taxonID	type	description	language	source
1336C156E0055A7D8D59839CE87A724A.taxon	diagnosis	Diagnosis. Representatives of the genus Coccidula with its general body shape may resemble Tetrabrachys Kapur, however, it can be separated based on the structure of the tarsi which are tetramerous in both genera but in Coccidula the first tarsomere is sub-triangularly broadened apically and the second is elongate and distinctly lobbed, while in Tetrabrachys both the first and second are narrow, elongate and without lobes. Moreover, in Tetrabrachys the apical maxillary palpomere is widely securiform, and beetles are brachypterous, while in Coccidula the apical maxillary palpomere is only slightly widened and the second pair of wings is functional. Coccidula is also externally similar to European species of Rhyzobius but it can be separated based on the following characters: body almost parallel sided, elytra covered with punctures of two sizes, larger punctures arranged in nine rows (in C. litophiloides some of them are reduced), base of the pronotum not bordered, while in Rhyzobius the lateral body outline is broadly rounded, the elytra are covered with single sized, randomly arranged punctures, and base of the pronotum with distinct bordering line.	en	Szawaryn, Karol, Nedved, Oldrich, Biranvand, Amir, Czerwinski, Tomasz, Nattier, Romain (2021): Revision of the genus Coccidula Kugelann (Coleoptera, Coccinellidae). ZooKeys 1043: 61-85, DOI: http://dx.doi.org/10.3897/zookeys.1043.65829, URL: http://dx.doi.org/10.3897/zookeys.1043.65829
1336C156E0055A7D8D59839CE87A724A.taxon	description	Description. Body elongate-oval, with sides parallel (Fig. 1 C-H), body flattened in lateral view, convex in cross-section; dorsum covered with setiferous punctures of two sizes (Figs 7 A, 9 A), hairs directed forwards on pronotum, backwards on elytra. Head partially withdrawn into prothorax (Fig. 1 C-H); ventral antennal grooves shallow and moderately long, extending to posterior border of an eye (Fig. 3 E). Eyes prominent, coarsely facetted (7 - 8 ommatidia per eye width), ocular canthus distinct, about as long as 4 - 5 ommatidium diameters; interocular distance about 3 x as eye diameter; interfacetal setae present only in basal part; temple behind eye distinctly longer than eye (Fig. 3 E). Antennal insertion placed laterally, invisible from above, distance between antennal insertions about same as between eyes; frons around antennal insertions slightly expanded, covering antennal insertions, anterior tentorial pits placed ventrally below antennal insertions. Antennae (Figs 5 A, E, 7 A, D) longer than maximum head width including eyes, composed of 11 antennomeres (AN); scape simple, without projections, slightly curved; pedicel distinctly narrower than scape, elongate (1.5 x longer than wide); AN 3 - 8 elongate (AN 3 ≈ 3.5 x; AN 8 ≈ 1.3 x longer than wide); AN 9 - 11 forming a loose, asymmetric club, ultimate AN truncate apically. Frontoclypeus short, transverse, anterior margin straight. Labrum entirely exposed, transverse, anterior margin straight. Mandibles asymmetric, bifid apically (Fig. 10 G), molar part with basal tooth; prostheca distinct. Maxillary stipes (Figs 2 B, 5 C, 9 C, 10 H) with distinct groove for reception of maxillary palp in repose; palpomere 2 shorter than terminal (4 th) one, slightly broadened apically; palpomere 3 about 2.3 x shorter than terminal one, subtriangular; terminal palpomere slightly securiform; lacinia with stiff setae on outer margin in apical half, with several additional spurs on surface (Fig. 10 H). Labial palps (Figs 3 E, 9 C) with 3 palpomeres, inserted ventrally on prementum; palpomere 1 very small, apical palpomere as long as and about as broad as penultimate; distance between palp insertions about 1.5 - 2 x as its width. Prementum subquadrate, transverse apically. Mentum trapezoidal, broadest in anterior part, with horseshoe impression at base (Figs 7 C, 9 C). Submentum broad, transverse, with suture invisible. Anterior margin of pronotum weakly, broadly emarginate (Figs 3 B, 5 B) with anterior corners broadly rounded; lateral margins with moderately (Figs 7 B, 9 B) to distinctly expanded lateral beads (Fig. 3 B), distinctly margined; hind corners sharply pointed; hind margin not bordered. Prothoracic hypomeron smooth, without delimited foveae (Figs 3 C, 7 C). Prosternum in front of coxae about as