taxonID	type	description	language	source
8A39CA255557517559ABAB98829EC306.taxon	description	“ Phryganea ferruginea. Length of 6 1 / 3 lines [supposedly French lines, that gives approximately 14.287 mm of length. M. A. Alonso Zarazaga, personal communication].	en	Valladolid, María, Arauzo, Mercedes, Chertoprud, Mikhail V., Chvojka, Pavel, Czachorowski, Stanisław, Dorda, Beatriz A., Hinić, Jelena, Ibrahimi, Halil, Karaouzas, Ioannis, Krpač, Vladimir, Kučinić, Mladen, Lodovici, Omar, Salokannel, Juha, Stamenković, Valentina Slavevska, Stojanović, Katarina (2021): The Rhyacophila fasciata Group in Europe: Rhyacophila fasciata Hagen 1859 and formerly synonymized species (Trichoptera: Rhyacophilidae), with new description of Rhyacophila fasciata and Rhyacophila septentrionis McLachlan 1865 (stat. prom.). Zootaxa 4975 (1): 1-57, DOI: https://doi.org/10.11646/zootaxa.4975.1.1
8A39CA255557517559ABAB98829EC306.taxon	diagnosis	Diagnosis. All rust colored; forewings reticulated dark; black eyes. In forests by the waters; month of May. Abdomen with lateral black dots, as many as segments of it. Antennae 6 lines long, rusty. The other sex with a round white common spot, and white spots (5 – 6) on the outer margin of the forewings. ” Hagen (1854), in his study of the species of Phryganea included in Scopoli’s Entomologia Carniolica stated that the description of P. ferruginea does not match the drawing, even with other Phryganea. Following the “ reticulate wings ” and “ white spots ” of wings, he proposed three options that could be one of the following: Chaetopteryx villosa (Fabricius, 1798) (based on body contour and reticulation of wings), one of two Hydropsyche Pictet 1834, and one Rhyacophila (reticulation of wings); the color (all rust) only matched with the mentioned Rhyacophila (maybe R. nebulosa Stephen), and, as he had only the female, he couldn’t check if males also had the white spots on the wings. In 1859 Hagen cited this species and R. fasciata, together with R. vulgaris, R. dorsalis and R. paupera: “ 4. R. ferruginea Scop. Anal appendage rounded, almost completely covering the lateral appendages. The smallest species. Carniola, Austria. Maybe dark colored females of my collection from Zurich belong to it. ” “ 5. R. fasciata Hag. The appendages are very similar to the previous species, but the size of the animal is much more significant and the upper wings have three dark cross-ties. A single male from Elberfeld ”. McLachlan (1865), at the end of his description of Rhyacophila septentrionis wrote: “ I have some doubt in announcing this as a new species, since the form of the appendices agrees very well with the figure of those of R. ferruginea, sent to me by Dr. Hagen. I submitted an example to that gentleman, and he says that it is larger and darker than his continental types of that species and may be distinct. R. ferruginea is described as ‘ die kleinste Art’ of the group (the true genus Rhyacophila) to which it belongs ”. In his publication of 1868, McLachlan said: “ R. ferruginea, (Scopoli) Hagen. I have not seen this insect, which, according to Hagen (in litt.) is smaller than septentrionis, but with similar lobes and appendices. Scopoli’s name cannot be said to apply to this species with certainty. ” Finally, in his publication of 1879, McLachlan concluded at the end of the paragraph about Rhyacophila septentrionis: “ I do not feel justified in adopting Scopoli’s name ferruginea (as was done by Hagen for the example from Carniola). Even supposing that Scopoli had a Rhyacophila before him (which is just possible), the particular species must remain uncertain. ” We agree with this statement, due to the difficulty of identification from the original species described, so we propose to change the status of Rhyacophila ferruginea (Scopoli 1763) to nomen dubium.	en	Valladolid, María, Arauzo, Mercedes, Chertoprud, Mikhail V., Chvojka, Pavel, Czachorowski, Stanisław, Dorda, Beatriz A., Hinić, Jelena, Ibrahimi, Halil, Karaouzas, Ioannis, Krpač, Vladimir, Kučinić, Mladen, Lodovici, Omar, Salokannel, Juha, Stamenković, Valentina Slavevska, Stojanović, Katarina (2021): The Rhyacophila fasciata Group in Europe: Rhyacophila fasciata Hagen 1859 and formerly synonymized species (Trichoptera: Rhyacophilidae), with new description of Rhyacophila fasciata and Rhyacophila septentrionis McLachlan 1865 (stat. prom.). Zootaxa 4975 (1): 1-57, DOI: https://doi.org/10.11646/zootaxa.4975.1.1
8A39CA255556517659ABABB487D3C04E.taxon	materials_examined	Type locality: Elberfeld (Germany). Other information: (in label) 57; E. - illegible; Hagen; 107; ML; type 11067; (vial); Lectotype Rhyacophila fasciata Hagen. (MCZBase, 2020). (Figs 2 a, 2 b, 30 a, 30 b) Ross (1952) included the species in his publication about the lectotypes of Hagen in the collection of the Museum of Comparative Zoology at Harvard University, Cambridge, Massachusetts. Description. This species has been described by several authors over many years. The first publication where this species is cited is that of Hagen (1859), where he provided a very short description: “ A. Lower appendages undivided at the top. 1. The tongue-like extention of the last abdominal segment with an enlarged end. 5. R. fasciata Hag. The app. are very similar to the previous species (R. ferruginea Scop.), but the size of the animal is much more significant, and the upper wings have three dark cross bands. A single male from Elberfeld ” (Fig 30 a). ” McLachlan (1868) described it as follows: “ R. fasciata Hagen, may be distinguished almost with certainty by the dark fasciae on the anterior wings. The form of the dorsal lobe is very distinct from the preceding (R. paupera), being broad and rounded, and nearly concealing the app. sup. The second joint of the app. inf. is obliquely truncate (Pl. XIV. fig. 16). I have three ♂ from Carinthia from Zeller. ” In 1879, McLachan described R. fasciata and R. septentrionis. In the description of R. fasciata he included the original specimen from Elberfeld, his specimens from Carinthia, saying that they should be referred to septentrionis, and 7 ♂ and 1 ♀ from the Pyrenees (France), writing “ it may be that these should form a distinct species ”. These specimens from the Pyrenees were described in 1916 by Navás as Rhyacophila sociata (Valladolid et al. 2018). On the other hand, in the description