taxonID	type	description	language	source
AD307FF061155B70B16ABF354B9C696C.taxon	description	Description of the HT ♂. TL mm 2.66. Body elongated, depigmented, yellow-testaceous; integument shiny, with evident microsculpture, covered with very short pubescence. Head large, slightly wider than the base of the pronotum. Labium without tooth. Antennae short, just exceeding the base of the pronotum when stretched backwards. Fronto-clypeal furrow of frontal clypeo slightly distinct; anterior margin of the epistome subrectilinear. Pronotum elongated (max. width / max. length ratio = 0.90), maximum width at the middle, basal border slightly narrower than the anterior border, pronotum sides poorly arcuate, distinctly crenelated at the posterior fourth, emarginated before the base. Anterior angles rounded, not prominent; posterior angles sharp, squared, not protruding. Disc slightly convex, with very sparse and short pubescence; median groove very shallow, slightly marked. Marginal groove wide and flat, enlarged near the base; anterior marginal setae inserted inside the marginal groove, approximately on the anterior fifth; basal setae inserted before the posterior angles. Legs long and slender, with metatrochanters normal and metafemora (Fig. 50) armed with a long, internal spur; two protarsomeres dilated and without adhesive phanerae in male. Elytra subrectangular, very elongated (max. length / max. width ratio = 2.28), with parallel sides, slightly emarginated before apex. Disc convex; integument shiny, with evident microsculpture and short, longitudinally aligned and upright pubescence. Humeri very marked, squared; post-humeral margin denticulate, with a distinct crenulation up to the 8 th pore of the umbilicate series; elytral apices separately rounded. Marginal groove narrow and evident up to the 8 th pore of the umbilicate series. Chaetotaxy: scutellar pore large, foveate. Umbilicate series with 1 st, 2 nd, and 3 rd pores of the humeral group almost equidistant; 4 th pore clearly farther from the 3 rd one and placed at the end of the basal third of the elytron; 5 th pore placed at the end of median third of the elytron; 5 th and 6 th pores spaced out ca. 1 / 4 of the distance from 6 th and 7 th pores; 7 th, 8 th, and 9 th pores not displaced onto the disc and not equidistant, 8 th pore placed after the 9 th one. One single discal seta in the central area of the disc and placed approximately at the level of the 9 th pore. Aedeagus (Fig. 51) relatively large, median lobe, in lateral view, long and restricted at apex, regularly curved and not restricted before the basal bulb, basal bulb of normal size. Ventral margin curved from basal bulb to apex, gently emarginated just before the apex; apical blade evident, but short. Endophallus without sclerified phanerae. Parameres relatively long and robust, bearing two setae: one apical and one subapical; right paramere shorter than the left one.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
AD307FF061155B70B16ABF354B9C696C.taxon	etymology	Etymology. The name comes from the Latin word armato (which means armed) to highlight the presence of a large spur on metafemora.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
AD307FF061155B70B16ABF354B9C696C.taxon	distribution	Distribution. Angustanillus armatus sp. nov. is known only from the type locality Mesa B, 38 km W of Pannawonica, Pilbara, WA.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
168EFD30E44959B98C041473AD35E32B.taxon	diagnosis	Diagnosis. Included species strongly characterised by: body very elongated, posterior supraorbital seta and longitudinal elytral grooves absent, pronotum with basal border narrower than anterior border, and sides posteriorly distinctly crenelated, basal seta of pronotum present, one or two elytral discal seta present, 8 th pore of the umbilicate series located after the 9 th pore (sensu Giachino and Vailati 2011), metafemora dentate in males (state unknown in females), labial tooth absent, median lobe of the aedeagus long, slender and curved, parameres long and slender.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
E6438A92A0B35CFEB0A46237EE488E15.taxon	diagnosis	Diagnosis. Genus of the " Illaphanus phyletic series " (sensu Giachino 2005), with species characterised by: absence of longitudinal elytral grooves, pronotum with basal border wider than anterior border and denticulate sides before basal angles, elytral disc with two setae, 8 th pore of the umbilicate series after the 9 th (sensu Giachino and Vailati 2011), labium toothless, median lobe of the aedeagus long and weakly curved, and parameres very long and slender.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
6DE489455B5D5E45B776701B8D02FDC8.taxon	description	Description of the HT ♂. TL mm 2.98. Body elongated, depigmented, fulvo-testaceous; integument shiny, with evident microsculpture and sparse and short pubescence. Head very small, much narrower than pronotum. Labium toothless. Antennae long and robust, not moniliform (with relatively elongated antennomeres), considerably exceeding the base of the pronotum when stretched backwards. Fronto-clypeal furrow indistinct; anterior margin of the epistome subrectilinear. Pronotum slightly transverse (max. width / max. length ratio = 1.13) with maximum width at the anterior third, and a very wide basal border, wider than the anterior border. Pronotum sides anteriorly clearly arcuate, weakly sinuate and denticulate before the base, base laterally distinctly emarginated. Anterior angles rounded, not prominent. Posterior angles protruding, acute at the vertex. Disc slightly convex, with a short and relatively dense pubescence; median groove very shallow, slightly marked. Marginal groove relatively wide and flat, very enlarged near the base; anterior marginal setae inserted inside the marginal groove, on approximately the anterior third; basal setae inserted almost on the posterior angles. Legs long and slender, with metatrochanters normal and metafemora unarmed. Two asymmetrically dilated protarsomeres, with one row of adhesive phanerae in male. Elytra subrectangular, very elongated (max. length / max. width ratio = 1.77), with maximum width in the middle, very slightly emarginated before apex; sides slightly sinuate at the base of the anterior fourth. Disc poorly convex; integument shiny, with evident microsculpture and short, dense, upright pubescence. Humeri extremely marked, almost right angle; post-humeral margin denticulate, with a distinct crenulation up to the base of the apical third of the elytron; elytral apices separately rounded. Marginal groove wide and evident up to over the 9 th pore of the umbilicate series. Chaetotaxy: scutellar pore large, foveate. Umbilicate series with pores of the humeral group not equidistant; 4 th pore slightly displaced towards the disc, not so far from the third pore and inserted just on the basal fourth of the elytron; 5 th pore placed just before the apical third of the elytron; 5 th and 6 th pores spaced out, ca. 2 / 3 of the 6 th and 7 th; 7 th, 8 th and 9 th pores almost equidistant and slightly displaced onto the disc, 8 th pore placed after the 9 th one. Two discal setae, first placed at the base of the apical third, second one placed at the level of the 9 th pore. Aedeagus (Fig. 37) large, median lobe long, slender, subrectilinear, with basal bulb poorly evident; ventral margin poorly and gently curved from the basal bulb to the apex; apical blade evident, but short and emarginated in the lower edge. Endophallus with a median, C-shaped, sclerified phanera. Parameres slender and very elongated, reaching the distal seventh of the median lobe, and bearing three setae only; right paramere slightly shorter than the left.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
6DE489455B5D5E45B776701B8D02FDC8.taxon	etymology	Etymology. The name of the species comes from the type locality of the Jinayri deposit.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
6DE489455B5D5E45B776701B8D02FDC8.taxon	distribution	Distribution. Austranillus jinayrianus sp. nov. is known only from the type locality, Jinayri deposit, in the Pilbara region, 110 km NW of Newman, WA.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
762D38FC597D56FAB9BA6B714E234FD1.taxon	description	Description of the HT ♀. TL mm 1.43. Body elongated, depigmented, yellow-testaceous; integument shiny, with evident microsculpture, covered with short pubescence. Head large, slightly narrower than the base of the pronotum. Labium without tooth. Antennae short, just exceeding the base of the pronotum when stretched backwards. Fronto-clypeal furrow slightly distinct; subrectilinear anterior margin of epistome. Pronotum subrectangular (max. width / max. length ratio = 1.06) maximum width at the base of the anterior fourth, with basal border narrower than the anterior border, pronotum sides, anteriorly poorly arcuate, posteriorly subrectilinear and slightly sinuated, laterally completely smooth from anterior seta to basal angles, not emarginated before the base. Anterior angles rounded, slightly prominent; posterior angles obtuse, rounded, not protruding. Disc slightly convex, with very sparse and relatively short pubescence; median groove very shallow, slightly marked. Marginal groove relatively narrow and flat, not particularly enlarged near the base; anterior marginal setae inserted inside the marginal groove, approximately on the anterior fifth; basal setae inserted approximately on posterior angles. Legs long and slender, with metatrochanters normal and metafemora unarmed. Elytra subrectangular, very elongated (max. length / max. width ratio = 1.98), with parallel sides, slightly emarginated before apex. Disc convex; integument shiny, with evident microsculpture and very short, very dense and upright pubescence. Humeri very marked, squared; post-humeral margin denticulate, with distinct crenulations up to the discal pore; elytral apices separately rounded. Marginal groove narrow and evident up to the discal pore. Chaetotaxy: scutellar pore large, foveate. Umbilicate series with 1 st, 2 nd, and 3 rd pores of the humeral group almost equidistant; 4 th pore clearly farther from the 3 rd one and placed at the end of the basal third of the elytron; 5 th pore placed well after the middle length of the elytron; 5 th and 6 th pores spaced out ca. 1 / 3 of the distance between 6 th and 7 th pores; 7 th, 8 th, and 9 th pores not equidistant, 8 th placed after the 9 th one; 7 th slightly displaced onto the disc. One single discal seta in the central area of the disc and placed at ca. 2 / 3 of the distance between the 6 th and 7 th pores. Male. Unknown.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
762D38FC597D56FAB9BA6B714E234FD1.taxon	etymology	Etymology. The name comes from the type locality: Cundaline Ridge.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
762D38FC597D56FAB9BA6B714E234FD1.taxon	distribution	Distribution. Bylibaraphanus cundalinianus sp. nov. is known only from the type locality Cundaline Ridge, in the Yarrie Mining Area, 200 km E of Port Hedland, Pilbara, WA.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
1E96F6076F5256B09F5F92FD1484D4E2.taxon	materials_examined	Material examined. 2 ♂♂ 3 ♀♀, WA, Pilbara, 65 km NW of Tom Price, Cloudbreak Mine, 22 ° 20 ' 26.40 " S, 119 ° 25 ' 50.70 " E (WGS 84), J. Cocking, M. Scanlon, 15 Mar. 2011, Trog. Net scrape (GNGC 05589), Western Australian Museum Entomology Reg. no. 8269 - 82673 (WAM, CGi).	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
1E96F6076F5256B09F5F92FD1484D4E2.taxon	distribution	Distribution. B. currani is known only from Cloudbreak Mine, 65 km NW of Tom Price, Pilbara, WA.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