long as longitudinal length of procoxal cavity; anterior margin straight or slightly emarginate with distinct border. Prosternal process about 0.4 times of coxal diameter, surface smooth (Fig. 3 C) or with lateral carinae (Figs 7 C, 9 E). Procoxal cavity oval, distinctly bordered anteriorly. Mesoventrite 1.3 x longer than its width at the level of mid coxae (Figs 1 D, 5 D, 7 D); mesal surface with deep emargination for receiving tip of prosternal process (Fig. 2 C); anterior margin with completely raised border. Meso-metaventral process narrow (Figs 1 D, 2 C, 5 D, 7 E), about 0.5 times of mesocoxal diameter, junction slightly arcuate (Figs 2 C, 3 D, 5 D, 7 E, 9 D), with suture visible. Metendosternite with stalk sub-quadrate, tendons long, separated by a distance of about width of stalk and situated closer to center (Fig. 10 I). Scutellar shield pentagonal (Figs 7 B, 9 B). Elytra at base wider than pronotum, lateral margins clearly visible from above throughout (Figs 2 D, 3 A, 5 A, 7 A) (except C. scutellata where it is obscured in basal part, Fig. 9 A), surface covered with punctures of double size, smaller irregularly distributed, larger punctures arranged in nine irregular longitudinal rows. Sutural stria absent. Elytral epipleuron narrow, incomplete, reaching base of ventrite 4 (Fig. 1 D), with complete bordering line, epipleural foveae absent. Hind wings fully developed or missing (in C. litophiloides). Metaventral postcoxal lines roundly joined medially, complete laterally, straight or descending (Figs 2 C, 3 D, 5 D, 7 E, 9 D). Metaventrite with discrimen visible in posterior 2 / 3. Trochanters simple, subtriangular, without projection (figs 7 E, 9 D). Tibiae slightly expanded apically with one apical spur on forelegs, and two in mid and hind legs. Tarsi consisting of four tarsomeres, second tarsomere truncate apically; tarsal claws cleft apically (Fig. 9 G) with single empodial seta present. Abdomen in both sexes with 6 ventrites (Fig. 1 D); ventrite 1 about as long as ventrites 2 - 4 combined, ventrite 2 longer than ventrite 3, ventrites 3 - 5 subequal in length. Abdominal postcoxal lines (Figs 7 E, 9 D) separate medially, recurved and complete, reaching anterior margin of ventrite, posteriorly reaching about half length of ventrite 1. Ventrite 5 in female posteriorly rounded (Fig. 7 F), in male truncate (Fig. 9 F). Ventrite 6 rounded in both sexes. Male terminalia. Tegmen (Figs 4 A, B, 8 A, B, 10 A, B) symmetrical; parameres articulated with penis guide. Penis (Figs 4 C, 8 C, 10 C) slender, pointed apically; penis capsule asymmetrical with outer arm reduced, inner arm well developed. Apodeme of male sternum IX simple, not broadened apically (Figs 8 D, 10 D). Tergite X broadly rounded, semicircular (Figs 8 D, 10 D). Female terminalia. Coxites (Figs 10 F) distinctly elongate, subtriangular; styli small but visible, bearing several short setae; infundibulum absent (Figs 4 D, 8 E, 10 E); sperm duct simple. Spermatheca (Figs 4 D, 8 E, 10 E) worm-like, without clear ramus or nodulus; spermathecal accessory gland small, elongate. Proctiger elongate, rounded apically (Fig. 10 F).	en	Szawaryn, Karol, Nedved, Oldrich, Biranvand, Amir, Czerwinski, Tomasz, Nattier, Romain (2021): Revision of the genus Coccidula Kugelann (Coleoptera, Coccinellidae). ZooKeys 1043: 61-85, DOI: http://dx.doi.org/10.3897/zookeys.1043.65829, URL: http://dx.doi.org/10.3897/zookeys.1043.65829
1336C156E0055A7D8D59839CE87A724A.taxon	distribution	Distribution. Holarctic: Asia, Europe, Africa (North), North America.	en	Szawaryn, Karol, Nedved, Oldrich, Biranvand, Amir, Czerwinski, Tomasz, Nattier, Romain (2021): Revision of the genus Coccidula Kugelann (Coleoptera, Coccinellidae). ZooKeys 1043: 61-85, DOI: http://dx.doi.org/10.3897/zookeys.1043.65829, URL: http://dx.doi.org/10.3897/zookeys.1043.65829
4CE4208FE9C651F693F60F46591A20FB.taxon	materials_examined	Material examined. United States of America: America b., 82, coll. Ruzicka et Vokal, (1: NMP); Alaska, Mi. 1249, Alaska Hwy., Dedman Lk., 6. - 7. VII. 1968, Campbell & Smetana (1: NMP); Vermont, Korschefsky det. (2: MIZ); Canada: SK, Harris Reservoir, Hwy 21, 10 km S, Maple Creek, April 19 2016, drift D. Larson (1 female: NMP). Type material not studied, deposited in Museum of Comparative Zoology, Cambridge, USA.	en	Szawaryn, Karol, Nedved, Oldrich, Biranvand, Amir, Czerwinski, Tomasz, Nattier, Romain (2021): Revision of the genus Coccidula Kugelann (Coleoptera, Coccinellidae). ZooKeys 1043: 61-85, DOI: http://dx.doi.org/10.3897/zookeys.1043.65829, URL: http://dx.doi.org/10.3897/zookeys.1043.65829
