identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
1336C156E0055A7D8D59839CE87A724A.text	1336C156E0055A7D8D59839CE87A724A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Coccidula Kugelann 1798	<div><p>Coccidula Kugelann, 1798</p> <p>Coccidula Kugelann, 1798: 421. Type species: Chrysomela scutellata Herbst, 1783, by subsequent designation by Crotch 1874.</p> <p>Strongylus Panzer, 1813: 114.</p> <p>Cacidula Dejean, 1821: 132. Type species: Chrysomela pectoralis Fabricius, 1792 (= Dermestes rufus Herbst, 1783).</p> <p>Cacicula Stephens, 1831: 397.</p> <p>Diagnosis.</p> <p>Representatives of the genus Coccidula with its general body shape may resemble Tetrabrachys Kapur, however, it can be separated based on the structure of the tarsi which are tetramerous in both genera but in Coccidula the first tarsomere is sub-triangularly broadened apically and the second is elongate and distinctly lobbed, while in Tetrabrachys both the first and second are narrow, elongate and without lobes. Moreover, in Tetrabrachys the apical maxillary palpomere is widely securiform, and beetles are brachypterous, while in Coccidula the apical maxillary palpomere is only slightly widened and the second pair of wings is functional. Coccidula is also externally similar to European species of Rhyzobius but it can be separated based on the following characters: body almost parallel sided, elytra covered with punctures of two sizes, larger punctures arranged in nine rows (in C. litophiloides some of them are reduced), base of the pronotum not bordered, while in Rhyzobius the lateral body outline is broadly rounded, the elytra are covered with single sized, randomly arranged punctures, and base of the pronotum with distinct bordering line.</p> <p>Description.</p> <p>Body elongate-oval, with sides parallel (Fig. 1C-H), body flattened in lateral view, convex in cross-section; dorsum covered with setiferous punctures of two sizes (Figs 7A, 9A), hairs directed forwards on pronotum, backwards on elytra.</p> <p>Head partially withdrawn into prothorax (Fig. 1C-H); ventral antennal grooves shallow and moderately long, extending to posterior border of an eye (Fig. 3E). Eyes prominent, coarsely facetted (7-8 ommatidia per eye width), ocular canthus distinct, about as long as 4-5 ommatidium diameters; interocular distance about 3 × as eye diameter; interfacetal setae present only in basal part; temple behind eye distinctly longer than eye (Fig. 3E). Antennal insertion placed laterally, invisible from above, distance between antennal insertions about same as between eyes; frons around antennal insertions slightly expanded, covering antennal insertions, anterior tentorial pits placed ventrally below antennal insertions. Antennae (Figs 5A, E, 7A, D) longer than maximum head width including eyes, composed of 11 antennomeres (AN); scape simple, without projections, slightly curved; pedicel distinctly narrower than scape, elongate (1.5 × longer than wide); AN 3-8 elongate (AN3 ≈ 3.5 ×; AN8 ≈ 1.3 × longer than wide); AN 9-11 forming a loose, asymmetric club, ultimate AN truncate apically. Frontoclypeus short, transverse, anterior margin straight. Labrum entirely exposed, transverse, anterior margin straight. Mandibles asymmetric, bifid apically (Fig. 10G), molar part with basal tooth; prostheca distinct. Maxillary stipes (Figs 2B, 5C, 9C, 10H) with distinct groove for reception of maxillary palp in repose; palpomere 2 shorter than terminal (4th) one, slightly broadened apically; palpomere 3 about 2.3 × shorter than terminal one, subtriangular; terminal palpomere slightly securiform; lacinia with stiff setae on outer margin in apical half, with several additional spurs on surface (Fig. 10H). Labial palps (Figs 3E, 9C) with 3 palpomeres, inserted ventrally on prementum; palpomere 1 very small, apical palpomere as long as and about as broad as penultimate; distance between palp insertions about 1.5-2 × as its width. Prementum subquadrate, transverse apically. Mentum trapezoidal, broadest in anterior part, with horseshoe impression at base (Figs 7C, 9C). Submentum broad, transverse, with suture invisible.