taxonID	type	description	language	source
03E487B6FFD5FFCEFF41886FFC4F791A.taxon	description	(Figs. 1 – 11, 40 – 41)	en	Janák, Jiří, Cz-, Petr Baňař (2021): Two new species and new records of the genus Astenus from Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 4980 (1): 127-141, DOI: https://doi.org/10.11646/zootaxa.4980.1.8
03E487B6FFD5FFCEFF41886FFC4F791A.taxon	materials_examined	Type locality. Madagascar, Ambohitantely Special Reserve Type material. Holotype ♂: “ ABT / Sept. 2011 / 24 MADAGASCAR, AMBOHITANTELY Spec. Res.; 5. ix. 2011; S 18 ° 11 ’ 43.3 ‘‘ E 47 ° 17 ’ 22.5 ‘‘, 1640 m; sifting of forest litter under palm tree, Winkler app. Extraction, L. S. Rahanitriniaina lgt. // HOLOTYPUS Astenus ambohitantelyensis sp. nov., Janák & Baňař det. 2020 ” (MMBC). Paratypes: 1 ♀: first label same as the holotype (MMBC); 1 ♂: “ ABT / Sept. 2011 / 14 MADAGASCAR, AMBOHITANTELY Spec. Res.; 3. ix. 2011; S 18 ° 10 ’ 57.6 ‘‘ E 47 ° 17 ’ 19.7 ‘‘, 1544 m; sifting forest litter and dry fern, Winkler app. extr.; L. S. Rahanitriniaina lgt. ” (JJRC); 1 ♂: “ ABT / 06 / 2011 MADAGASCAR 2011, Ambohitantely Spec. Res.; 19. iv., S 18 ° 11 ’ 02.5 ‘‘, E 47 ° 17 ’ 12.9 ‘‘; 1577 m, sifting forest litter, Winkler apparatus extraction; L. S. Rahanitriniaina lgt. ” (MMBC, aedeagus missing), 1 ♀: “ ABT / Sept. 2011 / 21 MADAGASCAR, Ambohitantely Spec. Res.; 5. ix. 2011; S 18 ° 11 ’ 23.1 ‘‘ E 47 ° 17 ’ 38.1 ‘‘, 1635 m; sifting of forest litter and dry fern; secodary forest; Winkler app. extraction; L. S. Rahanitriniaina lgt. ” (MMBC). All paratypes with an additional label “ PARATYPUS Astenus ambohitantelyensis sp. nov., Janák & Baňař det. 2020 ”.	en	Janák, Jiří, Cz-, Petr Baňař (2021): Two new species and new records of the genus Astenus from Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 4980 (1): 127-141, DOI: https://doi.org/10.11646/zootaxa.4980.1.8
03E487B6FFD5FFCEFF41886FFC4F791A.taxon	description	Description: Body length 4.7 – 6.4 mm (M = 5.4 mm, HT = 4.7 mm), forebody length 2.4 – 2.6 mm (M = 2.6 mm, HT = 2.4 mm). Black to dark brown, anterior part of head and abdomen dark brown, femora and tibiae brown, knees and tarsi light reddish yellow, antennomeres 1 – 7 dark reddish brown, antennomeres 8 – 11 and mouthparts light reddish yellow (Fig. 1). Head (Fig. 2) moderately convex, a fifth longer than wide (L / W = 1.17 – 1.22, M = 1.20, HT = 1.20). Surface with coarse and dense umbilicate punctures. Eyes moderately large, temples on average about 1.2 times as long as eyes (TL / EL 1.16 – 1.30, M = 1.20, HT = 1.18). Temples moderately narrowed posteriorly. Antennae moderately short, fifth antennomere about 2.5 times longer than wide (L / W = 2.50 – 2.73, M = 2.63, HT = 2.50), tenth antennomere about twice as long as wide (L / W = 1.93 – 2.11, M = 2.02, HT = 2.08). Pronotum (Fig. 2) egg-shaped, moderately convex, a quarter longer than wide (L / W = 1.19 – 1.26, M = 1.24, HT = 1.25) and moderately narrower than head (PW / HW = 0.87 – 0.89, M = 0.88, HT = 0.87). Sides markedly narrowed posteriorly. Surface, except for very narrow unpunctured midline with coarse and dense umbilicate punctures similar as on head. Elytra (Figs. 2, 3) elongate with rounded sides, concave along suture, about 1.3 times longer than wide (R = 1.26 – 1.34, M = 1.30, HT = 1.30). Surface moderately coarsely and densely punctured. Interstices shining, without microscupture. Shoulders with 2 tubercules bearing long, protruding fragile spines (Fig. 3) [in most specimens including HT broken and missing]. Sides with additional a few shorter and less protruding spines. Abdomen (Fig. 1) slightly widened to segment VI, finely and densely punctured. Interstices shining, without microsculpture. Male. Posterior margin of sternite VII very slightly, almost invisible emarginate and with almost 40 rod shaped moderately long spines (Fig. 10), sternite VIII long and narrow, with triangular emargination in posterior quarter (Fig. 11). Aedeagus (Figs. 4 – 9) 0.77 mm long. Median lobe in ventral view moderately wide, sides slightly emarginate in middle part and with small narrow sub-rounded point. Median lobe in lateral view with distinct subapical dent. Female. Sternite VIII largely rounded posteriorly.	en	Janák, Jiří, Cz-, Petr Baňař (2021): Two new species and new records of the genus Astenus from Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 4980 (1): 127-141, DOI: https://doi.org/10.11646/zootaxa.4980.1.8
