taxonID	type	description	language	source
95F9D46816955537BADB471CC1F34045.taxon	distribution	Distribution and habitat. Theprisa australis is broadly sympatric with T. montana in the mountains of Victoria east and southeast of Melbourne (Fig. 10). Localities where T. australis and T. montana have been collected syntopically include Gunyah, Mt. Baw Baw, Mt. Donna Buang, Sherbrooke Forest, Tarra Valley in Tarra-Bulga N. P., and Warburton. Individuals have been found in leaf and log litter associated with Eucalyptus regnans, E. delegatensis R. T. Baker, Nothofagus cunninghamii, and Blechnum ferns, and via application of pyrethrin fog insecticide to logs with fungal outgrowths (FMNH).	en	Liebherr, James K., Porch, Nick, Shaw, Matthew, Sinclair, Bronte E., Maddison, David R. (2021): Systematic revision of the trans-Bassian moriomorphine genus Theprisa Moore (Coleoptera, Carabidae). ZooKeys 1044: 339-373, DOI: http://dx.doi.org/10.3897/zookeys.1044.62335, URL: http://dx.doi.org/10.3897/zookeys.1044.62335
472CE67146395350A146D7C06A1AB419.taxon	distribution	Distribution and habitat. Theprisa convexa is known from the mountainous western portions of Tasmania (Fig. 9), within the West, King, Central Highlands, and Northern Slopes biogeographic regions (IBRA 2012). The beetles are terrestrial, with specimens recorded from under decaying logs, in leaf litter associated with Nothofagus cunninghamii (Hooker) (Nothofagaceae), and from riverine forest with Eucryphia (Cunoniaceae), Richea (Ericaceae), N. cunninghamii, and tree ferns. Recorded habitats range 80 - 1200 m elevation. We have not had the opportunity to examine specimens reported by Eberhard and Giachino (2011), however the geographical expanse of collecting sites they reported agrees totally with our data, with the exception of additional outlying records herein from Christmas Hills, 35 km SW Smithton (Kethley, FMNH), and Trowutta (Madden, TMAG), northwest Tasmania (Fig. 9).	en	Liebherr, James K., Porch, Nick, Shaw, Matthew, Sinclair, Bronte E., Maddison, David R. (2021): Systematic revision of the trans-Bassian moriomorphine genus Theprisa Moore (Coleoptera, Carabidae). ZooKeys 1044: 339-373, DOI: http://dx.doi.org/10.3897/zookeys.1044.62335, URL: http://dx.doi.org/10.3897/zookeys.1044.62335
A497B6DA8CCC5CA4BB4E3B6877792F9C.taxon	diagnosis	Diagnosis (n = 5). T. darlingtoni is most similar to T. montana based on large body size; 7.4 - 8.0 mm versus 7.0 - 8.9 mm for T. montana. In both species the elytra are broad basally, here HuW / MEW = 0.66 - 0.67, with the lateral margins subparallel (Figs 1 C, 2 B), and the elytral striae smooth. But T. darlingtoni exhibits a much more pronounced tubercle laterad the deep, arcuate depression bordering the pronotal median base, and the pronotal lateral margins are only briefly sinuate laterad the basal pronotal seta (Fig. 2 B), not concavely curved laterad the entire laterobasal depression as in T. montana (Fig. 1 C). The elytral lateral marginal depression is also much broader in T. darlingtoni, with the rufous coloration of the depression contrasted to the rufopiceous interval 8, whereas the narrow lateral depression of T. montana is only slightly paler than interval 8. Cuticular microsculpture is well developed, with the vertex covered by a shallow but evident transverse mesh, the pronotal disc bearing a shallow transverse mesh, sculpticell breadth 4 x length, and the elytral surface with silvery subiridescence due to the presence of dense transverse lines across the convex elytral intervals. The apical abdominal ventrite of both male and female bear three seta each side along the margin, the median pair in line with the outer two bilateral pairs.	en	Liebherr, James K., Porch, Nick, Shaw, Matthew, Sinclair, Bronte E., Maddison, David R. (2021): Systematic revision of the trans-Bassian moriomorphine genus Theprisa Moore (Coleoptera, Carabidae). ZooKeys 1044: 339-373, DOI: http://dx.doi.org/10.3897/zookeys.1044.62335, URL: http://dx.doi.org/10.3897/zookeys.1044.62335