of R. septentrionis McLachlan included his specimens from Scotland, as well as others from different countries of North and Central Europe. Based on results of our studies of morphology and genetics, we can say that most of the specimens cited belong to R. fasciata, keeping only those from Scotland as R. septentrionis. Klapálek (1893) described the larva and the pupa of R. septentrionis in Bohem, but the area matches with those of R. fasciata, so it should be this species. The same situation pertains to Silfvenius (1905), who described larvae and pupae from South Finland as R. septentrionis that should be R. fasciata. Ulmer (1909), based on previous authors, described the imagos and larvae of R. fasciata and R. septentrionis, first one from Elberfeld and the second one from different places in Germany. We consider that these specimens all belong to R. fasciata. There are more recent authors that have described R. fasciata and R. septentrionis specimens: Nielsen (1942) as R. septentrionis from Denmark; Nóvak (1963) as R. fasciata from Czechoslovakia; Hickin (1954) and Mackereth (1954) as R. septentrionis from United Kingdom; Fotius-Jaboulet (1964) as R. septentrionis from France; Lepneva (1964) as R. septentrionis from northwest, west and southwest Russia; Eidel (1974) as R. fasciata from Austria; Buholzer (1978) as R. fasciata from Switzerland; Sedlak (1985) as R. fasciata from Central Europe; Pitsch (1993) as R. fasciata from Central Europe. Of all these descriptions, only those from the United Kingdom could belong to R. septentrionis; the rest, from continental Europe, should be R. fasciata or some related species. Finally, Malicky & Sipahiler (1993) illustrated the differences among male specimens of R. fasciata from Europe, Turkey, and Lebanon that they considered as subspecies of the R. fasciata Group. Some of them have been promoted to species (R. denticulata McLachlan 1879, by Valladolid et al. 2018; R. kykladica Malicky & Sipahiler 1993, by Valladolid et al. 2019), one species has been resurrected (R. sociata Navás 1916, previously a synonym of R. denticulata, by Valladolid et al. 2018), and others could be considered also valid species (Valladolid, unpublished data).	en	Valladolid, María, Arauzo, Mercedes, Chertoprud, Mikhail V., Chvojka, Pavel, Czachorowski, Stanisław, Dorda, Beatriz A., Hinić, Jelena, Ibrahimi, Halil, Karaouzas, Ioannis, Krpač, Vladimir, Kučinić, Mladen, Lodovici, Omar, Salokannel, Juha, Stamenković, Valentina Slavevska, Stojanović, Katarina (2021): The Rhyacophila fasciata Group in Europe: Rhyacophila fasciata Hagen 1859 and formerly synonymized species (Trichoptera: Rhyacophilidae), with new description of Rhyacophila fasciata and Rhyacophila septentrionis McLachlan 1865 (stat. prom.). Zootaxa 4975 (1): 1-57, DOI: https://doi.org/10.11646/zootaxa.4975.1.1
8A39CA255555516859ABA87F87E4C63A.taxon	materials_examined	Type locality: Near Longniddry & Aberlady, Scotland. Other information: Captured 18 / vi / 1865, at a little stream running into the sea on the coast of the Firth of Forth, near Longniddry. Description. According to McLachlan (1865), “ Antennae testaceous, with indistinct paler annulations. Head, palpi and mesothorax pale brown; the former with scattered golden hairs. Anterior wings broad, the apex nearly elliptical, golden-grey, thickly reticulated with dark grey; dark greyish-fuscous spots round the apical margin, one at the termination of each apical sector; the pale dorsal blotch is ill defined, and is indicated by an irregular pale space, scarcely reticulated; a similar but still more indistinct paler space occupies the whole of the lower two-thirds of the apical portion of the wing; neuration fuscous. Posterior wings pale greyish, subhyaline; apical veins darker grey, sometimes margined with grey at their points of insertion in the apical margin. Legs testaceous; the tips of the anterior and intermediate tibiae externally fuscous; tibial spurs dark brown. Abdomen fuscous, ochreous beneath, testaceous at the apex above, and with testaceous appendices. In the male the superior lobe is short, rather attenuated at the base, but soon greately dilated, the apical portion being nearly orbicular (McLachlan, 1879, pl. XLVI, Rh. septentrionis, fig 1) (herein Fig 31 b); app. sup. nearly concealed under the lobe, the external edge rounded, slightly excised in front (herein Fig 31 b); app. inf. rather short, the basal joint broad, with the sides parallel, the apical joint scarcely shorter than the basal and of equal breadth, the apex very obliquely truncated (McLachlan, 1879, pl. XLVI, Rh. septentrionis fig 2); penis furnished above with a short broad dark chesnut-brown superior cover; lower sheaths bent slightly downwards (their direction varies in dead specimens) (Fig 31 c), dark shining chestnut-brown, clear testaceous at the extreme base (McLachlan, 1879, pl. XLVI, Rh. septentrionis fig 3). Expanse of fore-wings 12 – 13 lines ”.	en	Valladolid, María, Arauzo, Mercedes, Chertoprud, Mikhail V., Chvojka, Pavel, Czachorowski, Stanisław, Dorda, Beatriz A., Hinić, Jelena, Ibrahimi, Halil, Karaouzas, Ioannis, Krpač, Vladimir, Kučinić, Mladen, Lodovici, Omar, Salokannel, Juha, Stamenković, Valentina Slavevska, Stojanović, Katarina (2021): The Rhyacophila fasciata Group in Europe: Rhyacophila fasciata Hagen 1859 and formerly synonymized species (Trichoptera: Rhyacophilidae), with new description of Rhyacophila fasciata and Rhyacophila septentrionis McLachlan 1865 (stat. prom.). Zootaxa 4975 (1): 1-57, DOI: https://doi.org/10.11646/zootaxa.4975.1.1
