identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
E35487F53524FFBF0F93FB2EFD0CD608.text	E35487F53524FFBF0F93FB2EFD0CD608.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cryptonyssus Radovsky 1966	<div><p>Genus Cryptonyssus Radovsky, 1966</p> <p>Cryptonyssus Radovsky, 1966: 97.</p> <p>Type species: Cryptonyssus desultorius Radovsky, 1966.</p> </div>	http://treatment.plazi.org/id/E35487F53524FFBF0F93FB2EFD0CD608	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Orlova, M. V.;Klimov, P. B.;Orlov, O. L.;Smirnov, D. G.;Zhigalin, A. V.;Budaeva, I. V.;Emelyanova, A. A.;Anisimov, N. V.	Orlova, M. V., Klimov, P. B., Orlov, O. L., Smirnov, D. G., Zhigalin, A. V., Budaeva, I. V., Emelyanova, A. A., Anisimov, N. V. (2021): A checklist of bat-associated macronyssid mites (Acari: Gamasina: Macronyssidae) of Russia, with new host and geographical records. Zootaxa 4974 (3): 537-564, DOI: https://doi.org/10.11646/zootaxa.4974.3.4
E35487F53524FFBF0F93FBE9FD98D783.text	E35487F53524FFBF0F93FBE9FD98D783.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macronyssidae Oudemans 1936	<div><p>Family Macronyssidae Oudemans, 1936</p> <p>Liponyssinae Ewing, 1923: 2.</p> <p>Liponyssidae Vitzthum, 1931: 14.</p> <p>Macronyssidae Oudemans, 1936: 430.</p> <p>Type genus: Macronyssus Kolenati, 1858.</p> <p>Genus Cryptonyssus Radovsky, 1966</p> <p>Cryptonyssus Radovsky, 1966: 97.</p> <p>Type species: Cryptonyssus desultorius Radovsky, 1966.</p> </div>	http://treatment.plazi.org/id/E35487F53524FFBF0F93FBE9FD98D783	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Orlova, M. V.;Klimov, P. B.;Orlov, O. L.;Smirnov, D. G.;Zhigalin, A. V.;Budaeva, I. V.;Emelyanova, A. A.;Anisimov, N. V.	Orlova, M. V., Klimov, P. B., Orlov, O. L., Smirnov, D. G., Zhigalin, A. V., Budaeva, I. V., Emelyanova, A. A., Anisimov, N. V. (2021): A checklist of bat-associated macronyssid mites (Acari: Gamasina: Macronyssidae) of Russia, with new host and geographical records. Zootaxa 4974 (3): 537-564, DOI: https://doi.org/10.11646/zootaxa.4974.3.4
E35487F53524FFBE0F93FA58FE7AD2A5.text	E35487F53524FFBE0F93FA58FE7AD2A5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cryptonyssus flexus Radovsky 1967	<div><p>Cryptonyssus flexus Radovsky, 1967</p> <p>Cryptonyssus flexus Radovsky, 1967: 175</p> <p>Cryptonyssus flexus.— Radovsky, 2010: 93.</p> <p>Ornithonyssus flexus.— Micherdzinski, 1980: 73.</p> <p>Type locality: Biological station of University of Michigan, Cheboygan County, Michigan (USA) (Radovsky, 1967).</p> <p>Type host: Myotis lucifugus (Le Conte, 1831) (Radovsky, 1967).</p> <p>Material. 3 N1 ex Myotis brandtii, N1 ex M. mystacinus from galleries near village Staritsa, 29 February 2020, leg. A.A. Emelyanova; 2 FF ex Eptesicus nilssonii (Keyserling &amp; Blasius, 1839) from Kyshtym town, 13 July 2010, leg. O.L. Orlov, M. V. Orlova; 2 N1 ex Vespertilio murinus Linnaeus, 1758 from biological station “Kolarovo”, 09 August 2014 leg. A. V. Zhigalin; N1 from Myotis sibiricus, from Tigirekskiy State Natural Reserve, 13 July 2019, leg. O.L. Orlov; 8 N1 ex Plecotus ognevi from Okhotnichiya cave, 4 September 2015, leg. D. V. Kazakov; 6 N1 ex Myotis ikonnikovi Ognev, 1912, 20 July 2016, 3 N1 ex M. ikonnikovi, 29 July 2016; N1 ex Myotis petax, 17 July 2016, 3 N1 ex Plecotus ognevi, 24 July 2016, N1 ex Ep. nilssonii, 24 July 2016, from Zeyskiy State Natural Reserve, leg. D. V. Kazakov.</p> <p>Distribution in Russian Federation: Crimea (Stanyukovich, 1997), Tver’ Province (present paper), Dagestan Republic (Orlova et al., 2020a), Chelyabinsk Province (present paper), Tomsk Province (present paper), Altai Province (present paper), Irkutsk Province (present paper), Buryatiya Republic (Orlova et al., 2017), Amur Province (present paper).</p> <p>Distribution outside Russian Federation USA (Radovsky, 1967; Dood &amp; Kurta, 1988); Tajikistan (Stanyukovich, 1997).</p> <p>Hosts (Old World): Myotis davidii (as Myotis mystacinus — Stanyukovich, 1997), M. ikonnikovi (present paper), M. sibiricus (Orlova et al., 2017a), M. mystacinus (Stanyukovich, 1997), M. petax (present paper), Eptesicus nilssonii (present paper), Pipistrellus nathusii (Orlova et al., 2020a), Plecotus ognevi (present paper), Vespertilio murinus (present paper).</p> </div>	http://treatment.plazi.org/id/E35487F53524FFBE0F93FA58FE7AD2A5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Orlova, M. V.;Klimov, P. B.;Orlov, O. L.;Smirnov, D. G.;Zhigalin, A. V.;Budaeva, I. V.;Emelyanova, A. A.;Anisimov, N. V.	Orlova, M. V., Klimov, P. B., Orlov, O. L., Smirnov, D. G., Zhigalin, A. V., Budaeva, I. V., Emelyanova, A. A., Anisimov, N. V. (2021): A checklist of bat-associated macronyssid mites (Acari: Gamasina: Macronyssidae) of Russia, with new host and geographical records. Zootaxa 4974 (3): 537-564, DOI: https://doi.org/10.11646/zootaxa.4974.3.4