4CE4208FE9C651F693F60F46591A20FB.taxon	diagnosis	Diagnosis. Coccidula lepida is the only Nearctic species of the genus, and is similar in many characters to C. scutellata, but with the head and epipleura black. In the typical form (C. lepida described by LeConte), the black elytral pattern resembles an extension of the five fused black maculae on the elytra of C. scutellata, with shoulders and epipleura black. Shoulder tubercle distinct, prototum relatively narrow. Male genitalia with penis guide much shorter than parameres. Detailed description of morphology including variability in pattern can be found in Gordon (1985: 656 - 659).	en	Szawaryn, Karol, Nedved, Oldrich, Biranvand, Amir, Czerwinski, Tomasz, Nattier, Romain (2021): Revision of the genus Coccidula Kugelann (Coleoptera, Coccinellidae). ZooKeys 1043: 61-85, DOI: http://dx.doi.org/10.3897/zookeys.1043.65829, URL: http://dx.doi.org/10.3897/zookeys.1043.65829
4CE4208FE9C651F693F60F46591A20FB.taxon	description	Description. Length = 2.7 - 3.5 mm, BL / BW = 1.88 - 1.96, EL / BW = 1.40 - 1.42, PW / BW = 0.73. Body elongate (Fig. 1 C), slightly widening in posterior part. Head black. Elytra of typical form, light testaceous with black pattern covering scutellar shield and surrounding portion of elytra through shoulders to lateral margins, covering about 60 % of its anterior part; pair of maculae in posterior 3 / 4 of elytra near suture; in western population fused and connected to scutellar shield over suture. Ventral side (Fig. 1 D) black with hypomera and ventrites 3 - 6 testaceous. Head and pronotum covered with uniform small setiferous punctures arranged irregularly. Pronotum transverse, broadly rounded laterally, with lateral margin glabrous; pronotum covered with dense setiferous punctures. Posterior pronotal corners not produced (Fig. 2 A). Prosternum with anterior margin with bordering line complete. Prosternal process with complete lateral carinae, joined roundly and merged with anterior border of pronotum (Fig. 2 B). Scutellar shield pentagonal, covered with dense setiferous punctures. Elytra (Fig. 2 D) covered with two types of punctures, small setiferous punctures irregularly distributed throughout the elytral surface, some of these punctures surrounded by larger depressed circles forming nine irregular longitudinal rows along the whole length of elytra. Shoulder tubercles distinct, but lateral elytral margin of elytra visible from above throughout. Mesoventrite (Fig. 2 C) with anterior border interrupted in median part. Metaventrite (Fig. 2 C) with postcoxal lines transverse in median part and then descending laterally, continuous on the metaventral process in median part; covered with setiferous punctures very sparsely distributed in central part of sclerite, densely setose in anterolateral parts, with a single row of large punctures below postcoxal lines and above metacoxae. Abdominal postcoxal lines (Fig. 2 E) complete, widely rounded, reaching about half of the length of the ventrite 1 measured below metacoxa. Ventrites covered with dense setiferous punctures. Male genitalia. Tegmen in inner view with penis guide subtriangular with pointed apex; short, about two times shorter than parameres. Parameres elongate elliptical, inner surface smooth, with long setae on the inner side and in apical margin. Penis simple with pointed apex. [see Gordon 1985: 657, fig. 539 a-d] Female genitalia. Sperm duct long, much longer than length of spermatheca. Spermatheca vermiform, broadest in basal part. [see Gordon 1985: 657, fig. 539 e]	en	Szawaryn, Karol, Nedved, Oldrich, Biranvand, Amir, Czerwinski, Tomasz, Nattier, Romain (2021): Revision of the genus Coccidula Kugelann (Coleoptera, Coccinellidae). ZooKeys 1043: 61-85, DOI: http://dx.doi.org/10.3897/zookeys.1043.65829, URL: http://dx.doi.org/10.3897/zookeys.1043.65829
4CE4208FE9C651F693F60F46591A20FB.taxon	distribution	Distribution. North part of North America.	en	Szawaryn, Karol, Nedved, Oldrich, Biranvand, Amir, Czerwinski, Tomasz, Nattier, Romain (2021): Revision of the genus Coccidula Kugelann (Coleoptera, Coccinellidae). ZooKeys 1043: 61-85, DOI: http://dx.doi.org/10.3897/zookeys.1043.65829, URL: http://dx.doi.org/10.3897/zookeys.1043.65829