</p> <p>Anterior margin of pronotum weakly, broadly emarginate (Figs 3B, 5B) with anterior corners broadly rounded; lateral margins with moderately (Figs 7B, 9B) to distinctly expanded lateral beads (Fig. 3B), distinctly margined; hind corners sharply pointed; hind margin not bordered. Prothoracic hypomeron smooth, without delimited foveae (Figs 3C, 7C). Prosternum in front of coxae about as long as longitudinal length of procoxal cavity; anterior margin straight or slightly emarginate with distinct border. Prosternal process about 0.4 times of coxal diameter, surface smooth (Fig. 3C) or with lateral carinae (Figs 7C, 9E). Procoxal cavity oval, distinctly bordered anteriorly.</p> <p>Mesoventrite 1.3 × longer than its width at the level of mid coxae (Figs 1D, 5D, 7D); mesal surface with deep emargination for receiving tip of prosternal process (Fig. 2C); anterior margin with completely raised border. Meso-metaventral process narrow (Figs 1D, 2C, 5D, 7E), about 0.5 times of mesocoxal diameter, junction slightly arcuate (Figs 2C, 3D, 5D, 7E, 9D), with suture visible. Metendosternite with stalk sub-quadrate, tendons long, separated by a distance of about width of stalk and situated closer to center (Fig. 10I). Scutellar shield pentagonal (Figs 7B, 9B). Elytra at base wider than pronotum, lateral margins clearly visible from above throughout (Figs 2D, 3A, 5A, 7A) (except C. scutellata where it is obscured in basal part, Fig. 9A), surface covered with punctures of double size, smaller irregularly distributed, larger punctures arranged in nine irregular longitudinal rows. Sutural stria absent. Elytral epipleuron narrow, incomplete, reaching base of ventrite 4 (Fig. 1D), with complete bordering line, epipleural foveae absent. Hind wings fully developed or missing (in C. litophiloides). Metaventral postcoxal lines roundly joined medially, complete laterally, straight or descending (Figs 2C, 3D, 5D, 7E, 9D). Metaventrite with discrimen visible in posterior 2/3.</p> <p>Trochanters simple, subtriangular, without projection (figs 7E, 9D). Tibiae slightly expanded apically with one apical spur on forelegs, and two in mid and hind legs. Tarsi consisting of four tarsomeres, second tarsomere truncate apically; tarsal claws cleft apically (Fig. 9G) with single empodial seta present.</p> <p>Abdomen in both sexes with 6 ventrites (Fig. 1D); ventrite 1 about as long as ventrites 2-4 combined, ventrite 2 longer than ventrite 3, ventrites 3-5 subequal in length. Abdominal postcoxal lines (Figs 7E, 9D) separate medially, recurved and complete, reaching anterior margin of ventrite, posteriorly reaching about half length of ventrite 1. Ventrite 5 in female posteriorly rounded (Fig. 7F), in male truncate (Fig. 9F). Ventrite 6 rounded in both sexes.</p> <p>Male terminalia. Tegmen (Figs 4A, B, 8A, B, 10A, B) symmetrical; parameres articulated with penis guide. Penis (Figs 4C, 8C, 10C) slender, pointed apically; penis capsule asymmetrical with outer arm reduced, inner arm well developed. Apodeme of male sternum IX simple, not broadened apically (Figs 8D, 10D). Tergite X broadly rounded, semicircular (Figs 8D, 10D).</p> <p>Female terminalia. Coxites (Figs 10F) distinctly elongate, subtriangular; styli small but visible, bearing several short setae; infundibulum absent (Figs 4D, 8E, 10E); sperm duct simple. Spermatheca (Figs 4D, 8E, 10E) worm-like, without clear ramus or nodulus; spermathecal accessory gland small, elongate. Proctiger elongate, rounded apically (Fig. 10F).</p> <p>Immature stages.</p> <p>Larva as in Fig. 1A, pupa as in Fig. 1B.</p> <p>Distribution.</p> <p>Holarctic: Asia, Europe, Africa (North), North America.</p></div> 	http://treatment.plazi.org/id/1336C156E0055A7D8D59839CE87A724A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Szawaryn, Karol;Nedved, Oldrich;Biranvand, Amir;Czerwinski, Tomasz;Nattier, Romain	Szawaryn, Karol, Nedved, Oldrich, Biranvand, Amir, Czerwinski, Tomasz, Nattier, Romain (2021): Revision of the genus Coccidula Kugelann (Coleoptera, Coccinellidae). ZooKeys 1043: 61-85, DOI: http://dx.doi.org/10.3897/zookeys.1043.65829, URL: http://dx.doi.org/10.3897/zookeys.1043.65829