E35487F53525FFBE0F93FDB4FC1AD55D.text	E35487F53525FFBE0F93FDB4FC1AD55D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cryptonyssus pipistrelli (Oudemans 1904)	<div><p>Cryptonyssus pipistrelli (Oudemans, 1904)</p> <p>Liponyssus pipistrelli Oudemans, 1904: 111.</p> <p>Macronyssus pipistrelle.— Buitendijk, 1945: 306.</p> <p>Ornithonyssus pavlovskii Lange, 1959: 484; Zemskaya, 1966: 50 (synonymy by Radovsky, 1969).</p> <p>Cryptonyssus pipistrelle.— Radovsky, 1969: 462; 2010: 93.</p> <p>Ornithonyssus pipistrelli.— Mehl, 1979: 33; Micherdzinski, 1980: 104; Baker &amp; Craven, 2003: 7; Villegas-Guzman et al., 2005: 125; Švana et al., 2006: 40; Marchenko, 2007.</p> <p>Type locality: Bodegraven (Netherlands, neotype).</p> <p>Type host: Unknown.</p> <p>Material. N1 ex Myotis dasycneme (Boie, 1825) from biological station “Lukashino”, 13 July 2018, leg. O.L. Orlov, M. V. Orlova.</p> <p>Distribution in Russian Federation: Leningrad Province (as Ornithonyssus pipistrelli — Zemskaya, 1966), Penza Province (as Ornithonyssus pipistrelli — Stanyukovich, 1997), Kirov Province (as Ornithonyssus pipistrelli — Zemskaya, 1966), North Osetia (as Ornithonyssus pipistrelli — Stanyukovich, 1997), Sverdlovsk Province (as Ornithonyssus pipistrelli — Orlova &amp; Orlov, 2011), Altai Region (as Ornithonyssus pipistrelli — Marchenko, 2007), Khakassia Republic (Orlova et al., 2017a), Buryatia Republic (Orlova et al., 2016).</p> <p>Distribution outside Russian Federation: Europe (Great Britain— Evans &amp; Till, 1966, Spain— Quetglas, 2016, etc.), Asia (Georgia: as Ornithonyssus pipistrelli — Stanyukovich, 1997), Central America (as Ornithonyssus pipistrelli — Villegas-Guzman et al., 2005).</p> <p>Hosts (Old World): [Muscardinus avellanarius Linnaeus 1758] (Lange, 1959), Myotis dasycneme (Stanyukovich, 1997), M. daubentonii (Stanyukovich, 1997), M. brandtii (Stanyukovich, 1997), M. mystacinus (Stanyukovich, 1997), M. ikonnikovi (Stanyukovich, 1997), M. nattereri (Baker &amp; Craven, 2003), M. sibiricus (Orlova et al., 2017a), M. escalerai Cabrera, 1904 (Quetglas, 2016), Plecotus auritus (Evans &amp; Till, 1966), Pl. austriacus Fisher, 1829 (Baker &amp; Craven, 2003), Pipistrellus pipistrellus (Schreber, 1774) (Beron, 2014), Pi. nathusii (Keyserling &amp; Blasius, 1839) (Stanyukovich, 1997), Eptesicus serotinus Schreber, 1774 (Evans &amp; Till, 1966), Ep. nilssonii (Stanyukovich, 1997), Nyctalus noctula Schreber, 1774 (Stanyukovich, 1997).</p> </div>	http://treatment.plazi.org/id/E35487F53525FFBE0F93FDB4FC1AD55D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Orlova, M. V.;Klimov, P. B.;Orlov, O. L.;Smirnov, D. G.;Zhigalin, A. V.;Budaeva, I. V.;Emelyanova, A. A.;Anisimov, N. V.	Orlova, M. V., Klimov, P. B., Orlov, O. L., Smirnov, D. G., Zhigalin, A. V., Budaeva, I. V., Emelyanova, A. A., Anisimov, N. V. (2021): A checklist of bat-associated macronyssid mites (Acari: Gamasina: Macronyssidae) of Russia, with new host and geographical records. Zootaxa 4974 (3): 537-564, DOI: https://doi.org/10.11646/zootaxa.4974.3.4
E35487F53525FFBE0F93F9ADFD22D49F.text	E35487F53525FFBE0F93F9ADFD22D49F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ichoronyssus Kolenati 1858	<div><p>Genus Ichoronyssus Kolenati, 1858</p> <p>Ichoronyssus Kolenati, 1858: 5.</p> <p>Ichoronyssus.— Radovsky, 2010: 41.</p> <p>Type species: Dermanissus [sic] scutatus Kolenati, 1856.</p> <p>Spinolaelaps Radford, 1940: 96.</p> <p>Type species: Spinolaelaps jacksoni Radford, 1940.</p> <p>Neospinolaelaps Zumpt &amp; Patterson, 1952: 159.</p> <p>Type species: Neospinolaelaps miniopteri Zumpt &amp; Patterson, 1952.</p> <p>Plesiolaelaps Womersley, 1957: 68.</p> <p>Type species: Plesiolaelaps miniopterus Womersley, 1957.</p> </div>	http://treatment.plazi.org/id/E35487F53525FFBE0F93F9ADFD22D49F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Orlova, M. V.;Klimov, P. B.;Orlov, O. L.;Smirnov, D. G.;Zhigalin, A. V.;Budaeva, I. V.;Emelyanova, A. A.;Anisimov, N. V.	Orlova, M. V., Klimov, P. B., Orlov, O. L., Smirnov, D. G., Zhigalin, A. V., Budaeva, I. V., Emelyanova, A. A., Anisimov, N. V. (2021): A checklist of bat-associated macronyssid mites (Acari: Gamasina: Macronyssidae) of Russia, with new host and geographical records. Zootaxa 4974 (3): 537-564, DOI: https://doi.org/10.11646/zootaxa.4974.3.4