A7EA4BBFAA5A5FCC835B872CB6F7A3D2.taxon	diagnosis	Diagnosis. Included species strongly characterised by: longitudinal elytral grooves absent, pronotum with basal border as wide as or narrower than anterior border, and sides not or only slightly posteriorly sinuate, basal seta of pronotum present, one elytral discal seta present, 8 th pore of the umbilicate series placed after the 9 th pore (sensu Giachino and Vailati 2011), metafemora not dentate, labial tooth absent, median lobe of aedeagus long and curved, parameres long and slender. Bylibaraphanus differs from Magnanillus Baher, 2017 by elytra with a single discal seta (3 in Magnanillus) and smaller body size. It differs from Hesperanillus Baher & Main, 2016 by head narrower than pronotum, and pronotum less cordiform. It differs from Gracilanillus Baher & Main, 2016 by the absence of longitudinal elytral grooves.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
A7EA4BBFAA5A5FCC835B872CB6F7A3D2.taxon	description	Description. Genus with small size species (TL mm 1.43 - 1.50), anophthalmous. Depigmented integument, medium sclerified, with strong microsculpture and dense pubescence. Head wider than pronotum base; mandibles short and simple, without hyperplasias. Maxillary palps ovoidal, swollen. Labium transverse, articulated; mentum not fused with submentum. Labial tooth absent. Antennae moniliform. Pronotum squared, pronotum sides smooth, not or only slightly sinuate at the basal third. Basal angles acute or obtuse, sharp or rounded; basal border as wide as, or narrower, than anterior border; basal seta present. Elytra convex, subrectangular, elongated, separately rounded, not truncate, apically slightly emarginated, and without longitudinal grooves. Elytral striae missing (except for the sutural stria). Lateral margin, starting from the humeral area, hardly serrulate up to the level of the 7 th or 9 th pore of the umbilicate series. Scutellar pore present, large and umbilicate; umbilicate series of type B (sensu Jeannel 1963; Giachino and Vailati 2011) with the 8 th pore placed after the 9 th one; disc bearing one seta (posterior) in the central area of the disc. Legs relatively long and slender. Pro- and metafemora unarmed; metatrochanters normal; two protarsomeres slightly dilated and without adhesive phanerae in males. Aedeagus relatively large; median lobe relatively long, curved, not restricted before the basal bulb, basal bulb of normal size. Parameres long, slender, bearing two or three apical setae. Endophallus with sclerified phanera.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
A7EA4BBFAA5A5FCC835B872CB6F7A3D2.taxon	etymology	Etymology. Combined name formed by Bylibara (aboriginal noun for the Pilbara region) and the suffix - phanus taken from Illaphanus genus. The gender of the name is masculine.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
AEBF67BA5DD650E0B257D501BA4B4B1C.taxon	diagnosis	Diagnosis. Small species, with longitudinal elytral grooves; male protarsi with two dilated tarsomeres and profemora unarmed; metafemora dentate and metatrochanters short and stout in male and female. Head characterised by temples with a series of excess long setae. One elytral discal seta present. Aedeagus with right paramere bearing two apical setae; left bearing only one seta.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
AEBF67BA5DD650E0B257D501BA4B4B1C.taxon	description	Description. TL 1.40 - 1.42 mm. Body elongated, depigmented, fulvo-testaceous with elytra and abdomen lighter, yellow-testaceous; integument shiny with evident microsculpture and spread pubescence. Head robust, almost hypertrophic, slightly narrower than the pronotum. Labium toothless, mentum articulate. Antennae robust, moniliform, short, not exceeding the base of the pronotum when stretched backwards. Fronto-clypeal furrow indistinct; anterior margin of the epistome subrectilinear. Pronotum subquadrate (max. width / max. length ratio = 1.05), with the maximum width on the anterior fourth, narrowed at the base, with the sides poorly arcuate anteriorly, sinuate before the base. Anterior angles rounded, weakly prominent; posterior angles subsquare and acuminate at tips. Disc convex, with a long and sparse pubescence; median groove very shallow, hardly evident. Marginal groove wide and flat, slightly enlarged near the base; anterior marginal setae placed inside the marginal groove, almost on the anterior fifth; basal setae before the posterior angles. Legs short and stout, two protarsomeres dilated and without adhesive phanerae in male. Metatrochanters short and stout, metafemora dentate (Fig. 2). Elytra oval, slightly elongated (max. length / max. width ratio = 1.72), with maximum width in the middle, not emarginated before apex. Disc poorly convex, provided with an evident longitudinal groove running more or less between the scutellar pore and the 9 th pore of the umbilicate series; integument shiny, with evident microsculpture and short, dense, upright pubescence. Humeri poorly marked, rounded. Post-humeral margin denticulate, with a very thin but distinct crenulation to the apical third; elytral apices separately rounded. Marginal groove wide and evident almost to the 8 th pore of the umbilicate series. Chaetotaxy: scutellar pore large, foveate. Umbilicate series with the first three pores of the humeral group not equidistant, with 1 st and 2 nd closer together than 2 nd and 3 rd; 4 th pore clearly farther and placed at the end of the basal third of the elytron; 5 th pore placed before the apical third of the elytra; 5 th and 6 th ones spaced out ca. half distance between the 6 th and 7 th; 7 th slightly and 8 th nearly moved onto the disc; 7 th and 8 th slightly closer to each other than the 8 th and 9 th. One discal seta, placed before the 7 th pore of the umbilicate series. Aedeagus (Fig. 3) small, abruptly arcuate in the basal part; median lobe moderately elongated with ventral margin gently arcuate up to the acuminate apex, with apical blade very evident. Endophallus without an evident lamella copulatrix. Parameres stocky in the basal part and relatively poorly elongated, not reaching the distal third. Right paramere bearing two apical setae; left bearing only one seta.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
AEBF67BA5DD650E0B257D501BA4B4B1C.taxon	etymology	Etymology. Dedicated to the memory of Martin Baehr, renowned beetle taxonomist, in honour of his contributions to the knowledge of Australian ground beetles.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
AEBF67BA5DD650E0B257D501BA4B4B1C.taxon	distribution	Distribution. Erwinanillus baehri sp. nov. is known only from bore WWMB 4, 80 km E of Hyden, Forrestania, Southern Goldfields region, WA.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
0F079F93A8BA5D169D032A412253679E.taxon	diagnosis	Diagnosis. Included species strongly characterised by the presence of longitudinal elytral grooves; male protarsi with two dilated tarsomeres and profemora unarmed; metafemora dentate and metatrochanters short and stout in male and female. Labial tooth absent. Head characterised by temples with a series of long excess setae. One elytral discal seta present. Aedeagus with right paramere bearing two apical setae; left bearing only one seta. This genus can be distinguished from Externanilus Baehr & Main, 2016 by the presence of grooves on elytral disc.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
0F079F93A8BA5D169D032A412253679E.taxon	description	Description. Included species of small-medium size (TL mm 1.40 - 1.42), anophthalmous. Depigmented and poorly sclerified integument covered with a sparse pubescence. Head robust, almost hypertrophic, slightly narrower than the pronotum, with a series of excess long setae; labium toothless, mentum articulated, not fused with the submentum. Antennae moniliform, without particular features. Pronotum subquadrate, with sides distinctly sinuate in the basal third. Basal angles subsquare and acuminate; basal border as wide as the anterior border; presence of two marginal setae, the posterior one placed before the basal angles. Elytra oval and slightly elongated, separately rounded, not truncate and not emarginated apically; poorly convex and with a longitudinal groove. Elytral striae missing (except for the sutural stria). Lateral margin, distinctly crenelated from the humeral area to the apical third. Scutellar pore present, large and umbilicate; umbilicate series of type B (sensu Jeannel 1963; Giachino and Vailati 2011); disc bearing one discal seta. Legs relatively short and stumpy. Profemora unarmed, metafemora dentate, metatrochanters normal, two dilated protarsomeres in the male. Aedeagus small, median lobe moderately elongated, apex acuminate with apical blade very evident. Parameres stocky, bearing one apical seta (left) or two apical setae (right). Endophallus without sclerified phanerae.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
0F079F93A8BA5D169D032A412253679E.taxon	etymology	Etymology. Dedicated to the memory of Terry Lee Erwin, renowned carabid beetle specialist and explorer of world biodiversity. Name composed of Erwin and the genus name Anillus. The gender of the name is masculine.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
29EADDB5148656E2B03F27A50A3C8BEC.taxon	description	Description of the HT ♂. Estimated TL mm 1.30. Elytra (Fig. 41) subrectangular (total length = mm 0.77), very elongated (max. length / max. width ratio = 1.94), maximum width near the middle, slightly emarginated before apex. Disc poorly convex; shiny, integument testaceous, with evident microsculpture and very short, sparse and upright pubescence. Humeri hardly marked, almost right; post-humeral margin denticulate, with distinct crenulation up to the 8 th, 9 th pores of the umbilicate series; elytral apices not separately rounded. Marginal groove wide and evident up to the 7 th pore of the umbilicate series. Chaetotaxy: scutellar pore large, foveate. Umbilicate series with the 1 st, 2 nd and 3 rd pores of the humeral group almost equidistant; 4 th pore clearly farther from the 3 rd, placed after the end of the basal third of the elytron; 5 th pore placed just after the middle length of the elytron; 5 th and 6 th pores spaced out ca. the 1 / 3 of the distance from 6 th and 7 th ones; 7 th, 8 th and 9 th pores not equidistant, with 8 th and 9 th pores closest. Due to setae missing it is impossible to determine if the 8 th pore is placed after the 9 th one; 7 th pore displaced onto the disc. One single discal seta not laterally located towards the edge and placed at the level of the 7 th pore. Metatrochanters unarmed in the male. Aedeagus (Fig. 42) relatively small, median lobe long, slender, subrectilinear, with basal bulb well-shaped; ventral margin poorly and gently curved from basal bulb to apex; apical blade short and poorly evident. Endophallus with a subapical, bisinuate, sclerified phanera. Parameres missing.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
29EADDB5148656E2B03F27A50A3C8BEC.taxon	etymology	Etymology. The name comes from the Ethel Gorge type locality.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
29EADDB5148656E2B03F27A50A3C8BEC.taxon	distribution	Distribution. Gilesdytes ethelianus sp. nov. is known only from the type locality (Ethel Gorge), 11 km ENE of Newman, Pilbara, WA.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
7A915CA1F70D523D93F365AE9169B42C.taxon	diagnosis	Diagnosis. Species of this genus are strongly characterised by: lacking longitudinal elytral grooves, pronotum with basal border as wide as the anterior border, sides posteriorly not sinuate, and denticulate before, or near, the basal angles, elytral disc bearing one (posterior) seta placed at the level, or after, the 7 th umbilicate pore, 8 th pore of the umbilicate series located after the 9 th (sensu Giachino and Vailati 2011), toothed labium, median lobe of the aedeagus long and weakly curved with parameres very long and slender. Differs from Gracilanillus Baehr & Main, 2016 by bearing a single discal seta on the elytral disc (2 in Gracilanillus), and for the absence of longitudinal elytral grooves.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
