taxonID	type	description	language	source
AC7B87B09755FFD287A450186C85CFFA.taxon	type_taxon	TYPE SPECIES. — Limentinus aldabranus Distant, 1917, by original designation and monotypy.	en	Gnezdilov, Vladimir M. (2021): New species of the genera Limentinus Distant, 1917 and Calodia Nielson, 1982 (Hemiptera, Auchenorrhyncha, Cicadellidae, Coelidiinae) from the Makay Massif of Madagascar, with a key to Malagasy species. Zoosystema 43 (16): 297-310, DOI: 10.5252/zoosystema2021v43a16
AC7B87B09755FFD287A450186C85CFFA.taxon	distribution	DISTRIBUTION. — Outer Seychelles and Madagascar.	en	Gnezdilov, Vladimir M. (2021): New species of the genera Limentinus Distant, 1917 and Calodia Nielson, 1982 (Hemiptera, Auchenorrhyncha, Cicadellidae, Coelidiinae) from the Makay Massif of Madagascar, with a key to Malagasy species. Zoosystema 43 (16): 297-310, DOI: 10.5252/zoosystema2021v43a16
AC7B87B09755FFD287A450186C85CFFA.taxon	diagnosis	DIAGNOSIS. — Head short, together with eyes narrower than pronotum; anterior margin rounded. Face long; frontoclypeus almost twice as long as wide between the eyes, without carinae; lorae narrowly oval; anteclypeus distinctly (mushroomlike) enlarged apically, with straight basement (Fig. 6 A); antennal pedicel small, cylindrical; ocelli prominent, on margin of face and crown; rostrum with short 3 rd segment, reaching middle coxae. Crown with lateral margins diverging anteriorly with weak median carina. Pronotum without carinae, anterior margin convex; posterior margin concave. Mesonotum large, with scutellum well separated by transverse depression; disk of pronotum separated by carina from its lateral parts (paranotal lobes). Forewings elongate, with five apical and three anteapical cells, outer one closed; appendix well developed. Hind femora with 2 + 2 macrosetae apically. First metatarsomere as long as second and third ones combined. First metatarsomere with 2 + 1 macrosetae dorsally – two apically and one laterally. First metatarsomere with two spines laterally and two platellae between them ventroapically (Fig. 5 A). Second metatarsomere with two spines laterally and one or two platellae between them ventroapically. Claws wide, each with single long lateral seta. Aedeagus tubular, asymmetrical, with long and narrow shaft bearing spiny processes. Connective short and broad, arcuated.	en	Gnezdilov, Vladimir M. (2021): New species of the genera Limentinus Distant, 1917 and Calodia Nielson, 1982 (Hemiptera, Auchenorrhyncha, Cicadellidae, Coelidiinae) from the Makay Massif of Madagascar, with a key to Malagasy species. Zoosystema 43 (16): 297-310, DOI: 10.5252/zoosystema2021v43a16
AC7B87B09755FFD48799559F6F9DCC35.taxon	description	(Figs 1; 2; 5)	en	Gnezdilov, Vladimir M. (2021): New species of the genera Limentinus Distant, 1917 and Calodia Nielson, 1982 (Hemiptera, Auchenorrhyncha, Cicadellidae, Coelidiinae) from the Makay Massif of Madagascar, with a key to Malagasy species. Zoosystema 43 (16): 297-310, DOI: 10.5252/zoosystema2021v43a16
AC7B87B09755FFD48799559F6F9DCC35.taxon	materials_examined	TYPE MATERIAL. — Examined from photos taken by Mr Fauvre. Holotype. Madagascar • sex unknown as the abdomen is missed; Périnet, Sahamaloto; 13 - 17. I. 1949; Inst. Scient. Madagascar, PC; MNHN-EH-EH 2701. MATERIAL EXAMINED. — Madagascar • 1 ♂; Périnet; Seyrig; 12. II. 1931; ZIN.	en	Gnezdilov, Vladimir M. (2021): New species of the genera Limentinus Distant, 1917 and Calodia Nielson, 1982 (Hemiptera, Auchenorrhyncha, Cicadellidae, Coelidiinae) from the Makay Massif of Madagascar, with a key to Malagasy species. Zoosystema 43 (16): 297-310, DOI: 10.5252/zoosystema2021v43a16
