yellow; yellow pilose; with some shorter and thicker black pile near eye margin; yellow and white pollinose. Antenna, scape and pedicel reddish-brown; postpedicel black; antennal arista reddish-brown. Thorax. Scutum black with dorsally a pair of well-demarcated yellow vittae which are largely connected posteriorly. Scutum with faint lateral vitta; yellow and black pilose. Scutellum uniformly yellow-brown; with long yellow and shorter black pile. Legs. All legs chocolate-brown; tarsi chocolate-brown. Proleg (Fig. 151): Femur dorsoventrally flattened; posterodorsal side with sparse, long black and yellow pile in proximal half; with apical pile brush of long, dense and curved yellow thick pile, this yellow pile is longer in the distal 1 / 3 and is interspersed with equally long black pile; ventrally with long, black pile. Basitarsus without tuft of orange or black pile posteriorly. Mesoleg: Femur with long, yellow pile on proximal 2 / 3 and long black pile on distal 1 / 3. Tibia and tarsi black pilose. Metaleg (Fig. 186): Femur long and thin, slightly curved pile; pile on dorsal and anterior side inconspicuous, except for some long, black pile at anterior distal end; patch of long yellow pile on proximal part; with row of very short and thick black spines on ventral distal half; with 2 - 5 very long, thick black pile in ventral middle; posteriorly with loose, long yellow and black pile which becomes denser at distal end. Tibia with long, black pile, especially in distal 2 / 3; anteriorly with shallow excavation in the middle; posteriorly with deep and broad bare excavation in proximal 1 / 3. Tarsi black pilose dorsally, orange pilose ventrally. Wing (Fig. 130). Entire wing dark, uniformly dense microtrichose. Abdomen (Fig. 88). Tergite II with pair of very large, yellow rounded maculae; black markings hourglass-shaped; anterior and posterior black markings equal in size marking, but posterior black marking with stronger white pollinosity; yellow and black pilose, but black pile more conspicuous in posterior half of tergite and somewhat denser in posterolateral corners. Tergite III and IV with broad yellow fascia; black markings on posterior half vague because of white pollinosity, but more pronounced in medial part; black and yellow pilose, but black pile rare in anterior 1 / 4. Genitalia (Fig. 210). Epandrium: Dorsal lobe of surstylus short, broadly rounded; with short black spines on almost entire surface. Ventral lobe of surstylus straight; bare.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
7220A207FAF45ACBA7C83B7B13D2F214.taxon	description	Description female (Fig. 31). Body length: 13.3 - 16.3 mm. Wing length: 10.2 - 11.0 mm. Head (Fig. 72). Eye bare; dichoptic. Face yellow-orange with dark medial vitta; white pilose; white pollinose. Frons black; black pilose in dorsal half, black and yellow pilose on ventral half; weakly white pollinose. Distance between lateral ocellus and eye margin approx. width of ocellus. Occiput black; yellow pilose, with some black pile near eye margin; yellow-white pollinose. Frontal prominence shiny black, distal end orange-brown; scape and pedicel orange-brown; postpedicel black; antennal arista reddish-brown. Thorax. Scutum dark brown to black with dorsolateral a pair of vague, grey pollinose vittae which are connected posteriorly; grey pollinose lateral vitta very vague; yellow pilose with some black pile interspersed. Scutellum yellow-orange; yellow pilose with very sparse, shorter black pile interspersed. Legs (Figs 169, 201). All femora black, except for extreme distal ends which are orange-brown; yellow pilose. All tibia orange-brown, most distal tarsomere darkened; yellow and black pilose dorsally, yellow pilose ventrally. All tarsi orange-brown; black pilose, except for protarsus which is yellow-orange pilose ventrally. Wing. Entire wing uniformly dense microtrichose. Abdomen (Fig. 111). Tergite II with a pair of large, orange, rounded maculae; yellow pilose, with dense short thick black pile on posterior black marking. Tergite III with orange fascia in anterior half and black marking in posterior half; yellow pilose with dense short thick black pile on posterior black marking. Tergite IV with narrow orange fascia; yellow-orange pillose. Tergite V black; yellow pilose.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
7220A207FAF45ACBA7C83B7B13D2F214.taxon	distribution	Distribution. Benin, Democratic Republic of the Congo and Nigeria.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
7445843A2C3757C4903ACFFDF002A473.taxon	materials_examined	Examined material. Mesembrius copelandi Jordaens, Goergen & De Meyer: Holotype, male, " HOLOTYPUS " " Kenya // Nairobi Prov. // ICIPE campus // Kasarani, 1.22296 ° S, // 36.89704 ° E, 1600 m " " 6 m Malaise trap, near // stream, woodland // remnant, 17 - 25 JAN // 2017, R. Copeland " " Mesembrius copelandi // Det. K. Jordaens, 2019 " " DNA 1301 F 04 // K. Jordaens // RMCA 2020 " " ICIPE 1180 " [KMMA] Paratype: Kenya • 1 ♂; Sosoma area; 30 Jun- 6 Jul 2018; R. Copeland leg.; ICIPE 9544; ICIPE.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
7445843A2C3757C4903ACFFDF002A473.taxon	description	Description male (Fig. 9). Body length: 13.7 - 14.7 mm. Wing length: 9 - 10.5 mm. Head (Fig. 52). Eyes bare; slightly dichoptic, distance between eyes approx. 1 / 2 width of ocellus. Face white-yellow with dark medial vitta; white pollinose; white pilose. Vertical triangle black pilose, with some yellow pile near vertex; yellow pollinose until just before anterior ocellus. Distance between lateral ocellus and eye margin 1 / 2 width of ocellus. Occiput yellow; yellow pilose; yellow and white pollinose. Frontal triangle brownish; yellow pilose with a few black pile near antenna; yellow pollinose. Frontal prominence shiny black, dark brown at apex. Antenna, scape and pedicel reddish-brown; postpedicel black; antennal arista reddish-brown. Thorax. Scutum black; with a pair of well-demarcated yellow vittae and a yellow medial line dorsally, vitta and medial line are broadly connected posteriorly; with lateral, yellow vitta; yellow pilose with some black pile in the posterolateral corners. Scutellum uniformly yellow; yellow pilose. Legs. Tibia and tarsi entirely rufous, but metatarsi black dorsally. Proleg: Femur black, distal end rufous; without apical pile brush; pile ventrally black on distal 1 / 4, otherwise rufous. Tibia rufous; black and yellow pilose which is longer on posterior side. Basitarsus black and rufous pilose. Other tarsi black pilose. Mesoleg: Femur black, distal end rufous; yellow and black pilose. Tibia similar as in proleg, but black and yellow pile in proximal 1 / 4 of markedly longer than on remainder of tibia. Tarsi similar as in proleg. Metaleg (Fig. 199): Femur black, distal end rufous; with long, thin yellow pile, but shorter and less dense ventrally and posteroventrally; with a band of black pile on extreme proximal end ventrally. Tibia rufous; unmodified; long yellowish pilose, except for the posterior side where pile shorter and black; with a patch of posteroventral longer black pile. Basitarsus black dorsally, rufous ventrally; very long, almost as long as tibia; very long yellow pilose with some long black pile at distal end. Other tarsi black dorsally, rufous ventrally; very long yellow pilose with some long black pile at distal end. Wing (Fig. 131). Entire wing uniformly dense microtrichose. Abdomen (Fig. 89). Tergite II with a pair of large, yellow rectangular maculae; black marking hourglass-shaped; posterior black marking equal in size or somewhat narrower than anterior black marking. Posterior black part with short, stiff black setulae which to not extend to the lateral tergal sides. Tergite III and IV with yellow fascia of variable size, often occupying almost the entire tergite with mostly posterior, triangular black marking. Tergite V strongly white pollinose, except for a black medial zone; with short, black stiff setulae at posteriorly which do not reach the lateral tergal sides, these setulae are absent in specimens where the posterior black marking is strongly reduced. Genitalia (Fig. 211). Epandrium: Dorsal lobe of surstylus elongated, distally rounded, with characteristic small tooth-like projection; dorsally long yellow pilose, short black pilose at distal end. Ventral lobe of surstylus with one large black setula in middle section and a row of approximately ten long black setulae.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
7445843A2C3757C4903ACFFDF002A473.taxon	distribution	Distribution. Kenya.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
7445843A2C3757C4903ACFFDF002A473.taxon	etymology	Etymology. Named in honour of Robert Copeland (ICIPE) who collected both males. The specific epithet should be treated as a noun in the genitive case.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
B32BAEF0179C51C69292EAA6D1C2F139.taxon	materials_examined	Examined material. Helophilus cyanipennis Bezzi: Lectotype (hereby designated), male, " SYNTYPUS " " Syn- // type " " Hel. (Mes.) // ♂ Type // Helophilus cyanipennis // Bezzi " " Pres. By // Impl. Bureau Ent. // 1915 - 165. " " Uganda Prot., // Entebbe. // 1 - 11 Sep 1911. // S. A. Neave. " " Mesembrius // Mesembrius cyanipennis // n. sp // Type ♂ " " NHMUK 013428968 " [NHMUK]. Paralectotype, female, " Syn- // type " " Hel. (Mes.) // ♀ Type // Mesembrius cyanipennis // Bezzi " " Ashanti. // Obuasi. // 7. xiii. 06 // W. M. Graham. // 1907 - 74. " " Mesembrius // Mesembrius cyanipennis // Type ♀ n. sp. " " NHMUK 013428969 " [NHMUK]. Paralectotype, female, " Syn- // type ", " H. (M.) Mesembrius cyanipennis, Bezzi // Bezzi det. // 1915.) " " caught on flower " " Obuasi, // Ashanti, // 20. vii. 1907, // Dr. W. M. Graham. // 1908 - 245. " " NHMUK 013428970 " [NHMUK]. Paralectoype, female, " Syn- // type ", " H. (M.) Mesembrius cyanipennis, Bezzi // Bezzi det. // 1915.) " " caught on flower " " Obuasi, // Ashanti, // 21. vi. 1907, // Dr. W. M. Graham // 1908 - 245. " " NHMUK 013428971 " [NHMUK]. Paralectotype, female, " Syn- // type ", " H. (M.) Mesembrius cyanipennis, Bezzi // Bezzi det. // 1915.) " " Caught on umbelli- // ferus flowers in // swamp. " " Obuasi, // Ashanti, // 28. vi. 1907, // Dr. W. M. Graham // 1908 - 245. " " NHMUK 013428972 " [NHMUK].	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
B32BAEF0179C51C69292EAA6D1C2F139.taxon	materials_examined	Other material Benin • 1 ♂ 1 ♀; Ifangni-range; 6 Jun 2015; G. Goergen leg.; IITA • 1 ♀; Ifangni-range; 6 May 2016; G. Goergen leg.; IITA • 1 ♂ 1 ♀; Ifangni-range; 19 Mar 2017; G. Goergen leg.; IITA • 1 ♂; Lokossa; Oct 2005; G. Goergen leg.; IITA • 1 ♀; Niaouli; 10 Dec 2013; G. Goergen and K. Jordaens leg.; KMMA • 1 ♂ 3 ♀♀; Niaouli; 2 Feb 2014; G. Goergen leg.; IITA • 1 ♂; Pobe; 13 Jun 2015; G. Goergen leg.; IITA • 1 ♀; Pobe; 28 Jan 2016; G. Goergen leg.; IITA • 1 ♀; Pobe; 27 Jan 2016; G. Goergen leg.; KMMA. Cameroon • 1 ♀; Abong M'Bang District; 1 - 30 Apr 1936; F. G. Merfield leg.; NHMUK. Democratic Republic of the Congo • 1 ♀; Haut-Uele, Arebi; date unknown; J. Bequaert leg.; KMMA • 1 ♀; Bas-Uele, Bambesi; 15 Sep 1933; H. J. Bredo leg.; KMMA • 1 ♀; Bas-Uele, Bambili; date unknown; J. Rodhain leg.; KMMA • 1 ♀; Lulua, Kapanga; Nov 1932; G. F. Overlaet leg.; KMMA • 1 ♀; Katanga; Mar 1933; G. F. Overlaet leg.; KMMA • 1 ♀; Ituri, Kilo, Kere-Kere; Mar 1948; Turco leg.; KMMA • 1 ♀; Tshopo, Stanleyville [= Kisangani]; 1914; J. Bequaert leg.; KMMA • 1 ♀; Tshopo, Stanleyville [= Kisangani]; 4 Apr 1915; Lang and Chapin leg.; KMMA • 1 ♀; Tshopo, Stanleyville [= Kisangani]; 7 Apr 1915; Lang and Chapin leg.; KMMA • 1 ♀; Tshopo, Stanleyville [= Kisangani]; 8 Apr 1915; Lang and Chapin leg.; KMMA • 1 ♀; Tshopo, Stanleyville [= Kisangani]; 9 Apr 1915; Lang and Chapin leg.; KMMA • 1 ♂ 1 ♀; Tshopo, Stanleyville [= Kisangani]; Mar 1915; Lang and Chapin leg.; KMMA • 1 ♂ 1 ♀; Elisabethville [= Lubumbashi]; 11 Apr 1921; M. Bequaert leg.; KMMA • 1 ♀; Lomami, Mutombo; Mar 1931; P. Quarre leg.; KMMA • 1 ♀; Haut-Lomami, Sankisia; 4 Apr 1911; Dr. Bequaert leg.; KMMA • 1 ♂; Uele; date unknown; J. Rodhain leg.; KMMA • 1 ♂; Eala; Apr 1935; J. Ghesquiere leg.; RMNH • 1 ♀; Eala; Feb 1936; J. Ghesquiere leg.; RMNH • 1 ♀; Eala; Nov 1935; G. F. Overlaet leg.; RMNH • 1 ♀; Eala, Bambesa; 30 Oct 1933; J. Leroy leg.; RMNH • 1 ♀; Lulua, Kapanga; Nov 1932; G. F. Overlaet leg.; RMNH. Ghana • 1 ♂; Wati Waterfalls; Feb 2003; G. Goergen leg.; IITA. Nigeria • 1 ♀; Ibadan, IITA station; 3 Feb 2000; G. Goergen leg.; IITA • 1 ♂; Ikotobo; 10 Nov 1913; J. W. S. ScottMacie leg.; NHMUK. Sierra Leone • 1 ♂ 1 ♀; Kamakoni; 22 Apr 1912; J. J. Simpson leg.; NHMUK. South Africa • 1 ♂; KwaZulu-Natal, Manguzi Forest Reserve; 13 - 16 Dec 2010; J. G. H. Londt leg.; NMSA. Togo • 1 ♀; Kloto Forest; Dec 2007; G. Goergen leg.; IITA • 1 ♀; Kloto Forest; Nov 2007; G. Goergen leg.; IITA • 1 ♂; Kloto Forest; Feb 2016; G. Goergen leg.; IITA • 2 ♀♀; Kloto Forest; 21 - 24 Jun 2015; G. Goergen leg.; KMMA • 1 ♀; Kloto Forest; Feb 2016; G. Goergen leg.; KMMA • 1 ♀; Kloto Forest; Nov 2016; G. Goergen leg.; KMMA • 2 ♂♂ 2 ♀♀; Kloto Forest; Feb 2017; G. Goergen leg.; KMMA • 1 ♂; Kloto Forest; Mar 2017; G. Goergen leg.; KMMA • 1 ♀; Kuma Adame; 22 - 24 Jan 2016; G. Goergen leg.; IITA. Uganda • 2 ♂♂ 1 ♀; Entebbe; 9 Aug 1911; C. C. Gowdey leg.; NHMUK • 1 ♂; Entebbe; 11 Aug 1911; C. C. Gowdey leg.; NHMUK • 1 ♀; Entebbe; 14 Aug 1911; C. C. Gowdey leg.; NHMUK • 1 ♂ 1 ♀; Entebbe; 16 Aug 1911; C. C. Gowdey leg.; NHMUK • 2 ♂♂; Entebbe; 17 Aug 1911; C. C. Gowdey leg.; NHMUK • 4 ♂♂ 2 ♀♀; Entebbe; 21 Aug 1911; C. C. Gowdey leg.; NHMUK • 1 ♀; Entebbe; 31 Aug 1911; C. C. Gowdey leg.; NHMUK • 1 ♂; Entebbe; 17 Jul 1911; C. C. Gowdey leg.; NHMUK • 1 ♀ • Entebbe; 28 Jul 1911; C. C. Gowdey leg.; NHMUK • 1 ♂ 3 ♀♀; Entebbe; 12 - 13 Dec 1912; C. C. Gowdey leg.; NHMUK • 3 ♀♀; Entebbe; 3 - 4 Dec 1912; C. C. Gowdey leg.; NHMUK • 1 ♂; Entebbe; 27 May 1912; C. C. Gowdey leg.; NHMUK • 1 ♀; Entebbe; 3 Nov 1912; C. C. Gowdey leg.; NHMUK • 3 ♀♀; Entebbe; 14 Nov 1912; C. C. Gowdey leg.; NHMUK • 9 ♀♀; Entebbe; 18 - 20 Nov 1912; C. C. Gowdey leg.; NHMUK • 1 ♀; Entebbe; 16 Oct 1912; C. C. Gowdey leg.; NHMUK • 1 ♀; Entebbe; 3 Sep 1912; C. C. Gowdey leg.; NHMUK • 6 ♂♂ 9 ♀♀; Entebbe; 1 - 11 Sep 1911; S. A. Neave leg.; NHMUK • 1 ♂; Entebbe; 7 - 9 May 1912; S. A. Neave leg.; NHMUK • 1 ♀; Entebbe; 18 May 1912; collector unknown; NHMUK • 1 ♂ 1 ♀; Entebbe; 7 Mar 1973; H. Falke leg.; RMNH • 1 ♀; Entebbe; 7 Mar 1973; H. Falke leg.; RMNH • 1 ♀; Entebbe; 7 Oct 1971; H. Falke leg.; RMNH • 1 ♀; Entebbe, Kisubi Forest; 8 - 9 Jun 1976; M. K. Paulus leg.; CNC • 1 ♀; Kenya Coast, Gedi for Malindi; May 1973; H. Falke leg.; CNC • 1 ♀; W. shores of Vic. Nyanza; 19 - 25 Sep 1911; S. A. Neave leg.; NHMUK • 2 ♀♀; locality and date unknown; C. C. Gowdey leg.; NHMUK. Country unknown • 1 ♂; Ruwengo; 14 May 1911; collector unknown; NHMUK.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
