taxonID	type	description	language	source
03A187D11868483AA2ED53145E49F82A.taxon	description	Venation (Fig. 5). Forewing with cell longer than half length of wing, R 1 from middle of cell, R 3 from near anterior angle of cell, R 4 and R 5 stalked, R 5 to costa, M 2 almost parallel with M 1, M 3 and CuA 1 nearly connate, 1 A + 2 A biforked basally. Hindwing with cell nearly half length of wing, Rs and M 1 connate at anterior angle of cell, M 3 and CuA 1 connate at posterior angle of cell. Male genitalia (Figs 6 – 13). Eighth tergite tongue-shaped, with a pair of long corematae; eighth sternite shorter than eighth tergite, broad, fan-shaped, posterior margin broadened to rounded (Fig. 13). Uncus strongly developed, posterior margin usually with several incisions or emarginations of same or different depths, forming several branches of same or different lengths (Fig. 6); basal part with a pair of symmetric affiliated plates convex into different shapes (Figs 9 – 12). Tegumen nearly trapezoidal, anterior margin deeply concave to triangle, forming a distinct groove. Gnathos with lateral sclerites wedge-shaped or triangular, middle process well developed, normally hook-shaped, dilated at base in some species. Valva extending beyond tip of uncus, flagelliform and arched. Sacculus short, narrow, apex pointed or beak-like. Juxta usually as a pair of wedge-shaped sclerites. Saccus rectangular or subtriangular (Fig. 6, 36 b). Phallus often stout, slightly arched, narrowed from base to backward-curved and strongly sclerotized apex, usually with numerous microthorns on distal half of dorsal surface (Figs 6 – 7). Female genitalia (Figs 14 – 16). Papillae anales sub-rectangular, setose. Apophyses posteriores extremely long. Subgenital plate strongly sclerotized, consisting of two sclerites, outer one sub-semicircular or sub-triangular, usually with an emargination distomedially; inner sclerite with a longitudinal broad crack posteromedially, posteriorly fused with apophyses anteriores and forming two various-shaped sclerites which bearing one or several folds; sometimes subgenital plate crown-shaped, with a highly microtrichiate membrane (Figs 14 – 15). Apophyses anteriores short, rod-shaped. Antrum usually pentagonal. Ductus bursae broad and short, posteriorly with a sclerotized plate near antrum. Corpus bursae oval or rounded; signum cross-shaped, sub-hexagonal or sub-elliptical, with serrate edges and a horizontal axis, surface with dense denticles. Abdominal segment VII of Psoricoptera curva sp. nov. with tufts of modified scales on both sides (Fig. 16).	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D11868483AA2ED53145E49F82A.taxon	diagnosis	Diagnosis. Psoricoptera can be easily recognized by the uncus shovel-shaped, usually with a pair of middle and lateral branches in the male genitalia; in the female genitalia, the species of Psoricoptera can be recognized by the strongly sclerotized subgenital plate with the various-shaped sclerites at base of apophyses anteriores. The raised scales on upper surface of third segment of labial palpus is an additional diagnostic character for separating species of Psoricoptera from other genera of Gelechiini.	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D11868483AA2ED53145E49F82A.taxon	biology_ecology	Biology. Six plant families (Fagaceae, Salicaceae, Juglandaceae, Anacardiaceae, Rosaceae, Betulaceae) have been recorded as host plants of Psoricoptera species. Kaitila (1996) recorded the larva of P. speciosella feeds on Salix sp. in late June and early July in Europe; Arahou et al. (1991) reported larva of P. gibbosella lives in buds of Quercus ilex in Morocco; P. gibbosella has been reared from Quercus mongolica and Salix rorida Lacksch. in the Russian Far East (Omelko 1999), and Rhus chinensis is reported as the host plant of P. kawabei (Park & Karsholt 1999). In China, Juglans mandschurica, Quercus sp. and Salix sp. were reported as host plants of Psoricoptera species (Liu & Bai 1979), host plants of P. gibbosella have been recorded as Juglans mandshurica Maxim., Quercus robur L., Salix caprea L., Malus pumila Mill. Crataegus sp., Padus sp. Eriophyes sp. and Carpinus sp. (Li 2002), but Piskunov (1981) once thought records of larvae from Crataegus, Malus and galls of Eriophyes mites need confirmation. However, due to the inaccuracy of the previous identification of Psoricoptera species, the records of the above-mentioned host plants are doubtful.	