taxonID	type	description	language	source
6CE79A2D0ED25B828439A36BD3B559F6.taxon	diagnosis	Diagnosis. Body size small, SBL = 7.2 to 9.5 mm. Head width medium or slightly broadened, not constricted behind eyes; vertex with a pair of paramedial pale spots on vertex and one pair of supraorbital setae. Eyes slightly to moderately reduced in size, moderately convex. Antennae medium length to slightly elongate; antennal scape with one subapicodorsal seta (or two in Nebria roddi Dudko & Shilenkov, 2001 and Nebria baenningeri korgonica Dudko & Shilenkov, 2001); antennomeres 3 and 4 not laterally compressed, without extra setae. Labrum with three pairs of apical setae. Maxillary stipes typical for genus, with setae inserted flush on smooth surface. Penultimate labial palpomere with three setae. Pronotum moderately cordate in dorsal aspect, with lateral margination shallowly to moderately impressed and markedly narrow at middle; one midlateral seta and one basolateral seta present on each side. Elytral intervals smooth, slightly to moderately flattened, without macrosculpture, elytral interval 3 with one to seven setiferous pores, intervals 5 and 7 with zero to two setiferous pores; umbilicate series comprised of eight to ten setae; intervals bearing setae faintly to moderately catenate. Hindwings reduced to short, slender strap-like vestiges. Metasternum moderately to markedly short. Metepisterna smooth, impunctate. Protarsomeres 1 - 3 expanded in males; mesotarsomeres 2 - 4 longer than their apical width; tarsi dorsally glabrous (or with few sparse and fine setae in Nebria mellyi Gebler, 1847 specimens). Abdominal sternites IV-VI with from two to five or more pairs of posterior paramedial setae and with zero to three pairs of anterior paralateral setae; sternite VII with one or two pairs of paramedial apical setae in males, two to three or more pairs in females. Median lobe of male aedeagus sclerotized dorsally at least to midlength on shaft, symmetrical to slightly deflected right in dorsal aspect; basal bulb expanded, quadrate, broadly open basally and closed dorsally, without a sagittal aileron present at base or with only a lightly sclerotized collar; mid-shaft parallel-sided in lateral aspect, slightly to moderately compressed in cross-section, right lateral face with a shallow longitudinal groove or shallow indentation; apical orifice moderately (in Nebria lyubechanskii Dudko, 2008 specimens only) or extremely deflected right. Right paramere narrow and slightly shortened. Female hemisternites VIII with basal apodeme faintly to moderately emarginated. Female valvifer without vestiture; gonopods VIII fused to dorsomedial bases of gonocoxae; gonocoxae with ventral diagonal row of setiform setae and mediodorsal row of setae present, mediodorsal row oriented parallel to longitudinal axis of gonocoxa. Bursa copulatrix without dorsal sclerites in vestibular chamber and without a posterodorsal extension from its longitudinal axis in lateral aspect; spermathecal chamber broadly cordate in dorsal aspect, without dorsal sclerites (in most species) or with a simple, broad and convoluted plate anterior to spermathecal duct insertion (in Nebria sajana sarlyk Dudko & Shilenkov, 2001, N. s. dubatolovi Dudko & Shilenkov, 2001 and N. s. sitnikovi Dudko & Shilenkov, 2001); spermathecal duct slightly short in length and of uniform diameter throughout or nearly so, inserted basodorsally on spermathecal chamber or on a dorsal accessory lobe of that chamber (in Nebria sajana sajana Dudko & Shilenkov, 2001); spermathecal reservoir of medium length.	en	Kavanaugh, David H., Maddison, David R., Simison, W. Brian, Schoville, Sean D., Schmidt, Joachim, Faille, Arnaud, Moore, Wendy, Pflug, James M., Archambeault, Sophie L., Hoang, Tinya, Chen, Jei-Ying (2021): Phylogeny of the supertribe Nebriitae (Coleoptera, Carabidae) based on analyses of DNA sequence data. ZooKeys 1044: 41-152, DOI: http://dx.doi.org/10.3897/zookeys.1044.62245, URL: http://dx.doi.org/10.3897/zookeys.1044.62245