8A39CA25554B516E59ABAEE08525C44A.taxon	description	Description of the larva (Figs 4 – 9, 21). The description of the specimens examined will be complemented with descriptions of previous authors, adding or confirming features. Biometrics of last instar larva and prepupa: Length: 14.25 – 19.00 mm ( = 16.09, n = 13). Maximum width: 2.98 – 3.90 mm ( = 3.28, n = 13) (Lepneva 1964, 5 th instar larval length: 23.00 – 23.40 mm; Nielsen 1942, 5 th instar larval length: 12.00 – 23.40 mm; Ulmer 1909, larval length: 22.00 – 24.00 mm, larval width: 3.00 – 3.50 mm). Maximum width of larva at metathorax. Colour: Live specimen greenish, dorsally with dark areas and light stripes, pale ventrally. Specimens preserved in ethanol reddish brown, dorsum dark, ventrally pale; on abdomen, semicircular pale anterior dorsolateral areas, dorsocentral area of segments with irregular pale spots, connected with anterior dorsolateral areas. Sclerotized areas of head and thorax brown with pale areas and stripes, black in some places (Fig 21 a). Head (Figs 4, 21 b). Length: 1.49 – 2.20 mm ( = 1.90, n = 13). Width: 1.14 – 1.50 mm ( = 1.30, n = 13). Maximum width in posterior third of head, lateral margins almost parallel. Cephalic capsule (Figs 4 a, 4 b), dorsally and dorsolaterally light brown in anterior third, areas adjacent to frontoclypeal and anterior coronal sutures and anterior dorsolateral areas paler; ventral side variable (Silfvenius 1905), pale (Pitsch 1993), or with dark spots in middle central area (Sedlák 1985, p. 111, table 3, fig 10); anterior edge and submentum reddish brown, posterior edge black (Nielsen 1942); oval brown area with dark spots extending laterally from anterior edge of occipital foramen to anterior third of head (Fig 4 b) (Klapálek 1893; Lepneva 1964, p. 223, figs 181, 182 B). Frontoclypeal apotome (Fig 4 c) pale; 2 oval dark brown muscle attachment spots in central anterior third, between setae # 4; dark pigmentation in posterior third heart-shaped, including pits # 4 but not setae # 6, with 4 – 6 oval brown muscle attachment spots with dark borders (Klapálek 1893; Silfvenius 1905; Nielsen 1942; Lepneva 1964; Pitsch 1993). Two-thirds of posterior dorsal area of head light brown, marked with darker muscle attachment spots; centrally one pair of dark brown patches, wing-shaped, outside posterior edge of frontal suture; light circular area on juncture of frontal and coronal sutures; three pale spots around setae # 15, # 16, and # 19 – 21; dorsal head pits with dark border, central posterior occipital foramen black. Mandibles (Fig 5) asymmetrical, as in other rhyacophilids; left one bigger than right one. Inner blade of left mandible irregular, convex, right mandible with small tooth in middle (Nielsen 1942, pl. 2, fig 13; Lepneva 1964, p. 223, fig 184). Labrum and maxillolabium as in other Rhyacophila species (Nielsen 1942, pl. 2, figs 12 A, 12 B, 14 A, 14 B; Lepneva 1964, p. 223, figs 183 A, 183 B, 185). Mentum rectangular, its anterior half slightly sclerotized. Labrum with small transparent area in central anterior edge. Thorax (Figs 6, 21 c): Anterior half of pronotum pale or very light coloured. Sinuous posterior margin bordered by dark band, brown in middle of each half, black posteromesally and posterolaterally, light brown submesally; posterior half of pronotum (Fig 6 a) with three dark areas separated by light zones, central dark region tapered posteriorly and reaching posterior edge of sclerite, anteriorly truncate, wider that posteriorly, two lines of dark brown spots inside central dark region and reaching posterior edge, several dark brown spots along posterior edge; each pronotal plate with lateral dark area almost triangular, from anterior submesal region to posterolateral edge of sclerite, posterior area with two – three rows of brown spots with dark borders distributed in circular-oval shape and parallel to posterior border. Four – six brown spots around setae # 5; curved row of dark dots from setae # 5 toward anterior submesal corner, curving around circular spot on anterior half (Fig 6 a); pale posterior and median areas connected (Fig 6 b); with narrow black band in anterolateral angles, extending posterad laterally to posterolateral margins, inner to margins (Fig 6 b; Lepneva 1964); anterior margin light brown, with black spot near anterolateal angle; central area of posterior marginal band of each sclerite dark brown or black (Fig 6 b) (Nielsen 1942, pl. 3, fig 17 A; Lepneva 1964, p. 223, fig 188). Legs (Fig 9): Similar to those other Rhyacophila species. Colour yellowish, proximal regions of coxa, femur and distal region of trochanter of fore- and midlegs black or dark brown. Narrow pale reddish-brown band in proximal dorsolateral area of tibia and distal dorsolateral area of femur of each pair of legs, each inner and outer face of each femur with band of circular spots. Forelegs each ventrally with spicules, extending from distal half of trochanter, all of femur, and distal two-thirds of tibia and tarsus. Setae black or dark brown, some transparent in ventral area. Abdomen (Figs 7, 8, 21 d, 21 e): Similar to other Rhyacophila species. Colour as for meso- and metathoracic segments, greenish with dark dorsal areas and light stripes in living larvae, in preserved specimens semicircular pale anterior dorsolateral areas, dorsocentral area of segments with irregular pale spots, connected with anterior dorsolateral areas, pale yellow ventrally. Widths of anterior abdominal segments similar, narrower in posterior segments. Lateral abdominal gills in tufts of 25 – 30 filaments (20 – 30 filaments, Lepneva 1964). Dorsal sclerite of segment IX (Figs 7, 21 d) brown, with continuous black band on anterior edge and two small and widely separated black bands on posterior edge, under setae # 2 (Lepneva 1964); band of anterior edge narrower in center, with pair of narrow extensions backward beyond and surrounding pit # 1; with some brown spots submesally near posterior margin; in darker larvae, extensions of anterior edge and spots of posterocentral area connected by dark stripes, creating X-shape figure; three pale areas along posterior margin, on either side of two black bands and central area. Anal prolegs typical of group, each with long sword process (Figs 8 a, 8 b, 21 e). Anal claws each with three short teeth on ventral edge (Figs 8 a, 8 c, 21 e), basoventral hook black basally and reddish brown apically (Figs 8 a, 8 b). Description of the pupa Biometrics of pupa and cocoon. Pupal length (total): 9.70 – 15.70 mm ( = 12.20, n = 28); male pupa: 10.07 – 12.28 mm ( = 11.06, n = 11) (length: 11.20 – 12.20, Lepneva 1964; Nielsen 1942); female pupa: 9.70 – 15.70 mm ( = 12.94, n = 17) (length: 12.40 – 15.50, Lepneva 1964; Nielsen 1942; up to 16.00 mm, Silfvenius 1905). Cocoon length (total): 12.37 – 18.35 mm ( = 15.11, n = 68) (length: 12.50 – 18.00 mm, Nielsen 1942); male cocoon length: 12.39 – 16.27 mm ( = 14.43, n = 10); female cocoon length: 13.43 – 17.73 mm ( = 15.99, n = 16). Head: Distribution of dorsal setae: four before, two between and two in middle of antennal pads, three on posterior edge of each cephalic