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D11868483AA2ED53145E49F82A.taxon	distribution	Distribution. The genus Psoricoptera is predominantly recorded from the Palaearctic Region prior to this study (Park & Karsholt 1999; Li 2002). In this paper, we also find several Oriental species of Psoricoptera, which mainly are distributed in southwest China.	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D1186F4839A2ED54D25D28F891.taxon	description	(Figs 9, 14 – 17, 31)	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D1186F4839A2ED54D25D28F891.taxon	materials_examined	Type material. CHINA: Holotype ♂, Yunnan Province: Nankang Management Station (24.82 ° N, 98.78 ° E), Baoshan City, 2009 m, 9. viii. 2014, coll. Kaijian Teng and Xia Bai, genitalia slide No. ZML 19302. Paratype: Yunnan Province: 1 ♀, Mt. Jizu (25.96 ° N, 100.30 ° E), Dali Bai Autonomous Prefecture, 2228 m, 27. vii. 2014, coll. Kaijian Teng, Wei Guan, Xiuchun Wang and Shurong Liu, genitalia slide No. ZML 19301.	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D1186F4839A2ED54D25D28F891.taxon	description	Description. Adult (Fig. 17) wingspan 14.5 – 17.0 mm. Head shining blackish grey, sometimes light yellow medially. Labial palpus blackish brown except inner surface of second segment white; third segment with tuft of yellowish brown scales tipped with white, apex reddish orange. Antenna black. Thorax and tegula shining black. Forewing ground colour dark yellowish brown, scales tipped with blackish brown; costal spots at basal 2 / 5 and 3 / 5 respectively; four blackish brown tufts at distal 1 / 3 and apex of cell, basal 1 / 3 and 2 / 3 of fold respectively; basal 1 / 5 of dorsum black; fringe dark grey, intermixed with scales tipped with white or yellowish brown. Hindwing silver grey, darker toward apex; fringe grey. Female individual with forewing almost entirely blackish brown except distal 1 / 4 of costa tinged with a few yellowish brown scales. Fore and mid legs blackish brown; tibiae with tufts blackish brown mixed with yellowish brown, scales tipped with white; tarsi with first, second and fifth tarsomeres ringed with reddish brown apically; outer surface of hindleg blackish brown and white on inner surface, all tarsomeres ringed with yellowish white apically. Male genitalia (Figs 9, 31). Uncus single branch, basal 4 / 5 conic, distal 1 / 5 clavate, apex narrowly rounded; affiliated plates about half length of uncus, rodlike but excurved to S-shape. Gnathos with middle process strongly sclerotized, hook-like. Valva broad flagelliform, inner margin slightly convex from basal 1 / 6 to 1 / 3, distal 1 / 6 gently dilated to drop-shaped, then sharply narrowed to pointed and setose apex. Sacculus about 1 / 7 length of valva, beaklike, setose. Juxta double wave-shaped, innerside with a pair of leaf-shaped sclerites. Saccus less than half length of valva, rectangular, stout. Phallus accordance with the typical shape of this genus, dorsally without microthorns but with a small denticle apically. Female genitalia (Figs 14 – 16). Abdomen VII with two black tufts of modified scales laterally. Apophyses posteriores about 6 × length of subgenital plate. Subgenital plate crown-shaped, posteriorly connected with a highly microtrichiate membrane. Apophyses anteriores extending to end of subgenital plate and about 2 × length of it, with a pair of hemispherical plates posteriorly. Antrum with rugby-shape rough area anteromedially. Ductus bursae without sclerite. Corpus bursae about same length of ductus bursae, oval; signum broadly cross-shaped.	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D1186F4839A2ED54D25D28F891.taxon	diagnosis	Diagnosis. Psoricoptera curva sp. nov. can be recognized by the third segment of labial palpus with apex reddish orange externally; and by the uncus with affiliated plates excurved to S-shape, the valva with distal 1 / 6 gently dilated to drop-shape in the male genitalia; and can be further separated from its allies by the abdomen VII with a pair of black tufts of modified scales in female, and the apophyses anteriores with a pair of hemispherical plates posteriorly.	