03E487B6FFD5FFCEFF41886FFC4F791A.taxon	diagnosis	Differential diagnosis. The new species is to be arranged in the last key of Madagascan Astenus (Janák & Lecoq 2007) in the couplet 12 b near A. ambondrombe Janák & Lecoq, 2007 and A. ambrensis Janák, 2008 and differs from these species by very slightly, almost invisible emarginated posterior margin of the male sternite VII with moderately long rod shaped spines and different shape of aedeagus (see Figs. 38 – 44 in Janák & Lecoq, 2007 and Figs. 2 – 6 in Janák, 2008). There are also some external differences among these species – the color of A. ambohitantelyensis Janák & Baňař sp. nov. is much darker and the pronotum and elytra are on average longer than in remaining two species, but these characters are quite variable (see Fig. 39 in Janák & Lecoq (2007) and Fig. 1 in Janák (2008) and measurements in the descriptions for details).	en	Janák, Jiří, Cz-, Petr Baňař (2021): Two new species and new records of the genus Astenus from Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 4980 (1): 127-141, DOI: https://doi.org/10.11646/zootaxa.4980.1.8
03E487B6FFD5FFCEFF41886FFC4F791A.taxon	etymology	Etymology. This species is named the after the type locality – Ambohitantely Special Reserve.	en	Janák, Jiří, Cz-, Petr Baňař (2021): Two new species and new records of the genus Astenus from Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 4980 (1): 127-141, DOI: https://doi.org/10.11646/zootaxa.4980.1.8
03E487B6FFD5FFCEFF41886FFC4F791A.taxon	distribution	Distribution. Astenus ambohitantelyensis Janák & Baňař sp. nov. is currently recorded only from the Ambohitantely Special Reserve in the northern central part of Madagascar Bionomics. All specimens were collected in siftings of forest litter, palm trees and dry ferns in an indigenous or secondary forests at an altitude of 1540 – 1640 m a. s. l. (Figs. 40, 41).	en	Janák, Jiří, Cz-, Petr Baňař (2021): Two new species and new records of the genus Astenus from Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 4980 (1): 127-141, DOI: https://doi.org/10.11646/zootaxa.4980.1.8
03E487B6FFD7FFC8FF41880BFF2978C2.taxon	description	(Figs. 12 – 23, 42 – 45)	en	Janák, Jiří, Cz-, Petr Baňař (2021): Two new species and new records of the genus Astenus from Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 4980 (1): 127-141, DOI: https://doi.org/10.11646/zootaxa.4980.1.8
03E487B6FFD7FFC8FF41880BFF2978C2.taxon	materials_examined	Type locality. Madagascar, Anjozorobe-Angavo Type material. Holotype ♂: MADAGASCAR: “ MADAGAS., Anjozorob. [Anjozorobe], - 18.4118 47.9504, 1424 m, 8. i. 2019, sift [sifting], MD 36, V. Grebennikov // HOLOTYPUS Astenus grebennikovi sp. nov., Janák & Baňař det. 2020 ” (JJRC). Paratypes: 1 ♂: “ AJR / Jan. 2019 / sift 5 C MADAGASCAR, ANJOZOROBE forest, 15. i. 2019, sifting forest litter 1381 m, S 18 ° 24 ’ 36.2 ” E 47 ° 56 ’ 42.3 ”, Winkler app. extr. P. Baňař lgt. ” (JJRC); 1 ♂: “ AJR / Jan. 2019 / sift 8 C MADAGASCAR, ANJOZOROBE forest, 17. i. 2019, sifting litter by big Pandanus 1394 m, S 18 ° 24 ’ 35.5 ” E 47 ° 56 ’ 43.0 ”, Winkler app. extr. P. Baňař lgt. ” (MMBC); 1 ♂: “ AJR / Jan. 2019 / sift 7 C MADAGASCAR, ANJOZOROBE forest, 15. i. 2019, sifting forest litter by big tree 1331 m, S 18 ° 24 ’ 47.4 ” E 47 ° 56 ’ 33.1 ”, Winkler app. extr. P. Baňař lgt. ” (MMBC); 1 ♀: “ AJR / Jan. 2019 / sift 4 C MADAGASCAR, ANJOZOROBE forest, 15. i. 2019, sifting under Pandanus 1382 m, S 18 ° 24 ’ 35.2 ” E 47 ° 56 ’ 42.5 ”, Winkler app. extr. P. Baňař lgt. ” (MMBC). All paratypes with an additional label “ PARATYPUS Astenus grebennikovi sp. nov., Janák & Baňař det. 2020 ”.	en	Janák, Jiří, Cz-, Petr Baňař (2021): Two new species and new records of the genus Astenus from Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 4980 (1): 127-141, DOI: https://doi.org/10.11646/zootaxa.4980.1.8