B32BAEF0179C51C69292EAA6D1C2F139.taxon	description	Re-description male (Fig. 10). Body length: 11.0 - 13.2 mm. Wing length: 9.6 - 10.5 mm. Head (Fig. 53). Eyes bare; dichoptic, distance between eyes approx. 11 / 2 x width of ocellus. Face dark brown to black; white pilose; white pollinose. Vertical triangle black; black pilose, yellow pilose on vertex; yellow pollinose until just before anterior ocellus. Distance between lateral ocellus and eye margin 1 / 2 width of ocellus. Frontal triangle brown, area near eye margin yellow; white pilose; white pollinose. Frontal prominence shiny black, reddish-brown at apex; black pilose. Occiput black; white pilose with some shorter, thicker, black pile near dorsal eye margin. Antenna, scape and pedicel reddish-brown; postpedicel black; antennal arista reddish-brown. Thorax. Scutum black; without vitta; pile short, yellow and black. Scutellum, anterior half dark brown, posterior half lighter; yellow pilose with some shorter black pile interspersed. Legs. Legs predominantly dark brown to black. Pile on posterior side of pro- and mesofemur and on proximal distal 1 / 4 yellow, pile otherwise black; black pile on ventral side of femora gradually becoming longer towards distal end; ventral side of metatibia with carina. Wing (Fig. 132). Entire wing uniformly, very densely microtrichose; anterior medial part and posterior half of cell bm brownish. Abdomen (Fig. 90). Tergite II with large, yellow fascia, interrupted in anterior 2 / 3; with T-shaped black marking; yellow pilose, with short thick black pile interspersed, especially in posteromedial section. Tergite III and IV yellow-orange; yellow pilose. Genitalia (Fig. 212). Epandrium: Dorsal lobe of surstylus very long and thin, with pointed apex and with distal end bent upwards; with short, black spines at apex and long yellow pilose on dorsal side. Ventral lobe of surstylus convex.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
B32BAEF0179C51C69292EAA6D1C2F139.taxon	description	Re-description female (Fig. 32). Body length: 12.2 - 13.1 mm. Wing length: 8.6 - 11.0 mm. Head (Fig. 71). Eyes bare; dichoptic. Face black; black and white pilose; white pollinose. Frons black; black pilose; lower half white pollinose. Vertex black; black pilose; grey pollinose. Distance between lateral ocellus and eye margin approx. the width of ocellus. Occiput black; yellow and black pilose dorsally, yellow pilose ventrally; grey pollinose. Frontal prominence shiny brown-black; black pilose. Antenna black; arista reddish-brown. Thorax. Scutum and scutellum black; without vitta; short white and black pilose. Legs. Dark reddish-brown to black; short black and white pilose. Wing. Entire wing uniformly, very densely microtrichose; anterior medial part and posterior half of cell bm brownish. Abdomen (Figs 112, 113). Second tergite with a pair of large orange maculae and without (Fig. 112) or with (Fig. 113) distinct posterior black marking. Tergite III from largely orange with a vague medial black marking (Fig. 112) to black with a pair of large, orange maculae (Fig. 113). Tergite IV from entirely orange (Fig. 112) to orange with a posterior black fascia (Fig. 113). Tergite V orange. All tergites short yellow-white and black pilose.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
B32BAEF0179C51C69292EAA6D1C2F139.taxon	distribution	Distribution. Benin, Cameroon, Democratic Republic of the Congo, Ghana, Nigeria, Sierra Leone, South Africa, Togo and Uganda.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
579D924128DE5AFF974E995F3D727CA1.taxon	materials_examined	Examined material. Helophilus ingratus Loew: Holotype, male, " Helophilus // Helophilus ingratus " " 209 " " 209 " " HOLOTYPUS // Helophilus ingratus // Loew, 1858 " " design. Kassebeer 1993 " " NHRS-BYWS // 000002619 " [NRMS].	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
579D924128DE5AFF974E995F3D727CA1.taxon	materials_examined	Other material Malawi • 1 ♂; Mount Mulanje; 20 Oct 1912; S. A. Neave leg.; NHMUK. Senegal • 2 ♂♂; Dakar; 14 Jan 1945; collector unknown; RMNH. South Africa • 1 ♂; KwaZulu-Natal, Manguzi Forest Reserve; 13 - 16 Dec 2010; J. G. H. Londt leg.; NMSA • 1 ♂; KwaZulu-Natal, St. Lucia Park Reserve; 2 Feb 1988; J. G. H. Londt leg.; NMSA • 1 ♂; KwaZulu-Natal, Ngoya Forest Reserve; 26 Apr 1988; J. G. H. Londt leg.; NMSA.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
579D924128DE5AFF974E995F3D727CA1.taxon	description	Re-description male (Fig. 11). Body length: 10.9 - 11.1 mm. Wing length: 8.0 - 8.4 mm. Head (Fig. 54). Eyes bare; holoptic, length of eye contiguity approx. 1 / 3 the length of ocellar triangle. Face white with dark medial vitta; white pilose; white pollinose. Vertical triangle with black pile in ventral half and at ocellar triangle; yellow pilose in dorsal half; yellow pollinose. Distance between lateral ocellus and eye margin 1 / 2 width of ocellus. Frontal triangle and gena white; white pilose; white pollinose. Frontal prominence shiny black; black pilose. Occiput black, but strongly white-grey pollinose; yellow-brown pilose, with a row of almost equally long black pile at dorsal eye margin. Antenna reddish-brown. Thorax. Scutum dark brown with dorsally a pair of vague yellow vittae; yellow-brown pilose. Scutellum uniformly light yellow-brown; yellow pilose. Legs. All legs dark brown, except for pro- and mesotarsi which are yellow-brown. All femora and tibiae with long, loose yellow pile. Proleg (Fig. 158): Femur with a loose, yellow apical pile brush interspersed with some long black pile; very short thick pile and longer thin black pile at proximal 1 / 4 ventrally. Tibia with long, black pile, except dorsally. Basitarsus black pilose dorsally, with tuft of black pile on posterior side, short orange pilose ventrally. Other tarsi black pilose dorsally, short orange pilose ventrally. Mesoleg: Femur long yellow pilose posteriorly and posterodorsally, except at distal end where pile is black; short black pilose anteriorly and anterodorsally; ventrally with longer black pile. Tibia long black and short yellow pilose. Tarsi short black pilose dorsally; short yellow pilose ventrally with a few thick black spines. Metaleg (Fig. 190): Femur very slender; covered with long, thin yellow pile; shorter black pile on ventral 1 / 4 and at distal end. Tibia with a deep posterior depression in the proximal half which is extended as a groove on the ventral side, demarcated with short, dense black pile; proximal half dorsoventrally flattened; predominantly black pilose. Tarsi black pilose dorsally; orange pilose ventrally. Wing (Fig. 133). Entire wing uniformly dense microtrichose. Abdomen (Fig. 91). Tergite II with pair of very large, yellow triangular maculae; yellow pilose; black marking hourglass-shaped; posterior black marking equal in size or somewhat narrower than anterior black marking, but more vague because of medial white pollinosity; posterior black marking with black pile that posterolaterally extends into the yellow maculae. Tergite III with yellow fascia of variable size, occupying approx. the entire tergite; yellow pilose; with posterior, triangular black marking; black pilose; white pollinose. Tergite IV black; white pilose and white pollinose on anterior fascia, black and yellow pilose on remainder of tergite. Genitalia (Fig. 213). Epandrium: Dorsal lobe of surstylus somewhat elongated, broadly rounded, with short, black spines on almost entire surface; dorsally long yellow pilose. Ventral lobe of surstylus straight; bare.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
579D924128DE5AFF974E995F3D727CA1.taxon	distribution	Distribution. Malawi, Senegal and South Africa.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
EB0E1ADA963B5287991F97C8F5258A02.taxon	materials_examined	Examined material. Mesembrius longipilosus Jordaens, Goergen & De Meyer: Holotype, male " HOLOTYPUS " " Entebbe, UGANDA // 2. III. 1972 // H. Falke // In forest " " Mesembrius // sp. 7 // Det J. R. Vockeroth " " Mesembrius longipilosus // Det. K. Jordaens, 2019 " " Barcode of Life // DNA voucher specimen // Smple | CNC DIPTERA 102305 // BOLD Proc. ID: CNCDB 1109 - 11 " " CNC DIPTERA // # 102305 " [CNC]. Paratype: Uganda • 1 ♂; near Entebbe; 23 - 31 Jan 1972; 1160 m; H. Falke leg.; CNC Diptera 102306 (head and abdomen lost) [CNC].	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
EB0E1ADA963B5287991F97C8F5258A02.taxon	description	Description male (Fig. 12). Body length: 8.6 mm. Wing length: 7.2 - 7.7 mm. Head (Fig. 55). Eyes bare; dichoptic, distance between eyes approx. the width of ocellus. Face yellow with dark medial vitta; white pilose; white pollinose. Vertical triangle black with yellow and black pile; yellow pollinose on lower half. Distance between lateral ocellus and eye margin approx. 1 / 2 width of ocellus. Frontal triangle black; white pilose; white pollinose. Frontal prominence shiny black. Occiput yellow; yellow pilose with interspersed short, black setulae; yellow and white pollinose. Antenna black; postpedicel white pollinose, antennal arista reddish-brown. Thorax. Scutum black; white pilose, with three dorsal, well-demarcated yellow-white pollinose vitta which are connected at posterior end; with lateral, yellow-white pollinose vitta. Scutellum yellow-brown; yellow pilose. Legs. Femora dark chocolate-brown, tibia and tarsi orange-brown. Proleg (Fig. 162): Femur without apical pile brush; yellow pile ventrally long in proximal half, shorter in distal half; ventrally 3 - 4 black pile at basal 1 / 3. Tibia yellow and black pilose. Tarsi black pilose dorsally, yellow-orange pilose ventrally. Mesoleg: Similar to proleg; with black and yellow pile on basitarsus. Metaleg: Femur with long yellow pile anteriorly and shorter pile, posteriorly; ventral pile scarce, yellow and black, the black pile is longer at proximal half than at distal half. Tibia unmodified; with long, yellowish pile and scarce black pile on distal half. Tarsi black pilose dorsally, orange pilose ventrally. Wing (Fig. 134). Entire wing uniformly dense microtrichose. Abdomen (Fig. 92). Tergite II with pair of very large, yellow to orange rounded maculae; black marking hourglass-shaped, but the posterior part is smaller compared to the anterior part; yellow pilose. Tergite III almost entirely orange with small medial black maculae; yellow pilose. Tergite IV with anterior white pollinose band; black on posterior 1 / 3; yellow pilose. Genitalia (Fig. 214). Epandrium: Dorsal lobe of surstylus elongated, more or less rectangular with upwardly curved apex, with short, black spines in distal half; dorsolaterally with a few longer, black setulae; dorsally and laterally with long, yellow pile. Ventral lobe of surstylus with large expansion ventrally, with, on ventral side of the expansion, short black setulae.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
EB0E1ADA963B5287991F97C8F5258A02.taxon	distribution	Distribution. Uganda.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
EB0E1ADA963B5287991F97C8F5258A02.taxon	etymology	Etymology. The specific epithet Mesembrius longipilosus (Latin for hairy, covered with long pili) refers to the long, thin yellow pile on the metalegs. It is to be treated as an adjective (nominative singular masculine).	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
685FFD53922E5405B76892715A036447.taxon	materials_examined	Examined material. Mesembrius maculifera Hull: Holotype, female, " Oriental forest // Fanovana, Dist. // Fianarantsoa // Madagascar " " I-V, 1937 // C Lamberton " " TYPE 6595 // Mesembrius // Mesembrius maculifera // F. M. Hull " " Mesembrius // Mesembrius maculifera // Hull n. sp. " [ANSP] Paratype: Madagascar • 1 ♀; Oriental Forest, Fanovana; date and collector unknown; CNC.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
685FFD53922E5405B76892715A036447.taxon	description	Re-description female (Fig. 33). Body length: 11.2 mm. Wing length: 8.7 mm. (Only female paratype measured). Head (Fig. 73). Eyes bare, dichoptic. Face chocolate-brown to mahogany-red. Frons with ventral 2 / 3 light yellow-brown pilose. Face very short reddish pubescent; sparsely long pale pilose. Ocellar triangle bare. Frontal triangle with a transverse band of dark pile. Occiput chocolate-brown to mahogany-red; thick dark brown pilose. Antenna dark reddish-brown; arista paler. Thorax. Dull black; lateral margins and scutellum very dark red; pleurites black; reddish-orange pilose. Scutellum dull black; white pilose. Legs. Predominantly black, the distal part of the pro- and mesolegs dark red; all tibiae dark red; all tarsi black dorsally, the hind pair rather flattened. Pile pale whitish on tarsi and tibiae. Wing (Fig. 135). Proximal half of the wing deep brown, especially at the r-m crossvein. Abdomen (Fig. 114). Dull black. Tergite II with a pair of prominent light-yellow maculae; white pilose anteriorly, light reddish pilose posteriorly. Tergite III and IV dark brown to black; light reddish pilose, but blackish on the middle of the posterior part.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
685FFD53922E5405B76892715A036447.taxon	distribution	Distribution. Madagascar.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
D68C114EABBE50678F1529F49480749B.taxon	materials_examined	Examined material. Mesembrius madagascariensis Keiser: Holotype, male, Madagascar, Antananarivo, M, 13 Dec 1957, F. Keiser (MNHN: type not studied; see comments). Mesembrius madagascariensis Keiser: Allotype, female, " ALLO // TYPUS " " Madagascar. TAN. // Tananarive // 13. XII. 1957 // F. KEISER " " Mesembrius // madagascar- // iensis " [NMB]. Paratypes: Madagascar • 7 ♂♂ 21 ♀; Tananarive; 13 Dec 1975; F. Keiser leg.; NMB • 1 ♀; Lac Kavitaha, Ampefy; 20 Mar 1958; F. Keiser; NMB • 4 ♂♂ 2 ♀♀; Ambalavao, Fianarantsoa; 28 - 29 Jan 1958; F. Keiser; NMB • 9 ♂♂ 2 ♀♀; Tzimbazaza Park, Antananarivo; 7 - 8 Feb 1968; F. Keiser leg.; NMB • 1 ♀; Station Agric., Alaotra, District Anbatondrakzaka; 24 Dec 1957; B. R. Stuckenberg leg.; NMSA • 4 ♂♂ 2 ♀♀; Antananarivo; 1 Jan 1958; B. R. Stuckenberg leg.; NMSA.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
D68C114EABBE50678F1529F49480749B.taxon	materials_examined	Other material Madagascar • 1 ♀; Antananarivo, Tsimbazaza; 16 Oct 1993; M. Hauser leg.; CAS • 1 ♂; Antananarivo, Tsimbazaza; 16 - 22 Oct 1993; C. Kassebeer leg.; CAS • 1 ♂ 1 ♀; Antananarivo, Tsimbazaza; 16 - 22 Oct 1993; C. Kassebeer leg.; CNC • 1 ♂; Antananarivo, Tsimbazaza; 14 Oct 1993; C. Kassebeer leg.; NHMUK • 1 ♀; Antananarivo, Tsimbazaza; 26 Oct 1993; C. Kassebeer leg.; NHMUK • 1 ♀; Atsimo, Andrefana; 6 - 16 Jul 2012; M. Irwin and R. Harin'Hala leg.; CAS • 1 ♀; Atsimo, Andrefana; 31 May 2012; M. Irwin and R. Harin'Hala leg.; CAS • 1 ♀; Tananarive; 13 Dec 1957; F. Keiser leg.; NMB • 1 ♀; Tananarive; 15 Dec 1957; F. Keiser leg.; NMB • 1 ♂ 1 ♀; Tananarive; 29 Dec 1957; F. Keiser leg.; NMB • 1 ♀; Tananarive; 17 Apr 1958; F. Keiser leg.; NMB • 2 ♀♀; Tananarive; 26 - 30 Apr 1968; K. M. Guichard leg.; NHMUK.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