AC7B87B09755FFD48799559F6F9DCC35.taxon	diagnosis	DIAGNOSIS. — Aedeagal shaft with three spine-shaped straight processes subapically on right side and one spine-shaped straight subapical process on left side and with two short and two long spine-shaped slightly curved processes below on right and left sides (Fig. 5 E, F).	en	Gnezdilov, Vladimir M. (2021): New species of the genera Limentinus Distant, 1917 and Calodia Nielson, 1982 (Hemiptera, Auchenorrhyncha, Cicadellidae, Coelidiinae) from the Makay Massif of Madagascar, with a key to Malagasy species. Zoosystema 43 (16): 297-310, DOI: 10.5252/zoosystema2021v43a16
AC7B87B09755FFD48799559F6F9DCC35.taxon	description	DESCRIPTION Structure As mentioned for the genus. Coloration (Figs 1; 2) Face yellow, with two wide black stripes running from ocelli throughout frontoclypeus and fused in one stripe on anteclypeus (Figs 1 C; 2 C); apical enlarged part of anteclypeus with brown yellow margins; lorae black, except brown yellow outer margins; genae with wide longitudinal black stripe under each eye running to level of lorae; antennae with scapus, pedicel, and flagellum light yellow; ocelli dark brown; rostrum brownish yellow. Crown black, except brown yellow lateral margins (Figs 1 A; 2 A). Pronotum and mesonotum black. Thorax below and coxae mainly black. Forewings dark brown to black, with black veins; corium with light brown yellow area along the costal margin almost from the basal cell to outer anteapical cell and with light brown yellow spot below the apex of clavus (Evans 1953: fig. 44) (Figs 1 B; 2 B); apical cells with appendix brown cinereous. Fore and middle legs light yellow. Middle trochanters light yellow, with dark brown inner sides. Hind trochanters brown yellow. Hind femora dark brown to black in its basal 2 / 3 and light yellow in its apical third. Hind tibiae dark brown, with light setae. Hind tarsi brown yellow. Claws dark brown. Abdominal segments black, with light yellow hind margins. Genital block black, except light yellow lateral margin apically of subgenital plate. Male genitalia (Fig. 5) Anal tube 1.5 times as long as wide, flattened ventrally. Anal column short. Pygofer lobe triangularly elongate, with an additional small lobe on apical angle under anal tube and with ventral lobe in its middle (Fig. 5 B); apical lobe with seven macrosetae subapically on its inner side (Fig. 5 C). Subgenital plate long and narrow, not narrowing apically, with microsetae apically and with long setae on main surface, including several subapical macrosetae (Fig. 5 D). Aedeagus asymmetrical, shaft long and narrow, slightly curved, with small hook apically bearing marginal denticles (Fig. 5 E-G), with three (2 + 1) spine-shaped straight processes subapically on right side and one spine-shaped straight subapical process on left side and with two short and two long (one is damaged) spineshaped slightly curved processes below on right and left sides respectively (Fig. 5 E, F); gonopore probably subapical. Styles elongate, narrowing apically, with rounded apices (Fig. 5 H). Connective arcuate (Fig. 5 I). Measurement Total length from the apex of head to the apices of forewings: 7.0 mm.	en	Gnezdilov, Vladimir M. (2021): New species of the genera Limentinus Distant, 1917 and Calodia Nielson, 1982 (Hemiptera, Auchenorrhyncha, Cicadellidae, Coelidiinae) from the Makay Massif of Madagascar, with a key to Malagasy species. Zoosystema 43 (16): 297-310, DOI: 10.5252/zoosystema2021v43a16