6CE79A2D0ED25B828439A36BD3B559F6.taxon	etymology	Etymology. The subgeneric epithet is a noun of feminine gender and combination of the Greek word, palaios, meaning old, in reference to the Palearctic (Old World) distribution of the group, aptenos, meaning unable to fly, and the genus name, Nebria, in reference to observation that all known members of this clade have extremely reduced hindwings incapable of supporting flight.	en	Kavanaugh, David H., Maddison, David R., Simison, W. Brian, Schoville, Sean D., Schmidt, Joachim, Faille, Arnaud, Moore, Wendy, Pflug, James M., Archambeault, Sophie L., Hoang, Tinya, Chen, Jei-Ying (2021): Phylogeny of the supertribe Nebriitae (Coleoptera, Carabidae) based on analyses of DNA sequence data. ZooKeys 1044: 41-152, DOI: http://dx.doi.org/10.3897/zookeys.1044.62245, URL: http://dx.doi.org/10.3897/zookeys.1044.62245
E9B783A0DB205E80BD3B2DAA17E02A8C.taxon	diagnosis	Diagnosis. Body size small, SBL = 8.2 to 9.5 mm (1 - 2). Head width medium to slightly broadened, not constricted behind eyes; vertex with a pair of paramedial pale spots and a single pair of supraorbital setae (13). Eyes of moderate size, moderately to markedly convex. Antennal scape with one subapicodorsal seta; antennomeres 3 and 4 not laterally compressed, without extra setae. Labrum with three pairs of apical setae. Maxillary stipes typical for genus, with setae inserted flush on smooth surface. Penultimate labial palpomere with three setae. Pronotum with midlateral setae absent, a single pair of basolateral setae present. Elytral intervals smooth, without macrosculpture, interval 3 with five to eight or more setiferous pores, interval 5 without setae (except one seta present in a few specimens) and interval 7 with zero or two or more setiferous pores, intervals 3, 5 (if setose), and 7 (if setose) moderately catenate. Hindwings reduced to short membranous lobes with only vestigial venation evident. Metepisterna smooth, impunctate. Protarsomeres 1 - 3 expanded in males; mesotarsomeres 2 - 4 longer than their apical width; tarsi dorsally glabrous. Abdominal sternites IV-VI with two to four pairs of posterior paramedial setae, without paralateral setae. Median lobe of male aedeagus sclerotized dorsally at least to midlength on shaft, only faintly deflected right in dorsal aspect; basal bulb expanded, triangular or quadrate, broadly open basally and closed dorsally, without a sagittal aileron present at base or with only a lightly sclerotized collar; mid-shaft parallel-sided in lateral aspect, circular or only slightly compressed in cross-section, with right lateral face unmodified; apical orifice slightly to moderately deflected right. Right paramere slender and moderately long. Female valvifers without vestiture; gonopods VIII fused to dorsomedial bases of gonocoxae; gonocoxae with ventral diagonal row of setiform setae and mediodorsal row of setae present. Bursa copulatrix without dorsal sclerites in vestibular chamber and with longitudinal axis slightly arched basodorsally in lateral aspect; spermathecal chamber broadly cordate in dorsal aspect, and without dorsal or anterodorsal sclerites, insemination duct sclerotized as a long, narrow plate ventrally in the chamber; spermathecal duct slightly long, uniform in diameter throughout or nearly so, inserted sub-basodorsally on spermathecal chamber; spermathecal reservoir of medium length.	en	Kavanaugh, David H., Maddison, David R., Simison, W. Brian, Schoville, Sean D., Schmidt, Joachim, Faille, Arnaud, Moore, Wendy, Pflug, James M., Archambeault, Sophie L., Hoang, Tinya, Chen, Jei-Ying (2021): Phylogeny of the supertribe Nebriitae (Coleoptera, Carabidae) based on analyses of DNA sequence data. ZooKeys 1044: 41-152, DOI: http://dx.doi.org/10.3897/zookeys.1044.62245, URL: http://dx.doi.org/10.3897/zookeys.1044.62245
E9B783A0DB205E80BD3B2DAA17E02A8C.taxon	etymology	Etymology. The subgeneric epithet is a noun of feminine gender and a combination of the Latin word, insula, meaning island, and the genus name, Nebria, in reference to the occurrence of both known species of this clade in Kuril Island Archipelago of far eastern Asia.	en	Kavanaugh, David H., Maddison, David R., Simison, W. Brian, Schoville, Sean D., Schmidt, Joachim, Faille, Arnaud, Moore, Wendy, Pflug, James M., Archambeault, Sophie L., Hoang, Tinya, Chen, Jei-Ying (2021): Phylogeny of the supertribe Nebriitae (Coleoptera, Carabidae) based on analyses of DNA sequence data. ZooKeys 1044: 41-152, DOI: http://dx.doi.org/10.3897/zookeys.1044.62245, URL: http://dx.doi.org/10.3897/zookeys.1044.62245