4CDB34600B8458C19ED8645728163B8E.taxon	materials_examined	Material examined. Holotype. Azerbaijan, " Caucasus Araxesthal Leder Reitter / Coll. Reitter / Coccidula litophiloides 1890 / Holotypus 1890 Coccidula litophiloides Reitter ", male (HNHM) (Fig. 11 C). Holotype of L. naviauxi, Iran, Vannae, 30 - V- 77, leg. M. Rapilly, female (MNHN). Paratypes of C. litophiloides. Data same as for the holotype, (7: HNHM). Paratypes of L. naviauxi: Iran, Daran, 9 - VI- 77, M. Rapilly leg. (2 females: MNHN) (Figs 11 A, B). Other material. Armenia, Eczmiadzin Cauc, 22 IV 1946, 6399, W. Eichler (2: MIZ); Jerevan mesto, Razdan, 26 - 27.5.1988, J. Strejcek lgt., (1 male, 1 female: NMP); Iran, Lorestan, 1.1960 leg. A. Warchalowski (1: AJC); Iran, Khorramabad, 19 - V- 77, M. Rapilly leg. (1: MNHN).	en	Szawaryn, Karol, Nedved, Oldrich, Biranvand, Amir, Czerwinski, Tomasz, Nattier, Romain (2021): Revision of the genus Coccidula Kugelann (Coleoptera, Coccinellidae). ZooKeys 1043: 61-85, DOI: http://dx.doi.org/10.3897/zookeys.1043.65829, URL: http://dx.doi.org/10.3897/zookeys.1043.65829
4CDB34600B8458C19ED8645728163B8E.taxon	diagnosis	Diagnosis. Coccidula litophiloides is very distinctive among Coccidula species with large produced posterior pronotal angles, and a prosternal process without carinae (which are present in all remaining species). With its general body shape slightly widening posteriorly and pronotum distinctly widened laterally with broad lateral bead appearing glabrous, it is similar to C. scutellata. Male genitalia are distinctive with large, elliptical parameres possessing projections on their inner surfaces, which is also unique among Coccidula. Spermatheca, in female genitalia, is distinctly widening apically and has a very short sperm duct, about 1 / 4 of the length of spermatheca.	en	Szawaryn, Karol, Nedved, Oldrich, Biranvand, Amir, Czerwinski, Tomasz, Nattier, Romain (2021): Revision of the genus Coccidula Kugelann (Coleoptera, Coccinellidae). ZooKeys 1043: 61-85, DOI: http://dx.doi.org/10.3897/zookeys.1043.65829, URL: http://dx.doi.org/10.3897/zookeys.1043.65829
4CDB34600B8458C19ED8645728163B8E.taxon	description	Description. Length 3.0 - 3.5 mm, BL / BW = 1.95 - 1.97, EL / BW = 1.32 - 1.40, PW / BW = 0.81. Body elongate, slightly widening in posterior part. Dorsal and ventral side yellow to testaceous (Fig. 1 E). Head and pronotum covered with uniform small setiferous punctures arranged irregularly. Pronotum transverse, broadly rounded laterally (Figs 1 E, 3 B), with broad, glabrous lateral margin, covered with dense setriferous punctures, with a single row of larger punctures along lateral border. Posterior pronotal corner large, distinctly pointed (Fig. 3 B). Prosternum with complete anterior bordering line. Prosternal process without lateral carinae (Fig. 3 C). Scutellar shield pentagonal, covered with dense setiferous punctures. Elytra covered with two types of punctures, small setiferous punctures irregularly distributed throughout elytral surface, some of these punctures surrounded by larger depressed circles, forming irregular longitudinal rows; rows 2 and 3 reduced or missing (Fig. 3 A). Elytra more flattened in lateral view than in other Coccidula, without shoulder tubercle, lateral elytral margin visible throughout (Fig. 3 A). Hind wings missing. Mesoventrite with anterior border complete. Metaventrite with postcoxal lines transverse, descending only laterally, fused on metaventral process in median part, forming continuous arc (Fig. 3 D); covered with setiferous punctures very sparsely distributed in central part of sclerite, densely setose in lateral parts, without distinct rows of large punctures below postcoxal lines, large punctures above metacoxae present. Abdominal postcoxal lines complete, rounded, reaching slightly less than half of length of the ventrite 1 measured below metacoxa. Ventrites covered with dense setiferous punctures. Male genitalia. Tegmen in inner view with penis guide pentagonal with pointed apex (Fig. 4 B); short, slightly longer than half length of parameres (Fig. 4 A). Parameres large, elliptical, inner surface with distinct projections (Fig. 4 B), with fringe of long setae in apical margin. Penis simple with pointed apex (Fig. 4 C). Female genitalia. Sperm duct short (Fig. 4 D), about as long as 1 / 4 of spermatheca. Spermatheca vermiform, distinctly broadened apically. Accessory gland membranous, longer than sperm duct.	en	Szawaryn, Karol, Nedved, Oldrich, Biranvand, Amir, Czerwinski, Tomasz, Nattier, Romain (2021): Revision of the genus Coccidula Kugelann (Coleoptera, Coccinellidae). ZooKeys 1043: 61-85, DOI: http://dx.doi.org/10.3897/zookeys.1043.65829, URL: http://dx.doi.org/10.3897/zookeys.1043.65829