4CE4208FE9C651F693F60F46591A20FB.text	4CE4208FE9C651F693F60F46591A20FB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Coccidula lepida LeConte 1852	<div><p>Coccidula lepida LeConte, 1852 Figs 1C, D, 2A-E</p> <p>Coccidula lepida LeConte, 1852: 132.</p> <p>Coccidula occidentalis Horn, 1895: 114.</p> <p>Coccidula suturalis Weise, 1895: 132.</p> <p>Material examined.</p> <p>United States of America: America b., 82, coll. Růžička et Vokál, (1: NMP); Alaska, Mi.1249, Alaska Hwy., Dedman Lk., 6.-7.VII.1968, Campbell &amp; Smetana (1: NMP); Vermont, Korschefsky det. (2: MIZ); Canada: SK, Harris Reservoir, Hwy 21, 10 km S, Maple Creek, April 19 2016, drift D. Larson (1 female: NMP). Type material not studied, deposited in Museum of Comparative Zoology, Cambridge, USA.</p> <p>Diagnosis.</p> <p>Coccidula lepida is the only Nearctic species of the genus, and is similar in many characters to C. scutellata, but with the head and epipleura black. In the typical form (C. lepida described by LeConte), the black elytral pattern resembles an extension of the five fused black maculae on the elytra of C. scutellata, with shoulders and epipleura black. Shoulder tubercle distinct, prototum relatively narrow. Male genitalia with penis guide much shorter than parameres. Detailed description of morphology including variability in pattern can be found in Gordon (1985: 656-659).</p> <p>Description.</p> <p>Length = 2.7-3.5 mm, BL/BW = 1.88-1.96, EL/BW = 1.40-1.42, PW/BW = 0.73.</p> <p>Body elongate (Fig. 1C), slightly widening in posterior part. Head black. Elytra of typical form, light testaceous with black pattern covering scutellar shield and surrounding portion of elytra through shoulders to lateral margins, covering about 60% of its anterior part; pair of maculae in posterior 3/4 of elytra near suture; in western population fused and connected to scutellar shield over suture. Ventral side (Fig. 1D) black with hypomera and ventrites 3-6 testaceous.</p> <p>Head and pronotum covered with uniform small setiferous punctures arranged irregularly. Pronotum transverse, broadly rounded laterally, with lateral margin glabrous; pronotum covered with dense setiferous punctures. Posterior pronotal corners not produced (Fig. 2A). Prosternum with anterior margin with bordering line complete. Prosternal process with complete lateral carinae, joined roundly and merged with anterior border of pronotum (Fig. 2B).</p> <p>Scutellar shield pentagonal, covered with dense setiferous punctures. Elytra (Fig. 2D) covered with two types of punctures, small setiferous punctures irregularly distributed throughout the elytral surface, some of these punctures surrounded by larger depressed circles forming nine irregular longitudinal rows along the whole length of elytra. Shoulder tubercles distinct, but lateral elytral margin of elytra visible from above throughout. Mesoventrite (Fig. 2C) with anterior border interrupted in median part. Metaventrite (Fig. 2C) with postcoxal lines transverse in median part and then descending laterally, continuous on the metaventral process in median part; covered with setiferous punctures very sparsely distributed in central part of sclerite, densely setose in anterolateral parts, with a single row of large punctures below postcoxal lines and above metacoxae.</p> <p>Abdominal postcoxal lines (Fig. 2E) complete, widely rounded, reaching about half of the length of the ventrite 1 measured below metacoxa. Ventrites covered with dense setiferous punctures.</p> <p>Male genitalia. Tegmen in inner view with penis guide subtriangular with pointed apex; short, about two times shorter than parameres. Parameres elongate elliptical, inner surface smooth, with long setae on the inner side and in apical margin. Penis simple with pointed apex. [see Gordon 1985: 657, fig. 539 a-d]</p> <p>Female genitalia. Sperm duct long, much longer than length of spermatheca. Spermatheca vermiform, broadest in basal part. [see Gordon 1985: 657, fig. 539e]</p> <p>Type locality.</p> <p>Vermont (USA).</p> <p>Distribution.</p> <p>North part of North America.</p></div> 	http://treatment.plazi.org/id/4CE4208FE9C651F693F60F46591A20FB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Szawaryn, Karol;Nedved, Oldrich;Biranvand, Amir;Czerwinski, Tomasz;Nattier, Romain	Szawaryn, Karol, Nedved, Oldrich, Biranvand, Amir, Czerwinski, Tomasz, Nattier, Romain (2021): Revision of the genus Coccidula Kugelann (Coleoptera, Coccinellidae). ZooKeys 1043: 61-85, DOI: http://dx.doi.org/10.3897/zookeys.1043.65829, URL: http://dx.doi.org/10.3897/zookeys.1043.65829