A3FC909FCAF051619EA8722B9782BB36.taxon	diagnosis	Diagnosis. Body size small or very small, SBL = 8.0 to 9.9 mm. Head moderately wide, slightly constricted behind eyes; vertex with a single central pale spot or a larger pale area and with a single pair of supraorbital setae present. Eyes slightly to moderately reduced in size, moderately to markedly convex. Antennal scape with one or two subapicodorsal setae; antennomeres 3 and 4 not laterally compressed, without extra setae. Labrum with three or four pairs of apical setae. Maxillary stipes typical for genus, with setae inserted flush on smooth surface. Penultimate labial palpomere with three setae. Pronotum with one midlateral seta (a second lateral seta present unilaterally in a few specimens) and one basolateral seta present on each side. Elytral intervals smooth, without macrosculpture, interval 3 with one to seven setiferous pores, intervals 5 and 7 without setiferous pores, intervals 3, 5, and 7 without catenations. Hindwings reduced to short, membranous lobes with only faint vestiges of venation. Metepisterna smooth to sparsely and faintly punctate. Protarsomeres 1 - 3 expanded in males; mesotarsomeres 2 - 4 longer than their apical width; tarsi dorsally glabrous or with only a few fine setae. Abdominal sternites IV to VI without paralateral setae. Median lobe of male aedeagus sclerotized dorsally at least to midlength on shaft; basal bulb only slightly expanded relative to shaft diameter and broadly open basally and with lateral and dorsal basal collar, without or with only slightly developed laterobasal lobes; sagittal aileron present at base as a small midline fin attached to collar; mid-shaft slightly tapered apically and with lateral right face unmodified; apical orifice mid-dorsal. Right paramere slender and moderately long. Female valvifers with vestiture; gonopods VIII fused to dorsomedial bases of gonocoxae; gonocoxae with ventral diagonal row of setiform setae and mediodorsal row of setae present. Bursa copulatrix without or (in one species) with dorsal paramedial sclerotized plates in vestibular chamber; spermathecal chamber broadly wedge-shaped, triangular in dorsal aspect, without a posterodorsal extension from its longitudinal axis in lateral aspect and without dorsal, anterodorsal or ventral sclerites; spermathecal duct of medium length, slender and uniform in diameter throughout or nearly so or (in one species) distinctly and abruptly thicker proximally, inserted basodorsally on spermathecal chamber; spermathecal reservoir moderately long.	en	Kavanaugh, David H., Maddison, David R., Simison, W. Brian, Schoville, Sean D., Schmidt, Joachim, Faille, Arnaud, Moore, Wendy, Pflug, James M., Archambeault, Sophie L., Hoang, Tinya, Chen, Jei-Ying (2021): Phylogeny of the supertribe Nebriitae (Coleoptera, Carabidae) based on analyses of DNA sequence data. ZooKeys 1044: 41-152, DOI: http://dx.doi.org/10.3897/zookeys.1044.62245, URL: http://dx.doi.org/10.3897/zookeys.1044.62245
A3FC909FCAF051619EA8722B9782BB36.taxon	etymology	Etymology. The subgeneric epithet is a noun of feminine gender and a combination of the Greek word, para, meaning near, and the genus-group name, Epinebriola, in reference to the marked similarity of members of this group to those of subgenus Epinebriola Epinebriola.	en	Kavanaugh, David H., Maddison, David R., Simison, W. Brian, Schoville, Sean D., Schmidt, Joachim, Faille, Arnaud, Moore, Wendy, Pflug, James M., Archambeault, Sophie L., Hoang, Tinya, Chen, Jei-Ying (2021): Phylogeny of the supertribe Nebriitae (Coleoptera, Carabidae) based on analyses of DNA sequence data. ZooKeys 1044: 41-152, DOI: http://dx.doi.org/10.3897/zookeys.1044.62245, URL: http://dx.doi.org/10.3897/zookeys.1044.62245