AC7B87B09753FFD7855556DE6E0ECC96.taxon	description	(Figs 4 A, B; 6) urn: lsid: zoobank. org: act: 37784 A 8 A- 30 EE- 4 C 8 F- 91 B 2 - 8182 E 2256259	en	Gnezdilov, Vladimir M. (2021): New species of the genera Limentinus Distant, 1917 and Calodia Nielson, 1982 (Hemiptera, Auchenorrhyncha, Cicadellidae, Coelidiinae) from the Makay Massif of Madagascar, with a key to Malagasy species. Zoosystema 43 (16): 297-310, DOI: 10.5252/zoosystema2021v43a16
AC7B87B09753FFD7855556DE6E0ECC96.taxon	materials_examined	TYPE MATERIAL. — Holotype. Madagascar • ♂; Toliara Province, Massif du Makay; 21 ° 36.144 ’ S, 45 ° 06.738 ’ E; 11. I. 2011; V. M. Gnezdilov leg.; ZIN. Paratype. Madagascar • ♂; Toliara Province, Massif du Makay; 243 m; 21 ° 36 ’ 236 ” S, 45 ° 06 ’ 464 ” E; forêt sèche; 12. I. 2011; D. Ouvrard leg.; MNHN-EH-EH 24763.	en	Gnezdilov, Vladimir M. (2021): New species of the genera Limentinus Distant, 1917 and Calodia Nielson, 1982 (Hemiptera, Auchenorrhyncha, Cicadellidae, Coelidiinae) from the Makay Massif of Madagascar, with a key to Malagasy species. Zoosystema 43 (16): 297-310, DOI: 10.5252/zoosystema2021v43a16
AC7B87B09753FFD7855556DE6E0ECC96.taxon	etymology	ETYMOLOGY. — The species name is referring to similarity in face coloration with Oryx gazella (L.) (Mammalia).	en	Gnezdilov, Vladimir M. (2021): New species of the genera Limentinus Distant, 1917 and Calodia Nielson, 1982 (Hemiptera, Auchenorrhyncha, Cicadellidae, Coelidiinae) from the Makay Massif of Madagascar, with a key to Malagasy species. Zoosystema 43 (16): 297-310, DOI: 10.5252/zoosystema2021v43a16
AC7B87B09753FFD7855556DE6E0ECC96.taxon	diagnosis	DIAGNOSIS. — Aedeagal shaft with two spine-shaped processes subapically on right side and one spine-shaped subapical process on left side and with two longer spine-shaped processes below on left side (Fig. 6 D, E).	en	Gnezdilov, Vladimir M. (2021): New species of the genera Limentinus Distant, 1917 and Calodia Nielson, 1982 (Hemiptera, Auchenorrhyncha, Cicadellidae, Coelidiinae) from the Makay Massif of Madagascar, with a key to Malagasy species. Zoosystema 43 (16): 297-310, DOI: 10.5252/zoosystema2021v43a16
AC7B87B09753FFD7855556DE6E0ECC96.taxon	discussion	TAXONOMIC RELATIONSHIPS. — Very close to L. declinatus Wang, Dietrich & Zhang, 2018 in coloration, but well distinguished by lighter forewings and dense brown dots on pronotum and by fewer spiny processes of aedeagus – five processes in comparison to 12 in L. declinatus.	en	Gnezdilov, Vladimir M. (2021): New species of the genera Limentinus Distant, 1917 and Calodia Nielson, 1982 (Hemiptera, Auchenorrhyncha, Cicadellidae, Coelidiinae) from the Makay Massif of Madagascar, with a key to Malagasy species. Zoosystema 43 (16): 297-310, DOI: 10.5252/zoosystema2021v43a16
AC7B87B09753FFD7855556DE6E0ECC96.taxon	description	DESCRIPTION Structure Structure as mentioned for the genus. Crown narrow and long, 1.5 times as long at midline as wide basally. Pronotum nearly as long as crown medially. Mesonotum 1.5 times as long as pronotum medially. Second metatarsomere with two spines laterally and one platella between them ventroapically. Apodemes of second abdominal segment rather long and wide (Fig. 6 C). Coloration (Fig. 4 A, B) Face with frontoclypeus light yellow, with two longitudinal black stripes laterally; lorae black, with light yellow margins; anteclypeus light yellow, with wide lateral black stripes converging subapically; black spot under each eye; ocelli dark brown; antennae light yellow. Legs light yellow. Crown brown, with wide longitudinal black stripes on the sides of median carina. Pronotum black, with dense light brown dots, sometimes fused in the areas; paranotal lobes of pronotum light yellow, each with black spot in its lower