taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
038D87D3A805FF9DC1ECF994718BFE2A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4882934/files/figure.png	http://doi.org/10.5281/zenodo.4882934	FIGURE 2. Benthophilus persicus sp. nov. Reversibly stained paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea. Preopercle with dermal ossifications: T – tubercles with spinules on tubercles appear irregularly arranged, G1 - single granule, G2 and G3 - granules grouped two and three together, respectively. Photo by M. Kovačić.	FIGURE 2. Benthophilus persicus sp. nov. Reversibly stained paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea. Preopercle with dermal ossifications: T – tubercles with spinules on tubercles appear irregularly arranged, G1 - single granule, G2 and G3 - granules grouped two and three together, respectively. Photo by M. Kovačić.	2021-05-31	Kovačić, Marcelo;Esmaeili, Hamid Reza;Zarei, Fatah;Abbasi, Keyvan;Schliewen, Ulrich K.		Zenodo	biologists	Kovačić, Marcelo;Esmaeili, Hamid Reza;Zarei, Fatah;Abbasi, Keyvan;Schliewen, Ulrich K.			
038D87D3A805FF9DC1ECF994718BFE2A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4882936/files/figure.png	http://doi.org/10.5281/zenodo.4882936	FIGURE 3. Benthophilus persicus sp. nov. Reversibly stained paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea. Dermal ossifications on anterior dorsal body view: D - dorsal row of tubercles: T1- tubercle with one radial row of spinules and T2 - tubercle with two radial rows of spinules, DL - dorsolateral row of tubercles, G - granules. Photo by M. Kovačić.	FIGURE 3. Benthophilus persicus sp. nov. Reversibly stained paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea. Dermal ossifications on anterior dorsal body view: D - dorsal row of tubercles: T1- tubercle with one radial row of spinules and T2 - tubercle with two radial rows of spinules, DL - dorsolateral row of tubercles, G - granules. Photo by M. Kovačić.	2021-05-31	Kovačić, Marcelo;Esmaeili, Hamid Reza;Zarei, Fatah;Abbasi, Keyvan;Schliewen, Ulrich K.		Zenodo	biologists	Kovačić, Marcelo;Esmaeili, Hamid Reza;Zarei, Fatah;Abbasi, Keyvan;Schliewen, Ulrich K.			
038D87D3A805FF9DC1ECF994718BFE2A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4882938/files/figure.png	http://doi.org/10.5281/zenodo.4882938	FIGURE 4. Benthophilus persicus sp. nov. preserved specimens. Holotype, ZSM 47595, male, 45.2+9.9 mm, Iran, Gilan Province, Anzali, southern Caspian Sea: A: lateral view, B: urogenital papilla of adult male (marked by arrow). Paratype, ZSM 47599, female, 30.8+7.0 mm, Iran, Gilan Province, Anzali, southern Caspian Sea: C: lateral view, D: urogenital papilla of adult female (marked by arrow). Photos by M. Kovačić.	FIGURE 4. Benthophilus persicus sp. nov. preserved specimens. Holotype, ZSM 47595, male, 45.2+9.9 mm, Iran, Gilan Province, Anzali, southern Caspian Sea: A: lateral view, B: urogenital papilla of adult male (marked by arrow). Paratype, ZSM 47599, female, 30.8+7.0 mm, Iran, Gilan Province, Anzali, southern Caspian Sea: C: lateral view, D: urogenital papilla of adult female (marked by arrow). Photos by M. Kovačić.	2021-05-31	Kovačić, Marcelo;Esmaeili, Hamid Reza;Zarei, Fatah;Abbasi, Keyvan;Schliewen, Ulrich K.		Zenodo	biologists	Kovačić, Marcelo;Esmaeili, Hamid Reza;Zarei, Fatah;Abbasi, Keyvan;Schliewen, Ulrich K.			