4CDB34600B8458C19ED8645728163B8E.taxon	distribution	Distribution. Armenia, Azerbaijan, Iran	en	Szawaryn, Karol, Nedved, Oldrich, Biranvand, Amir, Czerwinski, Tomasz, Nattier, Romain (2021): Revision of the genus Coccidula Kugelann (Coleoptera, Coccinellidae). ZooKeys 1043: 61-85, DOI: http://dx.doi.org/10.3897/zookeys.1043.65829, URL: http://dx.doi.org/10.3897/zookeys.1043.65829
D80F83A50C5E5F05A570C7F38CA8AB88.taxon	materials_examined	Material examined. Holotype. Russia, " Quell. d. Jrbut Reitter. / Transbaikal leg. Leder / Coll. Reitter / / Coccidula scutellaris m 1896 / Coccidula reitteri Dodge Khnzorian det. / prep. genital R. Bielawski 1956 / Holotypus 1897 Coccidula suturalis Reitter / Photo ID: HNHM _ COL _ 574 ", female (HNHM). Other material. Russia, " Transbaikalien Leder Reitter / Coccidula suturalis Rtt. Coll. Reitter / Coccidula reitteri Dodge, det. Merkl 1984 / prep. genital R. Bielawski 1956 " (1 male: HNHM); Listvjanka pr. Bajkal, step, 29.6.1977, H. Karnecka lgt. (1 male, 1 female: NMP).	en	Szawaryn, Karol, Nedved, Oldrich, Biranvand, Amir, Czerwinski, Tomasz, Nattier, Romain (2021): Revision of the genus Coccidula Kugelann (Coleoptera, Coccinellidae). ZooKeys 1043: 61-85, DOI: http://dx.doi.org/10.3897/zookeys.1043.65829, URL: http://dx.doi.org/10.3897/zookeys.1043.65829
D80F83A50C5E5F05A570C7F38CA8AB88.taxon	diagnosis	Diagnosis. Coccidula reitteri is very similar to C. rufa in external appearance, however, it can be distinguished by the presence of a small black transverse macula on the pronotum just anterior to the scutellar shield, and a longitudinal brown to black macula on the posterior half of the elytra on the elytral suture. Male genitalia are very close to C. rufa, however, the upper margin of the penis guide in lateral view is relatively less emarginated and parameres are narrower than in C. rufa.	en	Szawaryn, Karol, Nedved, Oldrich, Biranvand, Amir, Czerwinski, Tomasz, Nattier, Romain (2021): Revision of the genus Coccidula Kugelann (Coleoptera, Coccinellidae). ZooKeys 1043: 61-85, DOI: http://dx.doi.org/10.3897/zookeys.1043.65829, URL: http://dx.doi.org/10.3897/zookeys.1043.65829
D80F83A50C5E5F05A570C7F38CA8AB88.taxon	description	Description. Length = 2.8 - 3.2 mm, BL / BW = 1.85 - 1.90, EL / BW = 1.33, PW / BW = 0.77. Body elongate, parallel sided (Fig. 5 A). Pronotum (Fig. 1 F) with black transverse macula in front of the scutellar shield. Scutellar shield black. Elytra brown with elongate, dark brown to black macula along the elytral suture in posterior half. Ventral side testaceous with prosternum, mesoventrite, metaventrite, most of the ventrite 1 (except lateral corners), and central part of ventrite 2 black. Head and pronotum covered with uniform small setiferous punctures arranged irregularly. Pronotum (Fig. 5 B) transverse, broadly rounded laterally, with moderately broad, lateral margin without glabrous area; pronotum covered with dense setriferous punctures, with single row of larger punctures along lateral border. Posterior pronotal corners not produced (Fig. 5 B). Prosternum with anterior margin with bordering line incomplete in median part, without small sub-rounded impression in center. Prosternal process with lateral carinae straight, joined together roundly at level of anterior border of procoxae, forming sub-triangular pattern (Fig. 5 C). Scutellar shield pentagonal, covered with dense setiferous punctures. Elytra covered with two types of punctures, small setiferous punctures irregularly distributed throughout the elytral surface, some of these punctures surrounded by larger depressed circles forming nine irregular longitudinal rows along whole length of elytra. Lateral elytral margin well visible throughout (Fig. 5 A). Mesoventrite with complete anterior border. Metaventrite with postcoxal lines descending laterally, fused on metaventral process in median part, forming continuous arc (Fig. 5 D); covered with setiferous punctures very sparsely distributed in central part of sclerite, densely setose