A497B6DA8CCC5CA4BB4E3B6877792F9C.taxon	description	Description. Head robust, ocular lobe protruded, its juncture with gena obtuse; eye diameter small relative to elongate head capsule, surface not projected beyond curve of ocular lobe, EyL / OLL = 0.69 - 0.71, but ocular ratio moderate, 1.40 - 1.46, and horizontal line across eye bisecting 21 - 25 ommatidia; antennal scape robust, diameter 1.4 x apical diameter of pedicel; antennomeres 2 and 3 glabrous except for one dorsal seta on the pedicel and apical ring of setae on 3; antennae elongate, antennomere 9 maximal breadth 2.14 x length; frontal groove broad, deep, the surface of impression irregularly strigose, the two grooves laterally arcuate, defining an ovoid raised area on frons, divergent from frontoclypeal suture to lateral margin of clypeus; broadly, deeply emarginate apical margin of labrum lined with six (rarely seven) setae, six smaller setae visible along anterolateral labral margin; mentum tooth narrowly rounded apically, sides acutely divergent; maxillary stipes trisetose, the three setae on the base in either a triangle with apex upward, or in an irregular horizontal line; ligula truncate apically, narrowed basally, trumpet shaped, its two apical setae separated by three setal diameters; paraglossae elongate, total length 3 x distance from paraglossal base to ligular apical margin. Pronotum moderately transverse, lateral margins straight to slightly concave anterad acutely projected margin at basal pronotal seta; basal margin smoothly, convexly curved across width, the unmargined median base not projected beyond curve defined by distinct lateral margins posterad laterobasal depressions; median base depressed below disc, smooth basally, an arcuate line of isolated punctures each side along juncture with disc; longitudinal depression consisting of medial, deep, arcuate impression and a nearly smooth lateral tubercle, a few small punctures laterad tubercle near lateral marginal depression inside hind angles; median longitudinal impression present on median base as a series of isolated punctures, on disc consisting of lenticular depression at median base-disc juncture, and an irregular, deeply incised impression on disc; anterior transverse impression obsolete, very shallow, the median longitudinal impression continued anterad halfway across flat anterior collar of pronotum; pronotal anterior margin smooth medially, a narrow marginal bead increasingly more well developed in outer 2 / 3 of width each side; front angles protruded, tightly rounded; lateral marginal depression moderately narrow, broad enough so that cuticular sculpticells visible mesad beaded, raised margin; margin of lateral pronotal seta articulatory socket adjoining lateral marginal depression. Prosternum deeply canaliculate from prosternal process halfway to anterior prosternal margin, smooth over much of surface but with a few punctures or strigae at proepisternal suture anterad coxal cavity; proepisternum smooth, sutural groove with proepimeron smooth and deep. Elytra broadly ellipsoid, MEW / EL = 0.75 - 0.78, moderately convex, sides meeting lateral marginal depression nearly vertically, disc between striae 4 flat at midlength; basal groove arcuate, pitted at bases of striae 1 - 5, a small acute tooth present at juncture of groove and lateral marginal depression; stria 8 very deep and continuous between anterior and posterior series of lateral setae; apical carina of interval 8 narrowly upraised along stria 7, interval 8 a vertical lateral carina there; subapical sinuation abrupt, the internal plica visible ventrad deepest part of sinuation. Mesepisternum broadly punctate, ~ 19 - 22 punctures in 3 - 4 confused vertical rows. Metepisternum trapezoidal, maximum width subequal to lateral length; metepisternal-metepimeral suture complete. Legs gracile, the femora elongate and meso- and metatibiae only slightly broadened in apical half; metatarsomere 1 moderately elongate, length 0.20 x tibial length, lateral sulci present on mesal and lateral faces just dorsad the ventrolateral setae. Abdominal ventrites 2 - 6 smooth laterally, but hind margins of ventrites 2 - 5 concavely sinuate laterally, the sutures deeper in association with sinuation; suture between ventrites 1 and 2 deep, slightly curved anteriorly at midlength; suture between ventrites 2 and 3 complete laterally. Male genitalia (n = 5). Aedeagal