4CDB34600B8458C19ED8645728163B8E.text	4CDB34600B8458C19ED8645728163B8E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Coccidula litophiloides Reitter 1890	<div><p>Coccidula litophiloides Reitter, 1890 Figs 1E, 3A-E, 4A-D</p> <p>Coccidula litophiloides Reitter, 1890: 176</p> <p>Lithophilus naviauxi Duverger, 1983: 83. syn. nov.</p> <p>Material examined.</p> <p>Holotype. Azerbaijan, "Caucasus Araxesthal Leder Reitter/ Coll. Reitter/ Coccidula litophiloides 1890/ Holotypus 1890 Coccidula litophiloides Reitter", male (HNHM) (Fig. 11C). Holotype of L. naviauxi, Iran, Vannae, 30-V-77, leg. M. Rapilly, female (MNHN). Paratypes of C. litophiloides. Data same as for the holotype, (7: HNHM). Paratypes of L. naviauxi: Iran, Daran, 9-VI-77, M. Rapilly leg. (2 females: MNHN) (Figs 11A, B). Other material. Armenia, Eczmiadzin Cauc, 22 IV 1946, 6399, W. Eichler (2: MIZ); Jerevan město, Razdan, 26-27.5.1988, J. Strejček lgt., (1 male, 1 female: NMP); Iran, Lorestan, 1.1960 leg. A. Warchałowski (1: AJC); Iran, Khorramabad, 19-V-77, M. Rapilly leg. (1: MNHN).</p> <p>Diagnosis.</p> <p>Coccidula litophiloides is very distinctive among Coccidula species with large produced posterior pronotal angles, and a prosternal process without carinae (which are present in all remaining species). With its general body shape slightly widening posteriorly and pronotum distinctly widened laterally with broad lateral bead appearing glabrous, it is similar to C. scutellata. Male genitalia are distinctive with large, elliptical parameres possessing projections on their inner surfaces, which is also unique among Coccidula. Spermatheca, in female genitalia, is distinctly widening apically and has a very short sperm duct, about ¼ of the length of spermatheca.</p> <p>Description.</p> <p>Length 3.0-3.5 mm, BL/BW = 1.95-1.97, EL/BW = 1.32-1.40, PW/BW = 0.81.</p> <p>Body elongate, slightly widening in posterior part. Dorsal and ventral side yellow to testaceous (Fig. 1E).</p> <p>Head and pronotum covered with uniform small setiferous punctures arranged irregularly. Pronotum transverse, broadly rounded laterally (Figs 1E, 3B), with broad, glabrous lateral margin, covered with dense setriferous punctures, with a single row of larger punctures along lateral border. Posterior pronotal corner large, distinctly pointed (Fig. 3B). Prosternum with complete anterior bordering line. Prosternal process without lateral carinae (Fig. 3C).</p> <p>Scutellar shield pentagonal, covered with dense setiferous punctures. Elytra covered with two types of punctures, small setiferous punctures irregularly distributed throughout elytral surface, some of these punctures surrounded by larger depressed circles, forming irregular longitudinal rows; rows 2 and 3 reduced or missing (Fig. 3A). Elytra more flattened in lateral view than in other Coccidula, without shoulder tubercle, lateral elytral margin visible throughout (Fig. 3A). Hind wings missing. Mesoventrite with anterior border complete. Metaventrite with postcoxal lines transverse, descending only laterally, fused on metaventral process in median part, forming continuous arc (Fig. 3D); covered with setiferous punctures very sparsely distributed in central part of sclerite, densely setose in lateral parts, without distinct rows of large punctures below postcoxal lines, large punctures above metacoxae present.</p> <p>Abdominal postcoxal lines complete, rounded, reaching slightly less than half of length of the ventrite 1 measured below metacoxa. Ventrites covered with dense setiferous punctures.</p> <p>Male genitalia. Tegmen in inner view with penis guide pentagonal with pointed apex (Fig. 4B); short, slightly longer than half length of parameres (Fig. 4A). Parameres large, elliptical, inner surface with distinct projections (Fig. 4B), with fringe of long setae in apical margin. Penis simple with pointed apex (Fig. 4C).</p> <p>Female genitalia. Sperm duct short (Fig. 4D), about as long as 1/4 of spermatheca. Spermatheca vermiform, distinctly broadened apically. Accessory gland membranous, longer than sperm duct.</p> <p>Type locality.</p> <p>Caucasus, Ordubad (Azerbaijan).</p> <p>Distribution.</p> <p>Armenia, Azerbaijan, Iran</p> <p>Remarks.</p> <p>Duverger (1983) described Lithophilus naviauxi from Iran. After examination of the type specimens (Fig. 1A, B) we noticed that this species does not belong to the genus Lithophilus Frölich (= Tetrabrachys Kapur). As drawn in the original publication (Duverger 1983), it has antennae with 11 antennomeres (10 in Tetrabrachys), and pseudotrimerous tarsi with tarsomere 3 very small and tarsomere 2 distinctly lobed, while in Tetrabrachys tarsi are distinctly tetramerous, with tarsomere 3 and 2 elongate, without distinct lobe. Duverger in his paper (1983) described L. naviauxi based on just three female specimens of which he illustrated the spermatheca (Duverger 1983: 89, figs 30, 31). However, C. litophiloides is also found in Iran. Comparison of the female genitalia of the type material of both taxa, and other available material, together with the lack of a second and third row of large punctures on the elytra, and other morphological features described in the original description of Duverger, led to the conclusion that L. naviauxi Duverger falls well within the definition of C. litophiloides; thus, we propose to synonymize both species.