in lateral parts, without distinct rows of large punctures below postcoxal lines, large punctures above metacoxae present. Abdominal postcoxal lines complete, arcuate, reaching half of length of ventrite 1 measured below metacoxa. Ventrites covered with sparse setiferous punctures. Male genitalia. Tegmen in inner view (Fig. 6 B) with penis guide broadly rounded in the median or apical part, with rounded apex; in lateral view (Fig. 6 C) moderately expanded medially, with upper surface moderately emarginate; long, much longer than parameres. Parameres elongate, parallel sided, with narrow base, inner surface smooth, with fringe of long setae in apical part. Penis simple with pointed apex, with small bump before apex. Female genitalia. Spermatheca vermiform, not distinctly broadened apically (Fig. 6 A).	en	Szawaryn, Karol, Nedved, Oldrich, Biranvand, Amir, Czerwinski, Tomasz, Nattier, Romain (2021): Revision of the genus Coccidula Kugelann (Coleoptera, Coccinellidae). ZooKeys 1043: 61-85, DOI: http://dx.doi.org/10.3897/zookeys.1043.65829, URL: http://dx.doi.org/10.3897/zookeys.1043.65829
D80F83A50C5E5F05A570C7F38CA8AB88.taxon	distribution	Distribution. Russia (East Siberia).	en	Szawaryn, Karol, Nedved, Oldrich, Biranvand, Amir, Czerwinski, Tomasz, Nattier, Romain (2021): Revision of the genus Coccidula Kugelann (Coleoptera, Coccinellidae). ZooKeys 1043: 61-85, DOI: http://dx.doi.org/10.3897/zookeys.1043.65829, URL: http://dx.doi.org/10.3897/zookeys.1043.65829
25B90092038B55CE86609DF331C51381.taxon	materials_examined	Material examined. Czech Rep., Zlin, 11.6.1999, lgt. L. Bures (1: NMP); Mlada Boleslav, 25.4.1987, lgt. Nedved (1 male USB); Dvoriste, 9.8.1989, lgt. Nedved (1: USB); Kokorinsky dul, 9.8.1995, lgt. J. Řehounek (1: USB); Kyrgyzstan, Toktogul, 26 VI 2003, leg. A. Lason, WJ 2870, (1 male: AJC); Montenegro, Skadar jez. - Virpazar, 5.6.1984, J. Strejcek lgt. (1: NMP); Poland, Kampinos Forest near Warsaw, 17.06.2020, leg. D. Marczak, (7: MIZ); Russia, Leningrad-Lachta, IX 1988, J. Strejcek lgt., (1: NMP); Ukraine, Kharkiv region, Dergachevsky district, Boliboki vill., 50 ° 9 ' 16.57 " N, 36 ° 3 ' 58.87 " E, 1.5.2017, lgt. A. Slutsky (1: ASC); Uzbekistan, Buchara / Coccidula unicolor Rtt. / Coll. Reitter / MIZ PAN Warszawa 27 / 1955 / 1 (1: MIZ). Type material not studied, deposited in Museum fuer Naturkunde, Berlin, Germany.	en	Szawaryn, Karol, Nedved, Oldrich, Biranvand, Amir, Czerwinski, Tomasz, Nattier, Romain (2021): Revision of the genus Coccidula Kugelann (Coleoptera, Coccinellidae). ZooKeys 1043: 61-85, DOI: http://dx.doi.org/10.3897/zookeys.1043.65829, URL: http://dx.doi.org/10.3897/zookeys.1043.65829
25B90092038B55CE86609DF331C51381.taxon	diagnosis	Diagnosis. Coccidula rufa is most similar in external appearance to C. reitteri, however it can be separated by the uniform testaceous coloration of the dorsal surface (C. reitteri possesses dark macula near the elytral suture). From uniformly colored specimens of C. scutellata it can be separated by the shape of carinae on the prosternal process. Male genitalia are also very distinctive: in C. scutellata penis guide is small, about half length of parameres, while in C. rufa it is longer than parameres. Spermatheca in female genitalia of C. rufa is vermiform, not widening apically, while in C. scutellata it is distinctly widened in apical part.	en	Szawaryn, Karol, Nedved, Oldrich, Biranvand, Amir, Czerwinski, Tomasz, Nattier, Romain (2021): Revision of the genus Coccidula Kugelann (Coleoptera, Coccinellidae). ZooKeys 1043: 61-85, DOI: http://dx.doi.org/10.3897/zookeys.1043.65829, URL: http://dx.doi.org/10.3897/zookeys.1043.65829
25B90092038B55CE86609DF331C51381.taxon	description	Description. Length = 2.5 - 3.2 mm, BL / BW = 1.88 - 2.00, EL / BW = 1.38 - 1.44, PW / BW = 0.80 - 0.82. Body elongate, parallel sided. Elytra of typical (European) form testaceous without maculae (Fig. 1 G), only scutellar shield dark brown to black. Ventral side testaceous with prosternal process, mesoventrite, metaventrite, most of the ventrite 1 (except lateral corners), and central part of ventrite 2 black. Head and pronotum covered with uniform small setiferous punctures arranged irregularly. Pronotum transverse, broadly rounded laterally, with moderately broad, lateral margin without glabrous area (Fig. 7 B); pronotum covered with dense setiferous punctures, with a single row of