7A915CA1F70D523D93F365AE9169B42C.taxon	description	Description. Included medium size species (TL mm 1.29 - 2.11), and anophthalmous. Integument depigmented but well sclerified, with strong microsculpture and covered by short and moderately dense pubescence. Head large, slightly narrower than pronotum; mandibles short and simple, without hyperplasias. Maxillary palps ovoidal, swollen. Labium transverse, articulated; mentum not fused with submentum. Labial tooth present. Antennae moniliform, without particular features. Pronotum squared, sides not sinuate on the basal third, denticulated on the basal third or just before the posterior angles. Basal angles right, sharp, not rounded; basal border as wide as the anterior border; two marginal setae, posterior seta placed near the basal angles. Elytra elongated and subrectangular, separately rounded, not truncate and not apically emarginated; convex, without longitudinal grooves. Elytral striae absent (except for sutural stria). Lateral margin starting from the humeral area, distinctly serrulate up to the level of 8 th- 9 th pores of the umbilicate series. Scutellar pore present, large and umbilicate; umbilicate series of type B (sensu Jeannel 1963; Giachino and Vailati 2011) with the 8 th pore placed after the 9 th; disc bearing one seta. Legs relatively long and slender. Pro- and metafemora unarmed; metatrochanters normal, two slightly dilated protarsomeres, without adhesive phanerae, in the male. Aedeagus relatively small, median lobe long, subrectilinear with basal bulb of normal size. Parameres long, bearing two apical setae. Endophallus with a sinuate, slightly sclerified, apical phanera.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
7A915CA1F70D523D93F365AE9169B42C.taxon	etymology	Etymology. Gilesdytes: name composed by two sections, the first one (Giles) dedicated to William Ernest Powell Giles (20 July 1835 - 13 November 1897), Australian explorer who in 1876 named the Ophthalmia Range (type locality of the genus); and the second one (dytes) for diver.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
58350F0DFB395420A21FA7B88BC2A649.taxon	description	Description of the HT ♀. TL 2.11 mm. Body elongated, depigmented, testaceous; integument shiny, with evident microsculpture, covered with very sparse and short pubescence. Head relatively large, narrower than pronotum. Labium bearing a median tooth. Antennae short and delicate, moniliform, just exceeding the base of the pronotum when stretched backwards. Fronto-clypeal furrow slightly distinct; anterior margin of the epistome subrectilinear. Pronotum subrectangular, slightly transverse (max. width / max. length ratio = 1.05), maximum width at the anterior third, with very wide basal border, slightly wider than anterior border, pronotum sides anteriorly poorly arcuate, and laterally-posteriorly denticulate before the basal angles. Anterior angles obtuse, slightly prominent; posterior angles sub squared not protruding, gently rounded at the vertex. Disc slightly convex, with very sparse and short pubescence; median groove very shallow, faintly marked. Marginal groove relatively wide and flat, very enlarged near the base; anterior marginal setae inserted inside marginal groove, approximately on the anterior fifth; basal setae inserted approximately on the posterior angles. Legs long and slender, with metatrochanters normal and metafemora unarmed. Elytra subrectangular, very elongated (max. length / max. width ratio = 1.92), maximum width at the anterior 3 rd, slightly emarginated before apex. Disc poorly convex; integument shiny, with evident microsculpture and very short, sparse and upright pubescence. Humeri hardly marked, almost right; post-humeral margin denticulate, with distinct crenulations up to the 9 th pore of the umbilicate series; elytral apices not separately rounded. Marginal groove wide and evident up to the 8 th pore of the umbilicate series. Chaetotaxy: scutellar pore large, foveate. Umbilicate series with 1 st, 2 nd, and 3 rd pores of the humeral group almost equidistant; 4 th pore slightly displaced onto the disc and clearly farther from the 3 rd pore, placed at the end of the basal third of the elytron; 5 th pore placed just after the middle length of the elytron; 5 th and 6 th ones spaced out ca. the 1 / 3 of the distance from 6 th and 7 th; 7 th, 8 th, and 9 th not equidistant, 8 th and 9 th pores closest, 8 th pore located after the 9 th one; 8 th slightly displaced onto the disc. One single discal seta laterally placed towards the edge, at the level of the 9 th pore. Male. Unknown.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
58350F0DFB395420A21FA7B88BC2A649.taxon	etymology	Etymology. The name of the species comes from the type locality Pardoo, in the Pilbara region (WA).	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
58350F0DFB395420A21FA7B88BC2A649.taxon	distribution	Distribution. Gilesdytes pardooanus sp. nov. is known only from the type locality (Pardoo Mine), 100 km E of Port Hedland, Pilbara, WA.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
5880E919986B510B922C5271E3CBE388.taxon	materials_examined	Material examined. 1 ♂, WA, Pilbara, 90 km NW of Newman, near Yandi mining area, Ministers North deposit, 22 ° 49 ' 26.54 " S, 119 ° 05 ' 23.89 " E (WGS 84), 20 Feb. 2009, P. Bell, G. Perina, Trap, (BHP 010 - EXR 1154 - MNL: 7056) Western Australian Museum Entomology Reg. no. 82619 (WAM).	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
5880E919986B510B922C5271E3CBE388.taxon	description	Description of the ♂. TL 1.29 mm. Body elongated, depigmented, yellow-testaceous; integument shiny, with evident microsculpture, and covered with sparse and relatively long pubescence. Head relatively large, narrower than the pronotum. Labium bearing a median tooth. Antennae short, robust, moniliform, just exceeding the base of the pronotum when stretched backwards. Fronto-clypeal furrow slightly distinct; subrectilinear anterior margin of epistome. Pronotum subrectangular, slightly transverse (max. width / max. length ratio = 1.04), maximum width at the anterior third, very wide basal border, slightly wider than anterior border. Pronotum sides anteriorly poorly arcuate, posteriorly subrectilinear, gently convergent, and laterally smooth, non-denticulate, but distinctly emarginated before the base. Anterior angles rounded, not prominent; posterior angles not protruding externally, rounded at the vertex. Disc slightly convex, with sparse and relatively long pubescence; median groove very shallow, slightly marked. Marginal groove relatively wide and flat, much enlarged near the base; anterior marginal setae inserted inside the marginal groove, approximately on the anterior third; basal setae inserted approximately on the posterior angles. Legs long and slender, with metatrochanters normal and metafemora unarmed. Males with two dilated protarsomeres without adhesive phanerae. Elytra subrectangular, very elongated (max. length / max. width ratio = 1.94), maximum width at the anterior 7 th, not emarginated before apex. Disc poorly convex; integument shiny, with evident microsculpture and relatively long, sparse, upright pubescence. Humeri marked, almost right; post-humeral margin denticulate, with distinct crenulations up to the base of the apical 5 th of the elytron; elytral apices not separately rounded. Marginal groove wide and evident up to the 8 th pore of the umbilicate series. Chaetotaxy: scutellar pore large, foveate. Umbilicate series with 1 st, 2 nd, and 3 rd pores of the humeral group not equidistant, 2 nd and 3 rd pores closest; 4 th pore clearly farther from the 3 rd one and placed at the end of the basal third of the elytron; 5 th pore placed just after the middle length of the elytron; 5 th and 6 th pores spaced out ca. 1 / 3 of the distance from 6 th and 7 th; 7 th, 8 th, and 9 th pores almost equidistant and slightly displaced onto the disc, 8 th pore placed after the 9 th one. A single discal seta placed laterally towards the edge of the disc and located between the 7 th and 9 th pores. Aedeagus (Fig. 39) relatively small, median lobe long, slender, subrectilinear, with evident basal bulb; ventral margin poorly and gently curved from the basal bulb to the apex; apical blade evident, but short. Endophallus with a median, sinuate, sclerified phanera. Left paramere slender and very elongated, reaching the distal fifth of the median lobe, and bearing two setae; right paramere shorter than the left one. Female. See Baehr and Main (2016).	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
5880E919986B510B922C5271E3CBE388.taxon	distribution	Distribution. Gilesdytes vixsulcatus is known from two localities approximately 85 - 90 km NW of Newman, Pilbara, WA. The distance between the two collection points is ca. 30 km.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
C7332E338BED5DA789E6B481AEF6CE03.taxon	description	Description of the HT ♀. TL 1.39 mm. Body elongated, depigmented, yellow-testaceous; shiny integument with evident microsculpture and pubescence. Head robust, hypertrophic, slightly narrower than pronotum; bearing a number of excess setae. Labium toothless, mentum articulate. Antennae robust, moniliform, and short, not exceeding the base of the pronotum when stretched backwards. Fronto-clypeal furrow indistinct; anterior margin of the epistome subrectilinear. Pronotum subquadrate (max. width / max. length ratio = 1.05), with maximum width in the middle, and with basal border as wide as anterior border; sides slightly but regularly arcuate from anterior to posterior, not sinuate, but strictly emarginated before the basal angles. Anterior angles rounded, poorly prominent; posterior angles subsquare. Disc convex, with long and sparse pubescence; median groove very shallow, hardly evident. Marginal groove wide and flat, slightly enlarged near the base; anterior marginal setae placed inside marginal groove, almost on anterior fifth; basal setae almost before the posterior angles. Legs short and stout, with metatrochanters short and stout and metafemora not dentate (Fig. 5). Elytra subrectangular, elongated (max. length / max. width ratio = 1.84), with maximum width at the base of the posterior third, not emarginated before apex. Disc slightly convex, with evident longitudinal groove running more or less between the scutellar pore and the 9 th pore of the umbilicate series; integument shiny with evident microsculpture and long, sparse, upright pubescence, longitudinally aligned. Humeri well marked; post-humeral margin denticulate, with distinct crenulations all the way to the apical third; elytral apices separately rounded. Marginal groove wide and evident almost up to the 9 th pore of the umbilicate series. Chaetotaxy: scutellar pore large, foveate. Umbilicate series with first three pores of humeral group equidistant; 4 th pore farther and placed at the end of the basal fourth of elytron; 5 th pore placed before the apical third of elytra; 5 th and 6 th ones spaced out ca. half distance of 6 th and 7 th; 8 th slightly displaced on the disc; 7 th and 8 th closer to each other than 8 th and 9 th. Two discal setae, first placed before the 5 th pore of the umbilicate series, second one placed just before 8 th pore. Male. Unknown.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
C7332E338BED5DA789E6B481AEF6CE03.taxon	etymology	Etymology. The name comes from the Latin Gracilanillus hirsutus (= hairy, hirsute) to recognise the presence of many excess setae on the head.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