6F8FB7FBDD9C5589AD6BE214F38A097F.taxon	diagnosis	Diagnosis. Body size small to medium, SBL = 9.4 to 12.8 mm. Head width medium or slightly to markedly broadened, not constricted behind eyes; vertex with a pair of paramedial pale spots on vertex and one pair of supraorbital setae. Eyes moderate in size or slightly reduced, moderately convex. Antennae slightly to moderately elongate; antennal scape with one subapicodorsal seta; antennomeres 3 and 4 not laterally compressed, without extra setae. Labrum with three pairs of apical setae. Maxillary stipes typical for genus, with setae inserted flush on smooth surface. Penultimate labial palpomere with three setae (except only two present in Nebria carri Kavanaugh, 1979 specimens). Pronotum markedly to extremely cordate in dorsal aspect, with lateral margination faintly impressed and slightly narrow at middle; one midlateral seta and one basolateral seta present on each side (except basolateral setae absent from N. carri, Nebria balli Kavanaugh, 1979 and Nebria kincaidi Schwarz, 1900 specimens). Elytral intervals smooth, moderately convex, without macrosculpture; elytral interval 3 with two to seven setiferous pores, interval 5 with zero or one setiferous pore, and interval 7 with zero to two or more setiferous pores; umbilicate series comprised of from ten to 17 setae; intervals bearing setae faintly to moderately catenate. Hindwings reduced to short, slender strap-like vestiges. Metasternum very markedly to extremely short. Metepisterna smooth, impunctate. Protarsomeres 1 - 3 expanded in males (except all five tarsomeres expanded in males from some populations of N. ovipennis LeConte, 1878); mesotarsomeres 2 - 4 longer than their apical width (except expanded in specimens from some populations of N. ovipennis); tarsi dorsally glabrous. Abdominal sternites IV-VI with from one (seen only in some specimens of N. balli) to five or more pairs of posterior paramedial setae and with from zero to five pairs of posterior paralateral setae; sternite VII with one pair of paramedial apical setae in males, two pairs in females. Median lobe of male aedeagus sclerotized dorsally at least to midlength on shaft, symmetrical to slightly deflected right in dorsal aspect; basal bulb expanded, quadrate, broadly open basally and closed dorsally, without a sagittal aileron present at base or with only a lightly sclerotized collar; mid-shaft parallel-sided in lateral aspect, slightly to moderately compressed in cross-section, with right lateral face with a distinctly and deeply invaginated pouch; apical orifice extremely deflected right. Right paramere narrow and slightly shortened. Female hemisternites VIII with basal apodeme very deeply emarginated. Female valvifer without vestiture; gonopods VIII fused to dorsomedial bases of gonocoxae; gonocoxae with ventral diagonal row of setiform setae and mediodorsal row of setae present, mediodorsal row oriented obliquely relative to longitudinal axis of gonocoxa. Bursa copulatrix without dorsal sclerites in vestibular chamber and without a posterodorsal extension from its longitudinal axis in lateral aspect; spermathecal chamber broadly cordate in dorsal aspect, without dorsal sclerites (in most species) or with a simple flat plate on dorsal surface of an accessory lobe (in N. kincaidi and N. balli); spermathecal duct medium to slightly short in length and of uniform diameter throughout or nearly so, inserted basodorsally on spermathecal chamber or on a dorsal accessory lobe of that chamber; spermathecal duct slightly short to medium length; spermathecal reservoir of medium length.	en	Kavanaugh, David H., Maddison, David R., Simison, W. Brian, Schoville, Sean D., Schmidt, Joachim, Faille, Arnaud, Moore, Wendy, Pflug, James M., Archambeault, Sophie L., Hoang, Tinya, Chen, Jei-Ying (2021): Phylogeny of the supertribe Nebriitae (Coleoptera, Carabidae) based on analyses of DNA sequence data. ZooKeys 1044: 41-152, DOI: http://dx.doi.org/10.3897/zookeys.1044.62245, URL: http://dx.doi.org/10.3897/zookeys.1044.62245