larger punctures along lateral border. Posterior pronotal corners not produced (Fig. 7 B). Prosternum with anterior margin with incomplete bordering line in median part, with a small sub-rounded impression in center. Prosternal process with lateral carinae straight, joined together roundly at level of anterior border of procoxae, forming sub-triangular pattern (Fig. 7 C). Scutellar shield pentagonal, covered with dense setiferous punctures. Elytra covered with two types of punctures, small setiferous punctures irregularly distributed throughout elytral surface, some of these punctures surrounded by larger depressed circles forming nine irregular longitudinal rows along whole length of elytra. Lateral elytral margin well visible throughout (Fig. 7 A). Mesoventrite with complete anterior border. Metaventrite with postcoxal lines descending laterally, fused on metaventral process in median part, forming continuous arc (Fig. 7 E), covered with setiferous punctures very sparsely distributed in central part of sclerite, densely setose in lateral parts, without distinct rows of large punctures below postcoxal lines, large punctures above metacoxae present. Abdominal postcoxal lines complete, arcuate, reaching half of length of ventrite 1 measured below metacoxa. Ventrites covered with sparse setiferous punctures. Male genitalia. Tegmen in inner view (Fig. 8 A) with penis guide sub-parallel to broadly rounded, with rounded apex; in lateral view (Fig. 8 B) expanded medially, with deeply emarginated upper margin; long, much longer than parameres. Parameres elongate, parallel sided, with just slightly narrower base, inner surface smooth, with fringe of long setae in apical part. Penis simple with sharply pointed and curved apex, with small bump before apex (Fig. 8 C). Female genitalia. Sperm duct long, longer than spermatheca (Fig. 8 E). Spermatheca vermiform, not distinctly broadened apically. Accessory gland membranous, much shorter than sperm duct.	en	Szawaryn, Karol, Nedved, Oldrich, Biranvand, Amir, Czerwinski, Tomasz, Nattier, Romain (2021): Revision of the genus Coccidula Kugelann (Coleoptera, Coccinellidae). ZooKeys 1043: 61-85, DOI: http://dx.doi.org/10.3897/zookeys.1043.65829, URL: http://dx.doi.org/10.3897/zookeys.1043.65829
25B90092038B55CE86609DF331C51381.taxon	distribution	Distribution. Europe (all countries), Africa: Morocco, Asia: Afghanistan, China, Russia (Siberia), Iran, Kazakhstan, Kyrgyzstan, Mongolia, Turkey, Uzbekistan.	en	Szawaryn, Karol, Nedved, Oldrich, Biranvand, Amir, Czerwinski, Tomasz, Nattier, Romain (2021): Revision of the genus Coccidula Kugelann (Coleoptera, Coccinellidae). ZooKeys 1043: 61-85, DOI: http://dx.doi.org/10.3897/zookeys.1043.65829, URL: http://dx.doi.org/10.3897/zookeys.1043.65829
69A8F176F3EC52D783FC33DB64C28042.taxon	materials_examined	Material examined. Armenia, Erevan, 9.06.1987, leg. V. Karasjov (5: AJC); Czech Rep., Praha-Kyje, 21.1.1945, lgt. Guennther, (1: NMP); Plzen, 20.7.1978, lgt. V. Mach, (2: USB); Kokorinsky dul, 28.8.1994, lgt. J. Řehounek (1: USB); Loucen, 17.8.1994, lgt. J. Řehounek (1: USB); France, St. Cucufa, VI 65, MD, Ch. ' Duverger det., J. P. Coutanceau det. 2004 ' (1: MNHN); Poland, Kampinos Forest near Warsaw, 17.06.2020, leg. D. Marczak (11: MIZ); Slovakia, Bratislava, 27.4.36, lgt. O. Kavan (1: NMP); Ukraine, Kharkiv region, Kharkiv district, Bobrovka vill., reserve " Aleshkina balka ", 2017 - 04 - 28, lgt. A. Slutsky (1: ASC). Type material not studied, deposited in Museum fuer Naturkunde, Berlin, Germany.	en	Szawaryn, Karol, Nedved, Oldrich, Biranvand, Amir, Czerwinski, Tomasz, Nattier, Romain (2021): Revision of the genus Coccidula Kugelann (Coleoptera, Coccinellidae). ZooKeys 1043: 61-85, DOI: http://dx.doi.org/10.3897/zookeys.1043.65829, URL: http://dx.doi.org/10.3897/zookeys.1043.65829