part. Mesonotum black, with sparse brown dots and with four black spots on brown yellowish background medially. Episternae and epimerae of thorax black, with light yellow margins. Scutellum with brown yellowish apex. Forewings with dark brown to black veins, brown to dark brown cells of corium and clavus in its basal 2 / 3 of length, with brown dots on veins in apical third of wing and with two separated light brown yellow areas – one narrow along the costal margin from the basal cell to apical third of inner anteapical cell and another one, wide, from costal margin throughout outer anteapical cell; appendix of forewings cinereous. Hindwings cinereous, with light spot on costal margin and dark brown veins. Hind tibiae with black apices of spines. Claws dark brown to black. Abdominal tergites dark brown to black, each with pair of large orange yellow spots laterally running from its anterior margin to its yellow posterior margin. Genital block dark brown. Abdominal sternites dark brown to black, with brown yellowish hind margins. Male genitalia (Fig. 6) Anal tube 1.5 times as long as wide, flattened ventrally (Fig. 6 F). Anal column short. Pygofer lobe triangularly elongate, with an additional small lobe on apical angle under the anal tube and with ventral lobe in its basal half (Fig. 6 B). Subgenital plate long and narrow, with around 12 microsetae apically and with long setae on the main surface. Aedeagus asymmetrical, shaft long and narrow, curved, with small hook apically (Fig. 6 D), with two spine-shaped slightly curved processes subapically on right side and one spine-shaped slightly curved subapical process on left side and with two longer slightly curved spine-shaped processes below on left side (Fig. 6 E), gonopore probably subapical. Styles elongate, narrowing apically, with rounded apices (Fig. 6 G). Connective arcuate (Fig. 6 H). Measurement Total length. 6.0 mm.	en	Gnezdilov, Vladimir M. (2021): New species of the genera Limentinus Distant, 1917 and Calodia Nielson, 1982 (Hemiptera, Auchenorrhyncha, Cicadellidae, Coelidiinae) from the Makay Massif of Madagascar, with a key to Malagasy species. Zoosystema 43 (16): 297-310, DOI: 10.5252/zoosystema2021v43a16
AC7B87B09750FFD78523557F6B66CD95.taxon	description	(Figs 3 A, B; 7) urn: lsid: zoobank. org: act: E 7 B 8 D 25 D- 6557 - 46 AC- 9 AB 3 - 12 CD 2999 CEE 7	en	Gnezdilov, Vladimir M. (2021): New species of the genera Limentinus Distant, 1917 and Calodia Nielson, 1982 (Hemiptera, Auchenorrhyncha, Cicadellidae, Coelidiinae) from the Makay Massif of Madagascar, with a key to Malagasy species. Zoosystema 43 (16): 297-310, DOI: 10.5252/zoosystema2021v43a16
AC7B87B09750FFD78523557F6B66CD95.taxon	materials_examined	TYPE MATERIAL. — Holotype. Madagascar • ♂; Toliara Province, Massif du Makay; 21 ° 36.415 ’ S, 45 ° 06.769 ’ E; 12. I. 2011; sweeping in dry forest; V. M. Gnezdilov leg.; ZIN.	en	Gnezdilov, Vladimir M. (2021): New species of the genera Limentinus Distant, 1917 and Calodia Nielson, 1982 (Hemiptera, Auchenorrhyncha, Cicadellidae, Coelidiinae) from the Makay Massif of Madagascar, with a key to Malagasy species. Zoosystema 43 (16): 297-310, DOI: 10.5252/zoosystema2021v43a16
AC7B87B09750FFD78523557F6B66CD95.taxon	etymology	ETYMOLOGY. — The species is named in honour of famous American specialist on coelidiine leafhoppers William Mervin Nielson (1927 - 2019) who described most of Malagasy Limentinus species.	en	Gnezdilov, Vladimir M. (2021): New species of the genera Limentinus Distant, 1917 and Calodia Nielson, 1982 (Hemiptera, Auchenorrhyncha, Cicadellidae, Coelidiinae) from the Makay Massif of Madagascar, with a key to Malagasy species. Zoosystema 43 (16): 297-310, DOI: 10.5252/zoosystema2021v43a16