E35487F53526FFBD0F93FF64FBAED04F.text	E35487F53526FFBD0F93FF64FBAED04F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ichoronyssus scutatus (Kolenati 1856)	<div><p>Ichoronyssus scutatus (Kolenati, 1856)</p> <p>Dermanissus [sic] scutatus Kolenati, 1856: 21.</p> <p>Ichoronyssus scutatus.— Kolenati, 1858: 6; 1859: 173; Radovsky, 1966: 93; 1967: 100; Beron, 1968: 159; 1974: 65; Uchikawa, 1976: 841; Haitlinger, 1978: 713; Rybin, 1983: 356; Gu &amp; Wang, 1985: 156; Whitaker et al., 1990; Cicek et al., 2007: 248; Stanyukovich &amp; Malinovskiy, 1992: 70; Radovsky, 2010: 41.</p> <p>Ichoronyssus (I.) scutatus.—Anciaux de Faveaux, 1985: 692.</p> <p>Type locality: Unknown.</p> <p>Type host: Unknown.</p> <p>Distribution in Russia: Crimea (as Spinolaelaps jacksoni — Vshivkov, 1963; Stanyukovich, 1997; Orlova &amp; Orlov, 2018), Primorskiy Region (Medvedev et al., 1991), [Far East] (Stanyukovich, 1997).</p> <p>Distribution outside Russia: Eurasia (Ogajanyan &amp; Arutyunyan, 1974; Stanyukovich, 1997). Africa (Beron, 2014).</p> <p>Hosts. Miniopterus schreibersi (Stanyukovich, 1997), Min. fuliginosus (as Miniopterus schreibersi — Medvedev et al., 1991), Myotis myotis (Borkhausen, 1797) (Stanyukovich, 1997), M. blythii Tomes, 1857 (Vshivkov, 1963; Stanyukovich, 1997; Orlova &amp; Orlov, 2018), M. macrodactylus (Radovsky, 1967), Vespertilio murinus (Radovsky, 1967), Plecotus auritus (Beron, 2014), Pipistrellus javanicus Gray, 1838 (Beron, 2014), Rhinolophus euryale Blasius, 1853 (Stanyukovich, 1997), Rh. ferrumequinum (Schreber, 1774) (Vshivkov, 1963).</p> </div>	http://treatment.plazi.org/id/E35487F53526FFBD0F93FF64FBAED04F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Orlova, M. V.;Klimov, P. B.;Orlov, O. L.;Smirnov, D. G.;Zhigalin, A. V.;Budaeva, I. V.;Emelyanova, A. A.;Anisimov, N. V.	Orlova, M. V., Klimov, P. B., Orlov, O. L., Smirnov, D. G., Zhigalin, A. V., Budaeva, I. V., Emelyanova, A. A., Anisimov, N. V. (2021): A checklist of bat-associated macronyssid mites (Acari: Gamasina: Macronyssidae) of Russia, with new host and geographical records. Zootaxa 4974 (3): 537-564, DOI: https://doi.org/10.11646/zootaxa.4974.3.4
E35487F53526FFBD0F93FC83FCD7D62A.text	E35487F53526FFBD0F93FC83FCD7D62A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macronyssus Kolenati 1858	<div><p>Genus Macronyssus Kolenati, 1858</p> <p>Macronyssus Kolenati, 1858: 5;</p> <p>Macronyssus.— Radovsky, 2010: 51.</p> <p>Type species: Caris longimana Kolenati, 1856.</p> <p>Lepronyssus Kolenati, 1858: 5.</p> <p>Type species: Lepronyssus leprosus Kolenati, 1858.</p> <p>Liponyssus of authors, in part, not Kolenati, 1858.</p> <p>Ichoronyssus of authors, in part, not Kolenati, 1858.</p> <p>Allonyssus Buitendijk 1945: 307 (synonymy by Evans &amp; Till, 1966).</p> <p>Hirstesia Fonseca, 1948: 293.</p> <p>Type species: Liponyssus sternalis Hirst, 1922.</p> <p>Langeonyssus Grokhovskaya &amp; Nguen-Huan-Hue, 1961: 1645.</p> <p>Type species: Langeonyssus tieni Grokhovskaya &amp; Nguen-Huan-Hue, 1961.</p> <p>Ichoronyssus (Bianyssus) Wen, 1975: 352.</p> <p>Type species: Ichoronyssus (Bianyssus) ventralis Wen, 1975.</p> </div>	http://treatment.plazi.org/id/E35487F53526FFBD0F93FC83FCD7D62A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Orlova, M. V.;Klimov, P. B.;Orlov, O. L.;Smirnov, D. G.;Zhigalin, A. V.;Budaeva, I. V.;Emelyanova, A. A.;Anisimov, N. V.	Orlova, M. V., Klimov, P. B., Orlov, O. L., Smirnov, D. G., Zhigalin, A. V., Budaeva, I. V., Emelyanova, A. A., Anisimov, N. V. (2021): A checklist of bat-associated macronyssid mites (Acari: Gamasina: Macronyssidae) of Russia, with new host and geographical records. Zootaxa 4974 (3): 537-564, DOI: https://doi.org/10.11646/zootaxa.4974.3.4
E35487F53526FFBC0F93FA3EFC49D131.text	E35487F53526FFBC0F93FA3EFC49D131.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macronyssus charusnurensis (Dusbabek 1966)	<div><p>Macronyssus charusnurensis (Dusbábek, 1966)</p> <p>Ichoronyssus charusnurensis Dusbábek, 1966: 44</p> <p>Ichoronyssus mirabilis Senotrusova &amp; Tagil’tsev, 1968: 134; Tagil’tsev, 1971: 383 (synonymy by Medvedev et al., 1991). Macronyssus sp. 7 Uchikawa, 1979a: 35.</p> <p>Macronyssus charusnurensis.— Medvedev et al., 1991: 33; Stanyukovich, 1997: 30.</p> <p>Type locality: Char-us-nuur (Mongolia) (Dusbábek, 1966).</p> <p>Type host: Myotis mystacinus (Dusbábek, 1966).</p> <p>Material. 