</p> </div>	http://treatment.plazi.org/id/4CDB34600B8458C19ED8645728163B8E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Szawaryn, Karol;Nedved, Oldrich;Biranvand, Amir;Czerwinski, Tomasz;Nattier, Romain	Szawaryn, Karol, Nedved, Oldrich, Biranvand, Amir, Czerwinski, Tomasz, Nattier, Romain (2021): Revision of the genus Coccidula Kugelann (Coleoptera, Coccinellidae). ZooKeys 1043: 61-85, DOI: http://dx.doi.org/10.3897/zookeys.1043.65829, URL: http://dx.doi.org/10.3897/zookeys.1043.65829
D80F83A50C5E5F05A570C7F38CA8AB88.text	D80F83A50C5E5F05A570C7F38CA8AB88.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Coccidula reitteri Dodge 1938	<div><p>Coccidula reitteri Dodge, 1938 Figs 1F, 5A-E, 6A-C</p> <p>Coccidula suturalis Reitter 1897: 127 nom. nud. (nec. C. suturalis Weise, 1895: 132).</p> <p>Coccidula reitteri Dodge, 1938: 222.</p> <p>Material examined.</p> <p>Holotype. Russia, "Quell. d. Jrbut Reitter./ Transbaikal leg. Leder/ Coll. Reitter/ / Coccidula scutellaris m 1896/ Coccidula reitteri Dodge Khnzorian det./ prep. genital R. Bielawski 1956/ Holotypus 1897 Coccidula suturalis Reitter/ Photo ID: HNHM_COL_574", female (HNHM). Other material. Russia, "Transbaikalien Leder Reitter/ Coccidula suturalis Rtt. Coll. Reitter/ Coccidula reitteri Dodge, det. Merkl 1984/ prep. genital R. Bielawski 1956" (1 male: HNHM); Listvjanka pr. Bajkal, step, 29.6.1977, H. Karnecka lgt. (1 male, 1 female: NMP).</p> <p>Diagnosis.</p> <p>Coccidula reitteri is very similar to C. rufa in external appearance, however, it can be distinguished by the presence of a small black transverse macula on the pronotum just anterior to the scutellar shield, and a longitudinal brown to black macula on the posterior half of the elytra on the elytral suture. Male genitalia are very close to C. rufa, however, the upper margin of the penis guide in lateral view is relatively less emarginated and parameres are narrower than in C. rufa.</p> <p>Description.</p> <p>Length = 2.8-3.2 mm, BL/BW = 1.85-1.90, EL/BW = 1.33, PW/BW = 0.77.</p> <p>Body elongate, parallel sided (Fig. 5A). Pronotum (Fig. 1F) with black transverse macula in front of the scutellar shield. Scutellar shield black. Elytra brown with elongate, dark brown to black macula along the elytral suture in posterior half. Ventral side testaceous with prosternum, mesoventrite, metaventrite, most of the ventrite 1 (except lateral corners), and central part of ventrite 2 black.</p> <p>Head and pronotum covered with uniform small setiferous punctures arranged irregularly. Pronotum (Fig. 5B) transverse, broadly rounded laterally, with moderately broad, lateral margin without glabrous area; pronotum covered with dense setriferous punctures, with single row of larger punctures along lateral border. Posterior pronotal corners not produced (Fig. 5B). Prosternum with anterior margin with bordering line incomplete in median part, without small sub-rounded impression in center. Prosternal process with lateral carinae straight, joined together roundly at level of anterior border of procoxae, forming sub-triangular pattern (Fig. 5C).</p> <p>Scutellar shield pentagonal, covered with dense setiferous punctures. Elytra covered with two types of punctures, small setiferous punctures irregularly distributed throughout the elytral surface, some of these punctures surrounded by larger depressed circles forming nine irregular longitudinal rows along whole length of elytra. Lateral elytral margin well visible throughout (Fig. 5A). Mesoventrite with complete anterior border. Metaventrite with postcoxal lines descending laterally, fused on metaventral process in median part, forming continuous arc (Fig. 5D); covered with setiferous punctures very sparsely distributed in central part of sclerite, densely setose in lateral parts, without distinct rows of large punctures below postcoxal lines, large punctures above metacoxae present.