D68C114EABBE50678F1529F49480749B.taxon	description	Re-description male (Fig. 13). Body length: 13.7 - 14.6 mm. Wing length: 9.6 - 10.5 mm. Head (Fig. 56). Eyes bare; holoptic, length of eye contiguity approx. length of ocellar triangle. Face white with dark medial vitta; white pilose; white pollinose. Vertical triangle black pilose on ventral half and at ocellar triangle, yellow pilose on dorsal half; yellow pollinose until just before anterior ocellus. Lateral ocellus touching eye margin. Occiput yellow; yellow pilose; yellow and white pollinose. Frontal prominence shiny black; black pilose. Antenna, scape and pedicel reddish-brown; postpedicel black; antennal arista reddish-brown. Thorax. Scutum black; dorsally with a pair of well-demarcated yellow vittae and a faint yellow medial line; vittae and medial line become faint posteriorly; yellow vague lateral vitta; yellow pilose. Scutellum uniformly yellow-brown; yellow pilose. Legs. All legs dark chocolate-brown to black, except for pro- and mesotibiae and basitarsi which are reddish-brown. Proleg: Femur dark chocolate-brown to black; without apical pile brush; pile inconspicuous, except for long yellow pile on the anterodorsal side which, dorsally, is bordered with a row of largely spaced black pile. Tibia reddish-brown; yellow pilose with some short black pile on distal end dorsally. Basitarsus orange-brown; black pilose with some longer yellow pile posteriorly. Other tarsi black; black pilose, with some longer yellow pile posteriorly, except on the two most distal tarsi. Mesoleg (Fig. 180): Femur with predominantly long yellow pile on ventral and posteroventral side, with shorter black pile on anterodorsal side. Tibia reddish-brown; proximal half with a comb of long, curved yellow pile on proximal half; with much shorter, yellow and black pile on distal half. Basitarsus orange-brown; black pilose with some longer yellow pile posteriorly. Other tarsi black; black pilose, with some longer yellow pile posteriorly, except on the two most distal tarsi. Metaleg: Femur dark chocolate-brown to black; with long, thin yellow pile, especially anteriorly; ventrally with less black pile. Tibia dark chocolate-brown to black; unmodified; with short, black pile throughout and short, yellow pile on anteroproximal half. Basitarsus and second tarsomere orange-brown, other tarsi black; black pilose. Wing (Fig. 136). Entire wing uniformly microtrichose. Abdomen (Fig. 93). Tergite II with a pair of yellow-orange, rounded maculae; black marking broadly hourglass-shaped; yellow pilose, except for posterior half of posterior black marking where pile is black; with a stretch of yellow pollinosity on anterior part of black marking. Tergite III with a pair of small, anterolateral maculae; yellow-white pilose; white pollinose on posterior half. Tergite IV dark chocolate-brown; yellow-white pilose; white pollinose. Genitalia (Fig. 215). Epandrium: Dorsal lobe of surstylus club-shaped; distal end densely covered with black spines; proximal half (' stalk') long yellow pilose dorsally and with some shorter, thicker black pile laterally. Ventral lobe of surstylus with one very long and thick black setula distally.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
D68C114EABBE50678F1529F49480749B.taxon	description	Re-description female (Fig. 34). Body length: 12.5 - 13.5 mm. Wing length: 8.7 - 9.3 mm. Head. Eyes bare; dichoptic. Face white with dark medial vitta; white pilose, white pollinose. Frons black, but strongly white pollinose on ventral 4 / 5; black pilose on ocellar triangle and just ventral of ocellar triangle; white pilose. Distance between lateral ocellus and eye margin approx. 11 / 2 x width of ocellus. Occiput yellow-white; yellow-white pilose; yellow-white pollinose. Frontal prominence shiny black, orange-brown at distal end. Antenna, scape and pedicel orange-brown; postpedicel black, white pollinose; antennal arista yellow-orange. Thorax. Scutum dark brown to black with a pair of dorsolateral white-grey pollinose vittae; with lateral grey pollinose vague vitta; yellow and black pilose. Scutellum yellow-brown; yellow and black pilose. Legs. Pro- and mesoleg: Femur black, distal end reddish-brown; short black and longer yellow pilose. Tibia reddish-brown; yellow and black pilose dorsally; yellow pilose ventrally. Basitarsus reddish-brown; black pilose dorsally; orange-yellow pilose ventrally. Other tarsi black; black pilose dorsally, orange-yellow pilose ventrally. Metaleg: Femur black, distal 1 / 5 reddish-brown; yellow-white pilose, with scarce very short black pile ventrally. Tibia reddish-brown; white pilose, short black pilose posteriorly. Basitarsus dark reddish-brown; black pilose dorsally, yellow and black pilose ventrally. Other tarsi black; predominantly black pilose dorsally, densely yellow-orange pilose ventrally. Wing. Entire wing uniformly microtrichose. Abdomen (Fig. 115). Dark brown to black; largely white pollinose, except for anterior border and medial area of tergite II and posterior half of tergite V; short black and white pilose, except for non-pollinose areas with white pilose only.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
D68C114EABBE50678F1529F49480749B.taxon	distribution	Distribution. Madagascar.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
918A816BE60C525D80994D5B7E1A1DDE.taxon	materials_examined	Examined material. Helophilus minor Bezzi: Lectotype (hereby designated), male, " LECTOTYPUS " " Hel. (Mes.) // ♂, Type // Helophilus minor // Bezzi " " Syn- // type " " Pres. By // Impl. Bureau Ent. // 1915 - 165. " " Chintechi // Nyasaland /; Dr. H. S. Stannus " " Mesembrius // Mesembrius minor n. sp. // Type ♂ " " NHMUK 013428949 " [NHMUK]. Female, " Syn- // type " " Hel. (Mes.) // ♀, Type // Mesembrius minor // Bezzi "; " Brit. E. Africa // N. of Mt. Kenia, // nr. crater lake, // 5700 ft. // T. J. Anderson. // 15. II. 1911 "; " Pres. By // Impl. Bureau Ent. // 1915 - 165. "; " Mesembrius // Mesembrius minor n. sp. // Type ♀ "; " NHMUK 013428950 " [NHMUK]. Female, " Syn- // type " " Brit. E. Africa // N. of Mt. Kenia, // nr. crater lake, // 5700 ft. // T. J. Anderson. // 15. II. 1911 " " H. (M.) Mesembrius minor Bezzi. // Bezzi det. // 1915. " " Pres. By // Impl. Bureau Ent. // 1915 - 165. " " NHMUK 013428951 " [NHMUK].	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
918A816BE60C525D80994D5B7E1A1DDE.taxon	materials_examined	Other material Benin • 5 ♀♀; Azaourisse; 7 Mar 2018; K. Jordaens leg.; KMMA • 1 ♂; Cotonou; Feb 2003; G. Goergen; IITA • 1 ♂; Cotonou; 14 Dec 2013; G. Goergen and K. Jordaens leg.; KMMA • 4 ♂♂ 5 ♀♀; Cotonou; 28 Jan 2016; K. Jordaens and G. Goergen leg.; KMMA • 1 ♀; Ouidah; Sep 2004; G. Goergen leg.; IITA • 1 ♂; Pobe; 16 Mar 2014; G. Goergen leg.; KMMA • 1 ♂; Sedje; Sep 2012; G. Goergen leg.; IITA. Cameroon • 1 ♀; Maroua, Meskina; 23 May 2018; M. Azo'o Ela leg.; MAPC • 1 ♂; Douala; 9 Jul 1974; J. A. W. Lucas leg.; RMNH. Chad • 1 ♀; Bebedja; date unknown; F. A. Bink and R. M. Bink-Moenen leg.; RMNH. Democratic Republic of the Congo • 1 ♂; Kongo-Central, Boma; 16 Jun 1915; Lang and Chapin leg.; KMMA • 1 ♂; Haut-Lomami, Kitompo, Fungwi; 18 Jun 1911; Dr. Bequaert leg.; KMMA. Mozambique • 1 ♂; Lower Shire R.; 24 Jun 1916; R. C. Wood leg.; NHMUK.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
918A816BE60C525D80994D5B7E1A1DDE.taxon	description	Re-description male (Fig. 14). Body length: 11.8 - 13.3 mm. Wing length: 8.0 - 9.8 mm. Head (Fig. 57). Eyes bare; slightly dichoptic, distance between eyes approx. the width of ocellus. Face white with dark medial vitta; white pilose; white pollinose. Vertical triangle black pilose; white pollinose in area before ocellar triangle. Distance between lateral ocellus and eye margin less than 1 / 2 width of ocellus. Occiput yellow; yellow pilose; yellow and white pollinose. Frontal triangle dark black pilose; strongly yellow pollinose on ventral half. Frontal prominence shiny black. Antenna black; antennal arista brown. Thorax. Scutum black with, dorsally, a pair of well-demarcated yellow vittae and a faint yellow medial line; vittae and line are connected anteriorly and posteriorly; with lateral, yellow vitta; yellow-white pilose. Scutellum uniformly yellow-brown; yellow pilose. Legs. Femora and entire metaleg brown; pro- and mesofemora and tarsi yellow-brown; small darkened medial patch in all tarsi, except for the basitarsus. Proleg: Femur without apical pile brush; yellow pilose on anterior and ventral side; with shorter, black pile on dorsal and posterior side. Mesoleg: Femur similar to profemur, but also with a row on longer black pile ventrally. Metaleg (Fig. 198): Femur with long, yellow pile on anterior side; with a basoventral patch of long, black pile and, somewhat more distally, a smaller patch of long, black pile on the posteroventral side; anterior to this black pile, a small swelling. Tibia unmodified; almost straight. Wing (Fig. 137). Entire wing uniformly dense microtrichose. Abdomen (Fig. 94). Tergite II with pair of very large yellow rounded maculae; black marking hourglass-shaped, with anterior black marking larger than posterior black marking, the latter not reaching the lateral sides of the tergite. Medial black vitta more or less parallel-sided. Black tergal marking with short stiff black pile which does not extend to the lateral sides. Tergite III with small triangular to rounded black marking. Tergite IV with very large rounded black marking occupying most of the tergite; narrowly white pollinose anteriorly. The yellow pile of the abdomen is very short, except on tergite V, where it is not appressed to the lateral sides of the tergite. Tergite V black; strongly white pollinose. Genitalia (Fig. 216). Epandrium: Dorsal lobe of surstylus short, nearly circular; with short black spines on almost entire surface; long yellow pilose dorsally. Ventral lobe of surstylus straight; bare. Hypandrium markedly downwardly curved.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
918A816BE60C525D80994D5B7E1A1DDE.taxon	description	Re-description female (Fig. 35). Body length: 10.6 - 13.5 mm. Wing length: 7.5 - 9.4 mm. Head. Eyes bare; dichoptic. Face white with dark medial vitta; white pilose, white pollinose. Frons black on dorsal 2 / 5, yellow-white on ventral 3 / 5; black and white pilose on ocellar triangle and just ventrally of ocellar triangle, otherwise white pilose; strongly white pollinose on ventral 3 / 5, weak white pollinose on dorsal 2 / 5. Distance between lateral ocellus and eye margin slightly less than width of ocellus. Occiput yellow-white; yellow-white pilose; yellow-white pollinose. Frontal prominence shiny black. Antenna dark brown to black; antennal arista reddish-brown. Thorax. Scutum dark brown with a pair of dorsolateral yellow pollinose vittae which are connected posteriorly; with lateral, yellow pollinose vitta; sometimes with a fine medial white to yellow pollinose vita; yellow pilose. Scutellum yellow-orange; yellow pilose. Legs. Proleg (Fig. 173): Femur black, distal end orange-brown; yellow pilose, with short black pile interspersed. Tibia orange, darkened in distal 1 / 2; yellow pilose on dorsal proximal half, yellow and black pilose otherwise. Tarsi orange-brown with darkened medial area; black pilose dorsally, yellow pilose ventrally; especially the posterior side has very conspicuous thick black pile. Mesoleg: Femur black, distal end orange-brown; black and white pilose. Tibia orange-brown; orange-yellow pilose on dorsal side, black pilose on ventral side. Tarsi orange-brown with darkened dorsal medial area; black pilose. Metaleg: Femur black, distal end orange-brown; orange-yellow pilose, with short, black pile on dorsal distal end, with shorter and thicker black pile on ventral distal half; with marked ventral swelling in middle (Fig. 204). Tibia orange-brown; orange-yellow pilose with some black pile interspersed at distal end. Tarsi black dorsally, orange ventrally; black pilose dorsally; densely orange pilose ventrally. Wing. Entire wing uniformly microtrichose. Abdomen (Fig. 116). Tergite II with a pair of very large, rounded yellow-orange maculae; black pilose on triangular posteromedial section, yellow pilose otherwise; posterior black marking does not reach the lateral tergal sides; medial part of black marking narrow, approx. 1 / 10 of tergal width; posterior black marking white pollinose. Tergite III with yellow-orange fascia which occupies entire tergal length on lateral sides and approx. 1 / 3 of tergal length in medial section; with triangular posterior black marking that does not reach the lateral tergal sides; black pilose on triangular posteromedial section, yellow pilose otherwise; posterior black marking white pollinose. Tergite IV as tergite III but with much narrower yellow-orange fascia (approx. 1 / 10 of tergal length in medial section). Tergite V with narrow anterior black marking; with a pair of yellow-orange maculae in anterolateral corners, otherwise black; yellow pilose; black marking strongly white pollinose, especially in anterior half.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
918A816BE60C525D80994D5B7E1A1DDE.taxon	distribution	Distribution. Benin, Cameroon, Chad, Democratic Republic of the Congo, Malawi and Mozambique.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
8B3481DFE2735475A417BB3469539B9A.taxon	materials_examined	Examined material. Helophilus morio Bezzi: Holotype, female, " Holo- // type " " Hel. (Mes.) // Type // Helophilus morio // Bezzi " " Neguelo, // Usambara, // German E. Africa // Purchd. From // H. Rolle. // 1904 - 117. " " Mesembrius // Mesembrius morio n. sp. // Type ♀ "; " NHMUK 013428952 " [NHMUK]. Paratype: Tanzania • 1 ♀; Usambara Mountains, Neguelo; date unknown; H. Rolle leg.; NHMUK.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
8B3481DFE2735475A417BB3469539B9A.taxon	materials_examined	Other material Democratic Republic of the Congo • 1 ♀; Eala; 24 Aug 1935; J. Ghesquiere leg.; KBIN. Malawi • 1 ♀; Mount Mulanje; 6 Nov 1913; S. A. Neave leg.; NHMUK. Tanzania • 2 ♀♀; Neguelo, Usambara Mountains; date unknown; H. Rolle leg.; NHMUK. Uganda • 2 ♀♀; Entebbe; 17 Jun 1972; H. Falke leg.; CNC.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
8B3481DFE2735475A417BB3469539B9A.taxon	description	Re-description female (Fig. 36). Body length: 12.7 - 13.5 mm. Wing length: 11.6 - 12.5 mm. Head (Fig. 74). Eyes bare; dichoptic. Face black; black and white pilose; white pollinose. Frons black; black pilose; lower half white pollinose. Vertex black; black pilose; grey pollinose. Distance between lateral ocellus and eye margin approx. the width of ocellus. Occiput black; yellow and black pilose dorsally, yellow pilose more ventrally; grey pollinose. Frontal prominence shiny brown-black; black pilose. Antenna black; arista reddish-brown. Thorax. Scutum and scutellum black; without vitta; short white and black pilose. Legs. Dark reddish-brown to black; short black and white pilose. Wing (Fig. 138). Entire wing uniformly, very densely microtrichose; dark brown in anterior half. Abdomen (Fig. 117). Entirely black; short yellow-white and black pilose. Male. Unknown.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