2 N1 ex Myotis petax from Arkheologicheskaya cave, 13 June 2016, leg. L.B. Kravchenko; 3 N1 ex Myotis frater (G. Allen, 1823), Argaley-3 cave, 21 May 2016, leg. D. V. Kazakov; N1 ex Plecotus ognevi from Argaley-3 cave, 19 September 2015, leg. D. V. Kazakov; 23 FF (inc. 20 with intrauterine larvae), 11 MM, 83 N1 ex Myotis petax, 2 N1 ex Murina hilgendorfi from Dolganskaya Yama cave, 16 August 2016, leg. D. V. Kazakov; F ex Eptesicus nilssonii from Babushkin town, 22 June 2016, leg. D. V. Kazakov; N1 ex Vespertilio sinensis Peters, 1880 from Nizhniy Tsasuchey, 17 July 2015, leg. D. V. Kazakov; 41 FF (inc. 37 with intrauterine larvae), 11 MM, 58 N1, 4 N2 ex Myotis petax from Zeyskiy natural reserve 17 July 2016, leg. D. V. Kazakov.</p> <p>Distribution in Russian Federation: Khanty-Mansi Autonomous Region (Orlova et al., 2014), Altai Region (Orlova et al., 2017a,b), Novosibirsk Province (Orlova et al., 2015c), Khakassia Republic (our data, present paper), Tuva Republic (Orlova et al., 2015a), Krasnoyarsk Region (Stanyukovich, 1997; Orlova et al., 2014), Irkutsk Province (Orlova et al., 2016), Buryatiya Republic (Orlova et al., 2016; present paper), Yakutiya Republic (Orlova et al., 2017a), Zabaykal’skiy Region (present paper), Amur Province (our data, present paper), Primorskiy Region (Medvedev et al., 1991), Sakhalin Province (Kunashir island) (Orlova et al., 2015b).</p> <p>Distribution outside Russian Federation: Kazakhstan (Stanyukovich, 1997). Mongolia (Dusbábek, 1966). Japan (Uсhikawa, 1979b).</p> <p>Hosts. Myotis petax (as Myotis daubentonii — Medvedev et al., 1991; Stanyukovich, 1997; Uсhikawa, 1979b), Myotis sibiricus (Orlova et al., 2017b; as M. mystacinus — Stanyukovich, 1997;), M. frater (present paper), M. ikonnikovi (Beron, 2014), M. frater (Beron, 2014), M. blythii (Orlova et al., 2017b), Vespertilio sinensis (Uсhikawa, 1979b), Murina hilgendorfi (present paper), Eptesicus nilssonii (this article; as Eptesicus parvus —Uсhikawa, 1979b), Plecotus ognevi (as Plecotus auritus — Stanyukovich, 1997).</p> </div>	http://treatment.plazi.org/id/E35487F53526FFBC0F93FA3EFC49D131	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Orlova, M. V.;Klimov, P. B.;Orlov, O. L.;Smirnov, D. G.;Zhigalin, A. V.;Budaeva, I. V.;Emelyanova, A. A.;Anisimov, N. V.	Orlova, M. V., Klimov, P. B., Orlov, O. L., Smirnov, D. G., Zhigalin, A. V., Budaeva, I. V., Emelyanova, A. A., Anisimov, N. V. (2021): A checklist of bat-associated macronyssid mites (Acari: Gamasina: Macronyssidae) of Russia, with new host and geographical records. Zootaxa 4974 (3): 537-564, DOI: https://doi.org/10.11646/zootaxa.4974.3.4
E35487F53527FFBC0F93FD40FD9DD535.text	E35487F53527FFBC0F93FD40FD9DD535.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macronyssus corethroproctus (Oudemans 1902)	<div><p>Macronyssus corethroproctus (Oudemans, 1902)</p> <p>Liponyssus corethroproctus Oudemans, 1902: 15.</p> <p>Macronyssus corethroproctus (Oudemans).— Buitendijk, 1945: 306; Radovsky, 1966: 94; 1967: 117; 2010: 56; Beron, 1969: 160; Haitlinger, 1978: 712.</p> <p>Type locality: Sneek (Netherlands) (Oudemans, 1902).</p> <p>Type host: Myotis dasycneme (Oudemans, 1902).</p> <p>Material. 3 FF, 31 N1 ex pond bat Myotis dasycneme, 1-31 November 2005, N1 ex Myotis mystacinus, 25 November 2001, 4 N1 from Natterer’s bat Myotis nattereri, from State Nature Reserve “Samarskaya Luka”, 25 November 2001, leg. D.G. Smirnov; F ex Eptesicus nilssonii, from Kuminskiy settlement, 7 August 2008, leg. K. V. Bernikov; 2 N1 ex Myotis dasycneme, from Mordovian State Natural Reserve, 23 June 2016, leg. D.G. Smirnov; 2 N1 ex Pipistrellus nathusii from Pogromnoye settlement 26 June 2003, leg. D.G. Smirnov; F ex Myotis dasycneme, from Barsukovskaya cave, 8 November 2013, leg. A. V. Zhigalin.</p> <p>Distribution in Russian Federation: Leningrad Province (Stanyukovich, 1997; Orlova et al., 2014), Penza Province (Stanyukovich, 1997), Samara Province (present paper), Mordovia Republic (present paper), Stavropol Region (Stanyukovich, 1997), Udmurtiya Republic (Orlova et al., 2011), Bashkiriya Republic (Orlova et al., 2014; Orlova &amp; Orlov, 2013), Orenburg Province (present paper), Sverdlovsk Province (Orlova et al., 2012; Vinarskaya &amp; Orlova, 2012), Chelyabinsk Province (Orlova et al., 2014), Khanty-Mansi Autonomous Region (Orlova et al., 2014; Orlova &amp; Tomishina, 2019; present paper), Altai Region (Orlova et al., 2014), Novosibirsk Province (present paper), [Urals, Western Siberia— Orlova et al., 2015d].</p> <p>Distribution outside Russian Federation: Europe (Radovsky, 1967).