</p> <p>Abdominal postcoxal lines complete, arcuate, reaching half of length of ventrite 1 measured below metacoxa. Ventrites covered with sparse setiferous punctures.</p> <p>Male genitalia. Tegmen in inner view (Fig. 6B) with penis guide broadly rounded in the median or apical part, with rounded apex; in lateral view (Fig. 6C) moderately expanded medially, with upper surface moderately emarginate; long, much longer than parameres. Parameres elongate, parallel sided, with narrow base, inner surface smooth, with fringe of long setae in apical part. Penis simple with pointed apex, with small bump before apex.</p> <p>Female genitalia. Spermatheca vermiform, not distinctly broadened apically (Fig. 6A).</p> <p>Type locality.</p> <p>Mongolia, Russia (Krasnoyarsk region, Irkutsk region, Tuva)</p> <p>Distribution.</p> <p>Russia (East Siberia).</p> <p>Remarks.</p> <p>Coccidula reitteri is very similar to C. rufa in external morphological characters as well as the structure of male and female genitalia (Fig. 6A) (Bielawski 1984); thus, further investigation, preferably of molecular markers, should be conducted to confirm whether it is a separate species or an eastern population of C. rufa.</p> </div>	http://treatment.plazi.org/id/D80F83A50C5E5F05A570C7F38CA8AB88	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Szawaryn, Karol;Nedved, Oldrich;Biranvand, Amir;Czerwinski, Tomasz;Nattier, Romain	Szawaryn, Karol, Nedved, Oldrich, Biranvand, Amir, Czerwinski, Tomasz, Nattier, Romain (2021): Revision of the genus Coccidula Kugelann (Coleoptera, Coccinellidae). ZooKeys 1043: 61-85, DOI: http://dx.doi.org/10.3897/zookeys.1043.65829, URL: http://dx.doi.org/10.3897/zookeys.1043.65829
25B90092038B55CE86609DF331C51381.text	25B90092038B55CE86609DF331C51381.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Coccidula rufa (Herbst 1783)	<div><p>Coccidula rufa (Herbst, 1783) Figs 1A, B, G, 7A-F, 8A-E</p> <p>Dermestes rufus Herbst, 1783: 22.</p> <p>Chrysomela pectoralis Fabricius, 1792: 328.</p> <p>Silpha rosea Marscham, 1802: 123.</p> <p>Coccidula conferta Reitter, 1890: 176.</p> <p>Coccidula rufa var. unicolor Reitter, 1890: 176.</p> <p>Coccidula rufa var. nigropunctata Reitter, 1900: 220.</p> <p>Coccidula rufa var. plagiata Gerhardt, 1910: 556.</p> <p>Material examined.</p> <p>Czech Rep., Zlín, 11.6.1999, lgt. L. Bureš (1: NMP); Mladá Boleslav, 25.4.1987, lgt. Nedvěd (1 male USB); Dvořiště, 9.8.1989, lgt. Nedvěd (1: USB); Kokořínský důl, 9.8.1995, lgt. J. Řehounek (1: USB); Kyrgyzstan, Toktogul, 26 VI 2003, leg. A. Lasoń, WJ 2870, (1 male: AJC); Montenegro, Skadar jez.- Virpazar, 5.6.1984, J. Strejček lgt. (1: NMP); Poland, Kampinos Forest near Warsaw, 17.06.2020, leg. D. Marczak, (7: MIZ); Russia, Leningrad-Lachta, IX 1988, J. Strejček lgt., (1: NMP); Ukraine, Kharkiv region, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.066353&amp;materialsCitation.latitude=50.154606" title="Search Plazi for locations around (long 36.066353/lat 50.154606)">Dergachevsky district</a>, Boliboki vill., 50°9'16.57"N, 36°3'58.87"E, 1.5.2017, lgt. A. Slutsky (1: ASC); Uzbekistan, Buchara/ Coccidula unicolor Rtt./ Coll. Reitter/ MIZ PAN Warszawa 27/1955/1 (1: MIZ). Type material not studied, deposited in Museum für Naturkunde, Berlin, Germany.</p> <p>Diagnosis.</p> <p>Coccidula rufa is most similar in external appearance to C. reitteri, however it can be separated by the uniform testaceous coloration of the dorsal surface (C. reitteri possesses dark macula near the elytral suture). From uniformly colored specimens of C. scutellata it can be separated by the shape of carinae on the prosternal process. Male genitalia are also very distinctive: in C. scutellata penis guide is small, about half length of parameres, while in C. rufa it is longer than parameres. Spermatheca in female genitalia of C. rufa is vermiform, not widening apically, while in C. scutellata it is distinctly widened in apical part.</p> <p>Description.</p> <p>Length = 2.5-3.2 mm, BL/BW = 1.88-2.00, EL/BW = 1.38-1.44, PW/BW = 0.80-0.82.</p> <p>Body elongate, parallel sided. Elytra of typical (European) form testaceous without maculae (Fig. 1G), only scutellar shield dark brown to black. Ventral side testaceous with prosternal process, mesoventrite, metaventrite, most of the ventrite 1 (except lateral corners), and central part of ventrite 2 black.