median lobe elongate, moderately robust, base broadly open on right side, basal margin heavily sclerotized dorsad basal opening (Fig. 3 E); apical face joined to slightly arcuate ventral margin at right angle, flattened tip not extended beyond ostium, lateral surfaces of apex smooth or with only several indistinct pits; internal sac largely membranous, with only very elongate flagellum and associated basal articulatory sclerites darker (Figs 3 E, 4 E); right paramere elongate, slightly angled at midlength, apex narrowly rounded, slightly asymmetrical in outside lateral view (Fig. 5 E), bearing 10 - 19 short and long setae along ventral margin in apical half, 2 - 4 setae on dorsal surface near apex, two or three setae at apex; left paramere broadest in basal half, narrowed in apical half but broadened apically to broadly rounded tip, ventral margin with 3 - 7 setae near apex, dorsal surface glabrous or with a single seta, and apex with two or three longer setae; antecostal apodeme of abdominal segment IX angulate distally, lateral arms gracile, their distal juncture only slightly broadened (Fig. 6 E). Female reproductive tract (n = 1). Bursa copulatrix columnar, length 1.5 x maximum breadth compressed under microslide cover slip, vagina translucent, apical portion of bursa staining more darkly with Chlorazol black (Fig. 11); helminthoid sclerite present, broadly rounded apically, not extended beyond juncture with spermathecal duct; spermathecal duct thick, sclerotized, coiled sinuously to spermatheca, length twice that of annulated spermathecal reservoir; spermathecal gland duct very thin, length half that of spermathecal reservoir which it joins at reservoir base; spermathecal gland comprising sclerotized stem plus membranous reservoir bearing numerous ductules; gonocoxa bipartite, basal gonocoxite 1 with single apical fringe seta, median surface glabrous, membranous ramus present (Fig. 8 E); apical gonocoxite 2 with base extended laterally, lateral margin arcuate, apex narrowly rounded; two lateral ensiform setae and one dorsal ensiform seta present; two apical nematiform setae set in fossa in apical 1 / 4 of apical gonocoxite length.	en	Liebherr, James K., Porch, Nick, Shaw, Matthew, Sinclair, Bronte E., Maddison, David R. (2021): Systematic revision of the trans-Bassian moriomorphine genus Theprisa Moore (Coleoptera, Carabidae). ZooKeys 1044: 339-373, DOI: http://dx.doi.org/10.3897/zookeys.1044.62335, URL: http://dx.doi.org/10.3897/zookeys.1044.62335
A497B6DA8CCC5CA4BB4E3B6877792F9C.taxon	etymology	Etymology. This species honors Professor Philip J. Darlington, Jr., who among his many roles, systematically collected carabid beetles across Australia, and served as the first post-doctoral supervisor for Dr. Terry L. Erwin (Kavanaugh 2020). Terry championed a Darlingtonian world view regarding the diversification of carabid beetles (Erwin 1981), never losing sight of those questions that tied both men to primary observations made over long careers spent in the field.	en	Liebherr, James K., Porch, Nick, Shaw, Matthew, Sinclair, Bronte E., Maddison, David R. (2021): Systematic revision of the trans-Bassian moriomorphine genus Theprisa Moore (Coleoptera, Carabidae). ZooKeys 1044: 339-373, DOI: http://dx.doi.org/10.3897/zookeys.1044.62335, URL: http://dx.doi.org/10.3897/zookeys.1044.62335
A497B6DA8CCC5CA4BB4E3B6877792F9C.taxon	distribution	Distribution and habitat. Theprisa darlingtoni is known from localities spanning northern and western Tasmania, from Blue Tier on the east to Corinna in the northwest, to the Frankland Range in southwest Tasmania (Fig. 10). Collecting localities lie within the Ben Lomond, Central Highlands, and West biogeographic regions (IBRA 2004). It has been collected syntopically with T. convexa at three localities: Corinna (Bouffard, CMNH), Whyte River at Corinna Road (Darlingtons, MCZ), and Zeehan (Simson, SAMA, and Darlingtons, MCZ), although these respective 1891 and 1957 collections were separated by 66 years. The five sites at which the Darlington family collected this species (Blue Tier, Corinna and nearby Whyte River, Mt. Ben Lomond, and Zeehan) were reported to support sclerophyll forest, wet sclerophyll forest, temperate rain forest with Nothofagus, and wet gullies, with specimens at all localities collected under logs and stones, along water sources, and via immersion of leaf litter in standing water (Darlington 1960).	en	Liebherr, James K., Porch, Nick, Shaw, Matthew, Sinclair, Bronte E., Maddison, David R. (2021): Systematic revision of the trans-Bassian moriomorphine genus Theprisa Moore (Coleoptera, Carabidae). ZooKeys 1044: 339-373, DOI: http://dx.doi.org/10.3897/zookeys.1044.62335, URL: http://dx.doi.org/10.3897/zookeys.1044.62335