tubercle. Mandibles brown, left one with two large teeth and right one with three, with numerous fine teeth on inner blade (Klapálek 1893; Nielsen 1942; Ulmer 1909). Antennae of variable length, reaching from middle of abdominal segment IV to posterior edge of segment VI (to anterior edge of segment IV in females and posterior edge of segment VI in males, Klapálek 1893; to abdominal segment VIII in males and V in females, Lepneva 1964; Nielsen 1942). Thorax: Tubercles of prothorax each with 5 setae, directed backwards, two central setae directed forwards. Hind wing pads variously reaching from anterior edge of abdominal segment III to posterior edge of segment IV (forewing pads reaching anterior edge of abdominal segment IV, Klapálek 1893; forewing pads reaching abdominal segment V in males and IV in females, Lepneva 1964; hind wing pads reaching end of segment IV in females, Silfvenius 1905). Abdomen: Paired anterior hook plates pedunculated, straight basally and dilated posteriorly, palmate, and present on abdominal terga IV to VII (Fig 10) (Lepneva 1964; Ulmer 1909), slightly bigger on segments V and VI, each with 15 – 20 spines; also, pair of inconspicuous small flat plates without spines on tergum III. Paired posterior hook plates sessile on terga III – V, slightly oblong tranversally (length: width = 1: 1.3) (Klapálek 1893: paired anterior hook plates present on abdominal terga III to VIII, paired posterior hook plates on terga III to V; Silfvenius 1905: paired anterior hook plates present on abdominal terga IV to VII, sometimes without plates on segment III, paired posterior hook plates present on terga III to V), each with variable number of spines, more than 25. Description of the imago Male body length: 7.04 – 10.46 mm ( = 9.23, n = 36) each forewing: 10.16 – 13.54 ( = 11.67, n = 36), each hind wing: 8.52 – 11.99 mm ( = 10.25, n = 36); females, length: 8.73 – 12.19 mm ( = 10.46, n = 19), each forewing: 10.05 – 13.90 mm ( = 12.14, n = 19), each hind wing: 8.24 – 12.03 mm ( = 10.68, n = 19) [Ulmer 1909: male length (R. fasciata) = 10.00 mm, wingspan = 27.00 mm; male length (R. septentrionis) = 6.50 – 9.00 mm, wingspan = 22.50 – 29.00 mm; Malicky 2004: male forewing length = 10.00 – 14.00 mm, female forewing length = 11.00 – 14.00 mm]. Colour: Specimens preserved in ethanol reddish brown dorsally, yellowish ventrally, spurs reddish, setae golden brown, females generally darker than males; small black or dark brown spots in dorsal area; in lateral view, some specimens with large black spots marking border between dorsal and ventral areas. Forewings dark with small pale spots, three transversal dark bands (fascias) in dry specimens (Fig 30 a), pale in preserved specimens, darker in females, hind wings pale: each with dark area on leading edge between subcostal (Sc) and second radial (R 2) veins in fore- and hind wings; each forewing hyaline in irregular zone connecting medial (M) and medio-cubital (m-cu) veins and in distal area of second cubital vein (Cu 2) (Holzenthal et al. 2007); in each hind wing proximal segment of medial veins and connection area (M 1 - 4) not conspicuous, second cubital (Cu 2) and first anal (A 1) veins not conspicuous either totally or partially, proximal portion of cubital vein (Cu) and distal portions of first cubital vein (Cu 1 a- 1 b) thicker. Male genitalia (Figs 2 b, 12, 23, 30 b): Apical segment of each inferior appendage (Figs 2 b, 12 A, 23 a) with basal and distal edges diverging, dorsal and ventral edges diverging, posterior edge of 2 nd segment of each inferior appendage slightly concave dorsally and slightly convex ventrally, ventral edge slightly concave (Figs 12 A, 23 a). Apicodorsal vertex slightly angular, apicoventral angle rounded, projecting as thick lobe narrowing progressively to apex. Parameres in ventral view curved posteromesad in apical half (Figs 12 BV p, 23 c). In lateral view (Fig 12 BL) each constricted at short distance beyond base, dilated at middle, with almost parallel dorsal and ventral margins in central area (S-shape), two files of long and thick spines on midventral margin, pointing outward; midlateral surface covered by sparse thin spicules or setae, from middle posteroventral to posterodorsal edge and covering most of apex, absents in middle anteroventral and dorsal edges. Aedeagus (phallicata) in lateral view (Fig 12 CL) with dorsal margin concave and posterior corner of concavity hooked anterad, posterior edge of aedeagus straight, projected posteroventrad, subapically rounded ventrally (Fig 12 CL); ventral lobe of aedeagus triangular (Fig 12 BV vl), lateral edges slightly rounded; lateroventral lobes of phallus straight, posterior edges strongly convex, apicolateral margins rounded (Fig 12 CD lvl). Apicodorsal lobe of segment IX slightly dilated subapicolaterally, almost round in shape (Figs 12 DD al, 23 b), with small apicomesal excision in some specimens; preanal appendages (Fig 12 DD pa) round, shorter than apicodorsal lobe, almost covered by it; in ventral view, apical band V-shaped (Fig 12 DV ab) nearly as wide as long, non-sclerotized inside area with posteromesal excision, posterior edges rounded (Fig 12 DV va), anal sclerites triangular (Fig 12 DV as). Female genitalia (Figs 13, 25): In lateral view (Figs 13 La, 13 Lb, 25 L), posterodorsal margin of segment VIII valves on each side irregular posteriorly, convex on ventral edge, with dorsolateral projection, in some specimens slightly indented posteriorly, posterolateral margin irregular, with 1 – 2 posterior projections, ventral projection more conspicuous (Fig 13 Lb 1 – 3). In dorsal view (Figs 13 Da, 13 Db, 25 D), with indentation between segment VIII valves, with rounded apical and subapical corners, delimiting oval space, small projection in apical corner in some specimens (Fig 13 Db). In ventral view (Fig 13 V, 25 V) segment VIII valves forming two elongate sclerites, basal 1 / 4 of mesal edges parallel, distal 3 / 4 of mesal edges irregular, slightly concave, diverging; intersegmental membrane with pair of dark, ovoid or slightly elongate sclerites.	en	Valladolid, María, Arauzo, Mercedes, Chertoprud, Mikhail V., Chvojka, Pavel, Czachorowski, Stanisław, Dorda, Beatriz A., Hinić, Jelena, Ibrahimi, Halil, Karaouzas, Ioannis, Krpač, Vladimir, Kučinić, Mladen, Lodovici, Omar, Salokannel, Juha, Stamenković, Valentina Slavevska, Stojanović, Katarina (2021): The Rhyacophila fasciata Group in Europe: Rhyacophila fasciata Hagen 1859 and formerly synonymized species (Trichoptera: Rhyacophilidae), with new description of Rhyacophila fasciata and Rhyacophila septentrionis McLachlan 1865 (stat. prom.). Zootaxa 4975 (1): 1-57, DOI: https://doi.org/10.11646/zootaxa.4975.1.1