03E487B6FFD7FFC8FF41880BFF2978C2.taxon	description	Description: Body length 5.5 – 7.5 mm (M = 6.4 mm, HT = 5.5 mm), forebody length 3.0 – 3.3 mm (M = 3.2 mm, HT = 3.0 mm). Dark brown, anterior part of head and sometimes also abdomen dark reddish brown, legs light yellow, antennomeres light reddish yellow with mostly slightly lighter first two antennomeres, mouthparts light reddish yellow (Fig. 12). Head (Fig. 13) moderately convex, a quarter longer than wide (L / W = 1.26 – 1.31, M = 1.28, HT = 1.27). Surface with coarse and dense umbilicate punctures. Eyes moderately large, temples on average about 1.3 times as long as eyes (TL / EL 1.23 – 1.43, M = 1.33, HT = 1.43). Temples markedly narrowed posteriorly. Antennae long and slender, fifth antennomere about 3 times as long as wide (L / W = 3.00 – 3.32, M = 3.16, HT = 3.00), tenth antennomere about two and half as long as wide (L / W = 2.38 – 2.56, M = 2.44, HT = 2.56). Pronotum (Fig. 13) egg-shaped, moderately convex, with shallow lateral impressions, about a quarter longer than wide (L / W = 1.23 – 1.30, M = 1.28, HT = 1.28) and moderately narrower than head (PW / HW = 0.86 – 0.90, M = 0.87, HT = 0.87). Sides markedly narrowed posteriorly, slightly emarginated in posterior half. Surface with coarse and dense umbilicate punctures similar as on head, unpunctured midline missing. Elytra (Fig. 13, 14) elongate with rounded sides, concave along suture, about 1.4 times longer than wide (R = 1.34 – 1.49, M = 1.41, HT = 1.49). Surface moderately coarsely and densely punctured. Interstices shining, without microscupture. Shoulders with 4 tubercules bearing long, protruding fragile spines (Fig. 14) [in most specimens including HT most spines broken and missing]. Sides with additional a few shorter and less protruding spines. Abdomen (Fig. 12) slightly widened to segment VI, finely and densely punctured. Interstices shining, without microsculpture. Male. Posterior margin of sternite VII broadly emarginate and with almost 25 moderately long spines (Fig. 22), sternite VIII long and narrow, with triangular emargination in posterior fifth (Fig. 23). Aedeagus (Figs. 15 – 21) long and slender, about 1 mm long (AL = 0.97 – 1.02 mm, M = 0.99 mm, HT = 0.99 mm). Median lobe in ventral view narrow, sides markedly emarginate in middle part and with relatively wide sub-rounded point. Median lobe in lateral view regularly bent. Female. Sternite VIII largely rounded posteriorly.	en	Janák, Jiří, Cz-, Petr Baňař (2021): Two new species and new records of the genus Astenus from Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 4980 (1): 127-141, DOI: https://doi.org/10.11646/zootaxa.4980.1.8
03E487B6FFD7FFC8FF41880BFF2978C2.taxon	diagnosis	Differential diagnosis. The new species is to be arranged in the last key of Madagascan Astenus species (Janák & Lecoq 2007) in the couplet 10 near A. duflosi Lecoq, 1996, but it differs externally by more rounded temples and in males by markedly emarginate posterior margin of sternite VII and different shape of the aedeagus (compare Figs. 31 – 39 for details).	en	Janák, Jiří, Cz-, Petr Baňař (2021): Two new species and new records of the genus Astenus from Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 4980 (1): 127-141, DOI: https://doi.org/10.11646/zootaxa.4980.1.8
03E487B6FFD7FFC8FF41880BFF2978C2.taxon	etymology	Etymology. This species is dedicated to Vasily Grebennikov, specialist in various groups of Coleoptera, who collected the holotype of the new species.	en	Janák, Jiří, Cz-, Petr Baňař (2021): Two new species and new records of the genus Astenus from Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 4980 (1): 127-141, DOI: https://doi.org/10.11646/zootaxa.4980.1.8