C7332E338BED5DA789E6B481AEF6CE03.taxon	distribution	Distribution. Gracilanillus hirsutus sp. nov. occurs only at the type locality in Mesa A Mine, Robe River Valley, 44 km W of Pannawonica, Pilbara, WA.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
A7FAEFEABA3A53A9BB715189EB98A1DC.taxon	description	Description of the HT ♀. TL 1.43 mm. Body elongated, depigmented, yellow-testaceous; shiny integument with evident microsculpture and pubescence. Head robust, almost hypertrophic, slightly narrower than pronotum, excess setae absent. Labium toothless, articulated mentum. Antennae robust, moniliform, short, not exceeding the base of the pronotum when stretched backwards. Fronto-clypeal furrow indistinct; anterior margin of the epistome subrectilinear. Pronotum subquadrate (max. width / max. length ratio = 1.07), maximum width at the basal anterior third, and basal border as wide as anterior border; sides slightly but regularly arcuate from anterior to posterior angles, strictly sinuate just before basal angles. Anterior angles rounded, slightly prominent; posterior angles subsquare and acuminate at vertex. Disc convex, with short and very sparse pubescence; median groove very shallow, hardly evident. Marginal groove wide and flat, slightly enlarged near the base; anterior marginal setae placed inside the marginal groove, almost on the anterior fifth; basal setae nearly before posterior angles. Legs short and stout, with metatrochanters short and stout and metafemora non dentate (Fig. 7). Elytra subrectangular, elongated (max. length / max. width ratio = 1.95), with maximum width in the middle, not emarginated in preapical zone. Disc slightly convex, with evident longitudinal groove running more or less between the scutellar pore and the 2 nd discal pore; integument shiny, with evident microsculpture, and short, very dense, and upright pubescence, not longitudinally aligned. Humeri well marked but rounded; post-humeral margin denticulate, with distinct crenulation up to the apical fourth; elytral apices separately and acutely rounded. Marginal groove wide and evident almost all the way to the 8 th pore of the umbilicate series. Chaetotaxy: scutellar pore large, foveate. Umbilicate series with the first three pores of the humeral group equidistant; 4 th pore farther and placed at the end of the basal third of the elytron; 5 th pore placed before the apical third of the elytra; 5 th and 6 th ones spaced out ca. half of the distance between 6 th and 7 th; 8 th slightly displaced onto the disc; 7 th and 8 th slightly closer to each other than the 8 th and 9 th; 9 th pore of the umbilicate series placed well forward (compare to the standard position). Two discal setae, first placed just after the 5 th pore of the umbilicate series, second one placed just after the 8 th pore. Male. Unknown.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
A7FAEFEABA3A53A9BB715189EB98A1DC.taxon	etymology	Etymology. The name of the species derives from the town Pannawonica near the type locality Mesa K Mine, in the Pilbara region.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
A7FAEFEABA3A53A9BB715189EB98A1DC.taxon	distribution	Distribution. Gracilanillus pannawonicanus sp. nov. is known so far only from the type locality Mesa K Mine, in the Robe River Valley, 11 km SSE of Pannawonica, Pilbara, WA.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
29F4C0B5ECE359A592A5B6EFD57D8101.taxon	diagnosis	Diagnosis. Genus characterised by normal metatrochanters, elytra reduced at tip and lacking longitudinal grooves, 9 th pore of the umbilicate series in normal position (placed after the 8 th one) and two discal setae; basal border of pronotum remarkably narrower than anterior border and sides distinctly sinuate before the basal angles; aedeagus with median lobe subrectilinear, basal bulb extremely reduced and parameres bearing one apical seta. Labial tooth lacking.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
29F4C0B5ECE359A592A5B6EFD57D8101.taxon	description	Description. Species small (TL mm 1.02 - 1.05) and anophthalmous. Integument depigmented, poorly sclerified, and covered with sparse pubescence. Head of normal size, narrower than pronotum; mandibles short and simple, without hyperplasias. Maxillary palpi ovoidal, swollen. Labium transverse, articulated; mentum not fused with submentum. Labial tooth absent. Antennae moniliform, without particular features. Pronotum slightly transverse, with sides distinctly sinuate at the basal third. Basal angles obtuse and rounded; basal border remarkably narrower than anterior border; presence of two marginal setae, the posterior one placed at the basal angles. Elytra subrectangular and elongated, separately rounded, depressed and without longitudinal groove, and slightly truncated but not emarginated at apex. Elytral striae absent (except sutural stria). Lateral margin starting from the humeral area, distinctly crenulate to at least half-length. Scutellar pore present, large and umbilicate; umbilicate series of type B (sensu Jeannel 1963; Giachino and Vailati 2011); disc bearing two discal setae. Legs relatively short and stumpy. Unarmed pro- and metafemora, normal metatrochanters, two dilated protarsomeres in the male. Aedeagus small, median lobe short and subrectilinear with basal bulb extremely reduced. Parameres long, bearing one apical seta. Endophallus without sclerified phanerae.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
29F4C0B5ECE359A592A5B6EFD57D8101.taxon	etymology	Etymology. Gregorydytes is a compound noun: in honour of Francis Thomas Gregory, the first European explorer who discovered Ophthalmia Range (type locality) in 1876, and dytes meaning diver in Greek. Gender name masculine.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
C8337997043B5A92A283BDEAC87F39D3.taxon	description	Description. TL mm 1.02 ♂ - 1.05 ♀. Body moderately elongated, depigmented, yellow-testaceous; integument shiny, with evident microsculpture and pubescence. Head robust, not hypertrophic, narrower than pronotum; excess setae absent. Labium toothless, mentum articulated. Antennae robust, moniliform, short, hardly exceeding the base of the pronotum when stretched backwards. Fronto-clypeal furrow indistinct; subrectilinear anterior margin of epistome. Pronotum transverse (max. width / max. length ratio = 1.25), with maximum width at the base of the anterior fourth, and basal border remarkably narrower than anterior border; sides distinctly and regularly arcuate anteriorly, sinuate before basal angles. Anterior angles rounded, not prominent; posterior angles obtuse and rounded. Disc convex, with very sparse pubescence of medium length; median groove very shallow, hardly evident. Marginal groove wide and flat, slightly enlarged near the base; anterior marginal setae placed inside the marginal groove, almost on the anterior fourth; basal setae at posterior angles. Legs short and stout, with metatrochanters short and stout and metafemora non dentate. Two protarsomeres dilated and without adhesive phanerae in male. Elytra subrectangular (max. length / max. width ratio = 1.55), with maximum width at the base of the posterior third, subtruncate but not emarginated before apex. Disc slightly convex, without longitudinal groove; integument shiny, with evident microsculpture and very sparse and upright pubescence of medium length, longitudinally aligned. Humeri well marked but rounded; post-humeral margin denticulate, with distinct crenulation up to the apical third; elytral apices separately rounded. Marginal groove wide and evident almost up to the 9 th pore of the umbilicate series. Chaetotaxy: large and foveate basal umbilicate pore. Umbilicate series with the first three pores of the humeral group very closed to each other and equidistant; 4 th pore farther and placed at the end of the basal third of the elytron; 5 th pore placed at the base the apical third of the elytron; 5 th and 6 th ones spaced from each other equidistant with 6 th and 7 th; 5 th, 7 th and 8 th displaced onto the disc; 7 th and 8 th spaced from each other as 8 th and 9 th. Two discal setae, the first placed at the level of the 3 rd pore of the umbilicate series, the second one placed just before the 8 th pore. Aedeagus (Fig. 9) small, median lobe short, stout, subrectilinear, with basal bulb extremely reduced; ventral margin gently bisinuated; apical blade evident. Endophallus without an evident lamella copulatrix. Left paramere elongated, reaching the aedeagal distal fourth and bearing only one seta; right paramere lost during the preparation of the specimen.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
C8337997043B5A92A283BDEAC87F39D3.taxon	etymology	Etymology. The name arises from the type locality Ophthalmia Range in the Pilbara region.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
C8337997043B5A92A283BDEAC87F39D3.taxon	distribution	Distribution. Gregorydytes ophthalmianus sp. nov. is known so far only from the type locality Ophthalmia Range, 25 km W of Newman, Pilbara, WA.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
E6DEB6FAC1E55E82B1BB65DFBE41FD77.taxon	diagnosis	Diagnosis. Species identified by the characters listed in the genus diagnosis.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
E6DEB6FAC1E55E82B1BB65DFBE41FD77.taxon	description	Description. TL mm 1.57 - 1.59 ♂♂. Body elongate, depigmented, testaceous; integument with medium length pubescence. Microsculpture evident and very strong: composed by isodiametric meshes on pronotal disc; scaly on basal part of elytral disc, and longitudinally oriented hollow points on apical elytral disc (Fig. 34). Head large, slightly narrower than pronotum; without excess setae. Labium without tooth, mentum articulated. Antennae robust, moniliform, short, reaching the base of the pronotum when stretched backwards. Fronto-clypeal furrow indistinct; anterior margin of the epistome subrectilinear. Pronotum trapezoidal (max. width / max. length ratio = 1.00), maximum width at the base of the anterior fourth, with basal border narrower than anterior border; sides subrectilinear, anteriorly slightly but regularly arcuate, not arcuate and not sinuate at the basal half, denticulated at the basal third. Anterior angles obtuse, rounded and not prominent; posterior angles obtuse, but evident. Disc convex, with sparse pubescence of medium length; median groove very shallow, hardly evident. Marginal groove wide and flat, enlarged near the base; anterior marginal setae placed inside the marginal groove, almost on the anterior fourth; basal setae not on the disk and placed at posterior angles. Legs long and slender, with metatrochanters normal and metafemora unarmed. Two poorly dilated protarsomeres, without adhesive phanerae in males. Elytra subrectangular, very elongated (max. length / max. width ratio = 2.20), maximum width at the base of the posterior third, not truncated and slightly emarginated before apex. Disc convex, without longitudinal grooves; integument with sparse and upright pubescence, longitudinally aligned. Humeri well marked; post-humeral margin denticulate, with distinct crenulations up to the 8 th and 9 th pores of the umbilicate series; elytral apices separately rounded. Marginal groove wide and evident almost up to the 9 th pore of the umbilicate series. Chaetotaxy: scutellar pore small and foveate. Umbilicate series with the first three pores of the humeral group very closed to each other and nearly equidistant; 4 th pore farther and placed at the end of the basal third of the elytron; 5 th pore placed just after the middle of the elytron; 5 th and 6 th ones spaced out ca. 1 / 4 of the distance between 6 th and 7 th; 7 th and 9 th placed onto the disc; 7 th and 8 th spaced out ca. double of the distance from 8 th and 9 th; 9 th placed at the level of the 8 th pore. Three discal setae, first placed at the level of the 2 nd pore of the umbilicate series, second one placed at the level of the base of the posterior fourth of elytron, third one placed just before the level of the 9 th pore. Aedeagus (Fig. 35) relatively small, median lobe long, slender, subrectilinear, with basal bulb tight and evident; ventral margin weakly curved from basal bulb to apex; apical blade evident, but short. Endophallus without any sclerified phanerae. Left paramere very elongated, reaching the distal fifth of the median lobe, and bearing only one seta; right paramere shorter than left one, bearing one apical seta.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