8A39CA25554D515859ABAC78820BC226.taxon	description	Description of the larva Colour: Live specimen golden-olive (Hickin 1954), pale ventrally. Specimens preserved in ethanol reddish brown, dorsum dark, ventrally pale; on abdomen, semicircular-triangular pale anterior dorsolateral areas, dorsocentral area of segments with darker circular pale spots, connected caudally with anterior dorsolateral areas through pale lines, forming W-shape. Sclerotized areas of head and thorax dark brown with pale areas, black in some places (Fig 22 a). Head (Figs 14, 22 b). Maximum width in posterior third of head, lateral margins almost parallel (tapering slightly towards anterior end, but not constricted at level of eyes, Mackereth 1954, fig 1 S; Hickin 1967, fig 40; tapering anteriorly, Edington & Hildrew 1981). Cephalic capsule (Figs 14 a, 14 b), dorsally and dorsolaterally dark brown, anterior third and areas adjacent to frontoclypeal and anterior coronal sutures and anterior dorsolateral areas paler (dorsal posterior region of head with conspicuous pattern of dark spots on lighter background, Edington & Hildrew 1981, Fig 33, arrow a); ventral side pale, with central and slightly darker U-shape area from middle to posterior end of protogula covered with pale spots (Fig 14 c), aboral margins of genae just before union with small black fold (Hickin 1954, fig 4 B, 1967, fig 54); in lateral view anterior edge and submentum reddish brown, posterior edge black; oval brown area with dark spots extending laterally from anterior edge of occipital foramen to anterior third of head (Fig 14 b). Frontoclypeal apotome (Fig 14 d) pale; 2 oval dark brown submesal muscle attachment spots in anterior third between setae # 4; dark brown pigmentation in posterior third heart-shaped (crescent shaped, Mackereth 1954, fig 1 S; Hickin 1967, fig 40), including pits # 4 but not setae # 6, with pair of oval lateral and 2 – 3 circular mesal paler brown muscle attachment spots with dark borders, anterocentral area pale with two oval pale brown spots in anteromesal position (Figs 14 a, 22 a, 22 b). Two-thirds of posterior dorsal area of head brown, marked with darker and paler muscle attachment spots; submesally one pair of dark brown triangular patches with paler spots with dark borders outside posterior edges of frontal sutures; light triangular area on juncture of frontal and coronal sutures, small pale area around pit # 9; three pale spots around setae # 15, # 16, and # 19 – 21, last one connected with posterior pale areas; dark brown dots in posterior area of genae (Mackereth 1954, fig 1 S; Hickin 1967, fig 40); dorsal head pits with dark brown borders, posteromesal edge of occipital foramen black. Mandibles (Fig 15) asymmetrical, as in other rhyacophilids; left one bigger than right one. Subapicomesal blade of left mandible slightly concave, right mandible with small tooth in middle, both with darker apices (Figs 15, 22 a). Labrum and maxillolabium as in other Rhyacophila species. Mentum rectangular, its anterior half slightly sclerotized. Labrum with small transparent area in central anterior edge. Thorax (Figs 16, 22 c): Anterior half of pronotum pale or very light coloured, posterior half with three darker areas, one median and two lateral (Edington & Hildrew 1981, fig 35; Hickin 1954, fig 5 B, 1967 fig 56), in darker specimens posterior half brown, with mesal area darker. Sinuous posterior margin bordered by black band, posteriorly and posterolaterally light brown submesally; posterior half of pronotum (Fig 16 a) dark, with narrow semicircular area extending anterolaterally and small area laterally, lateral area pale in each pronotal plate; central dark region triangular, tapered posteriorly and reaching posterior edge of sclerite, anteriorly truncate and wider than posteriorly, pale brown spots inside triangular region reaching posterior edge; each side of pronotal plate with two or three lines of pale brown spots with dark border of different shapes and parallel to posterior border; pale areas around setae 2 – 3 #, 5 # and 6 #, four – six brown spots around setae # 5; curved row of two or three dark dots extending from setae # 5 toward anterolateral corner, two pale brown spots with dark borders anterolateral and posterolateral of seta 6 #, circular and elongate respectively (Figs 16 b, 22 c); with narrow black band in anterolateral angles, extending posterad laterally above lateral margins to posterolateral corners (Fig 16 b); anterior margin light brown, with a line of black spots near anterior angle; mesal area of posterior marginal band black (Fig 16 b, 22 c). Legs: Similar to those of other Rhyacophila species. Colour pale yellowish, each with tarsus and claw darker, proximal parts of coxa, femur and distal parts of trochanters of fore- and midlegs black or dark brown. Narrow pale dark brown band in proximal dorsolateral area of tibia and distal dorsolateral area of femur of each pair of legs, each inner and outer face of each femur with band of dark brown circular spots. Forelegs each ventrally with spicules, extending from distal half of trochanter, all of femur, tibia and tarsus. Setae black or dark brown, some transparent in ventral area. Abdomen (Figs 17, 18, 22 d, 22 e): Similar to other Rhyacophila species. Colour as for meso- and metathoracic segments, in preserved larvae semicircular-triangular pale anterior dorsolateral areas, dorsocentral area of each segment with darker circular pale spots, connected caudally with anterior dorsolateral areas through pale lines, forming W-shape, pale yellow ventrally. Widths of anterior abdominal segments similar, narrower in posterior segments. Lateral abdominal gills in tufts of 25 – 30 filaments. Dorsal sclerite of segment IX (Figs 17, 22 d) light coloured, darker in central area, with continuous black band on anterior edge and two discontinuous black bands on posterior edge, posterior to setae # 2; band of anterior edge narrower in center, surrounding anteriorly pit # 1; pale spots inside central