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D1186F4839A2ED54D25D28F891.taxon	distribution	Distribution. China (Yunnan).	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D1186F4839A2ED54D25D28F891.taxon	etymology	Etymology. The specific name is derived from the Latin curvus, meaning curve, referring to the shape of affiliated plates of uncus.	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D118624834A2ED562B5D09FCDA.taxon	description	(Figs 19 – 20, 32 – 34)	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D118624834A2ED562B5D09FCDA.taxon	materials_examined	Material examined. CHINA: Shanxi Province: 1 ♂, Mt. Lingkong (36.61 ° N, 112.08 ° E), Qinyuan County, 1560 m, 2. viii. 2013, coll. Tengteng Liu and Peixin Cong; 1 ♂, Mt. Lingkong (36.61 ° N, 112.08 ° E), Qinyuan County, 1560 m, 26. vii. 2014, coll. Tengteng Liu, Meiqing Yang and Sihan Lu, genitalia slide No. ZML 19307. Xinjiang Uygur Autonomous Region: 2 ♂♂, Balian (43.26 ° N, 82.85 ° E), Mohe Town, Gongliu County, 1206 m, 5. viii. 2007, coll. Xinpu Wang et al., genitalia slide Nos. ZML 19286, ZML 19318.	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D118624834A2ED562B5D09FCDA.taxon	diagnosis	Diagnosis. Adult (Figs 19 – 20) wingspan 15.5 – 16.0 mm. This species is similar to P. speciosella both in adult and male genitalia, it can be distinguished from latter by the uncus with four branches about same height, middle incision normally not deeper than lateral incisions (Figs 32 – 34). In P. speciosella, the uncus with middle branches higher than lateral branches and middle incision deeper than lateral incisions (Figs 39 – 39 d).	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D118624834A2ED562B5D09FCDA.taxon	description	Variation. Uncus with middle incision variable, usually same depth with lateral incisions but can be deeper in some individual (Figs 32 – 34).	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D118624834A2ED562B5D09FCDA.taxon	distribution	Distribution. China (Shanxi, Xinjiang), Japan, Korea, Russia, Europe, Asia Minor, North Africa (Park & Karsholt 1999).	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D118624834A2ED562B5D09FCDA.taxon	discussion	Remarks. Huemer & Karsholt (1999) and Park & Karsholt (1999) mainly used incisions on posterior margin of uncus and number of microspines (which we called microthorns in the present study) on phallus to distinguish P. gibbosella and P. speciosella in male genitalia: they considered uncus of P. gibbosella with middle incision not deeper than lateral incisions, phallus with more than 20 microspines; while the uncus of P. speciosella with middle incision deeper than lateral incisions, phallus with less than 16 microspines. Kaitila (1996) considered biology and colour of larvae as other effective evidence in identification of the two species. In this study, many more specimens have been checked and dissected. As expected, more morphological variations on genitalia are observed, but the microthorns on phallus and vinculum seemingly are unstable, therefore we don't think the two characters can hold as diagnostic features for Psoricoptera species. We have not been able to collect or rear immature stages of the Psoricoptera species, the larvae can be associated with adults taking advantage of molecular techniques in the subsequent study.	