E6DEB6FAC1E55E82B1BB65DFBE41FD77.taxon	etymology	Etymology. The name derives from the name of one of the early European explorers to visit the region named as the Carr-Boyd Ranges (type locality). In 1883 William Henry James Carr-Boyd became second-in-command of an expedition led by W. J. O'Donnell on behalf of the Cambridge Downs Pastoral Association; their purpose was to explore the country around the Cambridge Gulf, and to establish a sheep station. The party of six men, including a cook and an Aboriginal boy, twenty-six horses and provisions for six months, left Katherine on 26 March 1883. O'Donnell named the impressive Carr Boyd Range after his second-in-command on 26 May.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
E6DEB6FAC1E55E82B1BB65DFBE41FD77.taxon	distribution	Distribution. Kimberleytyphlus carrboydianus sp. nov. occurs only at the type locality Carr-Boyd Ranges, 530 km SE Darwin, Kimberley, WA.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
B7E825B39D9B599CB651811C3E1B7D3C.taxon	diagnosis	Diagnosis. Genus of the " Illaphanus phyletic series " (sensu Giachino 2005), with species strongly characterised by normal metatrochanters; metafemora non dentate; elytra not reduced at tip; elytral disc without longitudinal grooves and bearing three setae, scaly microsculpture and 9 th pore of the umbilicate series in normal position (placed after the 8 th one); aedeagus with median lobe subrectilinear, size of basal bulb normal and parameres each bearing one apical seta. Labial tooth absent.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
B7E825B39D9B599CB651811C3E1B7D3C.taxon	description	Description. Species of a medium size (TL mm 1.57 - 1.59) and anophthalmous. Integument depigmented but well sclerified, with strong microsculpture and covered with short and sparse pubescence. Head large but slightly narrower than pronotum; mandibles short and simple, without hyperplasias. Maxillary palpi ovoidal, swollen. Labium transverse, articulated; mentum not fused with the submentum. Labial tooth absent. Antennae moniliform, without particular features. Pronotum trapezoidal, with sides not sinuate in the basal third, denticulated at the basal third. Basal angles obtuse, sharp, not rounded; basal border slightly narrower than anterior border; presence of two marginal setae, the posterior one placed near basal angles. Elytra subrectangular elongate, separately rounded, not truncated and apically only slightly emarginated, convex, without longitudinal grooves. Elytral striae absent (except sutural stria). Lateral margin starting from the humeral area and distinctly crenulate up to the level of 8 th- 9 th pores of the umbilicate series. Scutellar pore present, large and umbilicate; umbilicate series of type B (sensu Jeannel 1963; Giachino and Vailati 2011); disc bearing three setae. Legs relatively long and slender. Pro- and metafemora unarmed; metatrochanters normal, two slightly dilated protarsomeres, without adhesive phanerae in males. Aedeagus relatively small, median lobe long, subrectilinear with basal bulb of normal size. Parameres long, bearing one apical seta. Endophallus without any sclerified phanerae.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
B7E825B39D9B599CB651811C3E1B7D3C.taxon	etymology	Etymology. The name combines the name Kimberley (region where the type locality is located) and the Greek suffix - typhlos (meaning blind).	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
9B5AB93702305E309E7B7239586E527D.taxon	diagnosis	Diagnosis. This genus is characterised by: metatrochanters very long and sharp; metafemora dentate; elytra not reduced at tip; elytral disc with longitudinal grooves and bearing three setae; 9 th pore of the umbilicate series in normal position (placed after the 8 th one); aedeagus with median lobe gently curved, size of basal bulb normal and parameres bearing two apical setae. Labial tooth lacking. Differs from Pilbaraphanus by the presence of a longitudinal elytral grooves and the lack of labial tooth.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
CE565E4F410C52CB8199169747A80E79.taxon	description	Description. TL mm 1.44 - 1.46 ♂♂, 1.48 - 1.50 ♀♀. Body elongate, depigmented, yellow; integument shiny, with evident microsculpture, and short pubescence. Head robust, hypertrophic, slightly narrower than pronotum; without excess setae. Labium without tooth, mentum articulated. Antennae robust, moniliform, short, reaching the base of the pronotum when stretched backwards. Fronto-clypeal furrow indistinct; subrectilinear anterior margin of epistome. Pronotum sub-rectangular (max. width / max. length ratio = 1.14), with maximum width at the base of the anterior fifth, and with basal border remarkably narrower than anterior border; sides slightly and irregularly arcuate in the anterior part, subrectilinear at the basal half, gently sinuate and slightly dentate before basal angles. Anterior angles obtuse, slightly prominent; posterior angles squared, acute. Disc convex, with very sparse pubescence of medium length; median groove very shallow, hardly evident. Marginal groove wide and flat, enlarged near the base; anterior marginal setae placed inside the marginal groove, almost on the anterior fifth; basal setae not inside the disk, but placed before posterior angles. Legs long and slender, with metatrochanters long and acuminate, gently curved and metafemora dentate; metatrochanters (Fig. 16) slightly longer than femoral tooth. Two dilated protarsomeres in males. Elytra perfectly subrectangular (max. length / max. width ratio = 1.80), not truncated and not emarginated before the apex. Disc convex, with longitudinal grooves; integument shiny, with evident microsculpture, very short, sparse, and upright, pubescence, not longitudinally aligned. Humeri well marked, gently rounded; post-humeral margin denticulate, with distinct crenulation up to the apical third; elytral apices separately rounded. Marginal groove wide and evident almost up to the 8 th pore of the umbilicate series. Chaetotaxy: scutellar pore large and foveate. Umbilicate series with the first three pores of the humeral group very closed to each other and equidistant; 4 th pore farther and placed at the end of the basal third of the elytron; 5 th pore placed before the base of the apical third of the elytra; 5 th and 6 th ones spaced out ca. half of the distance between 6 th and 7 th; 7 th and 8 th displaced onto the disc; 7 th and 8 th spaced from each other as the 8 th and 9 th. Three discal setae, the first placed before the 4 th pore of the umbilicate series, second one placed just before the 5 th, the third one placed at the 7 th pore. Aedeagus (Fig. 17) large, median lobe long, stout, gently curved, with basal bulb small, but tight and evident; ventral margin gently curved from basal bulb to apex; apical blade evident but short. Endophallus without an evident lamella copulatrix, but with two small, apical, Y-shaped, crossing and slightly sclerified stripes. Left paramere elongated, not reaching the distal third and bearing two setae; right paramere shorter and bearing two apical setae.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
CE565E4F410C52CB8199169747A80E79.taxon	etymology	Etymology. The name comes from the type locality in the Solomon Mining Area, the Firetail deposit, in the Pilbara region.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
CE565E4F410C52CB8199169747A80E79.taxon	distribution	Distribution. Magnanillus firetailianus sp. nov. is known only from the type locality (Firetail deposit) 50 km N of Tom Price, Pilbara, WA.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
2B14A92FF7F85274B6E7DB03F931208C.taxon	description	Description of the HT ♂. TL mm 2.25. Body elongate, depigmented, yellow-testaceous; integument shiny with evident microsculpture and short pubescence. Head relatively large, narrower than pronotum; with two couples of excess setae on the vertex as in Fig. 24. Labium without tooth, mentum articulated. Antennae robust, submoniliform, very short, not reaching the base of the pronotum when stretched backwards. Fronto-clypeal furrow indistinct; anterior margin of the epistome subrectilinear. Pronotum sub-squared (max. width / max. length ratio = 1.14 maximum width at the base of the anterior fourth, and basal border slightly wider than anterior border; sides slightly and irregularly arcuate in anterior part, subrectilinear at the basal half, not sinuate but with an evident tooth before basal angles. Anterior angles obtuse, slightly prominent; posterior angles right, acute. Disc convex, with very sparse pubescence of medium length; median groove very shallow, hardly evident. Marginal groove wide and flat, enlarged near the base; anterior marginal setae placed inside the marginal groove, almost on the anterior fourth; basal setae slightly placed internally on the disk and before the posterior angles. Legs long and slender, with metatrochanters long, acuminate, subrectilinear and metafemora dentate; metatrochanters (Fig. 25) reaching the femoral tooth. Two dilated protarsomeres, without adhesive phanerae in males. Elytra subrectangular, elongate (max. length / max. width ratio = 1.83), not truncated and only slightly emarginated before apex. Disc convex, with longitudinal grooves; integument shiny, with evident microsculpture and very short, longitudinally aligned, upright pubescence. Humeri well marked, obtuse; post-humeral margin denticulate, with distinct crenulations up to the base of the apical third; elytral apices separately rounded. Marginal groove wide and evident almost up to the 8 th pore of the umbilicate series. Chaetotaxy: scutellar pore large and foveate. Umbilicate series with the first three pores of the humeral group very closed to each other and equidistant; 4 th pore farther and placed at the end of the basal third of the elytron; 5 th pore placed at the base the apical third of the elytron; 7 th pore very forward, placed near the 6 th pore; 6 th and 7 th pores closer than 5 th and 6 th; 8 th displaced onto the disc; 7 th and 8 th spaced out ca. the 5 th and 6 th. Three discal setae, first placed before the 4 th pore of the umbilicate series, second one placed just before the 5 th pore, third one placed after the 7 th pore. Aedeagus (Fig. 26) large, median lobe long, slender, gently curved, with basal bulb small but tight and evident; ventral margin gently curved from basal bulb to apex; apical blade poorly evident, short. Endophallus without an evident lamella copulatrix, but with small, Y-shaped, apical, slightly sclerified stripe. Left paramere elongate, reaching the distal third and bearing two setae; right paramere shorter and bearing two apical setae.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
2B14A92FF7F85274B6E7DB03F931208C.taxon	etymology	Etymology. The name comes from the Latin word Magnanillus regalis = royal, and it reminds the type locality " King " deposit in the Solomon Mining Area.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