03E487B6FFD7FFC8FF41880BFF2978C2.taxon	distribution	Distribution. Astenus grebennikovi Janák & Baňař sp. nov. is currently recorded only from the Anjozorobe-Angavo [Paysage Harmonieux Protègé du Complexe Anjozorobe-Angavo] in the northern central part of Madagascar. Bionomics. The type series was sifted from forest litter (Figs. 42 – 45). One teneral specimen was collected in January.	en	Janák, Jiří, Cz-, Petr Baňař (2021): Two new species and new records of the genus Astenus from Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 4980 (1): 127-141, DOI: https://doi.org/10.11646/zootaxa.4980.1.8
03E487B6FFD1FFC8FF418830FA987F7B.taxon	materials_examined	Type locality. North Madagascar, Montagne d’Ambre. Material examined (13 specimens). MADAGASCAR N, Montagne d’Ambre N. P.: 2 ♂♂: 1100 m, 30. x. 2010, sifting, P. Baňař (MMBC, JJRC); 1 spec.:, 1169 m, circuit „ Sommet “, S 12 ° 31 ’ 52 ‘‘, E 49 ° 10 ’ 17 ‘‘, sifting forest litter, 8.1.2015, P. Baňař & E. M. Rabotoson lgt. (MMBC); 1 spec.: 1169 m, circuit „ Sommet “, S 12 ° 31 ’ 52 ‘‘, E 49 ° 10 ’ 17 ‘‘, sifting, Pandanus litter, 8.1.2015, P. Baňař & E. M. Rabotoson lgt. (MMBC); 1 spec.: ~ 1120 m, circuit „ Sommet “, S 12 ° 32 ’ 14 ‘‘, E 49 ° 10 ’ 28 ‘‘, sifting Pandanus litter, 12.1.2015, P. Baňař & E. M. Rabotoson lgt. (JJRC); 13 spec.: ~ 950 m, circuit „ Ampijoroana “, sifting litter by big trees, 13.1.2015, P. Baňař & E. M. Rabotoson lgt. (MMBC, JJRC). 6 spec.: ~ 945 m, circuit „ Sommet “, S 12 ° 31 ’ 28 ‘‘, E 49 ° 09 ’ 52 ‘‘, sifting litter + rotten wood, 14.1.2015, P. Baňař & E. M. Rabotoson lgt. (MMBC, JJRC); 10 spec.: ~ 1100 m, sifting litter close to camp, 16.1.2015, P. Baňař & E. M. Rabotoson lgt. (MMBC, JJRC).	en	Janák, Jiří, Cz-, Petr Baňař (2021): Two new species and new records of the genus Astenus from Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 4980 (1): 127-141, DOI: https://doi.org/10.11646/zootaxa.4980.1.8
03E487B6FFD1FFC8FF418830FA987F7B.taxon	distribution	Distribution. The species is known only from Montagne d’Ambre National Park in Nord Madagascar.	en	Janák, Jiří, Cz-, Petr Baňař (2021): Two new species and new records of the genus Astenus from Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 4980 (1): 127-141, DOI: https://doi.org/10.11646/zootaxa.4980.1.8
03E487B6FFD1FFC8FF418E92FD4F7DDD.taxon	materials_examined	Type locality. Central Madagascar, Ankaratra Mts., Manjakatompo. Material examined (1 specimen). MADAGASCAR: 1 ♂: AMBOHITANTELY Spec. Res., S 18 ° 10 ’ 17.9 ‘‘, E 47 ° 16 ’ 36.9 ‘‘ 1584 m; Malaise trap, 17. - 25. xi. 2011, P. Baňař lgt. (MMBC).	en	Janák, Jiří, Cz-, Petr Baňař (2021): Two new species and new records of the genus Astenus from Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 4980 (1): 127-141, DOI: https://doi.org/10.11646/zootaxa.4980.1.8
03E487B6FFD1FFC8FF418E92FD4F7DDD.taxon	distribution	Distribution. The holotype was described from Manjakatompo in Ankaratra Mts. The second specimen (female) from Ambialialika was attributed originally with some doubts to this species (Janák & Lecoq 2007). The new records is situated far north of these localities. Note. The shape of red spots on the elytra is variable: limited on transverse bands at posterior sixth (type) or third of elytra (Ambialialika and Ambohitantely – Figs. 24 – 27). Aedeagus (Figs. 28, 29) and posterior margin of the male sternite VII (Fig. 30) of the specimen from Ambohitantely are very characteristic for this species and confirm identity of both specimens with the holotype.	en	Janák, Jiří, Cz-, Petr Baňař (2021): Two new species and new records of the genus Astenus from Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 4980 (1): 127-141, DOI: https://doi.org/10.11646/zootaxa.4980.1.8