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D118604832A2ED52F45E2EFD83.taxon	description	(Figs 2 – 3, 5 – 7, 18, 35 – 35 g, 40 – 43)	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D118604832A2ED52F45E2EFD83.taxon	materials_examined	Material examined. CHINA: Beijing: 1 ♀, Mid-Hill Pavilion (39.99 ° N, 116.21 ° E), 200 m, 23. vi. 2000, coll. Zhendong Li, genitalia slide No. ZML 19308; 1 ♂, Dakezhuang Village (40.52 ° N, 115.84 ° E), Mt. Song, 1. viii. 2010, coll. Aihuan Zhang and Xiaoyu Shi; 10 ♂♂, Dakezhuang Village (40.52 ° N, 115.84 ° E), Mt. Song, 1,19,23. viii. 2010, coll. Aihuan Zhang and Zheng Li, genitalia slide Nos. ZML 19335, ZML 19336; 5 ♂♂, Dakezhuang Village (40.52 ° N, 115.84 ° E), Mt. Song, 19. viii. 2010, coll. Aihuan Zhang and Jingxue Cai, genitalia slide Nos. ZML 19344 W, ZML 19346 W. Tianjin: 2 ♂♂, Mt. Baxian (40.16 ° N, 117.56 ° E), Ji County, 500 m, 7 – 9. ix. 2001, coll. Houhun Li, genitalia slide Nos. LZD 01283, SYW 05189; 12 ♂♂, 4 ♀♀, Mt. Baxian (40.16 ° N, 117.56 ° E), 500 m, 8 – 11. ix. 2005, Coll. Houhun Li et al., genitalia slide Nos. SYW 05455 ♂, ZML 19309 ♂, ZML 19340 W, ZML 19343 W; 1 ♀, Mt. Baxian, Ji County, 110 m, 22. ix. 2006, coll. Houhun Li et al.; 1 ♀, Mt. Baxian, Ji County, 600 m, 23. ix. 2006, coll. Houhun Li et al.; 1 ♂, 5 ♀♀, Heishui River (40.18 ° N, 117.56 ° E), Mt. Baxian, 550 m, 6,24,31. viii, 16. ix. 2009, coll. Zhipin Liang, genitalia slide No. ZML 19338 ♀; 1 ♀, Heishui River (40.18 ° N, 117.56 ° E), Mt. Baxian, 550 m, 24. viii. 2009, coll. Jiang Zhang; 7 ♂♂, 8 ♀♀, Heishui River (40.18 ° N, 117.56 ° E), Mt. Baxian, 600 m, 16. vii, 10,27. viii, 3 – 25. ix. 2010, coll. Yinghui Mou and Shurong Liu, genitalia slide Nos. ZML 19305 ♂, ZML 19333 ♂, ZML 19332 ♀, ZML 19306 ♀. Hebei Province: 1 ♂, Mt. Jinge (41.01 ° N, 115.88 ° E), Chicheng County, 850 m, 20. vii. 2001, coll. Yanli Du and Shulian Hao, genitalia slide No. SYW 05169; 1 ♂, Mt. Wuling (40.57 ° N, 117.5 ° E), Xinglong County, 800 m, 28. vii. 2010, coll. Aihuan Zhang and Xiaoyu Shi, genitalia slide No. ZML 19334. Shanxi Province: 2 ♂♂, 1 ♀, Mt. Li (35.45 ° N, 112.03 ° E), Jincheng City, 1520 m, 19. viii. 2006, coll. Xu Zhang and Haiyan Bai, genitalia slide Nos. ZH 07244 ♂, ZML 19325 ♀; 3 ♂♂, Xiachuan Village (35.45 ° N, 112.02 ° E), Mt. Li, Qinshui County, 11. viii. 2012, coll. Jun Tie and Chen Li, genitalia slide No. ZML 19331. Liaoning Province: 8 ♂♂, 1 ♀, Lvtuoshui (40.47 ° N, 119.56 ° E), Mt. Bailang, Jianchang County, 658 m, 9. viii. 2016, Coll. Mujie Qi, Juan Li and Yanyan Jia, genitalia slide Nos. ZML 19310 ♂, ZML 19311 ♂, ZML 19312 ♀, ZML 19313 ♂. Henan Province: 2 ♂♂, Baotianman (33.51 ° N, 111.92 ° E), Neixiang County, 1200 m, 11. viii. 2006, coll. Hui Zhen and Denghui Kuang, genitalia slide No. ZH 07417.	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D118604832A2ED52F45E2EFD83.taxon	description	Description. Adult (Figs 2 – 3, 5, 18) wingspan 16.0 – 24.0 mm. Female genitalia (Figs 40 – 43). Apophyses posteriores about 6 × length of subgenital plate. Subgenital plate moderately long, nearly triangular, extending posteriorly beyond base of apophyses anteriores, weakly emarginated distomedially. Apophyses anteriores about 5 / 6 length of subgenital plate, with a pair of wedge-shaped sclerites posteriorly. Ductus bursae with a weakly sclerotized plate accompanied by a triangular sclerite at distal 2 / 3. Corpus bursae oval; signum small, about 1 / 3 width of corpus bursae, rhomboidal, surface with dense denticles.	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D118604832A2ED52F45E2EFD83.taxon	diagnosis	Diagnosis. This species is unique in having the uncus with middle emargination extremely shallow; the gnathos with middle process short, not reaching anterior margin of tegumen, not hook-shaped but sub-triangular or only with an extremely short hook; the sacculus rather long and narrow, about 1 / 4 length of valva (Figs 6, 35 – 35 g). It can be further distinguished from its allies by the triangular subgenital plate in the female genitalia (Figs 40 – 43). In addition, this species with phallus bearing more microthorns (mostly more than thirty) than other species on average.	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D118604832A2ED52F45E2EFD83.taxon	description	Variation. The shapes of uncus in different specimens can be variable within a small range, the details are illustrated in the figures 35 – 35 g.	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D118604832A2ED52F45E2EFD83.taxon	biology_ecology	Host plants. Fagaceae: Quercus sp.; Juglandaceae: Juglans mandshurica Maxim.; Salicaceae: Salix sp. (Liu & Bai 1979).	