area, two pale areas laterally. Anal prolegs typical of group, each with long sword process (Figs 18 a, 18 b, 22 e). Anal claws each with three short teeth on ventral edge (Figs 18 a, 18 c, 22 e), basoventral hook black basally and reddish brown apically (Figs 18 a, 18 b). Description of the pupa Biometrics of pupa and cocoon. Male pupa: 10.00 mm (1 specimen); female pupa: 11.60 mm (1 specimen). Cocoon length: 14.40 mm (1 specimen). Head: Distribution of setae on dorsal head similar to R. fasciata. Mandibles dark brown, left one with two large teeth and right one with three, anterior tooth bigger, with numerous fine teeth on inner blade. Antennae of variable length, reaching posterior edge of segment VII and posterior edge of segment IV in male and female studied, respectively. Thorax: Tubercles of prothorax each with 6 setae on male and 4 setae on female studied. Hind wing pads reaching posterior edge of abdominal segment III of male and posterior edge of segment IV of female studied. Abdomen: Paired anterior hook plates pedunculated, almost circular and present from abdominal terga IV to VII (Fig 11), more elongated on segment VII, each with more than 30 spines; also tergum III with pair of small flat hook plates each with 10 – 15 spines. Paired posterior hook plates sessile on terga III – V, almost circular, each with more than 30 spines. Description of the imago Male body length: 7.70 mm (1 specimen) each forewing: 12.20 mm, each hind wing: 10.60 mm; female, length: 8 mm (1 specimen), each forewing: 11.50 mm, each hind wing: 10.30 mm. Colour: Specimens preserved in ethanol dark brown dorsally, yellowish ventrally, spurs yellowish brown, setae brown, females generally darker than males; small black or dark brown spots dorsally. Forewings dark with small pale spots, pale in preserved specimens, darker in females, hind wings pale; each with dark area on leading edge between subcostal (Sc) and middle space between first radial (R 1) and second radial (R 2) veins in fore- and hind wings, with some darker stripes; each forewing hyaline in circular-oval zone, connecting medial (M) and medio-cubital (m-cu) veins and in distal area of second cubital vein (Cu 2) (Holzenthal et al. 2007); in each hind wing proximal segment of medial veins and connection area (M 1 - 4), not conspicuous, second cubital (Cu 2) vein not conspicuous either totally or partially. Male genitalia (Figs 19, 24, 31 b, 31 c, 34): Apical segment of each inferior appendage (Figs 19 A, 24 a, 34 c) with basal and distal edges diverging, dorsal and ventral edges slightly diverging, posterior edge of 2 nd segment of each inferior appendage slightly concave, ventral edge slightly convex, 2.5 times longer that dorsal edge (Figs 19 A, 24 a). Apicodorsal vertex angular, apicoventral angle rounded, projecting as thick lobe narrowing progressively to apex. Parameres in ventral view curved posteromesad in apical half (Figs 19 BV p, 24 c, 34 d). In lateral view (Fig 19 BL) each constricted at short distance beyond base, dilated at middle, with rounded dorsal margin, without spines in midventral margin, few long and thick spines on lateroventral area; midlateral surface covered by fine spicules or setae from middle to ventral edge, absent in middle anteroventral and posterodorsal edges. Aedeagus (phallicata) in lateral view (Fig 19 CL) with dorsal margin concave and posterior corner of concavity hooked anterad, posterior edge of aedeagus straight, projected posteroventrad in small ventral apex (Fig 19 CL); ventral lobe of aedeagus semicircular (Fig 19 BV vl). Lateroventral lobes of phallus straight, posterior edges strongly convex, apicolateral margins semicircular (Fig 19 CD lvl). Apicodorsal lobe of segment IX dilated subapicolaterally (Figs 19 DD al, 24 b, 31 b, 34 b), preanal appendages (Fig 19 DD pa, 24 b, 31 b, 34 b) round, almost as long as apicodorsal lobe, covered partially by it, mainly posteriorly; in ventral view apical band (Fig 19 DV ab, 31 c) V-shaped, longer than wide, posterior edges rounded, almost parallel, non-sclerotized inside area (Fig 19 DV va) straight, anal sclerites triangular (Fig 19 DV as). Female genitalia (Figs 20, 26, 32 a, 35): In lateral view (Figs 20 L, 26 L, 32 a, 35 c), posterodorsal margin of segment VIII valves on each side round, posterolateral margin with small excision in middle, ventral edge convex, dorsal edge with dorsolateral triangular projection (Fig 20 L). In dorsal view (Figs 20 D, 26 D, 35 b), with indentation between segment VIII valves, projections triangular with rounded corners, delimiting oval space (Fig 20 D). In ventral view (Fig 20 V, 26 V, 35 d) segment VIII valves forming two elongate sclerites, distal 2 / 3 of proximal edges diverging, distal edges straight or slightly concave; intersegmental membrane with pair of deep-dark, slightly elongate sclerites.	en	Valladolid, María, Arauzo, Mercedes, Chertoprud, Mikhail V., Chvojka, Pavel, Czachorowski, Stanisław, Dorda, Beatriz A., Hinić, Jelena, Ibrahimi, Halil, Karaouzas, Ioannis, Krpač, Vladimir, Kučinić, Mladen, Lodovici, Omar, Salokannel, Juha, Stamenković, Valentina Slavevska, Stojanović, Katarina (2021): The Rhyacophila fasciata Group in Europe: Rhyacophila fasciata Hagen 1859 and formerly synonymized species (Trichoptera: Rhyacophilidae), with new description of Rhyacophila fasciata and Rhyacophila septentrionis McLachlan 1865 (stat. prom.). Zootaxa 4975 (1): 1-57, DOI: https://doi.org/10.11646/zootaxa.4975.1.1