03E487B6FFD2FFCAFF418D24FDE37E3A.taxon	materials_examined	Type locality. East Madagascar, Périnet. Material examined (25 specimens). MADAGASCAR: 1 ♀: Maromizaha, - 18.976 48.4651, 1014 m, 14. xi. 2018, sif, V. Grebennikov (JJRC); 1 ♂: border of ANDASIBE N. P., „ Parc d’Orchidées “; 916 m, S 18 ° 55´59.9 ´´ E 048 ° 24´46.5 ´´ sifting; 5. xi. 2010; P. Baňař lgt. (JJRC); 1 ♀: Réserv. Expérimentale de VOHIMANA, „ Circuit 4, bivouac “; 20. viii. 2012, S 18 ° 55 ’ 07.8 ‘‘ E 48 ° 29 ’ 26.6 ‘‘; 937 m, sifting litter; L. S. Rahanitriniaina & E. M. Rabotoson lgt. (JJRC); 3 ♂♂: Réserv. Expérimentale de VOHIMANA, „ Circuit 3 “; 23. viii. 2012, S 18 ° 55 ’ 02.3 ‘‘ E 48 ° 30 ’ 21.2 ‘‘; 909 m, sifting litter; L. S. Rahanitriniaina & E. M. Rabotoson lgt. (MMBC, JJRC); 1 ♂: Réserv. Expérimentale de VOHIMANA, „ Circuit 3 “; 23. viii. 2012, S 18 ° 55 ’ 04.0 ‘‘, E 48 ° 30 ’ 27.9 ‘‘; 882 m, sifting litter; L. S. Rahanitriniaina & E. M. Rabotoson lgt. (JJRC); 2 ♀♀: Réserv. Expérimentale de VOHIMANA, „ Circuit 2, PK 09 “; 25. viii. 2012, S 18 ° 55 ’ 15.8 ‘‘ E 48 ° 30 ’ 05.1 ‘‘; 922 m, sifting litter; L. S. Rahanitriniaina & E. M. Rabotoson lgt. (MMBC, JJRC); 1 ♂: Réserv. Expérimentale de VOHIMANA, „ Circuit 2, PK 03 “; 25. viii. 2012, S 18 ° 55 ’ 23.8 ‘‘, E 48 ° 30 ’ 20.6 ‘‘; 867 m sifting litter; L. S. Rahanitriniaina & E. M. Rabotoson lgt. (JJRC); 1 ♂: Réserv. Expérimentale de VOHIMANA, „ Circuit 5, PK 09 “; 26. viii. 2012, S 18 ° 55 ’ 50.3 ‘‘, E 48 ° 30 ’ 05.3 ‘‘; 922 m, sifting litter; L. S. Rahanitriniaina & E. M. Rabotoson lgt. (JJRC); 2 ♂♂, 2 ♀♀: Réserv. Expérimentale de VOHIMANA, „ Circuit 5, PK 06 “; 26. viii. 2012 S 18 ° 55 ’ 49.8 ‘‘, E 48 ° 30 ’ 13.1 ‘‘; 901 m, sifting litter; L. S. Rahanitriniaina & E. M. Rabotoson lgt. (MMBC, JJRC); 1 ♂: Réserv. Expérimentale de VOHIMANA, „ Circuit 1 “; 27. viii. 2012, S 18 ° 55 ’ 32.9 ‘‘, E 48 ° 30 ’ 20.9 ‘‘; 860 m, sifting litter; L. S. Rahanitriniaina & E. M. Rabotoson lgt. (JJRC); 1 ♂, 1 ♀: Réserv. Expérimentale de VOHIMANA, „ Circuit 3 “; 28. viii. 2012, S 18 ° 55 ’ 32.0 ‘‘, E 48 ° 30 ’ 28.8 ‘‘; 840 m, sifting litter; L. S. Rahanitriniaina & E. M. Rabotoson lgt. (MMBC, JJRC); 1 ♂: Réserv. Expérimentale de VOHIMANA, „ Circuit 3 “; 28. viii. 2012 ,, S 18 ° 55 ’ 42.2 ‘‘, E 48 ° 30 ’ 55.5 ‘‘; 782 m, sifting litter; L. S. Rahanitriniaina & E. M. Rabotoson lgt. (JJRC); 1 ♂: Réserv. Expérimentale de VOHIMANA, „ Circuit Principal “; 29. viii. 2012, S 18 ° 55 ’ 14.6 ‘‘, E 48 ° 30 ’ 54.7 ‘‘; 831 m, sifting litter; L. S. Rahanitriniaina & E. M. Rabotoson lgt. (MMBC); 1 ♀: Beforona commune, VOHIDRAZANA forest; 6. vi. 2012, 1153 m, sifting litter, S 18 ° 58 ’ 33.5 ‘‘, E 48 ° 30 ’ 57.2 ‘‘, L. S. Rahanitriniaina & E. M. Rabotoson lgt. (MMBC); 1 ♀: Beforona commune, VOHIDRAZANA forest; 6. vi. 2012, 1130 m, sifting litter, S 18 ° 58 ’ 30.6 ‘‘, E 48 ° 30 ’ 53.4 ‘‘, L. S. Rahanitriniaina & E. M. Rabotoson lgt. (JJRC); 2 ♂♂: Beforona commune, RANOMENA forest; 7. vi. 2012, 1103 m, sifting litter, S 18 ° 58 ’ 28.5 ‘‘, E 48 ° 30 ’ 25.4 ‘‘, L. S. Rahanitriniaina & E. M. Rabotoson lgt. (MMBC); 1 ♀: Beforona commune, 10. vi. 2012, AMBATONDRALANGA, sifting litter, S 18 ° 54 ’ 34.6 ‘‘, E 48 ° 30 ’ 10.6 ‘‘, 877 m, L. S. Rahanitriniaina & E. M. Rabotoson lgt. (MMBC).	en	Janák, Jiří, Cz-, Petr Baňař (2021): Two new species and new records of the genus Astenus from Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 4980 (1): 127-141, DOI: https://doi.org/10.11646/zootaxa.4980.1.8
03E487B6FFD2FFCAFF418D24FDE37E3A.taxon	distribution	Distribution. The species was hitherto known from Analamazaotra National Park [= Andasibe or Périnet], nearby situated Maromizaha [Réserve de Ressources Naturelles de Maromizaha], a locality 9 km South of Moramanga, Ambohiboatavo forest East of Mantasoa lake and Soa Camp in Anjozorobe. The new records are from forested areas East of Analamazaotra National Park.	en	Janák, Jiří, Cz-, Petr Baňař (2021): Two new species and new records of the genus Astenus from Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 4980 (1): 127-141, DOI: https://doi.org/10.11646/zootaxa.4980.1.8