</p> <p>Hosts. Myotis dasycneme (Stanyukovich, 1997; Orlova et al., 2011; Orlova et al., 2012; Orlova et al., 2012; Vinarskaya &amp; Orlova, 2012; Orlova et al., 2014), Myotis blythii (Beron, 2014) Myotis daubentonii (Orlova et al., 2011), Myotis nattereri (present paper), Myotis mystacinus (Orlova &amp; Orlov, 2013; present paper), Myotis brandtii (present paper), Pipistrellus nathusii (Stanyukovich, 1997; present paper), Eptesicus nilssonii (present paper), Vespertilio murinus (Stanyukovich, 1997).</p> </div>	http://treatment.plazi.org/id/E35487F53527FFBC0F93FD40FD9DD535	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Orlova, M. V.;Klimov, P. B.;Orlov, O. L.;Smirnov, D. G.;Zhigalin, A. V.;Budaeva, I. V.;Emelyanova, A. A.;Anisimov, N. V.	Orlova, M. V., Klimov, P. B., Orlov, O. L., Smirnov, D. G., Zhigalin, A. V., Budaeva, I. V., Emelyanova, A. A., Anisimov, N. V. (2021): A checklist of bat-associated macronyssid mites (Acari: Gamasina: Macronyssidae) of Russia, with new host and geographical records. Zootaxa 4974 (3): 537-564, DOI: https://doi.org/10.11646/zootaxa.4974.3.4
E35487F53527FFB30F93F944FEF6D07C.text	E35487F53527FFB30F93F944FEF6D07C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macronyssus crosbyi (Ewing & Stover 1915)	<div><p>Macronyssus crosbyi (Ewing &amp; Stover, 1915)</p> <p>Liponyssus crosbyi Ewing &amp; Stover, 1915: 109.</p> <p>Leiognathus crosbyi.— Ewing, 1923: 7.</p> <p>Ichoronyssus hasei Vitzthum, 1932: 23 (synonymy by Radovsky, 1966).</p> <p>Ichoronyssus kolenatii Fonseca, 1948: 280 (synonymy by Radovsky, 1966).</p> <p>Ichoronyssus crosbyi.— Fonseca, 1948: 299; Yunker &amp; Radovsky, 1966: 98.</p> <p>Ichoronyssus britanicus.— Furman, 1950: 480 (misidentification).</p> <p>Ichoronyssus quadridentatus Strandtmann &amp; Hunt, 1951: 462; Baker &amp; Strandtmann, 1959: 225 (synonymy by Radovsky, 1966).</p> <p>Macronyssus crosbyi.— Radovsky, 1966: 94; 1967: 119; Anciaux de Faveaux, 1971: 52; Hoffmann et al., 1972: 153; Whitaker, 1973: 1149; Yunker et al., 1975: 252; Whitaker et al., 1983; Whitaker &amp; Maser, 1985: 73; Stanjukovich, 1990: 197; Ritzi et al., 2001: tabl.; Reeves et al., 2007: 53; Jaunbauere et al., 2008: 40; Radovsky, 2010: 56.</p> <p>Type locality: Rockport, Missouri (Ewing &amp; Stover, 1915).</p> <p>Type host: “ Vesper subulatus ” (Ewing &amp; Stover, 1915).</p> <p>Material. M ex soprano pipistrelle Pipistrellus pygmaeus Leach, 1825 from biological station “Venevitinovo”, 4 June 2019, leg. O.L. Orlov, I. V. Budaeva.</p> <p>Distribution in Russian Federation: Karelia Republic (Lebedeva et al., 2020), Voronezh Province (present paper), Krasnoyarsk Region (Orlova et al., 2017a), Irkutsk Province (Orlova et al., 2017a), Buryatiya Republic (Orlova et al., 2017a), Yakutiya Republic (Orlova et al., 2017a), Khabarovsk Region (Medvedev et al., 1991), Magadan Province (Medvedev et al., 1991), [Urals, Altai, Russian Far East— Orlova et al., 2015d].</p> <p>Distribution outside Russian Federation: Europe (Baltic states— Stanyukovich, 1990a; Jaunbauere et al., 2008), North and South America (Radovsky, 1967; Hoffmann et al., 1972; Presley 2004).</p> <p>Hosts (Old World): Eptesicus nilssonii (Stanyukovich, 1990a; Lebedeva et al., 2020), Myotis daubentonii (Stanyukovich, 1990a), M. dasycneme (Stanyukovich, 1990a), M. mystacinus (Stanyukovich, 1990a), M. sibiricus (Orlova et al., 2017a), M. brandtii (Stanyukovich, 1990a), Pipistrellus nathusii (Stanyukovich, 1990a), Pipistrellus pygmaeus (present paper), Vespertilio murinus (Stanyukovich, 1990a), Plecotus ognevi (as Pl. auritus) (Medvedev et al., 1991).</p> </div>	http://treatment.plazi.org/id/E35487F53527FFB30F93F944FEF6D07C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Orlova, M. V.;Klimov, P. B.;Orlov, O. L.;Smirnov, D. G.;Zhigalin, A. V.;Budaeva, I. V.;Emelyanova, A. A.;Anisimov, N. V.	Orlova, M. V., Klimov, P. B., Orlov, O. L., Smirnov, D. G., Zhigalin, A. V., Budaeva, I. V., Emelyanova, A. A., Anisimov, N. V. (2021): A checklist of bat-associated macronyssid mites (Acari: Gamasina: Macronyssidae) of Russia, with new host and geographical records. Zootaxa 4974 (3): 537-564, DOI: https://doi.org/10.11646/zootaxa.4974.3.4