69A8F176F3EC52D783FC33DB64C28042.taxon	diagnosis	Diagnosis. Coccidula scutellata is the most variable species in body coloration. Typical forms with five black maculae on the elytra can be easily distinguished from other Coccidula species, however uniformly colored testaceous forms are externally similar to C. rufa. They can be easily distinguished by the shape of carinae on prosternal process, which are straight and form a sub-triangular pattern in C. rufa, and are sinuate and broadly rounded apically, and fused with anterior border of prosternum in C. scutellata. Moreover, C. scutellata has a more distinct shoulder tubercle, and relatively narrower protnotum. Also, the male genitalia are distinctive, with penis guide longer than parameres in C. rufa and much shorter in C. scutellata. Spermatheca, in female genitalia, is broadened apically in C. scutellata, while in C. rufa it is almost uniform in diameter.	en	Szawaryn, Karol, Nedved, Oldrich, Biranvand, Amir, Czerwinski, Tomasz, Nattier, Romain (2021): Revision of the genus Coccidula Kugelann (Coleoptera, Coccinellidae). ZooKeys 1043: 61-85, DOI: http://dx.doi.org/10.3897/zookeys.1043.65829, URL: http://dx.doi.org/10.3897/zookeys.1043.65829
69A8F176F3EC52D783FC33DB64C28042.taxon	description	Description. Length = 2.8 - 4.2 mm, BL / BW = 1.85 - 2.05, EL / BW = 1.36 - 1.46, PW / BW = 0.70 - 0.75. Body elongate, slightly widening in posterior part. Elytra of typical (European) form testaceous with five black maculae (Fig. 1 H), one large covering scutellar shield and surrounding portion of elytra, and four sub-oval maculae in the median part, two of which are placed close to elytral suture and remaining two, close to lateral margin. Sometimes macula surrounding scutellar shield extends along elytral suture, sometimes maculae placed in median part of elytra are fused, forming single band. Various forms with reductions of this pattern are also present to completely testaceous forms without any trace of black color. Ventral side testaceous with prosternal process, mesoventrite, metaventrite, most of ventrite 1 (except lateral corners), and central part of ventrite 2 black. Head and pronotum covered with uniform small setiferous punctures arranged irregularly. Pronotum transverse, broadly rounded laterally, with broad, glabrous lateral margin (Fig. 9 B); pronotum covered with dense setiferous punctures, with single row of larger punctures along lateral border. Posterior pronotal corners not produced. Prosternum with anterior margin with bordering line complete. Prosternal process with complete lateral carinae in form of sinuate line, joined roundly and merged with anterior border of pronotum (Fig. 9 E). Scutellar shield pentagonal, covered with dense setiferous punctures. Elytra covered with two types of punctures, small setiferous punctures irregularly distributed throughout elytral surface, some of these punctures surrounded by larger depressed circles forming nine irregular longitudinal rows along whole length of elytra. Shoulder tubercles distinct, lateral elytral margin of elytra not visible from above in anterior part (Fig. 9 A). Mesoventrite with anterior border interrupted in median part. Metaventrite with postcoxal lines transverse in median part and then descending laterally, not fused on metaventral process in median part (Fig. 9 D). Covered with setiferous punctures very sparsely distributed in central part of sclerite, densely setose in lateral parts, with single row of large punctures below postcoxal lines and above metacoxae. Abdominal postcoxal lines complete, rounded, reaching slightly more than half of length of ventrite 1 measured below metacoxa. Ventrites covered with dense setiferous punctures. Male genitalia. Tegmen in inner view (Fig. 10 A) with penis guide subtriangular with pointed apex; short, about two times shorter than parameres. Parameres elongate elliptical (Fig. 10 B), inner surface smooth, with long setae on inner surface and in apical margin. Penis simple with pointed apex (Fig. 10 C). Female genitalia. Sperm duct short, about as long as half of length of spermatheca (Fig. 10 E). Spermatheca vermiform, distinctly broadened apically. Accessory gland membranous, longer than sperm duct.	en	Szawaryn, Karol, Nedved, Oldrich, Biranvand, Amir, Czerwinski, Tomasz, Nattier, Romain (2021): Revision of the genus Coccidula Kugelann (Coleoptera, Coccinellidae). ZooKeys 1043: 61-85, DOI: http://dx.doi.org/10.3897/zookeys.1043.65829, URL: http://dx.doi.org/10.3897/zookeys.1043.65829
69A8F176F3EC52D783FC33DB64C28042.taxon	distribution	Distribution. Europe (all countries), Africa: Morocco, Asia: Kazakhstan, Russia (West Siberia).	en	Szawaryn, Karol, Nedved, Oldrich, Biranvand, Amir, Czerwinski, Tomasz, Nattier, Romain (2021): Revision of the genus Coccidula Kugelann (Coleoptera, Coccinellidae). ZooKeys 1043: 61-85, DOI: http://dx.doi.org/10.3897/zookeys.1043.65829, URL: http://dx.doi.org/10.3897/zookeys.1043.65829