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D118604832A2ED52F45E2EFD83.taxon	distribution	Distribution. China (Beijing, Hebei, Henan, Heilongjiang, Liaoning, Shanxi, Tianjin), Korea (Park & Karsholt 1999), Japan [?] (Park 2004; Park & Ponomarenko 2007; Park 2012).	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D118604832A2ED52F45E2EFD83.taxon	discussion	Remarks. A few of specimens with phallus with less than twenty microthorns on distal half, which is in contrast with other specimens, we refrain from considering the number of microthorns as a diagnostic character. In addition, Park & Karsholt (1999) pointed out that “ the male genitalia figured by Liu & Pai (1979: Fig. 12) under the name of gibbosella obviously belong to latignathosa ”, we agree with Park & Karsholt (1999), but another figure (Fig. 13) of male genitalia in Liu & Pai (1979) is distinctly belong to P. gibbosella, Liu & Pai (1979) also provided an illegible figure (Fig. 23) which was hand-painted of the female genitalia under the name P. gibbosella, we can’t confirm the female genitalia provided by Liu & Pai (1979) was actually belong to P. gibbosella or P. latignathosa, Park & Karsholt (1999) also indicated that the female of P. latignathosa was “ unknown ”, here we have given detailed description and diagnosis of the female genitalia under the name P. latignathosa. As for the distribution of P. latignathosa, Park (2004), Park & Ponomarenko (2007) and Park (2012) mentioned that Japan is one of distribution localities. But the specimen mentioned in these literatures was the same one, the holotype of P. latignathosa. At present, there is no information about Japanese specimens. Therefore, the distribution of P. latignathosa in Japan is doubtful.	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D118664833A2ED53CD5844FF1B.taxon	description	(Figs 11, 21 – 22, 36 – 36 b)	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D118664833A2ED53CD5844FF1B.taxon	materials_examined	Type material. CHINA: Holotype ♂, Yunnan Province: Nanla River (21.59 ° N, 101.58 ° E), Bubang Town, Mengla County, 652 m, 12. vii. 2013, coll. Shurong Liu, Yuqi Wang and Kaijian Teng, genitalia slide No. ZML 19287. Paratypes: Hainan Province: 2 ♂♂, Yaxing Village (19.02 ° N, 109.40 ° E), Nankai Town, Baisha County, 321 m, 21. vi. 2015, coll. Peixin Cong, Wei Guan and Sha Hu, genitalia slide Nos. ZML 19321, ZML 19322; 1 ♂, Jianfengling (18.75 ° N, 108.87 ° E), Ledong County, 810 m, 14. vi. 2018, coll. Ping Liu, Xia Bai and Shuai Yu, genitalia slide No. ZML 19320.	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D118664833A2ED53CD5844FF1B.taxon	description	Description. Adult (Figs 21 – 22) wingspan 8.0 – 13.0 mm. Head light yellow to light grey, laterally edged with dark yellow scales tipped with yellowish brown. Labial palpus alternately blackish brown and yellowish brown except inner surface of second segment yellowish white; tuft above third segment yellowish brown, scales tipped with white. Antenna blackish brown, serrate and ciliate in male. Thorax and tegula covered with dark yellow scales tipped with blackish brown except basal half of tegula blackish brown. Forewing ground colour blackish brown suffused with yellowish brown and reddish brown scales evenly; basal fascia occupying about 1 / 4 length of wing, black; cell with a yellowish brown tuft at posterior angel; fold with two indistinct yellowish brown tufts sometimes; fringe grey intermixed with black scales tipped with white. Hindwing shining light grey, darker toward distal half of costa, termen and dorsum; fringe light grey. Legs blackish brown on fore and mid legs and outer surface of hindleg except fore coxa yellowish white, yellowish white on inner surface of hindleg; fore and mid legs with tibiae with three rounds of brick-red tufts respectively, scales tipped with white, tarsi with first, second and fifth tarsomeres ringed with brick-red apically; hindleg with all tarsomeres ringed with yellowish white apically. Male genitalia (Figs 11, 36 – 36 b). Uncus nearly rectangle, posterior margin with three denticles of equidistance, sparsely setose; affiliated plates about 3 / 4 length of uncus, sub-triangular but with two sharp horn-shaped posterolateral processes respectively. Gnathos with middle process extremely short, drop-shaped, apex pointed. Valva narrowly band-like, with a sub-triangular process at basal 1 / 4 on inner margin, distal 1 / 4 dilated to rounded and setose apex. Sacculus about 1 / 8 length of valva, nearly quadrate. Juxta triangular. Saccus about half length of valva, long triangle, apex pointed or blunt. Phallus almost straight, basally with a short handle, basal half dilated to elliptical, distal half without microthorns, but with an irregular-shaped sclerite subapically. Female unknown.	