FAC55FB0571156529595116DA0E43957.taxon	distribution	Distribution and habitat. Theprisa montana ranges in Victoria from the Dandenong Mountains, southeast to Gunyah and Tarra Valley (Fig. 9). Philip Darlington and family recorded specimens on Mt. Donna Buang at elevations ranging 450 - 1200 m (MCZ). One specimen was sifted from litter along a stream in Nothofagus forest at 1000 m elevation on Mt. Bullfight in the Yarra Ranges (ZMUC).	en	Liebherr, James K., Porch, Nick, Shaw, Matthew, Sinclair, Bronte E., Maddison, David R. (2021): Systematic revision of the trans-Bassian moriomorphine genus Theprisa Moore (Coleoptera, Carabidae). ZooKeys 1044: 339-373, DOI: http://dx.doi.org/10.3897/zookeys.1044.62335, URL: http://dx.doi.org/10.3897/zookeys.1044.62335
5E538809D5A054AEA5E1699E3DA6C600.taxon	diagnosis	Diagnosis. This genus is diagnosable within Moriomorphini by: 1, mesosternum broad between mesocoxae; 2, mandibular scrobe present, margined ventrally by a lateral expansion; 3, mesotibia gracile, not expanded apically; 4, apical two maxillary palpomeres and apical labial palpomere apparently glabrous; 5, prosternal process not margined apicoventrally; 6, elytral striation nearly complete, striae 1 - 7 all developed apically, and striae 6 - 8 as deep or only slightly shallower than striae 1 - 5 near elytral midlength (Liebherr 2020). The mandibles are moderately elongate, with the mandibular length measured from the anterior condyle 1.7 - 1.9 x distance from anterior condyle to apicolateral margin of labrum. The ligula is broadly rounded to truncate apically, bisetose, with the two setae separated by 2 - 3 diameters of the setal articulatory sockets. The paraglossae are elongate and thin, their overall length 2 - 3 x the distance from the paraglossal base to the apical ligular margin. The mentum tooth is present, its apex narrowly rounded to subacuminate and its lateral margins upraised. Both lateral and basal pronotal setae are present. The elytra lack dorsal setae except for T. otway sp. nov., which has a single dorsal seta near midlength. The parascutellar seta is present. There are 13 lateral elytral setae associated with the eighth stria, arranged in an anterior series of seven setae and a posterior series of six, and both subapical and apical elytral setae are present. As in many moriomorphines, the apical portion of interval 8 immediately laterad stria 7 is upraised as a distinct carina. Also, the metathoracic flight wings are vestigial in all Theprisa, with the metepisternum foreshortened; its maximum width subequal to greater than the lateral length.	en	Liebherr, James K., Porch, Nick, Shaw, Matthew, Sinclair, Bronte E., Maddison, David R. (2021): Systematic revision of the trans-Bassian moriomorphine genus Theprisa Moore (Coleoptera, Carabidae). ZooKeys 1044: 339-373, DOI: http://dx.doi.org/10.3897/zookeys.1044.62335, URL: http://dx.doi.org/10.3897/zookeys.1044.62335
936E7F35464354FDA3AAE17365619711.taxon	diagnosis	Diagnosis (n = 5). This species is diagnosable among Theprisa spp. by the broad body and the presence of a dorsal elytral seta immediately mesad the third stria near elytral midlength (Fig. 1 A). The pronotum is broad, MPW / PL = 1.38 - 1.46. The pronotal median base is sparsely punctate medially, with the marginal bead greatly reduced, evidenced only as a broad, slightly depressed area along the medially expanded base, in contrast to the narrowly upraised, distinct marginal bead posterad the laterobasal depressions. Whereas elytral striae 1 - 4 are crenulate on the disc (i. e., lined by punctures that expand the strial breadth) striae 5 - 7 are progressively broader and smoother, 6 - 7 slightly wavering