03E487B6FFD3FFCAFF418FD0FC8A7FD0.taxon	materials_examined	Type locality. East Madagascar, Nosy Mangabe. Material examined (3 specimens). MADAGASCAR: 1 ♂: Nosy Mangabe, Grebennikov (JJRC, stored in 96 % alcohol); 2 ♂♂: Madagascar, Farank. [Farankaraina], - 15.4283 49.8456, 132 m, 6. xii. 2018, sift, MD 14, V. Grebennikov (JJRC, 1 specimen stored in 96 % alcohol).	en	Janák, Jiří, Cz-, Petr Baňař (2021): Two new species and new records of the genus Astenus from Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 4980 (1): 127-141, DOI: https://doi.org/10.11646/zootaxa.4980.1.8
03E487B6FFD3FFCAFF418FD0FC8A7FD0.taxon	distribution	Distribution. The species was hitherto known from the island Nosy Mangabe, surroundings of Maroansetra and Ambodiavoangy. The new records is also from this region.	en	Janák, Jiří, Cz-, Petr Baňař (2021): Two new species and new records of the genus Astenus from Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 4980 (1): 127-141, DOI: https://doi.org/10.11646/zootaxa.4980.1.8
03E487B6FFD3FFC5FF418D66FBC17B12.taxon	materials_examined	Type locality. East Madagascar, Périnet. Material examined (13 specimens). MADAGASCAR: 4 ♂♂, 2 ♀♀:; border of ANDASIBE N. P.;´Parc a Orchidées´; S 18 ° 55 ’ 55.7 ‘‘ E 48 ° 24 ’ 47.5 ‘‘, 4. v. 2011, 935 m, sifting forest litter under Pandanus; L. S. Rahanitriniaina lgt. (MMBC, JJRC); 1 ♀: Réserv. Expérimentale de VOHIMANA „ Circuit 3 “; 23. viii. 2012, S 18 ° 55 ’ 04.0 ‘‘ E 48 ° 30 ’ 27.9 ‘‘; 882 m, sifting litter; L. S. Rahanitriniaina & E. M. Rabotoson lgt. (JJRC); 1 ♂: Réserv. Expérimentale de VOHIMANA, „ Circuit 5, PK 09 “; 26. viii. 2012, S 18 ° 55 ’ 50.3 ‘‘ E 48 ° 30 ’ 05.3 ‘‘; 922 m, sifting litter; L. S. Rahanitriniaina & E. M. Rabotoson lgt. (JJRC); 1 ♂, 1 ♀: Beforona commune, VOHIDRAZANA forest; 6. vi. 2012, 1153 m, sifting litter, S 18 ° 58 ’ 33.5 ‘‘ E 48 ° 30 ’ 57.2 ‘‘, L. S. Rahanitriniaina & E. M. Rabotoson lgt. (JJRC); 1 ♂, 1 ♀: Beforona commune, RANOMENA forest; 7. vi. 2012, 1103 m, sifting litter, S 18 ° 58 ’ 28.5 ‘‘ E 48 ° 30 ’ 25.4 ‘‘, L. S. Rahanitriniaina & E. M. Rabotoson lgt. (MMBC, JJRC); 1 ♀: MADAGASCAR, Beforona commune, RANOMENA forest; 8. vi. 2012, 1035 m, sifting litter, S 18 ° 58 ’ 25.0 ‘‘ E 48 ° 30 ’ 32.0 ‘‘, L. S. Rahanitriniaina & E. M. Rabotoson lgt. (JJRC).	en	Janák, Jiří, Cz-, Petr Baňař (2021): Two new species and new records of the genus Astenus from Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 4980 (1): 127-141, DOI: https://doi.org/10.11646/zootaxa.4980.1.8
03E487B6FFD3FFC5FF418D66FBC17B12.taxon	distribution	Distribution. The species was hitherto known only in the holotype from Analamazaotra National Park [Andasibe or Périnet]. The new records are from surroundings area east to the type locality.	en	Janák, Jiří, Cz-, Petr Baňař (2021): Two new species and new records of the genus Astenus from Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 4980 (1): 127-141, DOI: https://doi.org/10.11646/zootaxa.4980.1.8
03E487B6FFD8FFC1FF418BC0FBEB7ED6.taxon	materials_examined	Type locality. West Madagascar, Suberbieville [Maevatanana]. Material examined (25 specimens). MADAGASCAR: 1 spec.: ANDASIBE N. P.; 9. iv. 2011; edge, Parc d’Orchidées‘; sifting forest litter, P. Baňař & L. S. Rahanitriniaina lgt. (JJRC); 1 ♂: Beforona commune, 13. vi. 2012, SANDRASOA, sifting litter, S 18 ° 55 ’ 44.2 ‘‘, E 48 ° 31 ’ 17.9 ‘‘, 741 m; L. S. Rahanitriniaina & E. M. Rabotoson lgt. (JJRC); 2 ♂♂: Beforona commune, 13. vi. 2012, SANDRASOA, sifting litter, S 18 ° 55 ’ 44.6 ‘‘, E 48 ° 31 ’ 18.8 ‘‘, 724 m, L. S. Rahanitriniaina & E. M. Rabotoson lgt. (MMBC, JJRC); 1 ♂ 1 ♀: NW, ANKARAFANTSIKA N. P.; 22. iv. 2011, S 16 ° 18 ’ 46.6 ‘‘, E 46 ° 48 ’ 58.8 ‘‘; 89 m, sifting forest litter L. S. Rahanitriniaina & R. Raveloson lgt. (MMBC); 1 ♂: ANKARAFANTSIKA N. P.; 23. iv. 