8B3481DFE2735475A417BB3469539B9A.taxon	distribution	Distribution. Democratic Republic of the Congo, Malawi, Tanzania and Uganda.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
7691ECD7246651228DCD1D9BE08E00EC.taxon	materials_examined	Examined material. Mesembrius nigriceps Curran: Holotype, male, " Mesembrius // TYPE // Mesembrius nigriceps // Curran " " Taken from Bembex " " Stanleyville, Cgo. // 25 ° 10 ' E, 0 ° 30 ' N // III. 1915 " " Lang & Chapin // Collectors " [AMNH]. Type studied from picture on website.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
7691ECD7246651228DCD1D9BE08E00EC.taxon	materials_examined	Other material Ghana • 1 ♂; Eastern Region, N of Kibi, Atewa Range Forest Reserve; 21 Jun 2006; K. - D. B. Dijkstra leg.; MZH. Togo • 1 ♂; Kloto Forest; Mar 2004; G. Goergen leg.; IITA.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
7691ECD7246651228DCD1D9BE08E00EC.taxon	description	Re-description male (Fig. 15). Body length: 11.0 mm. Wing length: 8.4 mm. Head (Fig. 58). Eyes bare; slightly dichoptic, distance between eyes approx. width of ocellus. Face white with dark medial vitta; white pilose; white pollinose. Vertical triangle black; black pilose; lower half weakly white pollinose. Distance between lateral ocellus and eye margin somewhat less than width of ocellus. Frontal triangle black; white pilose; grey pollinose at the sides. Frontal prominence shiny black; black pilose. Occiput black; yellow pilose with a stretch of black pile near the eye margin; grey-white pollinose. Antenna black; antennal arista brown. Thorax. Scutum black with, dorsally, a pair of faint yellow vittae which fade out posteriorly; lateral yellow vitta very faint; yellow-rufous pilose. Scutellum uniformly yellow-brown; yellow-rufous pilose, with some short black pile interspersed, especially in the posterior half. Legs. Femora and entire metaleg dark brown to black; pro- and mesofemora and tarsi dark brown; tarsi without a small darkened medial patch. Proleg: Femur without apical pile brush; short black pilose dorsally, long black pilose ventrally, long yellow pilose posteriorly. Tibia long yellow pilose and short black pilose, except for a row of long black pile posterodorsally. Tarsi black pilose dorsally, yellow-orange pilose ventrally. Mesoleg: Femur similar to profemur, but with long, black pile on posterior and posteroventral side and with black pile on anterodorsal side which is markedly longer in the proximal half. Tibia yellow pilose ventrally, except at extreme distal end, where it is also black; short black pilose dorsally; long black pilose anterordorsally, especially in proximal 1 / 2. Tarsi black pilose dorsally, yellow-orange pilose ventrally, with some thick black pile on ventral side. Metaleg (Fig. 192): Femur weakly curved; thickened in distal 1 / 3; with long yellow pile on anterior and anteroventral side; ventrally with dense, long black pile in proximal 1 / 3 and less thick and less dense black pile elsewhere; no swelling on the mid-section of the ventral side. Tibia strongly curved, especially from posterior view; flattened; with very long, black pile on dorsal and ventral side. Tarsi black pilose dorsally, yellow-orange pilose ventrally. Wing (Fig. 139). Entire wing uniformly dense microtrichose. Abdomen (Fig. 95). Tergite II with a pair of large, triangular, yellow maculae; anterior and posterior black markings equal in size and with broad medial black marking; black markings with short, stiff black setulae which do not extend to the lateral sides; strongly yellow-orange pilose. Tergite III with a pair of smaller, triangular to rounded yellow maculae in anterior half; strongly yellow-orange pilose. Tergite IV black; long yellow-orange pilose, especially on lateral sides. Genitalia (Fig. 217). Epandrium: Dorsal lobe of surstylus strongly bent, sickle-shaped, short yellow pilose on distal half; with long, thick black setulae at bend ventrally; distal half dorsally broadly convex; densely covered with long yellow pile and with some equally long, but thicker black pile interspersed. Ventral lobe of surstylus bare.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
7691ECD7246651228DCD1D9BE08E00EC.taxon	distribution	Distribution. Democratic Republic of the Congo, Ghana and Togo.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
42E7056D366851C986C5B613BCE4D966.taxon	materials_examined	Examined material. Holotype, male: Tanzania • Niussi; 17 Dec 1905; Chr. Schroeder leg. (type not found / studied).	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
42E7056D366851C986C5B613BCE4D966.taxon	materials_examined	Other material Benin • 1 ♂; Calavi; 11 Nov 1993; G. Goergen leg.; IITA • 1 ♂; Calavi; Oct 2001; G. Goergen leg.; IITA. Democratic Republic of the Congo • 1 ♂; Elisabethville [= Lubumbashi]; Apr 1930; M. Bequaert leg.; KMMA • 1 ♂; Elisabethville [= Lubumbashi]; Apr 1930; M. Bequaert leg.; RMNH • 1 ♂; Tshibinda; 21 - 27 Aug 1931; W. P. Cockerell leg.; NHMUK. Kenya • 1 ♂; Kakamega Forest, Isecheno Station; 24 Jan 1991; Earthwatch Team 2 leg.; NMK. Uganda • 1 ♂; Entebbe; 27 May 1912; C. C. Gowdey leg.; NHMUK • 4 ♂♂; Entebbe; 7 - 9 May 1912; C. C. Gowdey leg.; NHMUK • 1 ♂; Entebbe; 21 Aug 1911; C. C. Gowdey leg.; NHMUK • 1 ♂; Entebbe; 18 - 20 Nov 1912; C. C. Gowdey leg.; NHMUK • 3 ♂♂; Entebbe; 7 Oct 1971; H. Falke leg.; CNC • 1 ♂; Entebbe; 23 - 31 Jan 1973; H. Falke leg.; CNC • 1 ♂; Entebbe; 25 - 27 Mar 1973; H. Falke leg.; CNC.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
42E7056D366851C986C5B613BCE4D966.taxon	description	Re-description male (Fig. 16). Body length: 14.6 - 15.7 mm. Wing length: 10.5 - 11.5 mm. Head (Fig. 59). Eyes bare; slightly dichoptic, distance between the eyes approx. the width of ocellus. Face yellow with dark medial vitta; white pilose; white pollinose. Vertical triangle black; black pilose; yellow pollinose on lower half. Distance between lateral ocellus and eye margin less than 1 / 2 width of ocellus. Occiput black; yellow pilose with black pile in dorsal area; yellow and white pollinose. Frontal triangle short; black; with some long black pile; white pollinose. Frontal prominence shiny black with orange-brown apex. Antenna, scape and postpedicel black; pedicel dark orange-black; antennal arista reddish-brown. Thorax. Scutum black with, dorsally, a pair of faint grey pollinose vittae; lateral vitta faint, not well-demarcated; black pilose with long yellow pile on anterolateral part and postpronotum. Scutellum uniformly yellow-brown; with long yellow and black pile. Legs. Proleg (Figs 152, 163): Femur black; dorsoventrally flattened; with a black apical pile brush; proximoventral section with long, thick black setae; posterior side with long golden pile. Tibia dorsally brown, ventrally orange-brown; with, especially on the ventral side, long black pile. Basitarsus very broad; orange; with a tuft of orange pile on posterior side (Fig. 163). Other tarsi very broad; orange; becoming shorter distally; the most distal tarsal segment white; sparsely black pilose dorsally, but with denser short black pile in anterior half, short orange pilose ventrally. Mesoleg: Femur dark brown; with long yellow pile on ventroproximal side, scattered yellow pile posterodorsally, but black at distal end. Tibia dorsally brown, ventrally orange-brown; with a tuft of black curved pile on ventroproximal end. Tarsi orange; sparsely black pilose dorsally, orange pilose ventrally with some thick long black pile. Metaleg (Fig. 185): Femur long and slender; dark brown; black pilose ventrally, the pile gradually becomes longer towards distal end; pile otherwise yellow and less dense. Tibia dorsally dark brown, ventrally orange-brown; in anterior view with a strong carina on the ventral side in the middle; with three deep depressions on the posterior side of the proximal half which are bordered with long, black pile, especially dorsally. Tarsi dark brown; sparsely black pilose dorsally, short orange pilose ventrally. Wing (Fig. 140). Entire wing uniformly dense microtrichose. Abdomen (Fig. 96). Tergite II with pair of large, yellow triangular maculae; black marking hourglass-shaped, white pollinose on posterior end; yellow and black pilose, but black pile most conspicuous on posterior black marking of tergite. Tergite III with broad yellow fascia and a triangular black marking on posterior half; black marking strongly white pollinose, covered with short black spines; otherwise yellow pilose. Tergite IV with large triangular posterior black marking, otherwise yellow; strongly white pollinose. Genitalia (Fig. 218). Epandrium: Dorsal lobe of surstylus short, broadly rounded, with short, black spines on almost entire surface. Ventral lobe of surstylus straight; bare.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
42E7056D366851C986C5B613BCE4D966.taxon	distribution	Distribution. Benin, Democratic Republic of the Congo, Kenya, Tanzania and Uganda.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
48C3F554CBCC5493984B7E0CD53C6BB6.taxon	materials_examined	Examined material. Tityusia regulus Hull: Holotype, male, " Efufup // Kamerun, // W. Africa // VIII. 30.1919 " " Carn. Mus. // Acc. 6552 " " type " " Tityusia // Tityusia regulus // type Hull " " Monstromyia rex // Hull Curr. " [MCZ] [type studied from pictures].	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
48C3F554CBCC5493984B7E0CD53C6BB6.taxon	materials_examined	Other material Benin • 2 ♂♂ 1 ♀; Calavi; Apr 2014; G. Goergen leg.; IITA • 1 ♀; Calavi; Oct 2015; G. Goergen leg.; IITA • 1 ♀; Ifangni-range; 6 May 2016; G. Goergen leg.; KMMA • 1 ♂ 2 ♀♀; Ifangni-range; 19 Mar 2017; G. Goergen leg.; KMMA • 1 ♂ 1 ♀; Pobe; 27 Jan 2016; G. Goergen leg.; IITA • 1 ♀; Porto Novo; Mar 2003; G. Goergen leg.; IITA • 1 ♀; Porto Novo; Dec 2005; G. Goergen leg.; IITA • 1 ♀; Porto Novo; Jul 2005; G. Goergen leg.; IITA • 1 ♀; Porto Novo; Jan 2008; G. Goergen leg.; IITA • 1 ♂ 2 ♀♀; Porto Novo; Mar 2008; G. Goergen leg.; IITA • 2 ♂♂; Porto Novo; 31 Jan 2014; G. Goergen leg.; KMMA • 2 ♀♀; Porto Novo; 27 Jan 2016; G. Goergen leg.; KMMA • 3 ♀♀; Porto Novo; date unknown; K. Jordaens leg.; KMMA. Democratic Republic of the Congo • 1 ♀; Equateur, Eala; Oct 1935; J. Ghesquiere leg.; KMMA • 1 ♀; Equateur, Eala; Sep 1935; J. Ghesquiere leg.; RMNH • 1 ♂; Equateur, Eala; Aug 1935; J. Ghesquiere leg.; KBIN • 1 ♀; Equateur, Eala; J. Ghesquiere leg.; KBIN • Jan 1936; J. Ghesquiere leg.; KBIN • 1 ♀; Equateur, Lopri River; May-Jun 1927; J. Ghesquiere leg.; KMMA • 1 ♀; Terr. de Banningville, Kwilu, Panga; Aug 1945; Fain leg.; KMMA • 1 ♀; Tshuapa, Flandria [= Boteka]; 18 Oct 1945; P. Hulstaert leg.; KMMA • 1 ♀; Ubangi, Nzali; 3 - 4 Mar 1932; H. J. Bredo leg.; KMMA • 1 ♂; Uele, Tukpwo; Jul 1937; J. Vrijdagh leg.; KMMA. Nigeria • 1 ♀; Lagos; 22 Nov 1911; W. A. Lamborn leg.; OXUM • 1 ♀; Lagos; 20 Feb 1912; W. A. Lamborn leg.; OXUM • 2 ♂♂; Lagos; 21 Mar 1912; W. A. Lamborn leg.; OXUM. Togo; 1 ♂; Kloto Forest; Feb 2008; G. Goergen leg.; IITA.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
48C3F554CBCC5493984B7E0CD53C6BB6.taxon	description	Re-description male (Fig. 18). Body length: 21.5 - 24.2 mm. Wing length: 13.2 - 15.0 mm. Head (Fig. 61). Eyes bare; holoptic, eye contiguity as long as length of ocellar triangle. Face white with dark medial vitta; white pilose; white pollinose. Vertical triangle black; black pilose; yellow pollinose on lower half. Ocellus and eye touching. Occiput black; yellow pilose; black pilose dorsally; yellow and white pollinose. Frontal triangle short; black; with some long black pile; strongly white pollinose. Frontal prominence shiny black with orange apex. Antenna black; postpedicel strongly white pollinose; antennal arista orange-brown. Thorax. Scutum dark brown to black with, dorsally, a pair of very faint, grey pollinose vittae which fade out posteriorly; pile short, dense, black and yellow-white. Scutellum yellow-brown with darker anterior border; with dense yellow and, on the posterior half and centre, shorter, black pile. Legs. Proleg (Figs 153, 171): Femur dark brown; dorsoventrally flattened; with a dark brown apical pile brush; remainder of posterior side with less dense, long brown pile. Tibia orange-brown in proximal 1 / 3, but darker in distal 2 / 3; very broad; with brown to black pile which is longer posteriorly. Basitarsus orange-brown, longer than wide. Other tarsi orange-brown; progressively becoming shorter, wider and lighter; most distal tarsal segment greyish. Mesoleg: Femur dark brown; with long yellow pile on ventroposterior 4 / 5, black on distal 1 / 5; pile otherwise short and black. Tibia orange-brown; with long black pile ventrally and shorter, strongly curved black pile dorsally. Tarsi orange; with short, black pile. Metaleg (Figs 183, 202): Femur long and slender; orange-brown; with long yellow pile on all but ventral sides, except for long black pile at extreme distal end; pile much shorter and black ventrally. Tibia orange-brown; with brown to black long pile; unmodified. Basitarsus orange-brown; with a very conspicuous thick tuft of very long and very dense brown pile on distal dorsal half; with long brown pile at extreme proximal end ventrally. Second tarsomere orange-brown; with long brown pile posteriorly. Other tarsi orange-brown; sparsely black pilose dorsally, short orange-brown pilose ventrally. Wing (Fig. 142). Entire wing uniformly dense microtrichose. Abdomen (Fig. 98). Tergite II with a pair of large yellow, rounded maculae; black marking hourglass-shaped; yellow pilose in anterior half and along tergite margins, black pilose in posterior half; black marking white pollinose posteriorly. Tergite III with yellow fascia and a triangular black marking on posterior half which is strongly white pollinose; yellow pilose on anterior 1 / 3 and along tergite margins, black pilose on posterior 2 / 3. Tergite IV dark brown to black; yellow-white pilose, but with shorter, black pile medially; strongly white pollinose on anterior 1 / 3 to 1 / 2. Genitalia (Fig. 220). Epandrium: Dorsal lobe of surstylus short, broadly rounded; with short, black spines on almost entire surface; long yellow pilose dorsally, especially at proximal end. Ventral lobe of surstylus straight; bare.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