6F8FB7FBDD9C5589AD6BE214F38A097F.taxon	etymology	Etymology. The subgeneric epithet is a noun of feminine gender and combination of the Greek word, neos, meaning new, in reference to the Nearctic (New World) distribution of the group, the Greek word, aptenos, meaning unable to fly, and the genus name, Nebria, in reference to the observation that all known members of this clade have extremely reduced hindwings incapable of supporting flight.	en	Kavanaugh, David H., Maddison, David R., Simison, W. Brian, Schoville, Sean D., Schmidt, Joachim, Faille, Arnaud, Moore, Wendy, Pflug, James M., Archambeault, Sophie L., Hoang, Tinya, Chen, Jei-Ying (2021): Phylogeny of the supertribe Nebriitae (Coleoptera, Carabidae) based on analyses of DNA sequence data. ZooKeys 1044: 41-152, DOI: http://dx.doi.org/10.3897/zookeys.1044.62245, URL: http://dx.doi.org/10.3897/zookeys.1044.62245
6226343FCCF3514DB574B8B0341021C8.taxon	diagnosis	Diagnosis. Body size small to medium, SBL = 7.1 to 11.7 mm. Head width medium to slightly broadened, not constricted behind eyes; vertex with a pair of paramedial pale spots and a single pair of supraorbital setae. Eyes slightly reduced to moderate in size, moderately to markedly convex. Antennal scape with one (two in very few specimens) subapicodorsal seta; antennomeres 3 and 4 not laterally compressed, without extra setae. Labrum with three pairs of apical setae. Maxillary stipes typical for genus, with setae inserted flush on smooth surface. Penultimate labial palpomere with three setae. Pronotum with midlateral setae absent, a single pair of basolateral setae present. Elytral intervals smooth, without macrosculpture, interval 3 with three to eight or more setiferous pores, intervals 5 and 7 without setiferous pores, interval 3 not or only faintly catenate, intervals 5 and 7 not catenate. Hindwings full-sized or slightly to markedly reduced in size. Metepisterna smooth or faintly punctulate. Protarsomeres 1 - 3 expanded in males; mesotarsomeres 2 - 4 longer than their apical width; tarsi dorsally glabrous or with a few fine setae. Abdominal sternites IV-VI with two to four pairs of posterior paramedial setae, without paralateral setae. Median lobe of male aedeagus sclerotized dorsally at least to midlength on shaft, moderately to markedly deflected right in dorsal aspect; basal bulb expanded, quadrate, broadly open basally and closed dorsally, without a sagittal aileron present at base or with only a lightly sclerotized collar; mid-shaft parallel-sided or slightly to moderately narrowed basally in lateral aspect, slightly compressed in cross-section, with right lateral face unmodified; apical orifice markedly to extremely deflected right. Right paramere slender and moderately long. Female valvifers with vestiture (a few specimens of some species without vestiture); gonopods VIII fused to dorsomedial bases of gonocoxae; gonocoxae with ventral diagonal row of setiform setae and mediodorsal row of setae present. Bursa copulatrix without dorsal sclerites in vestibular chamber and with longitudinal axis of the bursa moderately to markedly sigmoid dorsally in lateral aspect; spermathecal chamber broadly cordate in dorsal aspect, without (in Nebria lyelli Van Dyke, 1925 and Nebria quileute Kavanaugh, 1979) or with a small to large midline sclerotized dorsal plate, insemination duct not sclerotized; spermathecal duct medium-length to slightly long, uniform in diameter throughout or nearly so, inserted basodorsally on spermathecal chamber; spermathecal reservoir of medium length.	en	Kavanaugh, David H., Maddison, David R., Simison, W. Brian, Schoville, Sean D., Schmidt, Joachim, Faille, Arnaud, Moore, Wendy, Pflug, James M., Archambeault, Sophie L., Hoang, Tinya, Chen, Jei-Ying (2021): Phylogeny of the supertribe Nebriitae (Coleoptera, Carabidae) based on analyses of DNA sequence data. ZooKeys 1044: 41-152, DOI: http://dx.doi.org/10.3897/zookeys.1044.62245, URL: http://dx.doi.org/10.3897/zookeys.1044.62245