</p> <p>Head and pronotum covered with uniform small setiferous punctures arranged irregularly. Pronotum transverse, broadly rounded laterally, with moderately broad, lateral margin without glabrous area (Fig. 7B); pronotum covered with dense setiferous punctures, with a single row of larger punctures along lateral border. Posterior pronotal corners not produced (Fig. 7B). Prosternum with anterior margin with incomplete bordering line in median part, with a small sub-rounded impression in center. Prosternal process with lateral carinae straight, joined together roundly at level of anterior border of procoxae, forming sub-triangular pattern (Fig. 7C).</p> <p>Scutellar shield pentagonal, covered with dense setiferous punctures. Elytra covered with two types of punctures, small setiferous punctures irregularly distributed throughout elytral surface, some of these punctures surrounded by larger depressed circles forming nine irregular longitudinal rows along whole length of elytra. Lateral elytral margin well visible throughout (Fig. 7A). Mesoventrite with complete anterior border. Metaventrite with postcoxal lines descending laterally, fused on metaventral process in median part, forming continuous arc (Fig. 7E), covered with setiferous punctures very sparsely distributed in central part of sclerite, densely setose in lateral parts, without distinct rows of large punctures below postcoxal lines, large punctures above metacoxae present.</p> <p>Abdominal postcoxal lines complete, arcuate, reaching half of length of ventrite 1 measured below metacoxa. Ventrites covered with sparse setiferous punctures.</p> <p>Male genitalia. Tegmen in inner view (Fig. 8A) with penis guide sub-parallel to broadly rounded, with rounded apex; in lateral view (Fig. 8B) expanded medially, with deeply emarginated upper margin; long, much longer than parameres. Parameres elongate, parallel sided, with just slightly narrower base, inner surface smooth, with fringe of long setae in apical part. Penis simple with sharply pointed and curved apex, with small bump before apex (Fig. 8C).</p> <p>Female genitalia. Sperm duct long, longer than spermatheca (Fig. 8E). Spermatheca vermiform, not distinctly broadened apically. Accessory gland membranous, much shorter than sperm duct.</p> <p>Type locality.</p> <p>Berlin (Germany)</p> <p>Distribution.</p> <p>Europe (all countries), Africa: Morocco, Asia: Afghanistan, China, Russia (Siberia), Iran, Kazakhstan, Kyrgyzstan, Mongolia, Turkey, Uzbekistan.</p></div> 	http://treatment.plazi.org/id/25B90092038B55CE86609DF331C51381	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Szawaryn, Karol;Nedved, Oldrich;Biranvand, Amir;Czerwinski, Tomasz;Nattier, Romain	Szawaryn, Karol, Nedved, Oldrich, Biranvand, Amir, Czerwinski, Tomasz, Nattier, Romain (2021): Revision of the genus Coccidula Kugelann (Coleoptera, Coccinellidae). ZooKeys 1043: 61-85, DOI: http://dx.doi.org/10.3897/zookeys.1043.65829, URL: http://dx.doi.org/10.3897/zookeys.1043.65829
69A8F176F3EC52D783FC33DB64C28042.text	69A8F176F3EC52D783FC33DB64C28042.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Coccidula scutellata (Herbst 1783)	<div><p>Coccidula scutellata (Herbst, 1783) Figs 1H, 9A-G, 10A-I</p> <p>Chrysomela scutellata Herbst, 1783: 58.</p> <p>Nitidula quinquepunctata Fabricius, 1787: 52.</p> <p>Silpha melanophthalma Gmelin, 1790: 1627.</p> <p>Nitidula bipunctata Gmelin, 1790: 1630</p> <p>Coccidula scutellata: Kugelann 1798: 421.</p> <p>Coccidula scutellata var. subrufa Weise, 1879: 131.</p> <p>Coccidula scutellata var. arquata Weise, 1879: 131.</p> <p>Coccidula scutellata var. aethiops Krauss, 1902: 92.</p> <p>Material examined.</p> <p>Armenia, Erevan, 9.06.1987, leg. V. Karasjov (5: AJC); Czech Rep., Praha-Kyje, 21.1.1945, lgt. Günnther, (1: NMP); Plzeň, 20.7.1978, lgt. V. Mach, (2: USB); Kokořínský důl, 28.8.1994, lgt. J. Řehounek (1: USB); Loučeň, 17.8.1994, lgt. J. Řehounek (1: USB); France, St. Cucufa, VI 65, MD, Ch. 'Duverger det., J.P. Coutanceau det. 2004' (1: MNHN); Poland, Kampinos Forest near Warsaw, 17.06.2020, leg. D. Marczak (11: MIZ); Slovakia, Bratislava, 27.4.36, lgt. O. Kavan (1: NMP); Ukraine, Kharkiv region, Kharkiv district, Bobrovka vill., reserve "Aleshkina balka", 2017-04-28, lgt. A. Slutsky (1: ASC). Type material not studied, deposited in Museum für Naturkunde, Berlin, Germany.</p> <p>Diagnosis.