48C3F554CBCC5493984B7E0CD53C6BB6.taxon	description	Description female (Fig. 38). Body length: 11.8 - 16.7 mm. Wing length: 11.2 - 12.5 mm. Head (Fig. 76). Eyes bare; dichoptic. Face white with dark medial vitta; white pilose; white pollinose. Distance between lateral ocellus and eye margin approx. width of ocellus. Occiput black; yellow and black pilose; yellow pollinose. Frons black; black pilose; yellow pollinose on ventral half, sometimes pollinosity almost absent. Frontal prominence shiny black, orange-brown at distal end; scape and pedicel orange-brown to black; postpedicel black; postpedicel white pollinose; antennal arista reddish-brown. Thorax. Scutum dark brown to black with, dorsally, a pair of very vague yellow pollinose vittae; short yellow and black pilose. Legs. All legs brown to black, protibia and protarsus lighter, yellow-brown; protarsus lighter than distal part of protibia; profemur predominantly black pilose, the pile is longer on the posterior and posterodorsal side than on the remainder of the profemur; pro- and mesotibia black pilose in distal 1 / 2 - 1 / 4, otherwise yellow and black pilose. Wing. Entire wing uniformly dense microtrichose. Wing cell r 1 nearly closed. Abdomen (Fig. 119). Tergite II with a pair of large, orange maculae; black medial marking narrow, approximately 1 / 9 of tergal width; orange pilose on anterior half, black pilose on posterior half; posterior black marking strongly white pollinose. Tergite III with orange fascia (approx. half of tergite length on lateral sides; approx. 1 / 5 of tergite length in medial area); orange pilose on anterior end, otherwise black pilose; posterior half white pollinose, especially in medial area. Tergite IV as tergite III, but yellow pilose throughout with black pile interspersed on black marking. Tergite V black with or without a pair of vague orange maculae in anterolateral corner; yellow pilose; white pollinose on anterior half.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
48C3F554CBCC5493984B7E0CD53C6BB6.taxon	distribution	Distribution. Benin, Cameroon, Democratic Republic of the Congo, Nigeria and Togo.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
251C351C606C5B46884DCE570E23B925.taxon	materials_examined	Examined material. Mesembrius rex Curran: Holotype, male, " Mesembrius // TYPE // Mesembrius rex Curran // No. " " Taken from Bembex " " Stanleyville, Cgo. // 25 ° 10 ' E, 0 ° 30 ' N // IV. 7.1915 " " Lang & Chapin // collectors " " Stanleyville // Congo // From Leg of // Type [♂] " " Mesembrius // Mesembrius rex // det. Curran // Det. C. H. Curran " [AMNH] [type studied from picture on website].	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
251C351C606C5B46884DCE570E23B925.taxon	materials_examined	Other material Democratic Republic of the Congo • 1 ♀; Bolongo; 23 Jun 1936; J. Ghesquiere leg.; KBIN • 1 ♀; Lulua, Kapanga; Nov 1928; Walker leg.; KMMA • 1 ♀; Basoko; Oct 1948; P. L. G. Benoit leg.; RMNH • 1 ♂; Eala; Oct 1935; J. Ghesquiere leg.; RMNH • 1 ♀; locality and date unknown; J. Ghesquiere leg.; KBIN. Malawi • 1 ♀; Mount Mulanje, Likhubula; 19 Nov 1912; S. A. Neave leg.; NHMUK • 1 ♂; Mount Mulanje; 25 Nov 1912; S. A. Neave leg.; NHMUK • 1 ♀; Mount Mulanje; 2 Dec 1912; S. A. Neave leg.; NHMUK • 1 ♀; Mount Mulanje; 16 Nov 1912; S. A. Neave leg.; NHMUK • 1 ♀; Mount Mulanje; 25 Nov 1912; S. A. Neave leg.; NHMUK. Togo • 1 ♀; Kloto Forest; Feb 2016; G. Goergen leg.; IITA. Uganda • 1 ♂; W. shores of Vic. Nyanza, Buddu; 19 - 25 Sep 1911; S. A. Neave leg.; NHMUK • 1 ♂; Barada; 16 Apr 1940; B. Lebied leg.; NHMUK • 1 ♂; Entebbe; 11 Nov 1912; C. C. Gowdey leg.; NHMUK • 2 ♂♂; Entebbe; 7 Oct 1971; H. Falke leg.; CNC • Country Unknown • 1 ♀; locality unknown; 1973; F. M. Hull leg.; CNC.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
251C351C606C5B46884DCE570E23B925.taxon	description	Re-description male (Fig. 19). Body length: 14.3 - 17.0 mm. Wing length: 10.0 - 11.3 mm. Head (Fig. 62). Eyes bare; holoptic, eye contiguity approximately as long as the length of the ocellar triangle. Face yellow with dark medial vitta; white pilose; white pollinose. Frontal triangle short, black, with a few long, black pile; white pollinose on dorsal half; vertical triangle black, black pilose, yellow pollinose on lower half. Distance between lateral ocellus and eye margin less than 1 / 2 width of ocellus. Frontal prominence shiny black with orange-brown apex. Occiput black; yellow pilose, with black pile interspersed dorsally; yellow and white pollinose. Antenna, scape and pedicel black; postpedicel brown; antennal arista orange-brown. Thorax. Scutum black with, dorsally, one pair of grey pollinose vittae; lateral vitta faint, not well-demarcated; pile, especially on the anterior half yellow, but with some black pile interspersed; pile on posterior half very short. Scutellum black in anterior 1 / 3, brown in middle 1 / 3, white-yellow in posterior 1 / 3, with long yellow and shorter black pile. Metasternum with very long, strongly curved golden pile. Legs. Proleg (Fig. 154): Femur black; dorsoventrally flattened; with an apical pile brush of long, black pile dorsally and long, yellow pile ventrally. Tibia orange-brown; with very long, yellow pile on anteroventral side. Basitarsus orange; with a tuft of orange pile on posterior side. Other tarsi very broad; orange; becoming shorter distally; most distal tarsal segment white. Mesoleg: Femur dark brown; with long yellow pile dorsally, except for black pile on dorsal 1 / 5. Tibia orange-brown; proximal half strongly compressed. Basitarsus orange-brown. Other tarsi dark brown. Metaleg (Fig. 184): Coxa with long yellow pile on anteroventrally. Femur long and slender; chocolate-brown; ventrally with a row of> 10 short, widely spaced black spines in the proximal 2 / 3; with denser, black spines on posterior 1 / 3; pile otherwise yellow and loose. Tibia orange-brown; with a deep invagination in the proximal 1 / 3 posteriorly which is bordered with long, black pile ventrally; remainder of ventral side with dense, black pile. Tarsi orange-brown. Wing (Fig. 143). Entire wing uniformly dense microtrichose; brown infuscated in dorsal half. Abdomen (Fig. 99). Tergite II with a pair of large, yellow rounded maculae; black markings hourglass-shaped, white pollinose on posterior end; yellow and black pilose, but black pile more conspicuous in posterior part of black marking. Tergite III with broad yellow fascia and a triangular black marking on posterior half which is strongly white pollinose and covered with short, black spines; yellow pilose otherwise. Tergite IV with large triangular posterior black marking; otherwise yellow with strong white pollinosity. Genitalia (Fig. 221). Epandrium: Dorsal lobe of surstylus short, broadly rounded, with short, black spines on almost entire surface; long brown pilose dorsally. Ventral lobe of surstylus straight; bare.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
251C351C606C5B46884DCE570E23B925.taxon	description	Description female (Fig. 39). Body length: 16.0 - 16.2 mm. Wing length: 11.2 - 12.5 mm. As M. regulus, but with the following differences: Pro- and mesotibia yellow pilose, with only short black pile on posterior side of mesotibia; protarsus chocolate-brown, concolourous with protibia (Fig. 170); wing cell r 1 distinctly open. Abdomen (Fig. 120) with orange pilosity somewhat more prominent. Head as in Fig. 77.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
251C351C606C5B46884DCE570E23B925.taxon	distribution	Distribution. Democratic Republic of the Congo, Malawi, Togo and Uganda.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
DF44BA57885C50E8951B8C915B0E3D6B.taxon	materials_examined	Examined material. Helophilus senegalensis Macquart: Lectotype (hereby designated), male, " SYNTYPE " " MNHN, Paris // ED 6788 " " 1 ♂ Helophilus // Helophilus senegalensis Macq // C. F. Kassebeer 1999 " [MNHN]. Paralectotype, female, " SYNTYPE " " MNHN, Paris // ED 6789 " [MNHN] [the female is indicated as male on MNHN website] [a paralectotype is present at the MNHN, but could only be studied from the pictures on the website; see comments]. Helophilus africanus Verrall: Lectotype (hereby designated), male, " S. W. ARABIA // 19 m. fr. Aden, // Haithalhim. // Capt. Mar. 23.95 // & press. 1899 by // J. W. Yerbury. // Trans. Ent. Soc., // 1898, page 413. " " S. W. ARABIA // 19 m. fr. Aden, // Haithalhim. // Capt. Mar. 23.95 // & press. 1899 by // J. W. Yerbury. // Trans. Ent. Soc., // 1898, page 416. " " TYPE. // G. H. VERRALL // Trans. Ent. Soc., // 1898, page 413. " " TYPE. // G. H. VERRALL // Trans. Ent. Soc., // 1898, page 416. " " TYPE Dipt: 105 1 / 4 // Helophilus // Mesembrius africanus // Verrall // HOPE DEPT. OXFORD " " 1899 // 7645 " " RMCA PIC // 00012 " " LECTOTYPUS " [OXUM]. Paralectotype, male, " S. W. ARABIA // 19 m. fr. Aden, // Haithalhim. // Capt. Mar. 23.95 // & press. 1899 by // J. W. Yerbury. // Trans. Ent. Soc., // 1898, page 413. " " S. W. ARABIA // 19 m. fr. Aden, // Haithalhim. // Capt. Mar. 23.95 // & press. 1899 by // J. W. Yerbury. // Trans. Ent. Soc., // 1898, page 416. " " TYPE. // G. H. VERRALL // Trans. Ent. Soc., // 1898, page 413. " " TYPE. // G. H. VERRALL // Trans. Ent. Soc., // 1898, page 416. " " TYPE Dipt: 105 3 / 4 // Helophilus // Mesembrius africanus // Verrall // HOPE DEPT. OXFORD " " 1899 // 7646 " " RMCA PIC // 00013 " " PARA- // LECTOTYPUS " [OXUM]. Paralectotype, female, " S. W. ARABIA // 19 m. fr. Aden, // Haithalhim. // Capt. Mar. 24.95 // & press. 1899 by // J. W. Yerbury. // Trans. Ent. Soc., // 1898, page 413. " " S. W. ARABIA // 19 m. fr. Aden, // Haithalhim. // Capt. Mar. 24.95 // & press. 1899 by // J. W. Yerbury. // Trans. Ent. Soc., // 1898, page 416. " " TYPE. // G. H. VERRALL // Trans. Ent. Soc., // 1898, page 413. " " TYPE. // G. H. VERRALL // Trans. Ent. Soc., // 1898, page 416. " " TYPE Dipt: 105 4 / 4 // Helophilus // Mesembrius africanus // Verrall // HOPE DEPT. OXFORD " " 1899 // 7650 " " RMCA PIC // 00015 " " PARA- // LECTOTYPUS " [OXUM]. Paralectotype, female, " S. W. ARABIA // 19 m. fr. Aden, // Haithalhim. // Capt. Mar. 23.95 // & press. 1899 by // J. W. Yerbury. // Trans. Ent. Soc., // 1898, page 413. " " S. W. ARABIA // 19 m. fr. Aden, // Haithalhim. // Capt. Mar. 23.95 // & press. 1899 by // J. W. Yerbury. // Trans. Ent. Soc., // 1898, page 416. " " TYPE. // G. H. VERRALL // Trans. Ent. Soc., // 1898, page 413. " " TYPE. // G. H. VERRALL // Trans. Ent. Soc., // 1898, page 416. " " TYPE Dipt: 105 5 / 4 // Helophilus // Mesembrius africanus // Verrall // HOPE DEPT. OXFORD " " 1899 // 7648 " " RMCA PIC // 00016 " " PARA- // LECTOTYPUS " [OXUM]. Paralectotype, female, " Haithalhim // 23.3.95 // Col. Yerb. " " VC-TYPE 33 // Helophilus ♀ // Mesembrius africanus // Verrall " " Haithalhim " " PARA- // LECTOTYPUS " [OXUM].	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
DF44BA57885C50E8951B8C915B0E3D6B.taxon	materials_examined	Other material Benin • 1 ♂; Cotonou; Feb 2003; G. Goergen leg.; IITA • 2 ♀♀; Cotonou; 14 Jan 2013; K. Jordaens and G. Goergen leg.; KMMA • 1 ♂ 1 ♀; Cotonou; 28 Jan 2016; K. Jordaens and G. Goergen leg.; KMMA • 1 ♂; Togbin; Dec 2005; G. Goergen leg.; IITA. Chad • 1 ♀; Bebedja; date unknown; F. A. Brink and R. M. Brink-Moenen leg.; RMNH. Kenya • 2 ♂♂ 7 ♀♀; Jipe, Taita-Taveta; 27 Jan 2017; M. Reemer leg.; RMNH • 2 ♂♂ 2 ♀♀; Jipe; Taita-Taveta; 27 Jan 2017; X. Mengual leg.; ZFMK • 1 ♀; Nairobi, ICIPE campus; 6 May 2014; R. Copeland leg.; ICIPE • 1 ♂; Taita Hills; 2017; A. Ssymank leg.; ASPC • 1 ♀; Makindu; 5 - 7 Apr 1911; S. A. Neave leg.; NHMUK. Oman • 1 ♂; Dhofar, Ayun pools; 8 Oct 1977; K. M. Guichard leg.; NHMUK.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
DF44BA57885C50E8951B8C915B0E3D6B.taxon	description	Re-description male (Fig. 20). Body length: 12.0 - 13.8 mm. Wing length: 9.2 - 10.2 mm. Head (Fig. 63). Eyes bare; slightly dichoptic, distance between eyes approx. 1 / 2 width of ocellus. Face yellow with dark medial vitta; yellow pilose; yellow pollinose. Vertical triangle with yellow pile and yellow pollinosity in lower half, black in upper half; distance between lateral ocellus and eye margin less than 1 / 2 width of ocellus. Occiput yellow; yellow pilose with interspersed short, black setulae; yellow and white pollinose. Frontal triangle yellow; yellow pilose; yellow pollinose. Frontal prominence shiny black; yellow pilose. Antenna black; antennal arista reddish-brown. Thorax. Scutum black with three dorsal, well-demarcated yellow vittae which are connected anteriorly and posteriorly; with lateral, yellow vitta; pile rufous. Scutellum yellow-brown; yellow pilose. Legs. All legs light- to dark brown. Proleg (Fig. 161) and mesoleg (Fig. 181): profemur without apical pile brush; yellow pilose, pile ventrally long in proximal half, shorter in distal half; with shorter black pile on distal half. Tibia yellow and black pilose. Tarsi black pilose dorsally, yellow pilose ventrally; with some thick black pile posterodorsally. Metaleg: Femur anteriorly and dorsally with long and posteriorly with shorter, yellow pile; ventral yellow pile scarce, except for a row of long, thin pale pile; with band of short black pile posteroventrally. Tibia unmodified; long yellow pilose; ventrally with much shorter and thicker black pile. Tarsi black pilose dorsally, yellow pilose ventrally. Wing (Fig. 144). Entire wing uniformly dense microtrichose. Abdomen (Fig. 100). Tergite II with a pair of very large, yellow to orange rectangular maculae; medial black markings very narrow and perpendicular to anterior and posterior narrow, black marking; posterior black marking with short, stiff black setulae which do not extend to the lateral tergite sides; white pollinose. Tergite III and IV with very large, yellow fascia; yellow pilose; with large medial black marking; with short black stiff setulae posterior to medial dark spot, these setulae not reaching the lateral tergite sides. Tergite III white pollinose in medial part. Tergite IV entirely white pollinose. Genitalia (Fig. 222). Epandrium: Dorsal lobe of surstylus distally broadly rounded, with characteristic upwardly pointed projection; long pilose dorsally; with shorter, dense black pile ventrally and laterally. Ventral lobe of surstylus with a row of approx. 10 long black setulae.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
DF44BA57885C50E8951B8C915B0E3D6B.taxon	description	Re-description female (Fig. 40). Body length: 11.5 - 14.4 mm. Wing length: 8.3 - 10.4 mm. As male, except for the following character states: Eyes dichoptic (Fig. 78). Frons yellow pilose in ventral 2 / 3, black and yellow pilose on dorsal 1 / 3 (ocellar triangle and surrounding area); strongly yellow pollinose. Pile on legs shorter (Figs 174, 200). Abdomen as in Fig. 121.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
DF44BA57885C50E8951B8C915B0E3D6B.taxon	distribution	Distribution. Benin, Chad, Kenya, Oman and Yemen.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
2B80D8D091985DA382C8F97213F40741.taxon	materials_examined	Examined material. Mesembrius simplicipes Curran: Holotype, male: " Mesembrius // TYPE // Mesembrius simplicipes // Curran. // No. " " Mesembrius // Mesembrius simplicipes // Curran " " Madagascar // Great Oriental // Forest " " California Academy // of Sciences // Type No. 11230 " [CAS] [date and collector unknown; type studied from pictures]. Mesembrius platytarsis Curran: Holotype, male: " Mesembrius // TYPE // Mesembrius platytarsis // Curran. // No. " " Madagascar // Great Oriental // Forest " " California Academy // of Sciences // Type No. 11229 " [CAS] [date and collector unknown; type studied from pictures].	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