2B14A92FF7F85274B6E7DB03F931208C.taxon	distribution	Distribution. Magnanillus regalis sp. nov. is known only from the type locality (Kings deposit) in the Solomon Mining Area, 50 km N of Tom Price, Pilbara, WA.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
1CE389B1CDDC53549EED6975F16CBA17.taxon	description	Description. TL mm 2.42 - 2.46 ♂♂, 2.48 - 2.51 ♀♀. Body elongated, depigmented, testaceous; integument shiny, with evident microsculpture and very short pubescence. Head robust, narrower than pronotum, excess setae absent. Labium without tooth, mentum articulated. Antennae robust, moniliform, short, reaching the base of the pronotum when stretched backwards. Fronto-clypeal furrow indistinct; anterior margin of the epistome subrectilinear. Pronotum sub-squared (max. width / max. length ratio = 1.14), with maximum width at the base of the anterior fourth, and basal border slightly wider than anterior border; sides slightly and irregularly arcuate in the anterior part, subrectilinear in the basal half, not sinuate and slightly denticulate before basal angles. Anterior angles obtuse, slightly prominent; posterior angles obtuse, blunted. Disc convex, with very short and sparse pubescence; median groove very shallow, hardly evident. Marginal groove wide and flat, enlarged near the base; anterior marginal setae placed inside the marginal groove, almost on the anterior fifth; basal setae not inside on the disk, but placed before the posterior angles. Legs long and slender, with metatrochanters long, acuminate and subrectilinear and metafemora dentate; metatrochanters (Fig. 19) shorter than femoral tooth. Two dilated protarsomeres in males. Elytra subrectangular, relatively short (max. length / max. width ratio = 1.63), not truncated and very slightly emarginated before apex. Disc convex, with longitudinal grooves; very short, sparse, and upright pubescence not longitudinally aligned. Humeri well marked, obtuse; post-humeral margin denticulate, with a distinct crenulation down to the apical third; elytral apices separately rounded. Marginal groove wide and evident almost up to the 7 th pore of the umbilicate series. Chaetotaxy: scutellar pore large and foveate. Umbilicate series with the first three pores of the humeral group very closed to each other and equidistant; 4 th pore farther and placed at the end of the basal third of the elytron; 5 th pore placed before the base of the apical third of the elytron; 5 th and 6 th ones spaced out ca. 1 / 3 of the distance from 6 th and 7 th; 7 th and 8 th displaced onto the disc; 7 th and 8 th spaced from each other as the 8 th and 9 th. Three discal setae, first placed before the 4 th pore of the umbilicate series, second one placed just before the 5 th, third one placed before the 7 th pore. Aedeagus (Fig. 20) large, median lobe long, stout, gently curved, with basal bulb small but evident; ventral margin gently curved from basal bulb to apex; apical blade poorly evident, very short. Endophallus without an evident lamella copulatrix, but with two small, apical, subparallel, and slightly sclerified stripes. Left paramere elongate, not reaching the aedeagal distal third and bearing two setae; right paramere shorter and bearing two apical setae.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
1CE389B1CDDC53549EED6975F16CBA17.taxon	etymology	Etymology. The name is to remind one of the Solomon mining area and the different deposits where this species occurs, and it originates from the mythological Queen of Sheba (in Latin Sheba = Saba). According to tradition, the Queen of Sheba visited the Kingdom of Solomon with valuable gifts for its king.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
1CE389B1CDDC53549EED6975F16CBA17.taxon	distribution	Distribution. Magnanillus sabae sp. nov. is known from different deposits of the Solomon Mining Area, 50 - 60 km N / NW of Tom Price, Pilbara, WA.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
49F878A1A4885017B069D32DBB88145A.taxon	description	Description. TL mm 2.20 - 2.22 ♂♂, 2.25 - 2.28 ♀♀. Body elongated, depigmented, testaceous; integument shiny with evident microsculpture and very short pubescence. Head relatively small, narrower than pronotum; excess setae absent. Labium without tooth, mentum articulated. Antennae robust, submoniliform, short, reaching the base of the pronotum when stretched backwards. Fronto-clypeal furrow indistinct; anterior margin of the epistome subrectilinear. Pronotum transverse (max. width / max. length ratio = 1.16), with maximum width at the base of the anterior fourth, and basal border wider than anterior border; sides slightly and irregularly arcuate in anterior part, subrectilinear in the basal half, not sinuate and denticulate before basal angles. Anterior angles obtuse, slightly prominent; posterior angles right, acute. Disc convex, with short and very sparse pubescence; median groove very shallow, hardly evident. Marginal groove wide and flat, enlarged near the base; anterior marginal setae placed inside the marginal groove, almost on the anterior fourth; basal setae slightly located internally on the disk and placed before the posterior angles. Legs long and slender, with metatrochanters long, acuminate, and gently curved and metafemora dentate; metatrochanters (Fig. 22) as long as the femoral tooth. Anterior legs missing in all male specimens. Elytra subrectangular, relatively short (max. length / max. width ratio = 1.69), not truncate and not emarginated before the apex. Disc convex, with longitudinal grooves; integument shiny with evident microsculpture, very short, sparse, and upright, pubescence, not longitudinally aligned. Humeri well marked, obtuse; post-humeral margin denticulate, with distinct crenulations up to the base of the apical third; elytral apices separately rounded. Marginal groove wide and evident almost up to the 8 th pore of the umbilicate series. Chaetotaxy: scutellar pore large and foveate. Umbilicate series with the first three pores of the humeral group very closed to each other and equidistant; 4 th pore farther and placed at the end of the basal third of the elytron; 5 th pore placed at the base of the apical third of the elytron; 5 th and 6 th ones spaced out ca. half of the distance between 6 th and 7 th; 7 th and 8 th displaced onto the disc; 7 th and 8 th spaced from each other as the 8 th and the 9 th. Three discal setae, first placed before the 4 th pore of the umbilicate series, second one placed in the middle of the elytron, the third one placed before the 7 th pore. Aedeagus (Fig. 23) large, median lobe long, stout, gently curved, with basal bulb small but tight and evident; ventral margin gently curved from basal bulb to apex; apical blade poorly evident, very short. Endophallus without an evident lamella copulatrix, but with two very small, apical, and slightly sclerified, ovoidal areas. Left paramere elongate, reaching the distal third and bearing two setae; right paramere shorter and bearing two apical setae.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
49F878A1A4885017B069D32DBB88145A.taxon	etymology	Etymology. The name comes from type locality " Solomon mining area " and the mythological King of Solomon (in Latin Salomon).	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
49F878A1A4885017B069D32DBB88145A.taxon	distribution	Distribution. Magnanillus salomonis sp. nov. is known from different deposits of the Solomon Mining Area, 50 - 60 km N / NW of Tom Price, Pilbara, WA.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
785280137B635A939EC0C1A799D9D09E.taxon	description	Description. TL mm 2.27 - 2.30 ♂♂, 2.32 ♀. Body elongate, depigmented, testaceous; integument shiny, with evident microsculpture and very short pubescence. Head relatively small, narrower than pronotum; without excess setae. Labium without tooth, mentum articulated. Antennae robust, submoniliform, short, reaching the base of the pronotum when stretched backwards. Fronto-clypeal furrow indistinct; anterior margin of the epistome subrectilinear. Pronotum transverse (max. width / max. length ratio = 1.23), maximum width at the base of the anterior third, and basal border slightly wider than anterior border; sides slightly and irregularly arcuate in anterior part, very poorly arcuate in the basal half, not sinuate and denticulate before the basal angles. Anterior angles obtuse, slightly prominent; posterior angles right, acute. Disc convex, with very short and sparse pubescence; median groove very shallow, hardly evident. Marginal groove wide and flat, enlarged near the base; anterior marginal setae placed inside the marginal groove, almost on the anterior fourth; basal setae slightly placed inside the disk and before the posterior angles. Legs long and slender, with metatrochanters long, acuminate, and curved and metafemora dentate; metatrochanters (Fig. 28) longer than femoral tooth. Two dilated protarsomeres, without adhesive phanerae in males. Elytra subrectangular, relatively short (max. length / max. width ratio = 1.67), not truncated and only very slightly emarginated before apex. Disc convex, with longitudinal grooves; shiny integument with evident microsculpture and very short, sparse, upright pubescence, not longitudinally aligned. Humeri well marked but rounded; post-humeral margin denticulate, with distinct crenulations up to the base of the apical fourth; elytral apices separately rounded. Marginal groove wide and evident almost down to the 7 th pore of the umbilicate series. Chaetotaxy: scutellar pore large and foveate. Umbilicate series with the first three pores of the humeral group very closed to each other and nearly equidistant; 4 th pore farther and placed at the end of the basal third of the elytron; 5 th pore placed at the base of the apical third of the elytron; 5 th and 6 th ones spaced out ca. half of the distance between the 6 th and the 7 th; 7 th and 8 th displaced onto the disc; 7 th and 8 th spaced from each other as the 8 th and 9 th. Three discal setae, first placed before the 4 th pore of the umbilicate series, second one placed after the midpoint of the elytron, third one placed before the 7 th pore. Aedeagus (Fig. 29) large, median lobe long, slender, gently curved, with basal bulb small but tight and evident; ventral margin gently curved from basal bulb to apex; apical blade poorly evident, very short. Endophallus without an evident lamella copulatrix, but with very small, V-shaped, apical, slightly sclerified stripe. Left paramere elongate, reaching the distal third and bearing two setae; right paramere shorter and bearing two apical setae.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
785280137B635A939EC0C1A799D9D09E.taxon	etymology	Etymology. The name comes from the Serenity Valley, type locality of the species, in the Pilbara region.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
785280137B635A939EC0C1A799D9D09E.taxon	distribution	Distribution. Magnanillus serenitatis sp. nov. is known only from two drill holes in the Serenity Valley, 65 km N / NW of Tom Price, Pilbara, WA.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
BEE8065ABB5B51C0A5891B538032E2F7.taxon	diagnosis	Diagnosis. Species identified by the characters listed in the genus diagnosis.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