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D118664833A2ED53CD5844FF1B.taxon	diagnosis	Diagnosis. This species can be easily recognized by the following unique characters in the male genitalia: the uncus with three denticles on posterior margin, affiliated plates about 3 / 4 length of uncus, the gnathos with middle process shortly drop-shaped and the sacculus quadrate.	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D118664833A2ED53CD5844FF1B.taxon	description	Variation. Specimens from different places vary slightly in morphology. The holotype, which from Yunnan Province is larger than paratypes from Hainan Province in imaginal size (Figs 21 – 22). In addition, variations also occur in the male genitalia: the holotype with the uncus with affiliated plates more sharper on posterior margin, and the saccus with apex blunt (Fig. 36); on the contrary, the paratypes with the uncus with affiliated plates blunt on posterior margin, while the saccus with apex pointed (Figs 36 a – 36 b).	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D118664833A2ED53CD5844FF1B.taxon	distribution	Distribution. China (Hainan, Yunnan).	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D118664833A2ED53CD5844FF1B.taxon	etymology	Etymology. The specific epithet of this species is derived from the Latin minutus, meaning small, and the Latin - gnathus, meaning gnathos, referring to the small gnathos in the male genitalia.	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D118674833A2ED50355F02F85B.taxon	description	(Figs 10, 23 – 26, 37 – 38, 44)	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D118674833A2ED50355F02F85B.taxon	materials_examined	Type material. CHINA: Holotype ♂, Yunnan Province: Mt. Xiaohei (24.52 ° N, 98.84 ° E), Longling County, 1974 m, 27. vii. 2015, coll. Kaijian Teng and Xia Bai, genitalia slide No. ZML 19288. Paratypes: 1 ♂, 2 ♀♀, 27 – 28. vii. 2015, genitalia slide Nos. ZML 19289 ♀, ZML 19326 ♂, ZML 19327 ♀, other data same as holotype; 1 ♂, 1 ♀, Rare Botanical Garden (24.08 ° N, 97.83 ° E), Ruili City, 1000 m, 06. viii. 2005, coll. Yingdang Ren, genitalia slide Nos. SYW 05377 ♀, SYW 05489 ♂; 1 ♂, Bubang Village (21.60 ° N, 101.59 ° E), Xishuangbanna, 656 m, 22. vii. 2014, coll. Kaijian Teng, Wei Guan, Xiuchun Wang and Shurong Liu, genitalia slide No. ZML 19314; 1 ♂, Taiyang River National Forest Park (22.68 ° N, 101.03 ° E), 1450 m, 03. vii. 2015, coll. Kaijian Teng, genitalia slide No. ZML 19299.	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D118674833A2ED50355F02F85B.taxon	description	Description. Adult (Figs 23 – 26) wingspan 12.0 – 15.0 mm. Head vertex grey to dark grey, scales tipped with blackish brown, frons light yellow. Labial palpus with second segment with inner surface yellowish white, outer surface blackish brown but decorated with pale yellowish brown scales tipped with white from basal 1 / 3 to 2 / 3, ventral surface yellowish brown tipped with blackish brown; third segment blackish brown except basal 1 / 3 to 2 / 3 covered with yellowish brown scales tipped with white. Antenna with scape blackish brown; flagellum blackish brown and yellowish brown alternated, serrate and thicker in male but unmodified in female. Thorax and tegula darkgoldenrod, scales tipped with blackish brown, paler or darker in some specimens. Forewing ground colour dark yellow to yellowish brown; costal patch indistinctly triangular, extending from between basal 1 / 4 and 1 / 2 of costal margin to fold, decorated with a few of white scales on inner margin sometimes, costal spot at distal 1 / 3, followed by some white scales; cell suffused with numerous