2B80D8D091985DA382C8F97213F40741.taxon	materials_examined	Other material Madagascar • 2 ♀♀; Alaotra, Station Agric.; 24 Dec 1957; B. R. Stuckenberg leg.; NMSA • 1 ♀; Analvony; 30 Mar 1958; F. Keiser leg.; NMB • 1 ♂; Antananarivo; Nov 1952; E. S. Brown leg.; NHMUK • 1 ♂; Antananarivo; 18 Oct 1957; F. Keiser leg.; NMB • 1 ♀; Antananarivo; 13 Dec 1957; F. Keiser leg.; NMB • 2 ♀♀; Antananarivo; 14 Dec 1957; F. Keiser leg.; NMB • 1 ♂; Antananarivo; 6 Sep 1958; F. Keiser leg.; NMB • 1 ♀; Antananarivo; 8 Feb 1970; L. and R. Blommers leg.; RMNH • 1 ♂; Antananarivo, Ampefy, Lake Kavitaha; 25 Mar 1959; F. Keiser leg.; NMB • 1 ♂; Antananarivo, Ampefy, Lake Kavitaha; 29 Mar 1959; F. Keiser leg.; NMB • 1 ♂; Antananarivo, Parc Tsimbazaza; 2 Feb 1968; J. W. Boyes leg.; CNC • 1 ♀; Antananarivo, Park Tsimbazaza; 14 Dec 1957; F. Keiser; NMB • 1 ♀; Antananarivo, Park Tsimbazaza; 15 - 22 Oct 1993; C. Kassebeer leg.; NHMUK • 1 ♂ 1 ♀; Antananarivo, Park Tsimbazaza; 16 - 22 Oct 1993; C. Kassebeer leg.; CNC • 2 ♂♂; Antananarivo, Park Tsimbazaza; 16 - 22 Oct 1993; C. Kassebeer leg.; CAS • 1 ♂ 1 ♀; Antananarivo, Park Tsimbazaza; 16 - 22 Oct 1993; C. Kassebeer leg.; NMK • 1 ♀; Antananarivo, Park Tsimbazaza; 26 Oct 1993; C. Kassebeer leg.; CNC • 1 ♀; Antananarivo, Park Tsimbazaza; 26 Oct 1993; C. Kassebeer leg.; CAS • 1 ♂; Antananarivo, Park Tsimbazaza; 6 Nov 1993; C. Kassebeer leg.; CNC • 1 ♂; Antananarivo, Park Tsimbazaza; 6 Nov 1993; C. Kassebeer leg.; NHMUK • 1 ♂; Antananarivo, Perinet; 30 Sep 1957; F. Keiser leg.; NMB • 1 ♀; Nosivola; date and collector unknown; RMNH.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
2B80D8D091985DA382C8F97213F40741.taxon	description	Re-description male (Figs 17, 21). Body length: 12.7 - 13.4 mm. Wing length: 8.8 - 10.4 mm. Head (Figs 60, 64). Eyes bare; holoptic, eye contiguity as long as length of ocellar triangle. Face white with dark medial vitta; white pilose; white pollinose. Vertical triangle black; black pilose; strongly yellow pollinose before anterior ocellus. Lateral ocelli nearly touching eye margin. Occiput black, yellow pilose with a few very short, stiff black setulae near dorsal eye margin; strongly white pollinose. Frontal triangle black; black pilose; strongly yellow pollinose. Frontal prominence shiny reddish-black. Antenna, scape and pedicel very dark reddish-black; postpedicel black; antennal arista reddish-brown. Thorax. Scutum black with dorsally a pair of yellow pollinose vittae which are less well demarcated posteriorly; with a lateral yellow pollinose vitta; yellow pilose. Scutellum uniformly yellow-brown; yellow pilose. Legs. Proleg (Figs 166, 167): Femur dorsoventrally flattened; dorsally brown, except proximal 1 / 5 and distal end; otherwise orange; yellow pile on posteroventral side longer than black pile on posterodorsal side; with a patch of very short black stiff spines on proximal ventral 1 / 5; pile entirely yellow anteriorly. Tibia orange; with long orange pile, but pile black on ventral distal end. Basitarsus whitish; laterally expanded; with a tuft of orange pile on the expansion; ventrally with some thick stiff pile on the expansion. Other tarsi orange; with orange and black short pile. Mesoleg: Femur as proleg, but without the posterodorsal black pile. Tibia orange; yellow pilose, with some short stiff black pile on distal end ventrally. Tarsi orange; with short black pile and some short stiff black pile ventrally. Metaleg (Figs 193, 194): Femur chocolate-brown, distal end orange; with loose yellow pile on proximal 1 / 2 to 2 / 3 and mostly black pilose on distal 1 / 3 to 1 / 2. Tibia orange; with short black pile. Metabasitarsus either deeply excavated anteriorly at proximal end and with a lobe (Fig. 193) or unmodified (Fig. 194); orange; black pilose. Other tarsi orange; dorsally black pilose, ventrally orange pilose. Wing (Figs 141, 145). Entire wing uniformly very dense microtrichose. Abdomen (Figs 97, 101). Tergite II with a pair of very large, yellow almost square maculae which expand into the anterolateral corners; black markings hourglass-shaped, posterior part reaching the lateral sides of the tergite; posterior part of black marking with some black pile in the centre which do not extend to the lateral tergite sides; otherwise yellow pilose. Tergite III with a yellow fascia which is almost interrupted by the medial broad black marking; the latter strongly white pollinose anteriorly; with a medial area of white pollinosity; posterior part of black marking with sparse black pile in the centre; yellow pilose otherwise. Tergite IV strongly white pollinose, except for a black medial area and a darker posterior border; yellow pilose. Genitalia (Figs 219, 223). Epandrium: Dorsal lobe of surstylus short, bent (as a boomerang); irregularly covered with short black spines which are denser at distal and proximal end and at dorsal bend. Ventral lobe of surstylus straight; bare.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
2B80D8D091985DA382C8F97213F40741.taxon	description	Re-description female (Figs 37, 41). Body length: 14.2 - 14.9 mm. Wing length: 10.0 - 10.6 mm. Head (Figs 75, 79). Eyes bare; dichoptic. Face yellow-white with dark medial vitta; white pilose, white pollinose. Frons black in dorsal half and medial part of ventral half, yellow-white on lateral parts of ventral half; black pilose on black parts, white pilose on yellow-white parts. Distance between lateral ocellus and eye margin approx. width of ocellus. Occiput yellow-white; yellow-white pilose; yellow-white pollinose. Frontal prominence shiny black, orange-brown at distal end; scape black; pedicel orange-brown; postpedicel black; antennal arista reddish-brown. Thorax. Scutum dark brown with one pair of dorsolateral yellow-white pollinose vittae which are vaguely connected posteriorly; with lateral, yellow-white pollinose vitta; short yellow pilose on anterior half, short yellow and black pilose on posterior half. Scutellum yellow-orange; yellow pilose with shorter, black pile interspersed in posterior half. Legs. Proleg: Femur orange brown with darker, central area on dorsal side; yellow-white pilose with black pile on posterodorsal side and posteroventral distal half. Tibia orange, slightly darkened on ventral distal end; yellow-white pilose. Tarsi orange-brown; basitarsus and second tarsomere yellow-white pilose, other tarsi yellow-white and black pilose. Femur orange brown with darker, central area on dorsal side; yellow-white pilose with black pile on ventral distal half. Tibia orange-brown; yellow-white pilose, with some short, thick black pile at ventral distal end. Basitarsus orange; yellow-white pilose, with short, thick black pile on ventral side. Other tarsi orange-brown; yellow-white and black pilose; with short, thick black pile ventrally, except on most distal tarsomere. Metaleg: Femur dark brown to black, reddish-brown at distal end; yellow-white pilose with shorter and thicker black pile on anteroventral and ventral distal 1 / 2. Tibia orange-brown; yellow-white and black pilose. Tarsi orange-brown; black and yellow-white pilose dorsally, yellow-orange pilose ventrally. Wing. Entire wing uniformly microtrichose. Abdomen (Figs 118, 122). Tergite II with a pair of large orange maculae; with an anterior and posterior black marking which are connected with a parallel-sided central black marking which is 1 / 5 the tergite width; yellow pilose on maculae and on anterior and central black marking, black pilose on posterior black marking; white pollinose on posterior black marking. Tergite III with orange fascia (approx. 1 / 2 of tergal length laterally; approx. 1 / 8 in medially); with posterior large black marking; yellow pilose on most of the orange fascia, black pilose on black marking and central area of orange fascia; strongly white pollinose on posterior half and medial anterior part. Tergite IV similar, but without orange fascia. Tergite V orange-brown with darker lateral sides; yellow-white pilose; white pollinose in anterolateral corners.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
2B80D8D091985DA382C8F97213F40741.taxon	distribution	Distribution. Madagascar.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
904BC299CC2E5B9EA5B5B35FABC01813.taxon	materials_examined	Examined material. Tubifera strigilata Bezzi: Lectotype (hereby designated), male, " LECTOTYPUS " " SYNTYPUS ♂ // Tubifera (Mesemb.) // Tubifera strigilata // Bezzi, 1912 " " Congo Francese // Fernand-Vaz // IX-X. 1902. L. fea " " RMCA PIC // 00033 " " Museo Civico // di Genova " " LECTOTYPUS " [MSNG]. Paralectotype, 4 males, " SYNTYPUS ♂ // Tubifera (Mesemb.) // Tubifera strigilata // Bezzi, 1912 " " Congo Francese " // Fernand-Vaz // IX-X. 1902. L. fea " " Museo Civico // di Genova " " PARA- // LECTOTYPUS " [MSNG]. Paralectotype, female, " SYNTYPUS ♀ " " Tubifera (Mesemb.) // Tubifera strigilata // Bezzi, 1912 " " Museo Civico // di Genova " " PARA- // LECTOTYPUS " [MSNG].	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
904BC299CC2E5B9EA5B5B35FABC01813.taxon	materials_examined	Other material Benin • 1 ♂ 1 ♀; Azaourisse; 7 Mar 2018; K. Jordaens leg.; KMMA • 1 ♂; Calavi; 27 Jan 2017; G. Goergen leg.; IITA • 1 ♀; Cotonou; 14 Dec 2013; G. Goergen and K. Jordaens leg.; KMMA • 1 ♀; Cotonou; Feb 2003; G. Goergen leg.; IITA • 1 ♀; Cotonou; Dec 2003; G. Goergen leg.; IITA • 1 ♂; Lokossa; Jun 2006; G. Goergen leg.; IITA • 1 ♂; Pahou; 11 Jan 2014; G. Goergen leg.; IITA • 1 ♂ 1 ♀; Pobe; 27 Jan 2016; G. Goergen leg.; IITA • 2 ♂♂; Porto Novo; 20 Jan 2018; G. Goergen leg.; IITA • 1 ♂ 1 ♀; Gblo Gblo; 11 Sep 2014; G. Goergen leg.; KMMA • 1 ♀; Pahou; 11 Jan 2014; G. Goergen leg.; KMMA • 1 ♂ 1 ♀; Porto Novo; 27 Jan 2016; K. Jordaens leg.; KMMA • 7 ♂♂; Porto Novo; 7 Mar 2018; K. Jordaens leg.; KMMA • 1 ♀; Serou; Nov 2016; G. Goergen leg.; KMMA • 1 ♂ 3 ♀♀; Pobe; 12 Dec 2013; G. Goergen and K. Jordaens leg.; KMMA • 4 ♂♂ 4 ♀♀; Pobe; 28 Jan 2016; G. Goergen and K. Jordaens leg.; KMMA. Burundi • 2 ♂♂; Bujumbura; 21 Feb 2017; G. Goergen leg.; KMMA • 1 ♂; Nyanza-Lac; 14 Oct 2013; L. Ndayikeza leg.; OBPE • 1 ♀; Rusizi River; 10 Nov 2010; L. Ndayikeza leg.; OBPE. Cameroon • 1 ♂; Batanga; collection date unknown; A. I. Good leg.; CNC • 5 ♂♂ 2 ♀♀; Douala; 9 Jul 1974; J. A. W. Lucas leg.; RMNH. Democratic Republic of the Congo • 2 ♀♀; Haut-Katanga; 4 Sep 1930; G. F. de Witte leg.; KMMA • 1 ♀; Equateur, Bamania; 21 Jul 1924; J. Bequaert leg.; KMMA • 1 ♀; Equateur, Bamania; 24 Jul 1924; J. Bequaert leg.; KMMA • 1 ♂; Basoko; Oct 1948; P. L. G. Benoit leg.; KMMA • 1 ♂; Bokuma; Jul 1962; R. P. Lootens leg.; KMMA • 1 ♀; Mai-Ndombe, Bololo, Makamendulu; 1938; H. Schouteden leg.; KMMA • 2 ♂♂; Boma; Jul 1915; Lang and Chapin leg.; KMMA • 1 ♀; Boma; 16 Jun 1915; Lang and Chapin leg.; KMMA • 1 ♀; Boma; 17 Jun 1915; Lang and Chapin leg.; KMMA • 1 ♀; Boma; 18 Jun 1915; Lang and Chapin leg.; KMMA • 4 ♀♀; Boma; 4 Dec 1920; H. Schouteden leg.; KMMA • 1 ♀; Boma; 11 Jul 1920; H. Schouteden leg.; KMMA • 1 ♂ 1 ♀; Boma; 12 Jul 1920; H. Schouteden leg.; KMMA • 1 ♀; Haut-Lomami, Bukama; 8 Jun 1911; J. Bequaert leg.; KMMA • 1 ♀; South-Kivu, Bukavu; May 1949; H. Bomans leg.; KMMA • 1 ♂; Equateur, Eala; Jul 1931; H. J. Bredo leg.; KMMA • 1 ♀; Equateur, Eala; Nov 1931; H. J. Bredo leg.; KMMA • 1 ♀; Equateur, Eala; 7 Oct 1931; H. J. Bredo leg.; KMMA • 1 ♂; Equateur, Eala; Jul 1932; A. Corbissier leg.; KMMA • 1 ♀; Equateur, Eala; Apr 1933; A. Corbissier leg.; KMMA • 1 ♂; Equateur, Eala; 1933; A. Corbissier leg.; KMMA • 1 ♀; South-Kivu, Kabare; 1 ♀; 31 Jul 1914; J. Bequaert leg.; KMMA • 1 ♀; Lomami, Kabinda; date unknown; Schwetz leg.; KMMA • 1 ♂; Kachichwe; 17 Jan 1912; Dr. Bequaert leg.; KMMA • 1 ♀; Kalemie; Dec 1918; R. Mayne; leg.; KMMA • 1 ♂; Lualaba; Kabombo; 29 Jun 1947; M. Poll leg.; KMMA • 1 ♀; Leopoldville [= Kinshasa]; 28 Oct 1951; mevr. Bequaert leg.; KMMA • 1 ♂; Tshopo, Stanleyville [= Kisangani]; Apr 1915; Lang and Chapin leg.; KMMA • 1 ♂; Equateur, Lukolela; 17 Jul 1926; J. Bequaert leg.; KMMA • 2 ♂♂; Lomami, Luputa; Mar 1935; Bouvier leg.; KMMA • 1 ♀; Natl. Parc Albert, Rwindi, St. Edouard; 17 Apr 1936; L. Lippens leg.; KMMA • 1 ♀; Tumbalunga, Dibaya; 8 Nov 1930; G. F. de Witte leg.; KMMA • 1 ♀; Ubani, Bosobolo; 8 - 11 Jan 1932; H. J. Bredo leg.; KMMA • 1 ♀; Ubangi, Tungu; 4 Mar 1932; H. J. Bredo leg.; KMMA • 1 ♀; Uele, Garamba; Jul 1912; Lang and Chapin leg.; KMMA • 1 ♂ 1 ♀; Mai-Ndombe, Wimbali; Jul 1913; P. Vanderijst leg.; KMMA • 1 ♀; 30 Sep 1913; P. Vanderijst leg.; KMMA. Gabon • 1 ♀; Lolo River; 22 May 1925; J. Rodhain leg.; KMMA. Ghana • 1 ♀; Nsakjam; 13 Sep 2016; G. Goergen leg.; KMMA • 1 ♂; Tema; 19 Dec 2004; G. Goergen leg.; IITA • 1 ♂; Kumasi; 6 Nov 1946; J. Bowden leg.; NMSA. Kenya • 1 ♂; Kabete; 12 Jun 1916; T. J. Anderson leg.; NHMUK • 1 ♂; Merifano; Nov 1932; McArthur leg.; NMK • 1 ♀; Mugura Forest; 2 May 1981; R. H. Markham leg.; NMK • 1 ♀; Zwani; date unknown; van Someren leg.; CNC. Madagascar • 1 ♂; Antananarivo; 28 Feb 2016; G. Goergen leg.; IITA. Malawi • 4 ♂♂ 2 ♀♀; Chiromo; date unknown; J. E. S. Old. leg.; NHMUK • 1 ♂; Mount Mulanje; 4 Oct 1913; S. A. Neave leg.; NHMUK • 1 ♂; Monkey Bay; 12 Dec 1980; J. H. G. Londt leg.; NMSA • 3 ♂♂; Senga Hills; 1 Dec 1980; B. R. Stuckenberg leg.; NMSA. Mozambique • 1 ♀; Lourenco-Marques [= Maputo]; date unknown; H. A. Junod leg.; NHMUK • 2 ♂♂; Sofala, Gorongosa National Park, Chitengo; 20 - 30 Apr 2004; M. Hauser leg. and H. Rung leg.; CAS • 2 ♂♂ 1 ♀; Sofala, Gorongosa National Park, Chitengo; 16 - 30 Apr 2015; M. Hauser and H. Rung leg.; CAS • 6 ♂♂; Luabo, Lower Zambezi River; 1 Jun 1957; B. R. Stuckenberg leg.; NMSA • 2 ♂♂; Luabo, Lower Zambezi River; 1 Jun 1957; P. Usher leg.; NMSA • 1 ♂; Luabo, Lower Zambezi River; 1 Aug 1957; P. Usher leg.; NMSA • 1 ♂; Siluwe Hills, W. of Beira; 3 Jun 1964; D. Cookson leg.; NMSA. Nigeria • 1 ♂; Ibadan; 14 Jun 1957; G. H. Caswell leg.; NHMUK • 1 ♂; Ibadan; Dec 1988; G. Goergen leg.; IITA. Senegal • 1 ♀; Dassilame, Serere; 30 Nov 2016; S. Cavailles leg.; SCPC. South Africa • 1 ♂; Durban; Jul 1903; G. Burn leg.; NMSA • 1 ♂; KwaZulu-Natal, Durban, Blue Lagoon; 25 May 1991; J. A. W. Lucas leg.; RMNH • 1 ♂; KwaZulu-Natal, Durban; 14 May 1903; G. F. Leigh leg.; NMSA • 2 ♂♂; KwaZulu-Natal, Nseleni Nature Reserve; 10 Jan 1994; Natal Museum Staff leg.; NMSA • 2 ♂♂; KwaZulu-Natal, St. Lucia Park Reserve; 2 Feb 1988; J. H. G. Londt leg.; NMSA • 5 ♂♂; KwaZulu-Natal, Dukuduku Forest; 18 Jul 1981; B. R. Stuckenberg leg.; NMSA • 1 ♂; KwaZulu-Natal, Dukuduku Forest, 4 ♂♂ W of St. Lucia; 26 Nov 1971; M. E. Irwin leg.; NMSA • 1 ♂; KwaZulu-Natal, Ndumu Game Reserve; 26 Oct 1972; M. E. Irwin leg.; NMSA. Tanzania • 1 ♂; Kahe, Usambara Mountains; 2 Jun 1916; T. J. Anderson leg.; NHMUK. Togo • 1 ♀; Kloto Forest; Mar 2004; G. Goergen leg.; IITA • 1 ♂; Kloto Forest; May 2016; G. Goergen leg.; KMMA. Uganda • 1 ♀; Busoga; Mar 1906; A. Hodges leg.; CNC • 1 ♂; Tero Forest, S. E. Buddu; 26 - 30 Sep 1911; S. A. Neave leg.; NHMUK. Zambia • 1 ♂ 1 ♀; Lusaka Province, 8.5 km NW Katondwe; 20 Apr 2016; M. Hauser leg.; CSCA.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