BEE8065ABB5B51C0A5891B538032E2F7.taxon	description	Description. TL mm 1.93 - 1.97 ♂♂, 2.00 - 2.04 ♀♀. Body elongated, depigmented, testaceous; integument shiny, with evident microsculpture and medium length pubescence. Head large, narrower than pronotum; without excess setae. Labium without tooth, mentum articulated. Antennae robust, moniliform, short, reaching the base of the pronotum when stretched backwards. Fronto-clypeal furrow distinct; anterior margin of the epistome subrectilinear. Pronotum cordiform (max. width / max. length ratio = 1.13), maximum width at the base of the anterior third, and basal border narrower than anterior border; sides slightly and regularly arcuate in anterior part, subrectilinear at the basal half, sinuate and denticulate before basal angles. Anterior angles obtuse, prominent; posterior angles right, sharp at tips. Disc convex, with sparse pubescence of medium length; median groove very shallow, hardly evident. Marginal groove wide and flat, poorly enlarged near the base; anterior marginal setae placed inside the marginal groove, almost on the anterior fifth; basal setae not on the disk and placed before posterior angles. Legs long and slender, with metatrochanters long and acuminate but not curved and metafemora unarmed; metatrochanters (Fig. 31) as long as 2 / 3 of the femoral length. Two poorly dilated protarsomeres, without adhesive phanerae in males. Elytra ovoidal, relatively short (max. length / max. width ratio = 1.8), maximum width at the base of the posterior third, not truncated and not emarginated in preapical zone. Disc convex, with longitudinal grooves; integument shiny with evident microsculpture and pubescence of medium length, sparse and upright, not longitudinally aligned. Humeri rounded; post-humeral margin denticulate, with distinct crenulations up to half-length; elytral apices separately rounded. Marginal groove wide and evident almost up to the 7 th pore of the umbilicate series. Chaetotaxy: scutellar pore large, foveate. Umbilicate series with the first three pores of the humeral group very closed to each other and nearly equidistant; 4 th pore farther and placed at the end of the basal third of the elytron; 5 th pore placed just before the base the apical third of the elytron; 5 th and 6 th ones spaced out ca. half of the distance between 6 th and 7 th; 7 th and 8 th displaced onto the disc; 7 th and 8 th spaced out ca. as the 8 th and 9 th. One discal seta placed just before the 7 th pore of the umbilicate series. Aedeagus (Figs 32, 33) large, median lobe long, slender, curved, with basal bulb tight and evident; ventral margin regularly curved from basal bulb to apex; apical blade evident, but short. Endophallus without an evident lamella copulatrix, but with very small, preapical, slightly sclerified stripe. Left paramere elongate, reaching the distal third of median lobe and bearing two setae; right paramere shorter and bearing two apical setae.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
BEE8065ABB5B51C0A5891B538032E2F7.taxon	etymology	Etymology. The name comes from the Callawa Ridge (type locality) in the NE of the Pilbara region.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
BEE8065ABB5B51C0A5891B538032E2F7.taxon	distribution	Distribution. Neoillaphanus callawanus sp. nov. is known only from a few drill holes on the Callawa Ridge, 200 km E of Port Hedland, Pilbara, WA.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
3B2B4B9D3C115DB0B14B11D6189B613F.taxon	diagnosis	Diagnosis. Genus characterised by: metatrochanters long and sharp; metafemora non dentate; elytra not reduced at tip; elytral disc with longitudinal grooves and bearing one seta; 9 th pore of the umbilicate series in normal position (placed after the 8 th one); aedeagus with median lobe curved, size of the basal bulb normal and parameres bearing two apical setae. Labial tooth absent.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
3B2B4B9D3C115DB0B14B11D6189B613F.taxon	description	Description. Species of medium size (TL mm 1.93 - 2.04) and anophthalmous. Integument depigmented but well sclerified and covered with sparse pubescence. Head large but narrower than pronotum; mandibles short and simple, without hyperplasias. Maxillary palpi ovoidal, swollen. Labium transverse, articulated; mentum not fused with submentum. Labial tooth absent. Antennae moniliform, without particular features. Pronotum cordiform, sides sinuate at the basal third, denticulated before basal angles. Basal angles right, sharp at tips, not rounded; basal border narrower than anterior border; presence of two marginal setae, the posterior one placed before basal angles. Elytra ovoidal short, separately rounded, not truncated and apically not emarginated; convex, with longitudinal grooves. Elytral striae absent (except sutural stria). Lateral margin starting from the humeral area, distinctly crenulate up to its half-length. Scutellar pore present, large and umbilicate; umbilicate series of type B (sensu Jeannel 1963; Giachino and Vailati 2011); disc bearing one seta. Legs relatively long and slender. Pro- and metafemora unarmed; metatrochanters long and sharp, two dilated protarsomeres in the male. Aedeagus relatively large, median lobe long and curved with basal bulb of normal size. Parameres long, each of them bearing two apical setae. Endophallus with very small and poorly sclerified phanerae.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
3B2B4B9D3C115DB0B14B11D6189B613F.taxon	etymology	Etymology. The name is derived by the combination of the Greek prefix Neo - (means new) and the name Illaphanus. Gender masculine.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
F9C87688C6FB570D9B479CFC4A2D27A1.taxon	description	Description of the HT ♂. TL mm 2.12. Body elongated, depigmented, yellow-testaceous; shiny integument, with evident microsculpture, covered with very sparse and short pubescence. Head small, narrower than the base of pronotum. Labium without tooth. Antennae missing. Fronto-clypeal furrow slightly distinct; subrectilinear anterior margin of epistome. Pronotum subrectangular, slightly transverse (max. width / max. length ratio = 1.22), maximum width at the base of the anterior third, very wide basal border, as wide as the anterior border, pronotum sides poorly arcuate anteriorly, subrectilinear and gently convergent posteriorly, laterally completely serrulate from the anterior seta to the basal angles, not emarginated before the base. Anterior angles rounded, only slightly prominent; posterior angles sharp and obtuse, not protruding. Disc slightly convex, with very sparse and short pubescence; median groove very shallow, slightly marked. Marginal groove wide and flat, very enlarged near the base; anterior marginal setae inserted inside the marginal groove, approximately on the anterior fifth; basal seta lacking. Legs missing. Elytra subrectangular, very elongated (max. length / max. width ratio = 1.79), maximum width at the middle, slightly emarginated in the pre-apical zone. Disc convex; shiny integument, with evident microsculpture and short, very sparse, upright pubescence. Humeri very marked, but rounded; post-humeral margin denticulate, with distinct crenulations up to 6 th pore of the umbilicate series; elytral apices not separately rounded. Marginal groove wide and evident up to the 8 th pore of the umbilicate series. Chaetotaxy: scutellar pore large, foveate. Umbilicate series with 1 st, 2 nd, and 3 rd pores of the humeral group not equidistant, 2 nd and 3 rd pores closest; 4 th pore clearly farther from the 3 rd one and placed at the end of the basal third of the elytron; 5 th pore placed well after the middle length of the elytron; 5 th and 6 th pores spaced out ca. the 1 / 3 of the distance from 6 th and 7 th ones; 7 th, 8 th, and 9 th pores almost equidistant, 8 th pore slightly displaced onto the disc and placed after the 9 th one. One single discal seta laterally placed near the edge, midway between the 7 th and the 9 th pores. Aedeagus (Fig. 44) relatively large, median lobe long, tubular, regularly curved, with basal bulb evident; ventral margin gently curved from the basal bulb to the apex; apical blade evident, but short. Endophallus without any sclerified phanerae. Left parameres slender and very elongated, reaching the distal fourth of the median lobe, and bearing two setae; right paramere stout and shorter than the left one, and bearing two setae. Female. Unknown.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
F9C87688C6FB570D9B479CFC4A2D27A1.taxon	etymology	Etymology. The name comes from the type locality Abydos deposit.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
F9C87688C6FB570D9B479CFC4A2D27A1.taxon	distribution	Distribution. Pilbaradytes abydosianus sp. nov. is known only from the type locality 100 km SE of Port Hedland, Pilbara, WA.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
C37247B2D72D59BFA50B0BD4B4872D7F.taxon	diagnosis	Diagnosis. Included species strongly characterised by: lacking longitudinal elytral grooves, pronotum basal border as wide as anterior border, with sides posteriorly not or only slightly sinuate, basal seta of pronotum absent, one elytral discal seta placed near elytral edge, 8 th pore of the umbilicate series placed after the 9 th one (sensu Giachino and Vailati 2011), metafemora not dentate, labium tooth absent, median lobe of the aedeagus long and curved with parameres long and slender. Differs from Magnanillus Baher, 2017 by the absence of elytral longitudinal grooves, and by a single elytral discal seta (3 setae in Magnanillus).	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
C37247B2D72D59BFA50B0BD4B4872D7F.taxon	description	Description. A genus of Anillini with species of a medium size (TL mm 2.12 - 2.23), anophthalmous. Integument depigmented but well sclerified, with strong microsculpture and covered with short and sparse pubescence. Head as wide as, or narrower, than the pronotum base; mandibles short and simple, without hyperplasias. Maxillary palps ovoidal, swollen. Labium transverse, articulated; mentum not fused with submentum. Labial tooth absent. Antennae not strictly moniliform (with relatively elongated antennomeres). Pronotum squared, with sides smooth or serrulate, not or only slightly sinuate towards the basal third. Basal angles right or obtuse, sharp, not rounded; basal border as wide as or wider than anterior margin; presence of only one marginal seta, posterior seta absent. Elytra subrectangular, elongated, not separately rounded, not truncate and slightly emarginated apically; convex, without longitudinal grooves. Elytral striae missing (except for the sutural stria). Lateral margin, starting from the humeral area, hardly serrulate up to the level of 6 th pore of the umbilicate series. Scutellar pore present, large and umbilicate; umbilicate series of type B (sensu Jeannel 1963; Giachino and Vailati 2011) with the 8 th pore placed after the 9 th one; disc bearing one seta strictly located near the elytral edge. Legs relatively long and slender. In female, unarmed pro- and metafemora; metatrochanters normal; male legs unknown. Aedeagus relatively large; median lobe long, curved, with basal bulb of normal size. Parameres long, slender, bearing two apical setae. Endophallus without sclerified phanerae.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
C37247B2D72D59BFA50B0BD4B4872D7F.taxon	etymology	Etymology. Composite name coming from the Pilbara region with the suffix - dytes (diver). Gender masculine.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