along length but impunctate. In contrast T. australis, T. darlingtoni, and T. montana have striae 1 - 7 smooth, and T. convexa has striae 1 - 7 punctate near midlength. Apical abdominal ventrite of male with a single seta each side along margin, female apical ventrite with two setae each side plus a median pair of subapical setae. Standardized body length is 5.9 - 6.3 mm.	en	Liebherr, James K., Porch, Nick, Shaw, Matthew, Sinclair, Bronte E., Maddison, David R. (2021): Systematic revision of the trans-Bassian moriomorphine genus Theprisa Moore (Coleoptera, Carabidae). ZooKeys 1044: 339-373, DOI: http://dx.doi.org/10.3897/zookeys.1044.62335, URL: http://dx.doi.org/10.3897/zookeys.1044.62335
936E7F35464354FDA3AAE17365619711.taxon	description	Description. Head narrow, ocular lobe little projected, juncture with gena very obtuse; eyes small, EyL / OLL = 0.65 - 0.70, not projected beyond curvature of ocular lobe, 17 - 20 ommatidia bisected by a horizontal line across eye; antennomeres 2 and 3 glabrous except for one dorsal seta on 2 and an apical ring of setae on 3; antennae moderately elongate, antennomere 9 maximal breadth 2 x length; frontal groove deeply, medially arcuate from anterior supraorbital seta to frontoclypeal suture and continued onto lateral reaches of clypeus, area laterad groove distinctly convex; slightly concave apical labral margin 6 - setose, with three smaller setae lining the lateroapical margin; mentum tooth narrowly rounded apically, sides subparallel; maxillary stipes trisetose, the three setae on the base in either a triangle with apex upward, or in an irregular horizontal line; ligula slightly convex apically, narrowed basally, trumpet shaped, its two apical setae separated by three setal diameters; paraglossae elongate, total length 2.5 x distance from paraglossal base to ligular apical margin. Pronotum transverse, lateral margins straight anterad projection defined by articulatory socket of basal seta; basal margin nearly straight, the obsoletely margined median base projected posterad only slightly beyond the lateral beaded margins posterad the laterobasal depressions; median base smooth at middle, ~ 10 small punctures each side mesad deepest point of laterobasal depression; pronotal disc extended to basal marginal bead defining a tubercle that divides laterobasal depression into a median longitudinal groove and a broad lateral marginal depression inside the basal seta; median longitudinal depression deepest just anterad median base, deeply incised to very shallow, broad anterior transverse impression; anterior callosity nearly flat, front margin smooth medially, margined in outer half each side; front angles broadly, slightly protruded, subangulate; lateral marginal depression very narrow from front angle to basal 1 / 4 of length where it expands to meet laterobasal depression; lateral pronotal seta positioned one setal diameter inside lateral marginal depression. Prosternum medially depressed mesad anterior margins of procoxal cavities, up to 16 punctures present in a transverse band across apical 1 / 4 of prosternum, ~ 10 punctures each side of prosternum in depressed area anterad coxal cavity; proepisternum smooth, sutural groove with proepimeron smooth and deep. Elytra broadly hemi-ovoid, MEW / EL = 0.81 - 0.86, convex, sides vertically meeting lateral marginal depression; basal groove arcuate, juncture with lateral marginal depression tightly rounded, a broad, blunt tooth on margin at juncture; lateral marginal depression narrow; stria 8 deep, continuous between anterior and posterior series of lateral setae; apical carina of interval 8 narrowly upraised along stria 7, interval 8 a vertical lateral carina there; subapical sinuation evident, the internal plica visible ventrad deepest part of sinuation. Mesepisternum with ~ 8 shallow punctures in two vertical rows. Metepisternum wider than long, maximum width 1.8 x lateral length; metepimeron fused to metepisternum laterally. Legs gracile, femora narrow, meso- and metatibiae little expanded apically, of consistent diameter throughout apical half of length; metatarsomere 1 elongate, length 0.22 x tibial length, lateral sulci present on mesal and lateral faces