904BC299CC2E5B9EA5B5B35FABC01813.taxon	description	Re-description male (Fig. 22). Body length: 11.2 - 12.5 mm. Wing length: 8.1 - 8.8 mm. Head (Fig. 65). Eyes bare; slightly dichoptic, distance between eyes approx. the width of ocellus. Face white with dark medial vitta; white pilose; white pollinose. Distance between lateral ocellus and eye margin 1 / 2 width of ocellus. Occiput yellow; yellow pilose; yellow and white pollinose. Frontal triangle black; black pilose; ventral half weakly white pollinose. Frontal prominence shiny black. Antenna black; antennal arista brown. Thorax. Scutum black with dorsally a pair of weak yellow vittae which fade out posteriorly; with very faint lateral yellow vitta; yellow-rufous pilose. Scutellum uniformly yellow-brown; yellow-rufous pilose, with some short black pile interspersed, especially in the posterior half. Legs. Femora and entire metaleg dark brown to black; pro- and mesofemora and tarsi yellow-brown; tarsi without a small darkened medial patch. Proleg: Femur without apical pile brush; yellow pilose ventrally; with long black pile on anterodorsally; with shorter, black pile dorsally. Mesoleg: Femur similar as profemur, but with long, black pile posteriorly and posteroventrally; with black pile anterodorsally which is markedly longer in the proximal half. Metaleg (Fig. 197): Femur with long, yellow pile anteriorly and anteroventrally; with long black pile on the posteroventral distal 1 / 3; thickened on distal 1 / 3; no swelling on the mid-section of the ventral side. Tibia strongly curved, especially from posterior view, flattened; with very long black pile dorsally and ventrally. Wing (Fig. 146). Entire wing uniformly dense microtrichose. Abdomen (Fig. 102). Tergite II with a pair of very large yellow, rounded maculae; anterior black marking larger in size than posterior black marking, which mostly reach to the lateral tergite sides; medial black marking more or less parallel-sided; long yellow pilose; posterior black marking with short, stiff black pile which extend to the lateral sides. Tergite III with small, triangular to rounded black marking; long yellow pilose with some short thick black pile in medial posterior area; black marking usually strongly white pollinose. Tergite IV entirely dark brown to black, except for anterior 1 / 5 which is strongly white pollinose; long yellow pilose, the pile is strongly appressed on the lateral sides. Tergite V dark brown. Genitalia (Fig. 224). Epandrium: Dorsal lobe of surstylus strongly bent, sickle-shaped, short yellow pilose on distal half; with long, thick black setulae at bend ventrally; distal half dorsally convex; densely covered with long yellow pile and with some equally long, but thicker black pile interspersed. Ventral lobe of surstylus bare.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
904BC299CC2E5B9EA5B5B35FABC01813.taxon	description	Re-description female (Fig. 42). Body length: 11.3 - 15.0 mm. Wing length: 8.1 - 9.4 mm. Head. Eyes bare; dichoptic. Face white with dark medial vitta; white pilose, white pollinose. Frons black on dorsal 2 / 5, yellow-white on ventral 3 / 5; black and white pilose on ocellar triangle and just ventrally of ocellar triangle, otherwise white pilose; strongly white pollinose on ventral 3 / 5, weak white pollinose on dorsal 2 / 5. Distance between lateral ocellus and eye margin slightly less than width of ocellus. Occiput yellow-white; yellow-white pilose; yellow-white pollinose. Frontal prominence shiny black. Antenna dark brown to black; antennal arista reddish-brown. Thorax. Scutum dark brown with a pair of dorsolateral yellow pollinose vittae which are connected posteriorly; with lateral, yellow pollinose vitta; sometimes with a fine medial white to yellow pollinose vitta; yellow pilose. Scutellum yellow-orange; yellow pilose. Legs. Proleg (Fig. 172): Femur black, distal end orange-brown; yellow pilose, with short black pile interspersed. Tibia orange, darkened in distal 1 / 2; yellow pilose on dorsal proximal half, yellow and black pilose otherwise. Tarsi uniformly dark brown; black pilose dorsally, yellow pilose ventrally; especially the posterior side has very conspicuous thick black pile. Mesoleg: Femur black, distal end orange-brown; black and white pilose. Tibia orange-brown; orange-yellow pilose on dorsal side, black pilose on ventral side. Tarsi orange-brown with darkened dorsal medial area; black pilose. Metaleg: Femur black, distal end orange-brown; orange-yellow pilose, with short, black pile on dorsal distal end, with shorter and thicker black pile on ventral distal half; without ventral swelling in middle (Fig. 203). Tibia orange-brown; orange-yellow pilose with some black pile interspersed at distal end. Tarsi black dorsally, orange ventrally; black pilose dorsally; densely orange pilose ventrally. Wing. Entire wing uniformly microtrichose. Abdomen (Fig. 123). Tergite II with a pair of very large, rounded yellow-orange maculae; black pilose on triangular posteromedial section, yellow pilose otherwise; posterior black marking extends to lateral tergite sides; medial part of black marking narrow, approx. 1 / 10 of tergal width; posterior black marking white pollinose. Tergite III with yellow-orange fascia which occupies entire tergal length on lateral sides and approx. 1 / 3 of tergal length in medial section; with triangular posterior black marking that extends to the lateral tergite sides; black pilose on triangular posteromedial section, yellow pilose otherwise; posterior black marking white pollinose. Tergite IV as tergite III, but with much narrower yellow-orange fascia (approx. 1 / 10 of tergal length in medial section). Tergite V with narrow anterior black board; with a pair of yellow-orange maculae in anterolateral corners, otherwise black; yellow pilose; black marking strongly white pollinose, especially in anterior half.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
904BC299CC2E5B9EA5B5B35FABC01813.taxon	distribution	Distribution. Benin, Burundi, Cameroon, Democratic Republic of the Congo, Gabon, Ghana, Guinea-Bissau, Kenya, Madagascar, Malawi, Mozambique, Nigeria, Senegal, South Africa, Tanzania, Togo, Uganda and Zambia.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
762E8C6BA67D5D0E831DFC9035491346.taxon	materials_examined	Examined material. Mesembrius sulcus Jordaens, Goergen & De Meyer: Holotype, male, " HOLOTYPUS " " MUSEE DU CONGO // Ituri: Nioka // - VII- 1934 // J. Leroy " " van Doesburg det., 1956 // Mesembrius // spec.? nov. ♂ " " RMCA ENT // 000030186 " [KMMA]. Paratypes: Malawi • 1 ♂; Mount Mulanje; 17 Oct 1913; S. A. Neave leg.; NHMUK 013428977 • 1 ♂; Zomba; Feb 1911; J. E. G. Old. Leg.; NHMUK • 2 ♂♂; Mount Mulanje; 17 Oct 1913; S. A. Neave leg.; NHMUK. Kenya • 3 ♂♂ 3 ♀♀; Nairobi, Karura Forest; 2 Dec 2017; K. Jordaens leg.; ICIPE. South Africa • 1 ♂; Port St. Johns; 1 - 31 Oct 1969; E. and W. Gess leg.; AMGS.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
762E8C6BA67D5D0E831DFC9035491346.taxon	description	Description male (Fig. 23). Body length: 14.0 - 15.6 mm. Wing length: 10.2 - 11.5 mm Head (Fig. 66). Eyes bare; holoptic, eye contiguity approx. as long as length of ocellar triangle. Face yellow to orange with dark medial vitta; white pilose; white pollinose. Vertical triangle black; black pilose; yellow pollinose on medium third. Distance between lateral ocellus and eye margin slightly less than width of ocellus. Occiput yellow; yellow pilose with some shorter and thicker black pile near eye margin; yellow and white pollinose. Frontal triangle short; yellow-white; with some long, yellow and black pile; white pollinose. Frontal prominence shiny black with orange-brown apex. Antenna, scape and pedicel reddish-brown; postpedicel black, white pollinose; antennal arista reddish-brown. Thorax. Scutum black with dorsally a pair of very faint white pollinose vittae; lateral white pollinose vitta very faint; yellow pilose. Scutellum uniformly yellow-brown; long yellow pilose with some very short black pile on posterior half. Legs. All legs chocolate-brown to black, but protarsus orange. Proleg (Figs 155, 164): Femur dorsoventrally flattened; with long yellow pile posterodorsally; with apical pile brush of thick black pile dorsally and thick yellow pile ventrally; with short and black thick pile at proximal end. Basitarsus orange; with a tuft of black pile posteriorly. Other tarsi orange; with sparse short black pile. Mesoleg (Fig. 176): Femur, ventrally with long yellow pile on proximal 2 / 3 and shorter black pile on distal 1 / 3, with some long black pile interspersed in middle section. Metaleg (Fig. 187): Femur long and thin, slightly curved; with yellow pile anterodorsally; with some denser shorter and black pile at distal ventral end. Tibia with long black pile; posteriorly with deep and broad excavation in proximal half which is bordered with long black pile. Wing (Fig. 147). Entire wing uniformly dense microtrichose. Abdomen (Fig. 103). Tergite II with a pair of very large, yellow, more or less triangular maculae; black marking hourglass-shaped; posterior and anterior black marking slightly connected in the middle; posterior marking with strong white pollinosity; yellow pilose, except for black pile at posterior border of posterior black marking. Tergite III and IV with orange fascia; black marking on posterior half strong white pollinose; black pilose in medial part of black marking, yellow pilose on orange fascia and lateral parts of black marking. Genitalia (Fig. 225). Epandrium: Dorsal lobe of surstylus broadly rounded; with short black spines on almost entire surface; dorsally long yellow pilose. Ventral lobe of surstylus straight; bare.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
762E8C6BA67D5D0E831DFC9035491346.taxon	description	Description female (Fig. 43). Body length: 11.2 - 13.4 mm. Wing length: 12.7 - 16.6 mm. Head (Fig. 80). Eyes bare; dichoptic. Face white with dark medial vitta; white pilose; white pollinose. Frons black; black pilose in dorsal half and medial ventral half, yellow pilose on lateral sides of ventral half; yellow-white pollinose, especially in ventral half. Distance between lateral ocellus and eye margin 11 / 2 x width of ocellus. Occiput black; yellow pilose, with some black pile near eye margin; yellow-white pollinose. Frontal prominence shiny black. Antenna, scape and pedicel orange-brown; postpedicel black; antennal arista reddish-brown. Thorax. Scutum dark brown to black with dorsally a pair of vague brown pollinose vittae which are connected posteriorly; yellow pilose with some black pile interspersed. Scutellum dark brown with lighter posterior border; yellow pilose. Legs. All femora, tibiae and metatarsi dark brown to black, except for extreme distal ends which are reddish-brown (as in M. tarsatus; Fig. 168); yellow pilose; metatibia with short black spines on posteroventral 1 / 3. Pro- and mesotarsi orange; most distal tarsomere darkened distally, sometimes all tarsi darkened; dorsally short black pilose, ventrally short yellow pilose. Wing. Entire wing uniformly dense microtrichose. Abdomen (Fig. 124). Tergite II black; yellow pilose with some very short thick black pile on black markings; with a pair of L-shaped, small orange maculae; white pollinose, especially in the central area of the posterior black marking. Tergite III black; yellow pilose with short thick black pile interspersed, especially in the posterior half; with a pair of small orange maculae in the anterolateral corners. Tergite IV as tergite III, but without orange maculae. Tergite V black; yellow pilose.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
762E8C6BA67D5D0E831DFC9035491346.taxon	distribution	Distribution. Democratic Republic of the Congo, Kenya, Malawi and South Africa.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
762E8C6BA67D5D0E831DFC9035491346.taxon	etymology	Etymology. The specific epithet Mesembrius sulcus (Latin) means groove (noun in apposition) and was chosen with reference to the deep groove on the metatibia. It is to be treated as an adjective (nominative singular masculine).	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
92B41985235A5D4C8A9900262C4BF2F1.taxon	materials_examined	Examined material. Prionotomyia tarsata Bigot: Lectotype (hereby designated), male, " LECTOTYPUS " " SYN- // TYPE " " Prionotomyia // Prionotomyia tarsata Big. " " Prionotomyia ♂ // Prionotomyia tarsata Bigot // Senegal " " ex. coll. Bigot, // Press. by // G. H. Verrall. // B. M. 1894234 " " BMNH (E) # // 230741 " " NHMUK 010369820 " [NHMUK]. Paralectotype, male, " SYN- // TYPE " " Prionotomyia // Prionotomyia tarsata Big. " " Prionotomyia ♂ // Prionotomyia tarsata Bigot // Senegal " " ex. coll. Bigot, // Press. by // G. H. Verrall. // B. M. 1894234 " " BMNH (E) # // 230742 " " NHMUK 010369821 " " PARA- // LECTOTYPUS " [NHMUK].	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