AC7B87B09750FFD78523557F6B66CD95.taxon	diagnosis	DIAGNOSIS. — Aedeagal shaft with three straight short processes apically, two spine-shaped slightly curved processes subapically on right and left sides (right processes longer) and two spine-shaped slightly curved processes (longest ones) below on left side (Fig. 7 D-F). DESCRIPTION Structure In structure and coloration very similar to L. oryx n. sp. Second metatarsomere with two spines laterally and two platellae between them ventroapically. Coloration (Fig. 3 A, B) Scutellum with brown yellowish margins. Abdominal tergites black, each with pair of large orange yellow spots laterally running from its anterior margin to its yellow posterior margin. Abdominal sternites black, with yellow hind margins. Male genitalia (Fig. 7) Anal tube 1.5 times as long as wide, flattened ventrally (Fig. 7 G, H). Anal column short. Pygofer covered by long setae; pygofer lobe triangularly elongate, with an additional small lobe on its apical angle under the anal tube and with ventral lobe in its basal half (Fig. 7 A, B); apical lobes of pygofer each with eight and more setae on its inner side (Fig. 7 B). Subgenital plate long and narrow, with microsetae apically and with long setae on the main surface. Aedeagus asymmetrical, shaft long and narrow, slightly curved (in ventral and lateral views), with small hook apically (in lateral view) (Fig. 7 D), with three straight short processes apically, two spine-shaped slightly curved processes subapically on right and left sides (right processes longer) and two spine-shaped slightly curved processes (longest ones) below on left side (Fig. 7 D-F), gonopore subapical, large (Fig. 7 F). Styles small, elongate, with rounded apices (Fig. 7 D). Connective arcuate (Fig. 7 C). Measurement Total length. 6.0 mm.	en	Gnezdilov, Vladimir M. (2021): New species of the genera Limentinus Distant, 1917 and Calodia Nielson, 1982 (Hemiptera, Auchenorrhyncha, Cicadellidae, Coelidiinae) from the Makay Massif of Madagascar, with a key to Malagasy species. Zoosystema 43 (16): 297-310, DOI: 10.5252/zoosystema2021v43a16
AC7B87B09750FFD987CD567E6B66CF76.taxon	description	(Figs 3 C-F; 8) urn: lsid: zoobank. org: act: C 0339619 - CA 8 C- 452 D-A 09 D- 0832 C 636880 E	en	Gnezdilov, Vladimir M. (2021): New species of the genera Limentinus Distant, 1917 and Calodia Nielson, 1982 (Hemiptera, Auchenorrhyncha, Cicadellidae, Coelidiinae) from the Makay Massif of Madagascar, with a key to Malagasy species. Zoosystema 43 (16): 297-310, DOI: 10.5252/zoosystema2021v43a16
AC7B87B09750FFD987CD567E6B66CF76.taxon	materials_examined	TYPE MATERIAL. — Holotype. Madagascar • ♂; Toliara Province, Massif du Makay; 21 ° 37.783 ’ S, 45 ° 00.286 ’ E; 170 m; 23. I. 2011; sweeping in dry forest; V. M. Gnezdilov leg.; ZIN. Paratype. Madagascar • ♂; Toliara Province, Massif du Makay, forêt d’Ambalamanga Sud; 21 ° 40.493 ’ S, 44 ° 59.603 ’ E; 157 m; 20. I. 2011; sweeping in dry forest; V. M. Gnezdilov leg.; MNHN.	en	Gnezdilov, Vladimir M. (2021): New species of the genera Limentinus Distant, 1917 and Calodia Nielson, 1982 (Hemiptera, Auchenorrhyncha, Cicadellidae, Coelidiinae) from the Makay Massif of Madagascar, with a key to Malagasy species. Zoosystema 43 (16): 297-310, DOI: 10.5252/zoosystema2021v43a16
AC7B87B09750FFD987CD567E6B66CF76.taxon	etymology	ETYMOLOGY. — The species name refers to the coloration of the face.	en	Gnezdilov, Vladimir M. (2021): New species of the genera Limentinus Distant, 1917 and Calodia Nielson, 1982 (Hemiptera, Auchenorrhyncha, Cicadellidae, Coelidiinae) from the Makay Massif of Madagascar, with a key to Malagasy species. Zoosystema 43 (16): 297-310, DOI: 10.5252/zoosystema2021v43a16