038D87D3A805FF9DC1ECF994718BFE2A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4882940/files/figure.png	http://doi.org/10.5281/zenodo.4882940	FIGURE 5. Benthophilus persicus sp. nov. Reversibly stained paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea. Dermal ossifications on dorsal view of head, TCO - tubercles along upper cheek, preopercle and opercle, TI - interorbital tubercles, G - granules. Tubercles not present on temporal and occipital head regions, i.e., all dermal ossifications there are definable as granules according to size and shape; and the first tubercles anteriorly to temporal region are located at interorbital space, i.e., anteriorly to rear edge of eyes. Photos by M. Kovačić.	FIGURE 5. Benthophilus persicus sp. nov. Reversibly stained paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea. Dermal ossifications on dorsal view of head, TCO - tubercles along upper cheek, preopercle and opercle, TI - interorbital tubercles, G - granules. Tubercles not present on temporal and occipital head regions, i.e., all dermal ossifications there are definable as granules according to size and shape; and the first tubercles anteriorly to temporal region are located at interorbital space, i.e., anteriorly to rear edge of eyes. Photos by M. Kovačić.	2021-05-31	Kovačić, Marcelo;Esmaeili, Hamid Reza;Zarei, Fatah;Abbasi, Keyvan;Schliewen, Ulrich K.		Zenodo	biologists	Kovačić, Marcelo;Esmaeili, Hamid Reza;Zarei, Fatah;Abbasi, Keyvan;Schliewen, Ulrich K.			
038D87D3A805FF9DC1ECF994718BFE2A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4882942/files/figure.png	http://doi.org/10.5281/zenodo.4882942	FIGURE 6. Benthophilus persicus sp. nov. Dermal fold (DF) and chin barbell (CB). Reversibly stained paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea: A: lateral detail of head, B: ventral view of head. Reversibly stained holotype, ZSM 47595, male, 45.2+9.9 mm, Iran, Gilan Province, Anzali, southern Caspian: C: chin close up with barbel. Photo by M. Kovačić.	FIGURE 6. Benthophilus persicus sp. nov. Dermal fold (DF) and chin barbell (CB). Reversibly stained paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea: A: lateral detail of head, B: ventral view of head. Reversibly stained holotype, ZSM 47595, male, 45.2+9.9 mm, Iran, Gilan Province, Anzali, southern Caspian: C: chin close up with barbel. Photo by M. Kovačić.	2021-05-31	Kovačić, Marcelo;Esmaeili, Hamid Reza;Zarei, Fatah;Abbasi, Keyvan;Schliewen, Ulrich K.		Zenodo	biologists	Kovačić, Marcelo;Esmaeili, Hamid Reza;Zarei, Fatah;Abbasi, Keyvan;Schliewen, Ulrich K.			
038D87D3A805FF9DC1ECF994718BFE2A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4882944/files/figure.png	http://doi.org/10.5281/zenodo.4882944	FIGURE 7. Benthophilus persicus sp. nov. Reversibly stained paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea. A: Dermal ossifications on lateral view of head, TCO - tubercles along upper cheek, preopercle and opercle, G - granules. B: Dermal ossifications on posterior dorsal body view: D - dorsal row of tubercles, DL - dorsolateral row of tubercles, T - tubercle, G - granules. C: Dermal ossifications on anterior lateral body view, D - dorsal row of tubercles, DL - dorsolateral row of tubercles, V - ventral row of tubercles, T - tubercles. D: Dermal ossifications on posterior lateral body view, D - dorsal row of tubercles, DL - dorsolateral row of tubercles, V - ventral row of tubercles, T - tubercles. Photos by M. Kovačić.	FIGURE 7. Benthophilus persicus sp. nov. Reversibly stained paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea. A: Dermal ossifications on lateral view of head, TCO - tubercles along upper cheek, preopercle and opercle, G - granules. B: Dermal ossifications on posterior dorsal body view: D - dorsal row of tubercles, DL - dorsolateral row of tubercles, T - tubercle, G - granules. C: Dermal ossifications on anterior lateral body view, D - dorsal row of tubercles, DL - dorsolateral row of tubercles, V - ventral row of tubercles, T - tubercles. D: Dermal ossifications on posterior lateral body view, D - dorsal row of tubercles, DL - dorsolateral row of tubercles, V - ventral row of tubercles, T - tubercles. Photos by M. Kovačić.	2021-05-31	Kovačić, Marcelo;Esmaeili, Hamid Reza;Zarei, Fatah;Abbasi, Keyvan;Schliewen, Ulrich K.		Zenodo	biologists	Kovačić, Marcelo;Esmaeili, Hamid Reza;Zarei, Fatah;Abbasi, Keyvan;Schliewen, Ulrich K.			