92B41985235A5D4C8A9900262C4BF2F1.taxon	materials_examined	Other material Democratic Republic of the Congo • 1 ♀; Banningville [= Bandundu], Kwilu River, Panga; Aug 1945; Fain leg.; RMNH • 1 ♀; Haut-Katanga, Kasenga; 5 Mar 1912; J. Bequaert leg.; KMMA • 1 ♂; South-Kivu, Musingiro; 8 Oct 1922; Ch. Seydel leg.; KMMA • 1 ♀; Tshibinda; 21 - 27 Aug 1931; T. D. A. Cockerell leg.; NHMUK • 1 ♀; Eala; 5 Oct 1935; J. Ghesquiere leg.; RMNH • 1 ♀; Eala; Aug 1936; J. Ghesquiere leg.; RMNH • 1 ♀; Eala; Mar 1935; J. Ghesquiere leg.; RMNH • 1 ♀; Eala; Oct 1936; J. Ghesquiere leg.; RMNH • 1 ♀; Lulua, Kapanga; Nov 1928; Walker leg.; RMNH • 1 ♂; Lulua, Kapanga; Aug 1932; F. G. Overlaet leg.; RMNH. Kenya • 1 ♂; Naivasha, Fisherman's Camp; 14 Mar 1993; M. De Meyer leg.; NMK • 4 ♂♂ 2 ♀♀; Nairobi; Mar 1928; van Someren leg.; KMMA • 1 ♂; Nairobi; 20 Mar 1921; A. F. J. Gedye leg.; NMK • 7 ♂♂ 2 ♀♀; Nairobi, Karura Forest; 23 Nov 2017; PINDIP course leg.; KMMA. Malawi • 1 ♂; Mulanje Mountain; 26 Nov 1912; S. A. Neave leg.; NHMUK • 1 ♂ 1 ♀; Zomba Plateau; 1 Dec 1911; collector unknown; NHMUK • 3 ♂♂; Zomba Plateau; 24 Nov 1980; B. R. Stuckenberg leg.; NMSA • 2 ♂♂; Zomba Plateau; date unknown; H. S. Stannus leg.; NHMUK • 1 ♂; Zomba Plateau; 24 - 27 Nov 1980; J. G. H. Londt and B. Stuckenberg leg.; NMSA • 7 ♂♂ 6 ♀♀; Zomba, Kuchawe Trout Farm; 8 - 11 Nov 2016; K. Jordaens leg.; KMMA. Senegal • 2 ♂♂; locality and date unknown; Bigot leg.; NHMUK • 1 ♂; Nema Ba; 10 Nov 2016; S. Cavailles leg.; SCPC. South Africa • 2 ♂♂; Barber Nature Reserve; 7 Oct 2015; A. Vujic et al. leg.; UNS • 1 ♂; KwaZulu-Natal, Dukuduku Forest Reserve; 18 - 19 Jul 1981; J. G. H. Londt and B. Stuckenberg leg.; NMSA • 1 ♂; KwaZulu-Natal, Umlalazi Nature Reserve; 8 Nov 1997; J. G. H. Londt and A. Londt leg.; NMSA • 1 ♂; KwaZulu-Natal, Bluff Nature Reserve; 3 Sep 2018; J. Midgley leg.; NMSA • 1 ♂; KwaZulu-Natal, Durban; 23 Apr 1920; C. N. Barker leg.; DMSA • 1 ♂; KwaZulu-Natal, Durban; 8 Feb 1919; C. N. Barker leg.; DMSA • 1 ♂; KwaZulu-Natal, Durban; Nov-Dec 1945; H. W. Bell Marley leg.; DMSA • 1 ♂; KwaZulu-Natal, Pietermaritzburg, Botanical Gardens; 15 Nov 2018; J. Midgley leg.; NMSA • 1 ♂; KwaZulu-Natal, Ngoye Forest Reserve; 29 Jan 1968; J. G. H. Londt leg.; NMSA • 1 ♂; KwaZulu-Natal, Ubombo Mountain Reserve; 11 Oct 2019; D. Brothers leg.; NMSA • 1 ♂; KwaZulu-Natal, Umlalazi Nature Reserve; 8 Nov 1997; J. G. H. Londt leg.; NMSA • 1 ♂; KwaZulu-Natal, Dukuduku Forest, 4 ♂♂ W of St. Lucia; 26 Nov 1971; M. E. Irwin leg.; NMSA. Uganda • 1 ♂; between Jinja and Bussia; 28 Jul- 1 Aug 1911; S. A. Neave leg.; NHMUK • 1 ♀; Jinja; Oct 1930; van Someren leg.; NHMUK • 2 ♂♂ 1 ♀; between Sewiza and Kampala; 27 - 31 Aug 1911; S. A. Neave leg.; NHMUK • 1 ♂; Entebbe; 17 Aug 1911; C. C. Gowdey leg.; NHMUK • 1 ♂ 2 ♀♀; Entebbe; 21 Aug 1911; C. C. Gowdey leg.; NHMUK • 1 ♀; Entebbe; 31 Aug 1911; C. C. Gowdey leg.; NHMUK • 1 ♂; Entebbe; 18 - 20 Nov 1911; C. C. Gowdey leg.; NHMUK • 1 ♀; Entebbe; 11 Aug 1912; C. C. Gowdey leg.; NHMUK • 1 ♂; Entebbe; 16 Aug 1912; C. C. Gowdey leg.; NHMUK • 2 ♂♂; Entebbe; 17 Aug 1912; C. C. Gowdey leg.; NHMUK • 2 ♂♂; Entebbe; 27 May 1912; C. C. Gowdey leg.; NHMUK • 3 ♂♂ 1 ♀; Entebbe; 7 - 9 May 1912; C. C. Gowdey leg.; NHMUK • 2 ♂♂ 2 ♀♀; Entebbe; 3 Nov 1912; C. C. Gowdey leg.; NHMUK • 1 ♂ 1 ♀; Entebbe; 14 Nov 1912; C. C. Gowdey leg.; NHMUK • 6 ♂♂ 1 ♀; Entebbe; 18 - 20 Nov 1912; C. C. Gowdey leg.; NHMUK • 1 ♂; Entebbe; 13 Oct 1912; C. C. Gowdey leg.; NHMUK • 1 ♂ 1 ♀; Entebbe; 16 Oct 1912; C. C. Gowdey leg.; NHMUK • 1 ♂; Entebbe; 3 Sep 1912; C. C. Gowdey leg.; NHMUK • 1 ♂; Entebbe; 18 - 20 Nov 1911; C. C. Gowdey leg.; NHMUK • 1 ♂; Entebbe; 1 - 14 Sep 1912; C. A. Wiggins leg.; NHMUK • 1 ♂; Entebbe; 1 - 11 Sep 1911; S. A. Neave leg.; NHMUK • 1 ♂; South of Maseka, Katera Forest; May 1972; E. B. Babyetagara leg.; CNC • 5 ♂♂ 2 ♀♀; N. W. shores of Vic. Nyanza; 12 - 15 Sep 1911; S. A. Neave leg.; NHMUK • 5 ♂♂ 3 ♀♀; Northern Buddu; 16 - 18 Sep 1911; S. A. Neave leg.; NHMUK • 1 ♂; Nsoje River; 2 Mar 1911; van Someren leg.; NHMUK • 1 ♂; N. Ankole, Nyrbthozi; 21 Jan 1975; M. K. Paulus leg.; CNC • 1 ♀; South of Lake George; 17 - 19 Oct 1911; S. A. Neave leg.; NHMUK • 1 ♂; S. E. Ankole; 4 - 8 Oct 1911; S. A. Neave leg.; NHMUK • 1 ♂; Tero Forest; 8 Jul 1912; C. C. Gowdey leg.; NHMUK • 2 ♂♂; S. E. Buddu, Tero Forest; 26 - 30 Sep 1911; S. A. Neave leg.; NHMUK • 1 ♀; Toro, Duro River; 6 Mar 1911; van Someren leg.; NHMUK • 1 ♂ 1 ♀; Toro, Duro River; 12 Mar 1911; van Someren leg.; NHMUK • 1 ♂; Tororo; 25 Jan 1967; F. K. Masasai leg.; NHMUK • 1 ♂; District West Uganda, W. Ankole; 19 - 24 Apr 1973; H. Falke leg.; CNC • 1 ♂; District West Uganda, W. Ankole; 30 Dec 1975; M. K. Paulus leg.; CNC • 1 ♀; locality and date unknown; R. C. Bradley leg.; NHMUK. Zambia • 1 ♂; Lake Bangweulu, Kapola, N. of Kapata; 27 Oct 1946; collector unknown; NHMUK. Zimbabwe • 1 ♂; Mporokoso; 2 Aug 1909; S. A. Neave leg.; OXUM.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
92B41985235A5D4C8A9900262C4BF2F1.taxon	description	Re-description male (Fig. 24). Body length: 13.1 - 15.4 mm. Wing length: 9.7 - 10.9 mm. Head (Fig. 67). Eyes bare; holoptic, eye contiguity somewhat shorter than length of ocellar triangle. Face white with dark medial vitta; white pilose; white pollinose. Vertical triangle black; black pilose; yellow pollinose on ventral half. Distance between lateral ocellus and eye margin 1 / 2 width of ocellus. Occiput yellow; yellow pilose, with some shorter and thicker black pile near eye margin; yellow and white pollinose. Frontal triangle short; yellow-white; with some long, black pile; yellow pollinose. Frontal prominence shiny black with orange-brown apex. Antenna black; postpedicel white pollinose; antennal arista reddish-brown. Thorax. Scutum black with, dorsally, a pair of weakly-demarcated yellow pollinose vittae which fade out posteriorly; lateral yellow pollinose vitta very faint to absent; yellow and black pilose. Scutellum dark brown to black with a lighter posterior border; with long yellow pile and shorter black pile, the latter most prominent in the posterior half. Legs (Fig. 168). All legs black, but protarsus reddish-brown; black pilose. Proleg (Fig. 159): Femur dorsoventrally flattened; posterodorsal side with yellow, long pile and some long black pile in proximal half; with apical pile brush of long, relatively loose and curved black pile; with long black pile anteroventrally. Basitarsus orange; with a tuft of black pile on posterior side and two black spots on the most distal tarsomere. Mesoleg: Femur with long yellow and black pile. Metaleg (Fig. 189): Femur with long and thin yellow pile; with some black pile towards distal end; with a patch of shorter and thicker black pile ventroproximally. Tibia with long black pile, especially in distal 2 / 3; ventrally with a swelling on distal 1 / 3. Wing (Fig. 148). Entire wing uniformly dense microtrichose. Abdomen (Fig. 104). Tergite II with a pair of very large yellow-orange rounded maculae; black marking hourglass-shaped; posterior black marking equal in size to anterior black marking and with a medial white pollinose area; yellow pilose, but black pilose on posterior black marking and on posterolateral corners. Tergite III and IV with broad yellow-orange fascia, with large black, white pollinose marking on posterior 2 / 3; black pilose on black marking and adjacent parts of yellow-orange fascia, yellow pilose on remainder of yellow fascia and on lateral sides. Genitalia (Fig. 226). Epandrium: Dorsal lobe of surstylus broadly rounded; with short black spines on almost entire surface; dorsally long yellow pilose. Ventral lobe of surstylus straight; bare.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
92B41985235A5D4C8A9900262C4BF2F1.taxon	description	Description female (Fig. 44). Body length: 11.0 - 13.2 mm. Wing length: 12.7 - 15.6 mm. Similar to the female of M. sulcus sp. nov., but the maculae on abdominal tergite II are larger and triangular (Fig. 125). Head as in Fig. 81.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
92B41985235A5D4C8A9900262C4BF2F1.taxon	distribution	Distribution. Democratic Republic of the Congo, Kenya, Malawi, Senegal, South Africa, Uganda, Zambia and Zimbabwe.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
6A80FA85551F5011BCC3E82CBBEEFAD3.taxon	materials_examined	Examined material. Mesembrius tibialis Jordaens, Goergen & De Meyer: Holotype, male " HOLOTYPUS " " Togo, Kloto Forest // II. 2017 // leg. G. Goergen " " Mesembrius tibialis // Det. K. Jordaens " " DNA 1149 A 04 // K. Jordaens // RMCA 2019 " [KMMA]. Paratypes: Togo • 2 ♂♂; Kloto Forest; Dec 2017; G. Goergen leg.; IITA.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
6A80FA85551F5011BCC3E82CBBEEFAD3.taxon	description	Description male (Fig. 25). Body length: 14. mm. Wing length: 12.7 mm. Head (Fig. 68). Eyes bare; holoptic, eye contiguity approx. as long as length of ocellar triangle. Face yellow to orange with dark medial vitta; white pollinose; yellow-white pilose. Vertical triangle black; black pilose; yellow pollinose on medium third. Distance between lateral ocellus and eye margin slightly less than width of ocellus. Occiput black; yellow pilose with some shorter and thicker black pile near eye margin; white pollinose. Frontal triangle short; yellow-white; with some long black pile; white pollinose. Frontal prominence shiny black. Antenna black; postpedicel white pollinose; antennal arista orange-brown. Thorax. Scutum black with dorsally, in the anterior half, a pair of very faint yellow pollinose vittae. Scutellum black in anterior half, yellow-brown in posterior half; yellow pilose with, in the posterior half, some shorter black pile interspersed. Legs. All legs chocolate-brown to black; distal ends black; other tarsi black. Proleg (Figs 156, 165): Femur dorsoventrally flattened; with apical pile brush of long dense and curved thick black pile dorsally and thick yellow pile ventrally; ventrally with long black and shorter yellow pile. Tibia black; with long black pile. Basitarsus black and orange; with tuft of black pile on posterior side. Tarsi 2 - 4 black, tarsomere 5 white. Mesoleg (Fig. 177): Femur with long yellow pile posterodorsally; short black pile ventroproximally. Tibia curved; proximal half compressed. Metaleg (Fig. 188): Femur with very long and thin yellow pile, especially on anterior and posterior side; with shorter black pile ventrally and posteriorly. Tibia with long black pile; without groove on posterior side. Wing (Fig. 149). Entire wing uniformly dense microtrichose. Abdomen (Fig. 105). Tergite II with a pair of very large yellow, rounded maculae; black marking hourglass-shaped; yellow pilose except for short, black pile on the posterior black marking; posterior marking white pollinose. Tergite III and IV with orange fascia; short orange pile in medial part of tergites; long yellow-orange pilose on lateral sides; with white pollinose triangular posterior area. Genitalia (Fig. 227). Epandrium: Dorsal lobe of surstylus short, broadly rounded; covered in short black spines and some longer pile. Ventral lobe of surstylus straight; bare.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
6A80FA85551F5011BCC3E82CBBEEFAD3.taxon	distribution	Distribution. Togo.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
6A80FA85551F5011BCC3E82CBBEEFAD3.taxon	etymology	Etymology. The specific epithet Mesembrius tibialis is derived from the Latin word tibia (pertaining to the tibia) and was chosen in reference to the mesotibia, which is curved and proximally compressed. It is to be treated as an adjective (nominative singular masculine).	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
2D27B621B82A5446BA6D65C5ACCF7564.taxon	materials_examined	Examined material. Mesembrius vockerothi Jordaens, Goergen & De Meyer: Holotype, male, " " UGANDA: // Kampala, // 12. xii. 1934, // F. W. Edwards. // B. M. 1935 - 203. " " HOLOTYPUS " " Mesembrius vockerothi // Jordaens & De Meyer 2019 " " NHMUK 010369964 " [NHMUK]. Paratypes: Democratic Republic of the Congo • 1 ♀; Kalembelembe, Baraka; Jul 1918; R. Mayne leg.; RMNH • 1 ♀; North-Kivu, Beni a Lesse; Jul 1911; Murtula leg.; KMMA. Kenya • 1 ♂; Jinja; Oct 1930; van Someren leg.; NHMUK • 1 ♀; Nyeri; Oct 1948; van Someren leg.; NHMUK. Uganda • 1 ♂ 1 ♀; Entebbe; 17 Aug 1911; C. C. Gowdey leg.; NHMUK • 2 ♂♂; Entebbe; 9 Nov 1971; H. Falke leg.; CNC • 1 ♂; Entebbe; 5 Jan 1972; H. Falke leg.; CNC • 1 ♀; Kampala; 12 - 20 Mar 1918; C. C. Gowdey leg.; NHMUK • 3 ♂♂; Kampala; 12 Dec 1934; F. W. Edwards leg.; NHMUK • 1 ♂ 1 ♀; Namanue; 13 Dec 1934; J. Ford leg.; NHMUK • 1 ♀; Tero Forest; 26 - 30 Sep 1911; S. A. Neave leg.; NHMUK • 1 ♀; Unyoro District; C. H. Marshall leg.; NMSA • 1 ♀; Central Region, Wakiso District, Mabamba Swamp; 16 Dec 2018; G. Stahls leg.; MZH.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
2D27B621B82A5446BA6D65C5ACCF7564.taxon	materials_examined	Other material 1 ♀ with locality and date unknown, D. Bruce leg. (NHMUK).	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
2D27B621B82A5446BA6D65C5ACCF7564.taxon	description	Description male (Fig. 26). Body length: 11.0 - 13.2 mm. Wing length: 9.6 - 10.5 mm. Head (Fig. 69). Eyes dichoptic; distance between eyes approx. the width of anterior ocellus. Face conical; white with dark medial vitta; white pilose. Vertical triangle black; black pilose; yellow pollinose on ventral half. Distance between lateral ocellus and eye margin 1 / 2 width of ocellus. Occiput yellow; yellow pilose; yellow and white pollinose. Frontal triangle short; yellow-white; with long, black pile medially, yellow pilose on gena; yellow pollinose. Frontal prominence shiny dark brown to black. Antenna dark brown, antennal arista reddish-brown. Thorax. Scutum black with, dorsally, a pair of well-demarcated white pollinose vittae which are connected posteriorly; lateral white pollinose vitta clear; yellow pilose. Scutellum yellow-brown; yellow pilose with shorter black pile interspersed on its entire surface. Legs. All femora dark brown to black, except for extreme distal ends which are orange-brown; femora yellow to orange. Pro- and mesoleg: Femur with black pile on anterior and dorsal side and with longer yellow pile on posterior and posterodorsal sides. Tarsi yellow to orange. Metaleg: Femur with long and thin yellow pile; with black pile ventrally on distal half. Tibia with yellow and black pile, of which the yellow pile is longer on posterodorsal side. Metatibia unmodified. Metatarsi dark brown. Wing (Fig. 150). Entire wing uniformly very dense microtrichose. Abdomen (Fig. 106). Tergite II with a pair of very large yellow-orange, rounded maculae; black marking hourglass-shaped; posterior black marking equal in size to anterior black marking and with a medial white pollinose area; yellow pilose, but black pilose on posterior half of black marking. Tergite III with a pair of large yellow-orange maculae; with large black marking on posterior 2 / 3; yellow pilose on maculae, black pilose on black marking. Tergite IV black, with a pair of small yellow maculae in anterolateral corners; white pilose and strongly white pollinose on anterior and lateral parts; predominantly black pilose on black marking. Genitalia (Fig. 228). Epandrium: Dorsal lobe of surstylus distally broadly rounded, with characteristic large tooth-like projection; entirely pilose, except on tooth and at basis (stalk). Ventral lobe of surstylus with one large black setula in middle section and a row of 4 - 5 long black setulae.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
2D27B621B82A5446BA6D65C5ACCF7564.taxon	description	Description female (Fig. 45). Body length: 14.0 - 15.1 mm. Wing length: 9.7 - 10.3 mm. As male, except for the following character states: Eyes dichoptic (Fig. 82). Frons white pilose, brown pilose on ocellar triangle and surrounding area; strongly white pollinose to just before ocellar triangle. Pile on legs shorter. Abdomen as in Fig. 126.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
2D27B621B82A5446BA6D65C5ACCF7564.taxon	distribution	Distribution. Democratic Republic of the Congo, Kenya and Uganda.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
2D27B621B82A5446BA6D65C5ACCF7564.taxon	etymology	Etymology. Named in honour of the Dipterist Dick Vockeroth (1928 - 2012), who already indicated on the labels that some specimens from Uganda probably belonged to a new species. The specific epithet should be treated as a noun in the genitive case.	en	Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott, Meyer, Marc De (2021): Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data. ZooKeys 1046: 1-141, DOI: http://dx.doi.org/10.3897/zookeys.1046.57052, URL: http://dx.doi.org/10.3897/zookeys.1046.57052