8A39CA25554D515859ABAC78820BC226.taxon	description	Head: In R. fasciata ventral side of head pale, in some specimens one or two rows of low conspicuous pale spots from posterior edge to halfway up the head. In R. septentrionis central area with slightly darker U-shape area from middle to posterior end of protogula, covered with pale spots. In R. fasciata the pair of dark brown patches outside posterior edge of frontal suture in the shape of open wings, with dark spots inside. In R. septentrionis the pair of dark brown patches more nearly triangular, with paler spots with dark borders inside. In R. fasciata mandibles dark reddish brown, uniformly coloured, left mandible with inner blade slightly convex. In R. septentrionis mandibles reddish brown, with darker apices, left mandible with inner blade slightly concave. Thorax: In R. fasciata on posterior half three dark areas, one median and two lateral, two lines of dark spots inside median area, circular spot on anterior half; posterior area with two to three rows of brown spots with dark border in circular-oval shape and parallel to posterior border. In R. septentrionis on posterior half three dark areas (in Hickin 1954) or dark base with darker mesal stripe with two lines of pale spots inside, two pale brown spots with dark borders anterolateral and posteriolaterally to seta 6 #, circular and elongate respectively; posterior area with two-three lines of darkly-bordered pale brown spots of different shapes and parallel to posterior border. In R. fasciata anterior margin with black spot near anterolateral angle; mesal area of posterior marginal band of sclerite dark brown or black posteriorly. In R. septentrionis anterior margin with a line of black spots near anterolateral angle, light brown in middle; mesal area of posterior marginal band of each sclerite black posteriorly. Abdomen: In R. fasciata dorsal sclerite of segment IX uniformly coloured, brown spots in posteromesal area, band of anterior edge with two extensions surrounding pit # 1, connected with dark stripes in darker larvae. In R. septentrionis sclerite of segment IX light coloured, mesal area slightly darker, with paler spots inside.	en	Valladolid, María, Arauzo, Mercedes, Chertoprud, Mikhail V., Chvojka, Pavel, Czachorowski, Stanisław, Dorda, Beatriz A., Hinić, Jelena, Ibrahimi, Halil, Karaouzas, Ioannis, Krpač, Vladimir, Kučinić, Mladen, Lodovici, Omar, Salokannel, Juha, Stamenković, Valentina Slavevska, Stojanović, Katarina (2021): The Rhyacophila fasciata Group in Europe: Rhyacophila fasciata Hagen 1859 and formerly synonymized species (Trichoptera: Rhyacophilidae), with new description of Rhyacophila fasciata and Rhyacophila septentrionis McLachlan 1865 (stat. prom.). Zootaxa 4975 (1): 1-57, DOI: https://doi.org/10.11646/zootaxa.4975.1.1
8A39CA25554D515859ABAC78820BC226.taxon	description	Abdomen: In R. fasciata paired anterior hook plates palmate, pair of small flat anterior plates without spines on tergum III; paired posterior hook plates slightly oblong transversally. In R. septentrionis paired anterior hook plates almost circular, pair of small flat anterior plates on tergum III with 10 – 15 spines; paired posterior hook plates almost circular.	en	Valladolid, María, Arauzo, Mercedes, Chertoprud, Mikhail V., Chvojka, Pavel, Czachorowski, Stanisław, Dorda, Beatriz A., Hinić, Jelena, Ibrahimi, Halil, Karaouzas, Ioannis, Krpač, Vladimir, Kučinić, Mladen, Lodovici, Omar, Salokannel, Juha, Stamenković, Valentina Slavevska, Stojanović, Katarina (2021): The Rhyacophila fasciata Group in Europe: Rhyacophila fasciata Hagen 1859 and formerly synonymized species (Trichoptera: Rhyacophilidae), with new description of Rhyacophila fasciata and Rhyacophila septentrionis McLachlan 1865 (stat. prom.). Zootaxa 4975 (1): 1-57, DOI: https://doi.org/10.11646/zootaxa.4975.1.1
8A39CA25554D515859ABAC78820BC226.taxon	description	Genitalia: In R. fasciata, posterior edge of 2 nd segment of each inferior appendage slightly concave dorsally and convex ventrally, ventral edge slightly concave and 2.0 times longer than dorsal edge, apicodorsal vertex slightly angular. In R. septentrionis, posterior edge of 2 nd segment of each inferior appendage completely concave, ventral edge slightly convex and 2.5 times longer than dorsal edge, apicodorsal vertex more conspicuously angular. In R. fasciata parameres in lateral view each slender near base, dilated in middle, with almost parallel dorsal and ventral margins in central area (S-shape), with two rows of long and thick spines on midventral margin, pointing outward; midlateral surface with sparse fine spicules or setae, from middle posteroventral to posterodorsal edge, covering most of apex, absent on middle anteroventral and dorsal edges. In R. septentrionis, parameres each slender near base, dilated in middle, with rounded dorsal margin, without spines in midventral margin, few long and thick spines on lateroventral area; midlateral surface covered by fine spicules or setae, from middle to ventral edge, absent on middle anteroventral and posterodorsal edges. In R. fasciata posterior edge of aedeagus straight, projected posteroventrad and rounded at posteroventral apex; ventral lobe of aedeagus triangular, lateral edges nearly straight; lateroventral lobes of phallus straight, posterior edges convex, apicolateral margins rounded. In R. septentrionis posterior edge of aedeagus straight, projected posteroventrad in small posteroventral protrusion; ventral lobe of aedeagus semicircular; lateroventral lobes of phallus straight, posterior edges convex, apicolateral margins semicircular. In R. fasciata apicodorsal lobe of segment IX slightly dilated subapicolaterally, almost round, with small apicomesal excision in some specimens; preanal appendages round, shorter than apicodorsal lobe, almost covered by it; in ventral view, apical band V-shaped and 1.8 times as long as wide, posterior arms widely separated, nonsclerotized area between arms of apical band heart-shaped and with posteromesal excision and posterior edges rounded. In R. septentrionis apicodorsal lobe of segment IX strongly dilated subapicolaterally; preanal appendages round, almost as long as apicodorsal lobe, covered partially by it, mainly posteriorly; in ventral view, apical band V-shaped and 1.2 times as long as wide, posterior arms close together and almost parallel, non-sclerotized area between arms of apical band straight and narrow and without posteromesal excision.	en	Valladolid, María, Arauzo, Mercedes, Chertoprud, Mikhail V., Chvojka, Pavel, Czachorowski, Stanisław, Dorda, Beatriz A., Hinić, Jelena, Ibrahimi, Halil, Karaouzas, Ioannis, Krpač, Vladimir, Kučinić, Mladen, Lodovici, Omar, Salokannel, Juha, Stamenković, Valentina Slavevska, Stojanović, Katarina (2021): The Rhyacophila fasciata Group in Europe: Rhyacophila fasciata Hagen 1859 and formerly synonymized species (Trichoptera: Rhyacophilidae), with new description of Rhyacophila fasciata and Rhyacophila septentrionis McLachlan 1865 (stat. prom.). Zootaxa 4975 (1): 1-57, DOI: https://doi.org/10.11646/zootaxa.4975.1.1