E35487F53528FFB30F93FC8DFD83D5D6.text	E35487F53528FFB30F93FC8DFD83D5D6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macronyssus cyclaspis (Oudemans 1906)	<div><p>Macronyssus cyclaspis (Oudemans, 1906b)</p> <p>Liponyssus pipistrelli Oudemans, 1906a: 37.</p> <p>Liponyssus cyclaspis Oudemans, 1906b: 61.</p> <p>Steatonyssus cyclaspis.— Buitendijk, 1945: 307; Fonseca, 1948: 316.</p> <p>Ichoronyssus štorkáni.— Dusbábek, 1964a: 97 (synonymy by Radovsky, 1966).</p> <p>Ichoronyssus cyclaspis.— Pinchuk, 1970: 79.</p> <p>Macronyssus cyclaspis.— Till &amp; Evans, 1964: 518; Radovsky, 1966: 94; 1967: 125; Beron, 1968: 159; Haitlinger, 1978: 709; Rybin, 1983: 358; Rybin et al., 1989: 425; Estrada-Peña et al., 1989: 350; Villegas-Guzman et al., 2005: 125; Scheffler, 2010; Radovsky, 2010: 56.</p> <p>Type locality: Arnhem (Netherlands) (Oudemans, 1906a).</p> <p>Type host: Pipistrellus pipistrellus (Oudemans, 1906a).</p> <p>Distribution in Russian Federation: [Russia] (Stanyukovich, 1997).</p> <p>Distribution outside Russian Federation: Europe (Radovsky, 1967). Asia (Kyrgyzstan— Rybin 1983). Central America (Mexico— Beron, 2014)</p> <p>Hosts (Old World). Barbastella barbastellus (Radovsky, 1967), Barbastella darjelingensis (Beron, 2014), Myotis myotis (Dusbábek, 1964), M. nattereri (Stanyukovich, 1997), M. brandtii (Stanyukovich, 1997), M. frater (Stanyukovich, 1997), M. daubentoni (Beron, 2014), Eptesicus serotinus (Dusbábek, 1964), Ep. nilssonii (Stanyukovich, 1997), Nyctalus noctula (Beron, 2014), Pipistrellus pipistrellus (Radovsky, 1967), Pi. kuhlii (Kuhl, 1817) (Stanyukovich, 1997), Plecotus auritus (Dusbábek, 1964), Pl. austriacus (Stanyukovich, 1997).</p> <p>Remark. The record of M. cyclaspis on Murina leucogaster (Stanyukovich, 1997) should be attributed to Macronyssus sibiricus (Orlova et al., 2017).</p> </div>	http://treatment.plazi.org/id/E35487F53528FFB30F93FC8DFD83D5D6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Orlova, M. V.;Klimov, P. B.;Orlov, O. L.;Smirnov, D. G.;Zhigalin, A. V.;Budaeva, I. V.;Emelyanova, A. A.;Anisimov, N. V.	Orlova, M. V., Klimov, P. B., Orlov, O. L., Smirnov, D. G., Zhigalin, A. V., Budaeva, I. V., Emelyanova, A. A., Anisimov, N. V. (2021): A checklist of bat-associated macronyssid mites (Acari: Gamasina: Macronyssidae) of Russia, with new host and geographical records. Zootaxa 4974 (3): 537-564, DOI: https://doi.org/10.11646/zootaxa.4974.3.4
E35487F53528FFB20F93F92BFF30D0BD.text	E35487F53528FFB20F93F92BFF30D0BD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macronyssus diversipilis (Vitzthum 1920)	<div><p>Macronyssus diversipilis (Vitzthum, 1920)</p> <p>Liponyssus diversipilis Vitzthum, 1920: 23.</p> <p>Ichoronyssus diversipilis.— Fonseca, 1948: 299.</p> <p>Ichoronyssus blatnensis Dusbábek, 1964: 91 (synonymy by Radovsky, 1967).</p> <p>Macronyssus blatnensis.— Radovsky, 1966: 94.</p> <p>Macronyssus diversipilis.— Radovsky, 1966: 94; 1967: 127; Haitlinger, 1978: 711; Martyn, 1986: 137; Baker &amp; Craven, 2003: 6; Bobkova, 2005: 75; Baker &amp; Beccaloni, 2006: 173; Radovsky, 2010: 56.</p> <p>Type locality: Dockenhuden nr. Hamburg (Germany) (Vitzthum, 1920).</p> <p>Type host: Myotis daubentonii (Vitzthum, 1920).</p> <p>Material. N1 ex Myotis nattereri from State Nature Reserve “Samarskaya Luka”, 25 November 2001, leg. D.G. Smirnov; F, M ex Myotis daubentonii from State Nature Reserve “Samarskaya Luka”, 1-31 November 2005, leg. D.G. Smirnov; F ex M. daubentonii from State Nature Reserve “Samarskaya Luka”, 1-31 July 2006, leg. D.G. Smirnov; F (with intrauterinal egg), N1 ex M. brandtii from biological station “Preduraliye”, 24 July 2011, leg. O.L. Orlov, M. V. Orlova; 2 FF, 3 N1 ex M. daubentonii from Bolshoy Kizil river, 27 August 2014, leg. A. V. Zhigalin; 6 N1 ex M. daubentonii from Maliy Kizil river, 24 August 2014, leg. A. V. Zhigalin; 6 N1 ex M. daubentonii from Yangelka river, 21 August 2014, leg. A. V. Zhigalin; 5 FF, 3 MM, 21 N1 ex ex M. daubentonii from village Elimbetovo, 25 August 2014, leg. A. V. Zhigalin; 3 N1 ex M. nattereri from Serpievskaya cave, 30 October 2012, leg. O.L. Orlov; F, M, 2 N1 ex M. daubentonii from Arakaevskaya cave, 17 January 2010, leg. O.L. Orlov, M. V. Orlova.</p> <p>Distribution in Russian Federation: Russian Federation: Leningrad Province (Stanyukovich, 1990a; Stanyukovich, 1997), Samara Province (present paper), Udmurtia Republic (Orlova et al., 2011), Dagestan Republic (Orlova et al., 2020a), Bashkiria Republic (present paper), Sverdlovsk Region (present paper), Chelyabinsk Region (present paper), [European part of Russia, Pre-Urals, Urals— Orlova et al., 2015d].</p> <p>Distribution outside Russian Federation: Europe (Radovsky, 1967).</p> <p>Hosts. Myotis daubentonii (Radovsky, 1967; Stanyukovich, 1990a; Orlova et al., 2011; Orlova et al., 2020a), M. nattereri (Radovsky, 1967), M. myotis (Dusbábek, 1964), M. blythii (Haitlinger &amp; Walter, 1997), M. bechsteinii (Kuhl, 1818) (Haitlinger &amp; Walter, 1997), M. dasycneme (Haitlinger &amp; Walter, 1997), M. brandtii (Haitlinger &amp; Walter, 1997), Vespertilio murinus (Radovsky, 1967), Plecotus auritus (Radovsky, 1967), Barbastella barbastellus Schreber, 1774 (Beron, 2014), Nyctalus noctula (Beron, 2014), Pipistrellus pipistrellus (Beron, 2014), Pi. nathusii (Beron, 2014), Eptesicus nilssonii (Stanyukovich, 1997), Ep. serotinus (Stanyukovich, 1997), Rhinolophus hipposideros Bechstein, 1800 (Haitlinger &amp; Walter, 1997). Rh. euryale (Beron, 2014), Miniopterus schreibersi (Beron, 2014).