87567848E1EA5A7EBBC62358334AEF81.taxon	description	Description of the HT ♀. TL mm 2.23. Body elongated, depigmented, yellow-testaceous; shiny integument, with evident microsculpture, covered with very sparse and short pubescence. Head small, approximately as wide as the base of the pronotum. Labium without tooth. Antennae with relatively elongated antennomeres, short, just exceeding the base of the pronotum when stretched backwards. Fronto-clypeal furrow slightly distinct; subrectilinear anterior margin of epistome. Pronotum subrectangular, slightly transverse (max. width / max. length ratio = 1.23), maximum width at the base of the anterior fifth, with very wide basal border, as wide as the anterior border, pronotum sides poorly arcuate anteriorly, subrectilinear and very slightly sinuated posteriorly, laterally completely smooth from anterior seta to basal angles, not emarginated before the base. Anterior angles rounded, only slightly prominent; posterior angles right, sharp, not protruding. Disc slightly convex, with very sparse and short pubescence; median groove very shallow, slightly marked. Marginal groove narrow and flat, not enlarged near the base; anterior marginal setae inserted inside marginal groove, approximately on the anterior seventh; basal seta absent. Legs relatively long and slender. In females, unarmed pro- and metafemora; normal metatrochanters. Elytra subrectangular, very elongated (max. length / max. width ratio = 1.85), maximum width at the middle, lateral sides slightly emarginated at the end of the basal third and in the pre-apical zone. Disc convex; shiny integument, with evident microsculpture and short, sparse and upright pubescence. Humeri very marked, but rounded; post-humeral margin denticulate, with distinct crenulations up to 6 th pore of the umbilicate series; elytral apices not separately rounded. Marginal groove wide and evident up to the 9 th pore of the umbilicate series. Chaetotaxy: scutellar pore large, foveate. Umbilicate series with 1 st, 2 nd, and 3 rd pores of the humeral group not equidistant, 2 nd and 3 rd pores closest; 4 th pore clearly farther from the 3 rd one and placed at the end of the basal third of the elytron; 5 th pore placed well after the middle length of the elytron; 5 th and 6 th pores spaced out ca. half of the distance from 6 th and 7 th; 7 th, 8 th, and 9 th pores almost equidistant, 8 th and 9 th pores closer to each other than 7 th and 9 th; 8 th pore placed after the 9 th one. One single discal seta laterally placed near the edge, midway between the 7 th and the 9 th pores. Male. Unknown.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
87567848E1EA5A7EBBC62358334AEF81.taxon	etymology	Etymology. The name comes from the type locality Mount Webber, in the Pilbara region.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
87567848E1EA5A7EBBC62358334AEF81.taxon	distribution	Distribution. Pilbaradytes webberianus sp. nov. is known only from Mount Webber, 150 km SE of Port Hedland, Pilbara, WA.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
7FB9AD69B2B1568FB7B0C8D494841646.taxon	description	Description of the HT ♀. TL mm 1.37. Body elongated and depigmented, yellow; integument shiny, with evident microsculpture and short pubescence. Head robust, hypertrophic, narrower than pronotum; excess setae absent. Labium with smooth tooth, mentum articulated. Antennae robust, moniliform, short, reaching the base of the pronotum when stretched backwards. Fronto-clypeal furrow indistinct; anterior margin of the epistome subrectilinear. Pronotum subsquare (max. width / max. length ratio = 1.03), with the maximum width at the base of the anterior fourth, and with basal border remarkably wider than anterior border; sides poorly and not regularly arcuate in the anterior part, gently sinuate in the basal half and slightly dentate before basal angles. Anterior angles obtuse, prominent; posterior angles squared, gently rounded. Disc convex, with very sparse pubescence of medium length; median groove very shallow, hardly evident. Marginal groove wide and flat, enlarged near the base; anterior marginal setae placed inside the marginal groove, almost on the anterior fourth; basal setae not placed inside on the disk, but before the posterior angles. Legs long and slender, with metatrochanters long and acuminate, gently curved and metafemora dentate; metatrochanters (Fig. 14) as long as femoral tooth. All left legs missing in the HT ♀. Elytra perfectly subrectangular (max. length / max. width ratio = 1.91), not truncated, only slightly emarginated before the apex. Disc convex, longitudinal grooves absent; integument shiny with evident microsculpture, and very short, sparse, upright pubescence not longitudinally aligned. Humeri well marked, gently rounded; post-humeral margin denticulate, with distinct crenulation up to the apical third; elytral apices separately rounded. Marginal groove wide and evident almost up to the 9 th pore of the umbilicate series. Chaetotaxy: scutellar pore large and foveate. Umbilicate series with the first three pores of the humeral group very closed to each other and equidistant; 4 th pore farther and placed at the end of the basal third of the elytron; 5 th pore placed before the base of the apical third of the elytron; 5 th and 6 th ones spaced from each other as half distance from 6 th and 7 th; 7 th and 8 th displaced onto the disc; 7 th and 8 th spaced from each other as the 8 th and 9 th. Three discal setae, the first placed before the 4 th pore of the umbilicate series, the second one placed just before the 5 th, and the third one placed at the level of the 7 th pore of the umbilicate series. Male. Unknown.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
7FB9AD69B2B1568FB7B0C8D494841646.taxon	etymology	Etymology. The species name derives from Bilybara, aboriginal name that refers to the Pilbara region.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
7FB9AD69B2B1568FB7B0C8D494841646.taxon	distribution	Distribution. Pilbaraphanus bilybarianus sp. nov. is known only from the type locality (Kings deposit, which is part of the Solomon Mining Area), 60 km N of Tom Price, Pilbara, WA.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
C5DB148CEEA254299DEC5B5DD39C604E.taxon	description	Description of the HT ♂. TL mm 2.18. Body elongated and depigmented, yellow-testaceous; integument shiny, with evident microsculpture and short pubescence. Head robust, hypertrophic, slightly narrower than pronotum; without excess setae. Labium with a smooth tooth, mentum articulated. Antennae robust, moniliform, short, not reaching the base of the pronotum when stretched backwards. Fronto-clypeal furrow indistinct; anterior margin of the epistome subrectilinear. Pronotum subsquare (max. width / max. length ratio = 1.12), with maximum width at the base of the anterior fifth, and with basal border remarkably wider than anterior border; sides slightly and irregularly arcuate in the anterior part, subrectilinear before basal angles. Anterior angles acute, prominent; posterior angles squared and not rounded. Disc convex, with very sparse pubescence of medium length; median groove very shallow, hardly evident. Marginal groove wide and flat, slightly enlarged near the base; anterior marginal setae placed inside the marginal groove, almost on the anterior fifth; basal setae placed inside the disk and before posterior angles. Legs long and slender, with metatrochanters long and acuminate, but not curved, and metafemora dentate; metatrochanters (Fig. 11) shorter than femoral tooth. Two protarsomeres dilated and without adhesive phanerae in males. Left pro-, left and right meso-, and left metalegs missing in the HT ♂. Elytra perfectly subrectangular (max. length / max. width ratio = 1.80), not truncate and not emarginated before apex. Disc convex, longitudinal grooves absent; integument shiny, with evident microsculpture, and very short, sparse and upright, pubescence longitudinally aligned. Humeri well marked, gently rounded; post-humeral margin denticulate, with distinct crenulation up to the apical third; elytral apices separately rounded. Marginal groove wide and evident almost up to the 8 th pore of the umbilicate series. Chaetotaxy: scutellar pore large and foveate. Umbilicate series with the first three pores of the humeral group very closed to each other and equidistant; 4 th pore farther and placed at the end of the basal third of the elytron; 5 th pore placed before the base of the apical third of the elytron; 5 th and 6 th ones spaced from each other half the distance from 6 th and 7 th; 7 th and 8 th displaced onto the disc; 7 th and 8 th spaced to each other as the 8 th and 9 th. Three discal setae, the first placed before the 4 th pore of the umbilicate series, the second and third ones placed respectively just before the 5 th and 8 th umbilicate pores. Aedeagus (Fig. 12) large, median lobe long, stout, subrectilinear, with basal bulb extremely reduced; ventral margin gently curved; apical blade evident but short. Endophallus without an evident lamella copulatrix, but with two crossing apical slightly sclerified stripes. Left paramere elongated, reaching the aedeagal distal third and bearing two setae; right paramere lost during the preparation of the specimen.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
C5DB148CEEA254299DEC5B5DD39C604E.taxon	etymology	Etymology. The species name comes from the Chichester Range, where the type locality (Quarry 8) is located.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
C5DB148CEEA254299DEC5B5DD39C604E.taxon	distribution	Distribution. Pilbaraphanus chichesterianus sp. nov. is known so far only from the type locality Quarry 8 in the Chichester Range, 100 km E of Port Hedland, Pilbara, WA.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
09074B2F2DE85F688FC4283571657045.taxon	diagnosis	Diagnosis. Genus characterised by very long and sharp metatrochanters; dentate metafemora; elytra not reduced at tip; elytral disc bearing three setae and longitudinal grooves absent; 9 th pore of the umbilicate series in the normal position (placed after the 8 th); aedeagus with median lobe not very curved, size of basal bulb normal and parameres bearing two apical setae. Labial tooth present and smooth. Differs from Magnanillus Baehr, 2017 by the absence of longitudinal elytral grooves.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
09074B2F2DE85F688FC4283571657045.taxon	description	Description. Species of medium size (TL mm 1.37 - 2.18) and anophthalmous. Integument depigmented but well sclerified and covered with sparse pubescence. Head from normal to large size, slightly narrower than pronotum; mandibles short and simple, without hyperplasias. Maxillary palpi ovoidal and swollen. Labium transverse and articulated; mentum not fused with submentum. Labial tooth present and smooth. Antennae moniliform, without particular features. Pronotum subquadrate, with sides not or only slightly sinuate at the basal third, smooth or denticulated before basal angles. Basal angles right, sharp, not rounded; basal border as wide as, or only slightly narrower, than the anterior border; presence of two marginal setae, the posterior one placed just before basal angles. Elytra subrectangular and elongated, separately rounded and depressed, not truncate and not emarginated apically, with longitudinal groove absent. Elytral striae absent (except sutural stria). Lateral margin starting from humeral area, distinctly crenulate up to at least half-length. Scutellar pore present, large and umbilicate; umbilicate series of type B (sensu Jeannel 1963; Giachino and Vailati 2011); disc bearing three discal setae. Legs relatively short and stumpy. Unarmed profemora; posterior edge of metafemora dentate, metatrochanters very long and sharp, two dilated protarsomeres in the male. Aedeagus relatively large, median lobe long and not very curved, with basal bulb of normal size. Parameres long and bearing two apical setae. Endophallus with poorly sclerified phanerae.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
09074B2F2DE85F688FC4283571657045.taxon	etymology	Etymology. The genus name combines the Pilbara name with the suffix - phanus to recognise the genus Illaphanus. Gender masculine.	en	Giachino, Pier Mauro, Eberhard, Stefan, Perina, Giulia (2021): A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia. ZooKeys 1044: 269-337, DOI: http://dx.doi.org/10.3897/zookeys.1044.58844, URL: http://dx.doi.org/10.3897/zookeys.1044.58844