2011, S 16 ° 18 ’ 09.9 ‘‘ E 46 ° 48 ’ 39.3 ‘‘; 87 m, sifting forest litter, L. S. Rahanitriniaina & R. Raveloson lgt. (MMBC); 1 ♀: ANKARAFANTSIKA N. P.; 24. iv. 2011, S 16 ° 18 ’ 09.0 ‘‘, E 46 ° 48 ’ 55.6 ‘‘; 83 m, sifting forest litter, L. S. Rahanitriniaina & R. Raveloson lgt. (MMBC); 1 ♂: ANKARAFANTSIKA N. P.; 25. iv. 2011, S 16 ° 17 ’ 32.9 ‘‘ E 46 ° 48 ’ 38.1 ‘‘; 75 m, sifting forest litter, L. S. Rahanitriniaina & R. Raveloson lgt. (MMBC); 2 ♂♂, 1 ♀: ANKARAFANTSIKA N. P.; 24.1.2015, circuit „ Source de vie “, sifting forest litter by big tree, S 16 ° 18 ’ 12.0 ‘‘, E 46 ° 48 ’ 47.4 ‘‘; 81 m a. s. l., P. Baňař & E. M. Rabotoson lgt. (MMBC, JJRC); 1 ♀: ANKARAFANTSIKA N. P.; 24.1.2015, close to main entrance, sifting forest litter, ~ 75 m a. s. l., very wet material, P. Baňař & E. M. Rabotoson lgt. (MMBC); 1 ♀: Réservation Expérimentale de VOHIMANA; 27. xi. 2011; S 18 ° 55 ’ 13.7 ‘‘ E 48 ° 30 ’ 50.8 ‘‘; 776 m; sifting forest litter, L. S. Rahanitriniaina & P. Baňař lgt. (MMBC); 1 ♀: Réservation Expérimentale de VOHIMANA; 27. xi. 2011; S 18 ° 55 ’ 13.7 ‘‘, E 48 ° 30 ’ 50.8 ‘‘; 776 m; sifting forest litter, L. S. Rahanitriniaina & P. Baňař lgt. (MMBC); 1 ♀: Réservation Expérimentale de VOHIMANA; 27. xi. 2011; S 18 ° 55 ’ 13.6 ‘‘, E 48 ° 30 ’ 57.9 ‘‘; 772 m; sifting forest litter, in rock ravine; L. S. Rahanitriniaina & P. Baňař lgt. (MMBC); 2 ♂♂: ZOMBITSE N. P. „ Circuit Mandresy “, S 22 ° 52 ’ 57.5 ‘‘, E 44 ° 41 ’ 34.1 ‘‘; 25. i. 2013, 805 m, sifting litter; L. S. Rahanitriniaina & E. M. Rabotoson lgt. (MMBC, JJRC); 3 ♀♀: ZOMBITSE N. P. „ Circuit Mandresy “, S 22 ° 52 ’ 43.8 ‘‘ E 44 ° 41 ’ 36.5 ‘‘; 25. i. 2013, 807 m, sifting litter; L. S. Rahanitriniaina & E. M. Rabotoson lgt. (MMBC); 1 ♂: ZOMBITSE N. P. „ Circuit Mandresy “ S 22 ° 52 ’ 46.6 ‘‘, E 44 ° 41 ’ 43.9 ‘‘; 25. i. 2013, 806 m, sifting litter; L. S. Rahanitriniaina & E. M. Rabotoson lgt. (MMBC); 1 ♂, 2 ♀♀: ZOMBITSE N. P.; Andalabiby, 772 m, S 22 ° 53 ’ 04.6 ‘‘, E 44 ° 42 ’ 25.6 ‘‘; 26. i. 2013, sifting litter; L. S. Rahanitriniaina & E. M. Rabotoson lgt. (MMBC, JJRC).	en	Janák, Jiří, Cz-, Petr Baňař (2021): Two new species and new records of the genus Astenus from Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 4980 (1): 127-141, DOI: https://doi.org/10.11646/zootaxa.4980.1.8
03E487B6FFD8FFC1FF418BC0FBEB7ED6.taxon	distribution	Distribution. The species is widely distributed in various parts of Madagascar	en	Janák, Jiří, Cz-, Petr Baňař (2021): Two new species and new records of the genus Astenus from Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 4980 (1): 127-141, DOI: https://doi.org/10.11646/zootaxa.4980.1.8
03E487B6FFD8FFC1FF418E77FB857C5A.taxon	materials_examined	Type locality. Madagascar Est, Nosy Mangabe. Material examined (2 specimens). MADAGASCAR: 2 ♂♂: Farank. [Farankaraina], - 15.4283 49.8456, 132 m, 6. xii. 2018, sift [sifting], MD 14, V. Grebennikov (JJRC, 1 spec. stored in 96 % alcohol).	en	Janák, Jiří, Cz-, Petr Baňař (2021): Two new species and new records of the genus Astenus from Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 4980 (1): 127-141, DOI: https://doi.org/10.11646/zootaxa.4980.1.8
03E487B6FFD8FFC1FF418E77FB857C5A.taxon	distribution	Distribution. The species was hitherto known from the island Nosy Mangabe, Mt. Andrangoatra, Ambohitsitondroina and Ambodiavoangy. The new record is situated near this region.	en	Janák, Jiří, Cz-, Petr Baňař (2021): Two new species and new records of the genus Astenus from Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 4980 (1): 127-141, DOI: https://doi.org/10.11646/zootaxa.4980.1.8