</p> </div>	http://treatment.plazi.org/id/E35487F53528FFB20F93F92BFF30D0BD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Orlova, M. V.;Klimov, P. B.;Orlov, O. L.;Smirnov, D. G.;Zhigalin, A. V.;Budaeva, I. V.;Emelyanova, A. A.;Anisimov, N. V.	Orlova, M. V., Klimov, P. B., Orlov, O. L., Smirnov, D. G., Zhigalin, A. V., Budaeva, I. V., Emelyanova, A. A., Anisimov, N. V. (2021): A checklist of bat-associated macronyssid mites (Acari: Gamasina: Macronyssidae) of Russia, with new host and geographical records. Zootaxa 4974 (3): 537-564, DOI: https://doi.org/10.11646/zootaxa.4974.3.4
E35487F53529FFB20F93FBCCFD4DD4ED.text	E35487F53529FFB20F93FBCCFD4DD4ED.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macronyssus ellipticus (Kolenati 1856)	<div><p>Macronyssus ellipticus (Kolenati, 1856)</p> <p>Caris ellipticus Kolenati, 1856: 16.</p> <p>Liponyssus ellipticus.— Hirst, 1922: 795.</p> <p>Ichoronyssus mohrae Vitzthum, 1932: 32 (synonymy by Radovsky, 1966).</p> <p>Ichoronyssus ellipticus.— Pinchuk, 1970: 82.</p> <p>Macronyssus ellipticus.—Fonseca, 1941: 263; Radovsky, 1966: 94; 1967: 128; Beron, 1968: 159; Haitlinger, 1978: 707; Haitlinger &amp; Ruprecht, 1985: 616; Baker &amp; Beccaloni, 2006: 173; Scheffler, 2010; Radovsky, 2010: 56.</p> <p>Type locality: unknown.</p> <p>Type host: Myotis myotis (Kolenati, 1856).</p> <p>Material. F, 4 N1 ex Natterer’s bat Myotis nattereri, from Staritsa, 29 February 2020, leg. O.L. Orlov, A.A. Emelyanova; 9 N1 ex Myotis nattereri, N1 ex Myotis mystacinus, 3 N1 ex Myotis brandtii, from Samara, 25 November 2001, leg. D.G. Smirnov; N1 ex M. brandtii, M ex Plecotus auritus, 5 N1 ex M. daubentonii, 4 N1 ex M. dasycneme, from SNR “Samarskaya Luka”, 1-31 May 2005 leg. D.G. Smirnov.</p> <p>Distribution in Russian Federation: Leningrad Province (Stanyukovich, 1990a; Orlova et al., 2015d), Tver Province (present paper), Samara Province (present paper), Northern Osetiya Republic (Stanyukovich, 1997), Sverdlovsk Province (Orlova, 2011), Chelyabinsk Province (Orlova &amp; Orlov, 2013), Altai Region (Orlova et al., 2017a), Primorskiy Region (Tiunov et al., 2021), [Far East] (Stanyukovich, 1997), [Urals, Altai] (Orlova et al., 2015d).</p> <p>Distribution outside Russian Federation: Europe (Radovsky, 1967), Central Asia (Stanyukovich, 1997).</p> <p>Hosts. Myotis myotis (Radovsky, 1967), M. blythii (Radovsky, 1967), M. brandtii (Orlova, 2011), M. dasycneme (Stanyukovich, 1990a), M. daubentonii (Stanyukovich, 1990a), M. nattereri (Stanyukovich, 1997; present paper), M. brandtii (Stanyukovich, 1990a), M. macrodactylus (present paper), M. mystacinus (Orlova &amp; Orlov, 2013), Vespertilio murinus (Beron, 2014), Plecotus auritus (Orlova &amp; Orlov, 2013), Murina hilgendorfi (as M. leucogaster — Stanyukovich, 1997), Rhinolophus ferrumequinum (Radovsky, 1967), Rh. hipposideros (Radovsky, 1967), Miniopterus schreibersi (Beron, 2014).</p> </div>	http://treatment.plazi.org/id/E35487F53529FFB20F93FBCCFD4DD4ED	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Orlova, M. V.;Klimov, P. B.;Orlov, O. L.;Smirnov, D. G.;Zhigalin, A. V.;Budaeva, I. V.;Emelyanova, A. A.;Anisimov, N. V.	Orlova, M. V., Klimov, P. B., Orlov, O. L., Smirnov, D. G., Zhigalin, A. V., Budaeva, I. V., Emelyanova, A. A., Anisimov, N. V. (2021): A checklist of bat-associated macronyssid mites (Acari: Gamasina: Macronyssidae) of Russia, with new host and geographical records. Zootaxa 4974 (3): 537-564, DOI: https://doi.org/10.11646/zootaxa.4974.3.4
E35487F5352AFFB10F93FF64FDC6D5CD.text	E35487F5352AFFB10F93FF64FDC6D5CD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macronyssus flavus (Kolenati 1856)	<div><p>Macronyssus flavus (Kolenati, 1856)</p> <p>Dermanissus [sic] flavus Kolenati, 1856: 19.</p> <p>? Dermanissus lobatus Kolenati, 1856: 20.</p> <p>Liponyssus lobatus.— Kolenati, 1856: 22; Porta, 1914: 481.</p> <p>Lepronyssus flavus.— Kolenati, 1858: 6; Fonseca, 1948: 306; Turk, 1953: 13; Bogdanov, 1953: 31.</p> <p>Lepronyssus lobatus.— Kolenati, 1858: 6; Turk, 1953: 13.</p> <p>Liponyssus flavus.— Hirst 1922: 791.</p> <p>Ichoronyssus britannicus Fonseca, 1948: 299 (synonymy by Radovsky, 1966).</p> <p>Ichoronyssus flavus.— Arzamasov &amp; Kurskov, 1962: 203; Vshivkov, 1963: 324; Bregetova, 1953: 330; Morozova et al., 1964: 162; Dusbábek, 1964a: 81; 1966: 44; Zemskaya, 1965: 316; Dubovchenko, 1968: 8; Pinchuk, 1970: 73; Mahnert, 1971: 926.