AC7B87B09750FFD987CD567E6B66CF76.taxon	diagnosis	DIAGNOSIS. — Aedeagal shaft with one short subapical process on left side, two long spine-shaped processes below it, and two long spine-shaped processes on opposite right side (Fig. 8 D, E).	en	Gnezdilov, Vladimir M. (2021): New species of the genera Limentinus Distant, 1917 and Calodia Nielson, 1982 (Hemiptera, Auchenorrhyncha, Cicadellidae, Coelidiinae) from the Makay Massif of Madagascar, with a key to Malagasy species. Zoosystema 43 (16): 297-310, DOI: 10.5252/zoosystema2021v43a16
AC7B87B09750FFD987CD567E6B66CF76.taxon	discussion	TAXONOMIC RELATIONSHIPS. — In general the male genitalia structure is close to L. aldabranus Distant sensu Nielson (1982), but is well differentiated by its black face and details of the aedeagus.	en	Gnezdilov, Vladimir M. (2021): New species of the genera Limentinus Distant, 1917 and Calodia Nielson, 1982 (Hemiptera, Auchenorrhyncha, Cicadellidae, Coelidiinae) from the Makay Massif of Madagascar, with a key to Malagasy species. Zoosystema 43 (16): 297-310, DOI: 10.5252/zoosystema2021v43a16
AC7B87B09750FFD987CD567E6B66CF76.taxon	description	DESCRIPTION Structure General structure as mentioned for the genus. Crown wider than in other species described, as long as wide basally. Second metatarsomere with two spines laterally and two platella between them ventroapically. Coloration (holotype) (Fig. 3 C, D) Head and body from below black. Legs black, with burgundy tint. Crown with brown yellowish median line. Pronotum with brown yellowish areas. Mesonotum black. Scutellum brown yellowish. Forewings cinereous to black, with black to dark brown veins. Hindwings cinereous, with brown to dark brown veins. First metatarsomere brown yellowish in its basal half and black in its apical half. Abdominal segments including genital block black, with light yellow hind margins. Coloration (paratype) (Fig. 3 E, F) Face black; anteclypeus with wide brown yellowish median line; lorae with brown yellowish margins; genae yellow along frontoclypeus and under antennae; rostrum with black 1 st visible segment and brown yellowish 2 nd one. Crown brown yellowish, with black longitudinal areas. Pronotum brown yellowish, except black paranotal lobes. Mesonotum brown yellowish. Forewings brown yellowish, with dark brown veins. Episternae and epimerae with yellow margins. Katepisternae yellow, with dark brown to black central part. Fore and middle coxae dark brown. Hind coxae black. Fore and middle femora and tibiae dark brown, with red tint. Hind femora and tarsomeres brown yellowish. Male genitalia (Fig. 8) Anal tube 1.4 times as long as wide, flattened ventrally (Fig. 8 B). Anal column short. Pygofer covered by long setae; pygofer lobe triangularly elongate, with an additional small lobe on apical angle under anal tube and with ventral lobe in its basal half (Fig. 8 A, C); apical pygofer lobe apically with macrosetae on its inner side (Fig. 8 C). Subgenital plate long and narrow, with long setae on the main surface. Aedeagus asymmetrical; shaft long and narrow, weakly curved, with large hook apically bearing five small lateral denticles on each side (in ventral and lateral views) (Fig. 8 D-G), with one short subapical process on left side, two long spine-shaped slightly curved processes below it, and two long spine-shaped slightly curved processes in opposite on right side, gonopore situated below long processes of shaft, small (Fig. 8 D). Styles small, rounded (Fig. 8 D, E). Connective arcuate. Measurement Total length. 6.0 mm.	en	Gnezdilov, Vladimir M. (2021): New species of the genera Limentinus Distant, 1917 and Calodia Nielson, 1982 (Hemiptera, Auchenorrhyncha, Cicadellidae, Coelidiinae) from the Makay Massif of Madagascar, with a key to Malagasy species. Zoosystema 43 (16): 297-310, DOI: 10.5252/zoosystema2021v43a16