</p> <p>Coccidula scutellata is the most variable species in body coloration. Typical forms with five black maculae on the elytra can be easily distinguished from other Coccidula species, however uniformly colored testaceous forms are externally similar to C. rufa. They can be easily distinguished by the shape of carinae on prosternal process, which are straight and form a sub-triangular pattern in C. rufa, and are sinuate and broadly rounded apically, and fused with anterior border of prosternum in C. scutellata. Moreover, C. scutellata has a more distinct shoulder tubercle, and relatively narrower protnotum. Also, the male genitalia are distinctive, with penis guide longer than parameres in C. rufa and much shorter in C. scutellata. Spermatheca, in female genitalia, is broadened apically in C. scutellata, while in C. rufa it is almost uniform in diameter.</p> <p>Description.</p> <p>Length = 2.8-4.2 mm, BL/BW = 1.85-2.05, EL/BW = 1.36-1.46, PW/BW = 0.70-0.75.</p> <p>Body elongate, slightly widening in posterior part. Elytra of typical (European) form testaceous with five black maculae (Fig. 1H), one large covering scutellar shield and surrounding portion of elytra, and four sub-oval maculae in the median part, two of which are placed close to elytral suture and remaining two, close to lateral margin. Sometimes macula surrounding scutellar shield extends along elytral suture, sometimes maculae placed in median part of elytra are fused, forming single band. Various forms with reductions of this pattern are also present to completely testaceous forms without any trace of black color. Ventral side testaceous with prosternal process, mesoventrite, metaventrite, most of ventrite 1 (except lateral corners), and central part of ventrite 2 black.</p> <p>Head and pronotum covered with uniform small setiferous punctures arranged irregularly. Pronotum transverse, broadly rounded laterally, with broad, glabrous lateral margin (Fig. 9B); pronotum covered with dense setiferous punctures, with single row of larger punctures along lateral border. Posterior pronotal corners not produced. Prosternum with anterior margin with bordering line complete. Prosternal process with complete lateral carinae in form of sinuate line, joined roundly and merged with anterior border of pronotum (Fig. 9E).</p> <p>Scutellar shield pentagonal, covered with dense setiferous punctures. Elytra covered with two types of punctures, small setiferous punctures irregularly distributed throughout elytral surface, some of these punctures surrounded by larger depressed circles forming nine irregular longitudinal rows along whole length of elytra. Shoulder tubercles distinct, lateral elytral margin of elytra not visible from above in anterior part (Fig. 9A). Mesoventrite with anterior border interrupted in median part. Metaventrite with postcoxal lines transverse in median part and then descending laterally, not fused on metaventral process in median part (Fig. 9D). Covered with setiferous punctures very sparsely distributed in central part of sclerite, densely setose in lateral parts, with single row of large punctures below postcoxal lines and above metacoxae.</p> <p>Abdominal postcoxal lines complete, rounded, reaching slightly more than half of length of ventrite 1 measured below metacoxa. Ventrites covered with dense setiferous punctures.</p> <p>Male genitalia. Tegmen in inner view (Fig. 10A) with penis guide subtriangular with pointed apex; short, about two times shorter than parameres. Parameres elongate elliptical (Fig. 10B), inner surface smooth, with long setae on inner surface and in apical margin. Penis simple with pointed apex (Fig. 10C).</p> <p>Female genitalia. Sperm duct short, about as long as half of length of spermatheca (Fig. 10E). Spermatheca vermiform, distinctly broadened apically. Accessory gland membranous, longer than sperm duct.</p> <p>Type locality.</p> <p>Pomerania (Germany, Poland)</p> <p>Distribution.</p> <p>Europe (all countries), Africa: Morocco, Asia: Kazakhstan, Russia (West Siberia).</p></div> 	http://treatment.plazi.org/id/69A8F176F3EC52D783FC33DB64C28042	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Szawaryn, Karol;Nedved, Oldrich;Biranvand, Amir;Czerwinski, Tomasz;Nattier, Romain	Szawaryn, Karol, Nedved, Oldrich, Biranvand, Amir, Czerwinski, Tomasz, Nattier, Romain (2021): Revision of the genus Coccidula Kugelann (Coleoptera, Coccinellidae). ZooKeys 1043: 61-85, DOI: http://dx.doi.org/10.3897/zookeys.1043.65829, URL: http://dx.doi.org/10.3897/zookeys.1043.65829