blackish brown scales especially on mesial 1 / 3, posterior angle with a yellowish brown tuft; fold with two yellowish brown tufts below basal 1 / 3 and 2 / 3 respectively; apical angle with scales tipped with blackish brown; fringe dark grey intermixed with yellowish brown scales tipped with blackish brown. Hindwing and fringe dark grey. Fore and mid legs blackish brown scattered with white, tufts on tibiae yellowish brown tipped with white, tarsi with first, second and fifth tarsomeres ringed with yellowish brown apically; hind leg with inner surface yellowish white, outer surface greyish white, scales tipped with blackish brown, all tarsomeres ringed with yellowish white apically. Male genitalia (Figs 10, 37 – 38). Uncus single branch, basal half conic, distal half rodlike, apex narrowly rounded; affiliated plates about 3 / 4 length of uncus, strongly produced to wedge-shaped. Gnathos with middle process entirely hook-shaped, not inflated basally. Valva apically far exceeding tip of uncus, flagelliform, distal 1 / 5 slightly dilated to drop-shape, setose. Sacculus leaf-shaped, apex inwardly curved in beak-shaped. Juxta doublewave shaped. Saccus tapered from base to apex. Phallus nearly straight, basally with a short handle, then narrowed from dilated base to narrowly rounded apex, without microthorns. Female genitalia (Fig. 44). Apophyses posteriores about 7 × length of subgenital plate. Subgenital plate rather narrow, sub-triangular, with a pair of rhomboidal posterolateral lobes of equal length of itself. Apophyses anteriores about 1 / 6 length of subgenital plate. Antrum indistinctly trapezoidal. Ductus bursae broadened anteriorly, with a small sclerotized plate near antrum. Corpus bursae oval; signum greatly long, almost same length of corpus bursae, cross-shaped, hardly with denticles except on outer edge.	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D118674833A2ED50355F02F85B.taxon	diagnosis	Diagnosis. This species is unique from its congeners in appearance and female genitalia. It can be recognized externally by the distinct costal patch, and by the unique subgenital plate which with a pair of posterolateral lobes, and the extremely big cross-shaped signum which about same length of corpus bursae in the female genitalia. In the male genitalia, it is similar to P. kawabei Park & Karsholt, 1999, but can be recognized by the gnathos thicker, the valva with distal 1 / 5 dilated to drop-shaped, the phallus nearly straight; in P. kawabei, the gnathos appears more slender, the valva with apex pointed, the phallus arched (Park & Karsholt 1999: 46, Figs 42 – 43).	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D118674833A2ED50355F02F85B.taxon	distribution	Distribution. China (Yunnan).	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D118674833A2ED50355F02F85B.taxon	etymology	Etymology. The specific name is derived from the Latin proximus, meaning similar, and the specific name of P. kawabei, referring to the similarity between the two species.	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D11865482FA2ED51E95FCFF854.taxon	description	(Figs 4, 8, 12 – 13, 27 – 30, 39 – 39 d, 45 – 47)	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D11865482FA2ED51E95FCFF854.taxon	materials_examined	Material examined. CHINA: Inner Mongolia Autonomous Region: 4 ♂♂, Nantianmen (40.44 ° N, 111.99 ° E), Horinger County, Hohhot City, 1400 m, 13. viii. 2007, coll. Houhun Li and Oleksiy Bidzilya, genitalia slide Nos. ZML 19294, ZML 19295. Liaoning Province: 1 ♀, Mt. Qian (41.03 ° N, 123.12 ° E), 5. vii. 2010, coll. Jiayu Liu and Yanpeng Cai, genitalia slide No. ZML 19293; 1 ♂ Taihe Palace (41.00 ° N, 123.10 ° E), Mt. Qian, Anshan City, 340 m, 5. viii. 2016, coll. Mujie Qi, Juan Li and Yanyan Jia. Heilongjiang Province: 4 ♂♂, 1 ♀, Tafeng Management Station (52.35 ° N, 124.83 ° E), Great Khingan, 474 m, 29. vii. 2016, coll. Mujie Qi, Juan Li and Yanyan Jia, genitalia slide Nos. ZML 19339 ♂, ZML 19298 ♀, ZML 19328 ♂, ZML 19341 ♂, ZML 19342 ♂; 1 ♂, Tahe Forest Centre (52.37 ° N, 124.67 ° E), Great Khingan, 429 m, 30. vii. 2016, coll. Mujie Qi, Juan Li and Yanyan Jia, genitalia slide No. ZML 