038D87D3A805FF9DC1ECF994718BFE2A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4882946/files/figure.png	http://doi.org/10.5281/zenodo.4882946	FIGURE 8. Head sensory papillae pattern of lateral line system of Benthophilus persicus sp. nov. Paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea. Terminology in text. Drawing by M. Kovačić.	FIGURE 8. Head sensory papillae pattern of lateral line system of Benthophilus persicus sp. nov. Paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea. Terminology in text. Drawing by M. Kovačić.	2021-05-31	Kovačić, Marcelo;Esmaeili, Hamid Reza;Zarei, Fatah;Abbasi, Keyvan;Schliewen, Ulrich K.		Zenodo	biologists	Kovačić, Marcelo;Esmaeili, Hamid Reza;Zarei, Fatah;Abbasi, Keyvan;Schliewen, Ulrich K.			
038D87D3A805FF9DC1ECF994718BFE2A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4882948/files/figure.png	http://doi.org/10.5281/zenodo.4882948	FIGURE 9. Benthophilus persicus sp. nov. Sensory papillae pattern of lateral line system in reversibly stained paratypes, anterior part of preoperculo-mandibular rows in PMR VP4680, male, 47.0+8.6 mm, Iran, Gilan Province, Anzali, southern Caspian Sea. Terminology in text. Photo by M. Kovačić.	FIGURE 9. Benthophilus persicus sp. nov. Sensory papillae pattern of lateral line system in reversibly stained paratypes, anterior part of preoperculo-mandibular rows in PMR VP4680, male, 47.0+8.6 mm, Iran, Gilan Province, Anzali, southern Caspian Sea. Terminology in text. Photo by M. Kovačić.	2021-05-31	Kovačić, Marcelo;Esmaeili, Hamid Reza;Zarei, Fatah;Abbasi, Keyvan;Schliewen, Ulrich K.		Zenodo	biologists	Kovačić, Marcelo;Esmaeili, Hamid Reza;Zarei, Fatah;Abbasi, Keyvan;Schliewen, Ulrich K.			
038D87D3A805FF9DC1ECF994718BFE2A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4882950/files/figure.png	http://doi.org/10.5281/zenodo.4882950	FIGURE 10. Benthophilus persicus sp. nov. Sensory papillae pattern of lateral line system in reversibly stained paratypes, sensory papillae rows on anterior flank in PMR VP4681, male, 35.1+8.5 mm, Iran, Gilan Province, Anzali, southern Caspian Sea. Sensory papillae rows marked, terminology text. The transversal ltm rows starting anteriorly behind axilla, alternating with three longitudinal llm rows making anterior beginning pattern of -II-I-I. Photo by M. Kovačić.	FIGURE 10. Benthophilus persicus sp. nov. Sensory papillae pattern of lateral line system in reversibly stained paratypes, sensory papillae rows on anterior flank in PMR VP4681, male, 35.1+8.5 mm, Iran, Gilan Province, Anzali, southern Caspian Sea. Sensory papillae rows marked, terminology text. The transversal ltm rows starting anteriorly behind axilla, alternating with three longitudinal llm rows making anterior beginning pattern of -II-I-I. Photo by M. Kovačić.	2021-05-31	Kovačić, Marcelo;Esmaeili, Hamid Reza;Zarei, Fatah;Abbasi, Keyvan;Schliewen, Ulrich K.		Zenodo	biologists	Kovačić, Marcelo;Esmaeili, Hamid Reza;Zarei, Fatah;Abbasi, Keyvan;Schliewen, Ulrich K.			
038D87D3A805FF9DC1ECF994718BFE2A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4882932/files/figure.png	http://doi.org/10.5281/zenodo.4882932	FIGURE 1. Bathymetric map of the Caspian Sea, showing the type locality (yellow star) of Benthophilus persicus sp. nov. near Anzali, southern Caspian Sea. The image was produced using Surfer ver. 10.1.561 mapping software package (Golden Software, LLC).	FIGURE 1. Bathymetric map of the Caspian Sea, showing the type locality (yellow star) of Benthophilus persicus sp. nov. near Anzali, southern Caspian Sea. The image was produced using Surfer ver. 10.1.561 mapping software package (Golden Software, LLC).	2021-05-31	Kovačić, Marcelo;Esmaeili, Hamid Reza;Zarei, Fatah;Abbasi, Keyvan;Schliewen, Ulrich K.		Zenodo	biologists	Kovačić, Marcelo;Esmaeili, Hamid Reza;Zarei, Fatah;Abbasi, Keyvan;Schliewen, Ulrich K.			