03E487B6FFD8FFC0FF418DCBFAE97ADB.taxon	materials_examined	Type locality. Madagascar West, Suberbieville [Maevatanana]. Material examined (3 specimens). MADAGASCAR: 1 ♀: border of ANDASIBE N. P.; 27. xi. - 3. xii.; sifting leaf litter, L. S. Rahanitriniaina lgt (MMBC); 1 ♂: Réserv. Expérimentale de VOHIMANA, „ Circuit Principal “; 29. viii. 2012, S 18 ° 55 ’ 14.6 ‘‘, E 48 ° 30 ’ 54.7 ‘‘; 831 m, sifting litter; L. S. Rahanitriniaina & E. M. Rabotoson lgt. (JJRC); 1 ♂: Beforona commune, 13. vi. 2012, SANDRASOA, sifting litter, S 18 ° 55 ’ 44.6 ‘, ‘ E 48 ° 31 ’ 18.8 ‘‘, 724 m, L. S. Rahanitriniaina & E. M. Rabotoson lgt. (JJRC).	en	Janák, Jiří, Cz-, Petr Baňař (2021): Two new species and new records of the genus Astenus from Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 4980 (1): 127-141, DOI: https://doi.org/10.11646/zootaxa.4980.1.8
03E487B6FFD8FFC0FF418DCBFAE97ADB.taxon	distribution	Distribution. The species was hitherto known from Comoro islands (Mohéli), central (Isalo), western (Maevatanana, Antsingy forest) and eastern (Ambohimanarivo, south of Ifanadiana) parts of Madagascar.	en	Janák, Jiří, Cz-, Petr Baňař (2021): Two new species and new records of the genus Astenus from Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 4980 (1): 127-141, DOI: https://doi.org/10.11646/zootaxa.4980.1.8
03E487B6FFD9FFC0FF4189FAFB067E0A.taxon	materials_examined	Type locality. East Madagascar, east of Vondrozo. Material examined (3 specimens). MADAGASCAR: 3 ♂♂: RANOMAFANA N. P.; 6. x. 2012, S 21 ° 17 ’ 51.8 ‘‘ E 47 ° 26 ’ 35.1 ‘‘; 1008 m, VALOHOAKA, sifting litter, L. S. Rahanitriniaina lgt. (MMBC, JJRC).	en	Janák, Jiří, Cz-, Petr Baňař (2021): Two new species and new records of the genus Astenus from Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 4980 (1): 127-141, DOI: https://doi.org/10.11646/zootaxa.4980.1.8
03E487B6FFD9FFC0FF4189FAFB067E0A.taxon	distribution	Distribution. The species was hitherto known from the type locality situated east of Vondrozo and from Mt. Ambondrombe. The new locality is situated north of the type locality. Note. The size of dark spots on the elytra is variable, in some specimens the spots are absent.	en	Janák, Jiří, Cz-, Petr Baňař (2021): Two new species and new records of the genus Astenus from Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 4980 (1): 127-141, DOI: https://doi.org/10.11646/zootaxa.4980.1.8
03E487B6FFD9FFC0FF418A71FDCD7854.taxon	materials_examined	Type locality. West Madagascar, Suberbieville [Maevatanana]. Material examined (1 specimen). MADAGASCAR NE: 1 ♂:, ANKARAFANTSIKA N. P., S 16 ° 18 ’ 43.7 ‘‘ E 4 6 ° 48 ’ 59.4 ‘‘, 23. iv. 2011; 100 m; at light close to small lake; P. Baňař lgt. (MMBC).	en	Janák, Jiří, Cz-, Petr Baňař (2021): Two new species and new records of the genus Astenus from Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 4980 (1): 127-141, DOI: https://doi.org/10.11646/zootaxa.4980.1.8
03E487B6FFD9FFC0FF418A71FDCD7854.taxon	distribution	Distribution. The species was hitherto known only in type series from Maevatanana. The new locality is situated north of the type locality.	en	Janák, Jiří, Cz-, Petr Baňař (2021): Two new species and new records of the genus Astenus from Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 4980 (1): 127-141, DOI: https://doi.org/10.11646/zootaxa.4980.1.8
03E487B6FFD9FFC0FF418F00FEA57FBF.taxon	materials_examined	Type locality. East Madagascar, Navana east of Maroansetra. Material examined (1 specimen). MADAGASCAR: 1 ♂: Ranomafana Nat. Park, near Ranomafana vill., M. Trýzna leg., 26. - 31. i. 2007 (JJRC).	en	Janák, Jiří, Cz-, Petr Baňař (2021): Two new species and new records of the genus Astenus from Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 4980 (1): 127-141, DOI: https://doi.org/10.11646/zootaxa.4980.1.8
03E487B6FFD9FFC0FF418F00FEA57FBF.taxon	distribution	Distribution. The species was hitherto known from Navana and Ambodivoangy east of Maroansetra, Anandrivola 70 km SSE of Maroansetra, from Nosy Bohara and Tampolo. The new locality is situated markedly south of this region.	en	Janák, Jiří, Cz-, Petr Baňař (2021): Two new species and new records of the genus Astenus from Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 4980 (1): 127-141, DOI: https://doi.org/10.11646/zootaxa.4980.1.8