8A39CA25554D515859ABAC78820BC226.taxon	description	Genitalia: In R. fasciata in lateral view with dorsal margin of segment VIII 3 / 4 as long as venter and with prominent dorsolateral projection, posterolateral margin of valve on each side irregular, with 1 or 2 posterior projections, posteroventral projection more conspicuous, ventral margin slightly concave near base and straight or convex distally, in some specimens slightly indented posteriorly. In R. septentrionis in lateral view, with dorsal margin of segment VIII half as long as venter and with prominent dorsolateral projection, posterolateral margin of valve on each side round and with small excision in middle, ventral margin convex. In R. fasciata in dorsal view with indentation between segment VIII valves nearly enclosed by pair of thick posterolateral projections, with rounded apical and subapical corners, small projection on apical corner of some specimens. In R. septentrionis with indentation between segment VIII valves nearly enclosed by pair of triangular projections, with more-nearly acute apical corners. In R. fasciata in ventral view segment VIII valves forming two elongate sclerites, basal third of proximal edges parallel, distal edges irregular, straight, diverging; intersegmental membrane with pair of dark ovoid or slightly elongate sclerites. In R. septentrionis in ventral view segment VIII valves forming two elongate sclerites, basal 1 / 3 of proximal edges converging, distal edges straight or slightly concave, diverging; intersegmental membrane with pair of deep-dark slightly elongate sclerites.	en	Valladolid, María, Arauzo, Mercedes, Chertoprud, Mikhail V., Chvojka, Pavel, Czachorowski, Stanisław, Dorda, Beatriz A., Hinić, Jelena, Ibrahimi, Halil, Karaouzas, Ioannis, Krpač, Vladimir, Kučinić, Mladen, Lodovici, Omar, Salokannel, Juha, Stamenković, Valentina Slavevska, Stojanović, Katarina (2021): The Rhyacophila fasciata Group in Europe: Rhyacophila fasciata Hagen 1859 and formerly synonymized species (Trichoptera: Rhyacophilidae), with new description of Rhyacophila fasciata and Rhyacophila septentrionis McLachlan 1865 (stat. prom.). Zootaxa 4975 (1): 1-57, DOI: https://doi.org/10.11646/zootaxa.4975.1.1
8A39CA25554D515859ABAC78820BC226.taxon	discussion	Rhyacophila fasciata has a wide area of distribution in northern and central Europe and there is no GeneBank entry from the type locality (Elberfeld, Germany), so we have included some sequences from localities around it (Belgium and Germany). In the first analysis, we used these sequences, together with others from localities that previously have been checked as R. fasciata from Bosnia and Herzegovina, Belgium, Croatia, Czech Republic, Estonia, Finland, Germany, Kosovo, Norway, Poland, Republic of Macedonia, Russia, Slovakia and Sweden, together with one sequence of R. cf. nubila from Russia and three of R. cf. obliterata from Poland and Russia. In the second analysis, to build the tree, we selected a total of 52 sequences: a maximum of two sequences of R. fasciata per country (25 in total), as well as 3 sequences of R. septentrionis, 3 sequences of R. denticulata, 4 sequences of R. sociata from Spain and France (promoted to species by Valladolid et al. 2018), 3 sequences of R. kykladica (promoted to species by Valladolid et al. 2019), 7 sequences of R. fasciata delici Kučinić & Valladolid (in Valladolid et al. 2020) and 3 sequences of R. fasciata viteceki Valladolid & Kučinić 2020 (in Valladolid et al. 2020), 1 sequence of R. cf. nubila from Russia and 3 of R. cf. obliterata from Poland and Russia (see Table 1 for references). The intra- and interspecific distances were calculated with all the sequences of each species (162 in total), as well as the bootstrap values of main branches in the second tree. The second tree (as a graphic) includes only some selected specimens and the bootstrap values cited. The first tree (Fig 27) shows that, despite the wide distribution of the specimens, the group is quite homogeneous genetically, with three well-supported branches, one for the R. fasciata specimens (bootstrap value of 98) (Fig 27 a), a second for R. septentrionis specimens (bootstrap value of 99) (Fig 27 b), and the last for the cf. obliterata specimens (b. v. of 99) (Fig 27 d). Intraspecific distances (maximum composite likelihood model) are lower than 1 % (0.0041 in the R. fasciata Group, 0.0014 in the R. septentrionis Group and 0.0053 in the R. cf. obliterata Group), while interspecific distances are higher than 1 % (R. fasciata-R. septentrionis = 0.0290, R. fasciata-R. cf. obliterata = 0.1040, R. fasciata-R. cf. nubil a = 0.0787, R. septentrionis-R. cf. obliterata = 0.0867, R. septentrionis-R. cf. nubila = 0.0618). The second tree (Fig 28) shows clear differences between R. fasciata and the other species included, that previously were considered as R. fasciata or subspecies of R. fasciata. Based on the divergence sequences, there are seven well-supported branches, corresponding to R. fasciata (bootstrap value of 99) (Fig 28 a), R. f. delici (b. v. of 98) (Fig 28 c), R. septentrionis (b. v. of 90) (Fig 28 d), R. denticulata (b. v. of 99) (Fig 28 e), R. sociata (b. v. of 97) (Fig 28 f), R. kykladica (b. v. of 100) (Fig 28 g), all clearly different from R. cf. nubila (Fig 28 h) and R. cf. obliterata (b. v. of 100) (Fig 28 i) branches. Rhyacophila f. viteceki (Fig 28 b) appears as an intermediate group between R. f. delici and R. fasciata groups. Intraspecific distances (maximum composite likelihood model) are lower than 1 % (Table 2 a), being the lowest in R. septentrionis, R. f. viteceki, and R. kykladica (0.0014, 0.0017, and 0.0019, respectively) while interspecific distances are higher than 1 % (Table 2 b) among all of these species. The lower distances are between R. septentrionis and R. f. viteceki (0.0101) and between R. f. delici and R. f. viteceki (0.0203).	en	Valladolid, María, Arauzo, Mercedes, Chertoprud, Mikhail V., Chvojka, Pavel, Czachorowski, Stanisław, Dorda, Beatriz A., Hinić, Jelena, Ibrahimi, Halil, Karaouzas, Ioannis, Krpač, Vladimir, Kučinić, Mladen, Lodovici, Omar, Salokannel, Juha, Stamenković, Valentina Slavevska, Stojanović, Katarina (2021): The Rhyacophila fasciata Group in Europe: Rhyacophila fasciata Hagen 1859 and formerly synonymized species (Trichoptera: Rhyacophilidae), with new description of Rhyacophila fasciata and Rhyacophila septentrionis McLachlan 1865 (stat. prom.). Zootaxa 4975 (1): 1-57, DOI: https://doi.org/10.11646/zootaxa.4975.1.1