AC7B87B0975EFFD987D7551F6B19CEB3.taxon	type_taxon	TYPE SPECIES. — Calodia multipectinata Nielson, 1982, by original designation.	en	Gnezdilov, Vladimir M. (2021): New species of the genera Limentinus Distant, 1917 and Calodia Nielson, 1982 (Hemiptera, Auchenorrhyncha, Cicadellidae, Coelidiinae) from the Makay Massif of Madagascar, with a key to Malagasy species. Zoosystema 43 (16): 297-310, DOI: 10.5252/zoosystema2021v43a16
AC7B87B0975EFFD987D7551F6B19CEB3.taxon	distribution	DISTRIBUTION. — Oriental Region and Madagascar.	en	Gnezdilov, Vladimir M. (2021): New species of the genera Limentinus Distant, 1917 and Calodia Nielson, 1982 (Hemiptera, Auchenorrhyncha, Cicadellidae, Coelidiinae) from the Makay Massif of Madagascar, with a key to Malagasy species. Zoosystema 43 (16): 297-310, DOI: 10.5252/zoosystema2021v43a16
AC7B87B0975EFFD987D7551F6B19CEB3.taxon	diagnosis	DIAGNOSIS. — As given by Nielson (1982, 2015). In general characters close to Limentinus mentioned above, but well differentiated in male genitalia structure by the aedeagal shaft with fewer processes and larger styles (Fig. 9 C, F-H).	en	Gnezdilov, Vladimir M. (2021): New species of the genera Limentinus Distant, 1917 and Calodia Nielson, 1982 (Hemiptera, Auchenorrhyncha, Cicadellidae, Coelidiinae) from the Makay Massif of Madagascar, with a key to Malagasy species. Zoosystema 43 (16): 297-310, DOI: 10.5252/zoosystema2021v43a16
AC7B87B0975FFFDF8523571E6E0ECFB7.taxon	description	(Figs 4 C, D; 9) urn: lsid: zoobank. org: act: 064 F 1 C 60 - 0996 - 483 D- 9 C 69 - FE 68 DEF 923 EB	en	Gnezdilov, Vladimir M. (2021): New species of the genera Limentinus Distant, 1917 and Calodia Nielson, 1982 (Hemiptera, Auchenorrhyncha, Cicadellidae, Coelidiinae) from the Makay Massif of Madagascar, with a key to Malagasy species. Zoosystema 43 (16): 297-310, DOI: 10.5252/zoosystema2021v43a16
AC7B87B0975FFFDF8523571E6E0ECFB7.taxon	materials_examined	TYPE MATERIAL. — Holotype. Madagascar • ♂; Toliara Province, Massif du Makay; 21 ° 33.901 ’ S, 45 ° 04.456 ’ E; 312 m; 15. I. 2011; sweeping in wet forest; V. M. Gnezdilov leg.; ZIN.	en	Gnezdilov, Vladimir M. (2021): New species of the genera Limentinus Distant, 1917 and Calodia Nielson, 1982 (Hemiptera, Auchenorrhyncha, Cicadellidae, Coelidiinae) from the Makay Massif of Madagascar, with a key to Malagasy species. Zoosystema 43 (16): 297-310, DOI: 10.5252/zoosystema2021v43a16
AC7B87B0975FFFDF8523571E6E0ECFB7.taxon	etymology	ETYMOLOGY. — The species is named after the Makay Massif.	en	Gnezdilov, Vladimir M. (2021): New species of the genera Limentinus Distant, 1917 and Calodia Nielson, 1982 (Hemiptera, Auchenorrhyncha, Cicadellidae, Coelidiinae) from the Makay Massif of Madagascar, with a key to Malagasy species. Zoosystema 43 (16): 297-310, DOI: 10.5252/zoosystema2021v43a16
AC7B87B0975FFFDF8523571E6E0ECFB7.taxon	diagnosis	DIAGNOSIS. — Aedeagal shaft with only pair of different in shape and length spine-shaped subapical processes (Fig. 9 G, H).	en	Gnezdilov, Vladimir M. (2021): New species of the genera Limentinus Distant, 1917 and Calodia Nielson, 1982 (Hemiptera, Auchenorrhyncha, Cicadellidae, Coelidiinae) from the Makay Massif of Madagascar, with a key to Malagasy species. Zoosystema 43 (16): 297-310, DOI: 10.5252/zoosystema2021v43a16