6226343FCCF3514DB574B8B0341021C8.taxon	etymology	Etymology. The subgeneric epithet is a noun of feminine gender and a combination of the surname of Terry L. Erwin, in whose honor we name this subgenus, and the genus name, Nebria.	en	Kavanaugh, David H., Maddison, David R., Simison, W. Brian, Schoville, Sean D., Schmidt, Joachim, Faille, Arnaud, Moore, Wendy, Pflug, James M., Archambeault, Sophie L., Hoang, Tinya, Chen, Jei-Ying (2021): Phylogeny of the supertribe Nebriitae (Coleoptera, Carabidae) based on analyses of DNA sequence data. ZooKeys 1044: 41-152, DOI: http://dx.doi.org/10.3897/zookeys.1044.62245, URL: http://dx.doi.org/10.3897/zookeys.1044.62245
E1AC85327E405F19924E8D07745F9C82.taxon	diagnosis	Diagnosis. Body size small to medium, SBL = 8.7 to 11.3 mm. Head markedly broadened, not constricted behind eyes; vertex with a single medial pale area and a single pair of supraorbital setae. Eyes moderate in size, moderately convex. Antennal scape with one subapicodorsal seta; antennomeres 3 and 4 not laterally compressed, without extra setae. Labrum with three pairs of apical setae. Maxillary stipes typical for genus, with setae inserted flush on smooth surface. Penultimate labial palpomere with three setae. Pronotum with one midlateral and one basolateral seta present on each side. Elytral intervals smooth, without macrosculpture, interval 3 with one to seven setiferous pores, interval 5 with zero to two and interval 7 with zero to two or more setiferous punctures, intervals with setae only faintly catenate; elytral medial margins diverge slightly and arcuate from the midline apically. Hindwings reduced to short, slender strap-like vestiges. Metepisterna smooth, impunctate. Protarsomeres 1 - 3 expanded in males; mesotarsomeres 2 - 4 longer than their apical width; tarsi dorsally glabrous. Abdominal sternites IV-VI with one to four pairs of posterior paramedial setae, without paralateral setae. Median lobe of male aedeagus sclerotized dorsally at least to midlength on shaft, symmetrical in dorsal aspect; basal bulb expanded, quadrate, broadly open basally and closed dorsally, without a sagittal aileron present at base or with only a lightly sclerotized collar; mid-shaft parallel-sided in lateral aspect, slightly compressed in cross-section, with right lateral face unmodified; apical orifice markedly deflected right. Right paramere slender and very long. Female valvifers without vestiture; gonopods VIII fused to dorsomedial bases of gonocoxae; gonocoxae with ventral diagonal row of setiform setae and mediodorsal row of setae present. Bursa copulatrix with a pair of large dorsal paramedial sclerotized plates in vestibular chamber, with its longitudinal axis moderately sigmoid dorsally in lateral aspect; spermathecal chamber broadly cordate in dorsal aspect, without dorsal sclerites (in some N. paradisi) or with a small midline plate (other N. paradisi) or large and broad dorsal plate (in N. turmaduodecima); spermathecal duct very long and of uniform diameter throughout or nearly so, inserted basodorsally on spermathecal chamber; spermathecal reservoir of medium length.	en	Kavanaugh, David H., Maddison, David R., Simison, W. Brian, Schoville, Sean D., Schmidt, Joachim, Faille, Arnaud, Moore, Wendy, Pflug, James M., Archambeault, Sophie L., Hoang, Tinya, Chen, Jei-Ying (2021): Phylogeny of the supertribe Nebriitae (Coleoptera, Carabidae) based on analyses of DNA sequence data. ZooKeys 1044: 41-152, DOI: http://dx.doi.org/10.3897/zookeys.1044.62245, URL: http://dx.doi.org/10.3897/zookeys.1044.62245
E1AC85327E405F19924E8D07745F9C82.taxon	etymology	Etymology. The subgeneric epithet is a noun of feminine gender and a combination of the Latin word, Nebria nivalis, meaning snow, and the genus name, Nebria, in reference to the occurrence of all known members of this clade in the vicinity of montane snowfields and glaciers.	en	Kavanaugh, David H., Maddison, David R., Simison, W. Brian, Schoville, Sean D., Schmidt, Joachim, Faille, Arnaud, Moore, Wendy, Pflug, James M., Archambeault, Sophie L., Hoang, Tinya, Chen, Jei-Ying (2021): Phylogeny of the supertribe Nebriitae (Coleoptera, Carabidae) based on analyses of DNA sequence data. ZooKeys 1044: 41-152, DOI: http://dx.doi.org/10.3897/zookeys.1044.62245, URL: http://dx.doi.org/10.3897/zookeys.1044.62245