19297. Hubei Province: 1 ♀, Shuangping (31.57 ° N, 109.87 ° E), Zhuxi County, 1201 m, 6. vii. 2017, coll. Wanding Qi et al., genitalia slide No. ZML 17545. Sichuan Province: 1 ♂, Shuzheng (33.2 ° N, 103.9 ° E), Jiuzhaigou County, Aba Tibetan and Qiang Autonomous Prefecture, 2300 m, 17. viii. 2002, coll. Shulian Hao, genitalia slide No. ZML 19290. Gansu Province: 2 ♂♂, Mt. Xinglong (35.8 ° N, 104.07 ° E), Yuzhong County, 2178 m, 20. viii. 2007, coll. Feng Yang and Hanguang Gao, genitalia slide No. ZML 19319. Qinghai Province: 1 ♂, Xining Botanical Garden (36.63 ° N, 101.75 ° E), Xining City, 2280 m, 20. vii. 1995, coll. Houhun Li and Shuxia Wang, genitalia slide No. L 95281.	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D11865482FA2ED51E95FCFF854.taxon	description	Redescription. Adult (Figs 4, 27 – 30) wingspan 17.0 – 21.0 mm. Head grey to pale yellowish brown, scales tipped with black. Labial palpus almost entirely blackish brown except inner surface of second segment yellowish white, scales tipped with white. Antenna with scape blackish brown; flagellum blackish brown and yellowish brown alternated, serrate. Forewing dark grey, scales densely tipped with blackish brown or yellowish brown; costal spots at middle and distal 1 / 3 respectively; three blackish brown tufts at posterior angle of cell, basal 1 / 3 and 2 / 3 of fold respectively; fringe grey, intermixed with scales tipped with blackish brown or white. Hindwing and fringe grey. Legs blackish brown except inner surface of hindleg yellowish white; fore and mid legs with tibiae with tufts brown, scales tipped with white, tarsi ringed with brown on first, second and fifth tarsomeres apically; hindleg with all tarsomeres ringed with yellowish white apically. Male genitalia (Figs 8, 12 – 13, 39 – 39 d). Uncus shovel-shaped, posterior margin with three incisions, middle incision about double length of lateral incisions, middle branches longer than lateral branches; affiliated plates small, slightly produced to semicircular. Gnathos with middle process hook-shaped, basal half inflated to heart-shaped. Valva-vinculum complex and phallus showing typical appearance of this genus but with vinculum densely with microthorns posterolaterally. Female genitalia (Figs 45 – 47). Apophyses posteriores about 6.5 – 7 × length of subgenital plate. Subgenital plate sub-semicircular or sub-triangular. Apophyses anteriores extending to beyond end of subgenital plate and about 2 × length of it, posteriorly with a pair of irregular-shaped sclerites bearing several wrinkles. Ductus bursae and corpus bursae densely covered with micro-granules and showing rough appearances, ductus bursae with a weakly sclerotized plate accompanied by a triangular sclerite. Corpus bursae oval; signum rhomboidal.	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D11865482FA2ED51E95FCFF854.taxon	diagnosis	Diagnosis. This species is similar to P. gibbosella both in adult and male genitalia, and the differences between them have been stated in the diagnosis of the latter species.	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D11865482FA2ED51E95FCFF854.taxon	description	Variation. The length of middle incision of uncus, the shape of gnathos and the size of signum can be variable, details are illustrated in the Figures 39 – 39 d and 45 – 47.	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D11865482FA2ED51E95FCFF854.taxon	biology_ecology	Host plants. Salicaceae: Larva on Salix sp., Salix caprea (Park & Karsholt 1999).	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D11865482FA2ED51E95FCFF854.taxon	distribution	Distribution. China (Gansu, Heilongjiang, Hubei, Inner Mongolia, Liaoning, Qinghai, Sichuan), Japan, Russia, Europe (Park & Karsholt 1999).	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
03A187D11865482FA2ED51E95FCFF854.taxon	discussion	Remarks. This species is recorded from China for the first time.	en	Zheng, Meiling, Li, Houhun (2021): Taxonomic revision of the genus Psoricoptera Stainton, 1854 (Lepidoptera Gelechiidae: Gelechiinae) from China, with descriptions of three new species. Zootaxa 4975 (2): 253-272, DOI: https://doi.org/10.11646/zootaxa.4975.2.2