just dorsad the ventrolateral setae. Abdominal ventrites 2 - 6 with 1 - 2 longitudinal wrinkles laterally; suture between ventrites 1 and 2 slightly curved anteriorly at midlength, ventrite 2 wrinkled within curve; suture between ventrites 2 and 3 complete laterally. Male genitalia (n = 2). Aedeagal median lobe robust, base broadly open on right side, basal margin thickened dorsad basal opening (Fig. 3 A); median lobe apex broadly rounded, slightly extended beyond ostium with a large apical divot-like concavity at tip, surface of apex densely covered with pits; internal sac bearing a stout flagellum, with apex concavely scooped (Fig. 4 A), a basal articulatory sclerite associated with flagellum; right paramere elongate, parallel sided in apical half with apex narrowly rounded (Fig. 5 A), bearing 12 - 15 short setae along ventral margin in apical half, 2 - 4 setae on dorsal surface near apex, though with apex glabrous; left paramere broadest near midlength, apex narrower, parallel sided with rounded tip, ventral margin bearing 2 - 4 setae in apical 1 / 3, dorsal surface with two or three setae near apex, and apex with none or one seta; antecostal apodeme of abdominal segment IX rounded distally, the apical juncture of lateral arms broad (Fig. 6 A). Female reproductive tract (n = 2). Bursa copulatrix columnar, length 1.25 x maximum breadth compressed under microslide cover slip, vagina translucent, broader apical portion of bursa staining more darkly with Chlorazol black (Fig. 7 A); helminthoid sclerite present, rounded apically, not extended beyond juncture with spermathecal duct; spermathecal duct stout, sinuously recurved to meet spermatheca, length subequal to length of annulated spermathecal reservoir; spermathecal gland duct very thin, length twice that of spermathecal reservoir which it joins at reservoir base; spermathecal gland comprising sclerotized stem plus membranous reservoir bearing numerous ductules; gonocoxa bipartite, basal gonocoxite 1 with two apical fringe setae in one dissected specimen, two setae on right gonocoxa and one on left in second dissected specimen, median surface glabrous, membranous ramus present (Fig. 8 A); apical gonocoxite 2 with base extended laterally, lateral margin arcuate, apex falciform; two lateral ensiform setae and one dorsal ensiform present; two apical nematiform setae set in fossa at apical 1 / 4 of apical gonocoxite length.	en	Liebherr, James K., Porch, Nick, Shaw, Matthew, Sinclair, Bronte E., Maddison, David R. (2021): Systematic revision of the trans-Bassian moriomorphine genus Theprisa Moore (Coleoptera, Carabidae). ZooKeys 1044: 339-373, DOI: http://dx.doi.org/10.3897/zookeys.1044.62335, URL: http://dx.doi.org/10.3897/zookeys.1044.62335
936E7F35464354FDA3AAE17365619711.taxon	etymology	Etymology. The species epithet is the mountain range from which this species is described, and based on current knowledge, precinctive. The epithet is to be treated as a noun in apposition.	en	Liebherr, James K., Porch, Nick, Shaw, Matthew, Sinclair, Bronte E., Maddison, David R. (2021): Systematic revision of the trans-Bassian moriomorphine genus Theprisa Moore (Coleoptera, Carabidae). ZooKeys 1044: 339-373, DOI: http://dx.doi.org/10.3897/zookeys.1044.62335, URL: http://dx.doi.org/10.3897/zookeys.1044.62335
936E7F35464354FDA3AAE17365619711.taxon	distribution	Distribution and habitat. Theprisa otway is restricted to the Otway Ranges (Fig. 9), with specimens collected in wet to mesic sclerophyll forest from 80 - 525 m elevations. The beetles are terrestrial, being discovered via raking litter, in a Berlese extraction of leaf and log litter, and via application of pyrethrin fog to logs of Eucalyptus regnans F. Mueller covered with fungus. The MCZ specimens were collected by the Darlington family in sclerophyll forest approaching rain forest in ravines, with some Nothofagus (Darlington 1960).	en	Liebherr, James K., Porch, Nick, Shaw, Matthew, Sinclair, Bronte E., Maddison, David R. (2021): Systematic revision of the trans-Bassian moriomorphine genus Theprisa Moore (Coleoptera, Carabidae). ZooKeys 1044: 339-373, DOI: http://dx.doi.org/10.3897/zookeys.1044.62335, URL: http://dx.doi.org/10.3897/zookeys.1044.62335