AC7B87B0975FFFDF8523571E6E0ECFB7.taxon	discussion	TAXONOMIC RELATIONSHIPS. — According to the shape of aedeagus with right subapical process longer then left one and aedeagal apex with carina below gonopore (Fig. 9 G, H) the species is closely related to Indian Calodia deergha Viraktamath et Meshram, 2019 (Viraktamath & Meshram 2019: figs 15 A-H) and to C. ostenta (Distant, 1918) (Nielson 1982, figs 469, 470; Viraktamath & Meshram 2019, figs 20 A-H), however, it is well differentiated by the pygofer lobes without ventral lobes and long styles (Fig. 9 A, C). The peculiar long style, with median process (Fig. 9 F), separates the new species from other Calodia species as well as from all other Coelidiini. Long and narrow styles, but without median processes, are known also for the African genera Africocoelidia Nielson, 1982 and Amplicoelidia Nielson, 1991 (Nielson 1991, figs 11, 17).	en	Gnezdilov, Vladimir M. (2021): New species of the genera Limentinus Distant, 1917 and Calodia Nielson, 1982 (Hemiptera, Auchenorrhyncha, Cicadellidae, Coelidiinae) from the Makay Massif of Madagascar, with a key to Malagasy species. Zoosystema 43 (16): 297-310, DOI: 10.5252/zoosystema2021v43a16
AC7B87B0975FFFDF8523571E6E0ECFB7.taxon	description	DESCRIPTION Structure Crown slightly longer than wide basally. Pronotum 1.3 times as long as coryphe medially. Mesonotum 1.5 times as long as pronotum. Second metatarsomere with two spines laterally and two platellae between them ventroapically. Coloration (Fig. 4 C, D) Face with frontoclypeus light yellow, with dense brown dots; genae light brownish yellow; lorae, anteclypeus, legs, and body from below light yellow; ocelli burgundy; crown light yellow. Pronotum and mesonotum dark brown, with dense light yellow dots; paranotal lobes of pronotum light yellow. Forewings ochre, with dark brown to black veins, bearing light dots. Hindwings cinereous, with dark brown veins. Claws dark brown to black. Hind tibiae with brown to dark brown apices of spines. Abdominal tergites brown to dark brown, with light yellow hind margins. Pygofer lobes with light brown yellowish basal and apical angles and dark brown medially and from above; subgenital plate light brown yellowish basally and dark brown laterally and apically; anal tube dark brown. Male genitalia (Fig. 9) Anal tube 1.5 times as long as wide, flattened dorso-ventrally, slightly enlarged from base to apex (in dorsal view) (Fig. 9 A, B). Anal column short. Pygofer covered by long setae; pygofer lobe triangularly elongate, with an additional small lobe on apical angle under anal tube, without ventral lobe (Fig. 9 A); pygofer lobe apically with 4 - 5 macrosetae on its inner side (Fig. 9 D). Subgenital plate long and narrow, rounded apically (Fig. 9 C). Aedeagus asymmetrical; shaft long and narrow, straight (in ventral view), wider in its basal third, with pair of different in shape and length spine-shaped subapical processes (Fig. 9 G, H); aedeagal apex with carina above the gonopore (in ventral view), hook-shaped and pointed (in lateral view); gonopore subapical, small. Connective arcuate (in dorsal view) (Fig. 9 E, G). Style large, nearly as long as subgenital plate, with narrow, pointed apex and triangular process near midlength (Fig. 9 C, F, G). Measurement Total length. 6.0 mm.	en	Gnezdilov, Vladimir M. (2021): New species of the genera Limentinus Distant, 1917 and Calodia Nielson, 1982 (Hemiptera, Auchenorrhyncha, Cicadellidae, Coelidiinae) from the Makay Massif of Madagascar, with a key to Malagasy species. Zoosystema 43 (16): 297-310, DOI: 10.5252/zoosystema2021v43a16