</p> <p>Macronyssus flavus.— Evans et al., 1961: 140; Radovsky, 1966: 94; 1967: 131; Juvara, 1967: 187; Beron, 1968: 159; Uchikawa, 1976: 841; Rybin, 1983: 357; Rybin et al., 1989: 425; Baker &amp; Beccaloni, 2006: 173; Haitlinger &amp; Lupicky, 2008: 124; Radovsky, 2010: 56.</p> <p>Type locality: Czech Republic (“Böhmen, Mähren ”) (Kolenati, 1856).</p> <p>Type host: Nyctalus noctula (Kolenati, 1856).</p> <p>Material. N1 ex Nyctalus noctula from Akhshtyrskaya cave, 1-31 August 2001, leg. D.G. Smirnov; 2 MM, N1 ex Nyctalus noctula from NP “Nurgush”, 24 June 2006, leg. D.G. Smirnov; 2 FF, 9 N1 ex N. noctula from Mordovskiy State Natural Reserve, 21 June 2016, leg. D.G. Smirnov; 6 FF, 3 MM, 5 N1 ex N. noctula from Shaytan-Kazak, 19 May 2019, leg. D.G. Smirnov, A.M. Lukyanenko; N1 ex Pipistrellus pipistrellus from neighborhood of Khasavyurt town, 20 May 2019, leg. D.G. Smirnov, A.M. Lukyanenko; 18 FF, 10 N1, N2 ex N. noctula from State Nature Reserve “Samarskaya Luka”, 1-31 June 2006, leg. D.G. Smirnov; 6 FF, M, N2 ex N. leisleri from SNR “Samarskaya Luka”, 1-31 June 2006, leg. D.G. Smirnov; 42 FF, 2 MM, 203 N1, 7 N2 ex N. lasiopterus from SNR “Samarskaya Luka”, 25 July 2006, leg. D.G. Smirnov; 5 FF ex Myotis daubentonii from SNR “Samarskaya Luka”, 1-31 November 2005, leg. D.G. Smirnov; F, M ex Myotis daubentonii from SNR “Samarskaya Luka”, 1-31 July 2006, leg. D.G. Smirnov; 4 MM, 5 N1 ex Eptesicus serotinus from Biological station “Venevitinovo”, 17 June 2019, leg. I. V. Budaeva.</p> <p>Distribution in Russian Federation: Crimea (as Ichoronyssus flavus — Bregetova, 1956; Vshivkov, 1963; Stanyukovich, 1997), Pskov Province (Medvedev et al., 2000), Leningrad Province (Bregetova, 1956; Stanyukovich, 1997), Penza Province (Stanyukovich, 1997), Voronezh Province (present paper), Nizhniy Novgorod Province (Stanyukovich, 1997), Samara Province (Stanyukovich, 1997; present paper), Volgograd Province (Bregetova, 1956; Stanyukovich, 1997), Rostov Province (Zabashta et al., 2019), Krasnodar Region (present paper), Dagestan Republic (present paper), Kirov Province (present paper), Udmurtia Republic (Orlova et al., 2011), Khabarovsk Region (Bregetova, 1956; as Far East— Stanyukovich, 1997).</p> <p>Distribution outside Russian Federation: Europe (Radovsky, 1967). Asia (Stanyukovich, 1997).</p> <p>Hosts. Nyctalus noctula (Vshivkov, 1963; Stanyukovich, 1997; Medvedev et al., 2000; present paper), N. leisleri (Kuhl, 1817) (Stanyukovich, 1997; present paper), N. lasiopterus (Schreber, 1780) (Stanyukovich, 1997; present paper), Myotis daubentonii (Stanyukovich, 1997), M. myotis (Stanyukovich, 1997), M. blythii (Stanyukovich, 1997), M. mystacinus (Stanyukovich, 1997), M. nattereri (Stanyukovich, 1997), Pipistrellus pipistrellus (Vshivkov, 1963; Stanyukovich, 1997; present paper), P. nathusii (Stanyukovich, 1997; Medvedev et al., 2000), Vespertilio murinus (Stanyukovich, 1997), Eptesicus nilssonii (Stanyukovich, 1997), Ep. serotinus (Beron, 2014), Eptesicus bottae ognevi Bobrinskoy, 1918 (Beron, 2014), Barbastella barbastellus (Dusbábek, 1964), Rhinolophus ferrumequinum (Vshivkov, 1963).</p> </div>	http://treatment.plazi.org/id/E35487F5352AFFB10F93FF64FDC6D5CD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Orlova, M. V.;Klimov, P. B.;Orlov, O. L.;Smirnov, D. G.;Zhigalin, A. V.;Budaeva, I. V.;Emelyanova, A. A.;Anisimov, N. V.	Orlova, M. V., Klimov, P. B., Orlov, O. L., Smirnov, D. G., Zhigalin, A. V., Budaeva, I. V., Emelyanova, A. A., Anisimov, N. V. (2021): A checklist of bat-associated macronyssid mites (Acari: Gamasina: Macronyssidae) of Russia, with new host and geographical records. Zootaxa 4974 (3): 537-564, DOI: https://doi.org/10.11646/zootaxa.4974.3.4
E35487F5352AFFB00F93F91DFE6FD724.text	E35487F5352AFFB00F93F91DFE6FD724.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macronyssus granulosus (Kolenati 1856)	<div><p>Macronyssus granulosus (Kolenati, 1856)</p> <p>Dermanissus [sic] granulosus Kolenati, 1856: 20.</p> <p>Dermanissus [sic] glutinosus Kolenati, 1856: 20.</p> <p>Lepronyssus granulosus.— Kolenati, 1858: 6; Fonseca, 1948: 307.</p> <p>Lepronyssus glutinosus.— Kolenati, 1858: 6.</p> <p>Lepronyssus leprosus Kolenati, 1858: 5 (synonymy by Radovsky, 1966).</p> <p>Liponyssus granulosus.— Hirst, 1921: 794. Hirstesia transvaalensis Zumpt, 1950: 89 (synonymy by Radovsky, 1966).</p> <p>Hirstesia transvaalensis.— Keegan, 1956: 213.</p> <p>Bdellonyssus pollerae Lombardini, 1957: 284 (synonymy by Radovsky, 1966).</p> <p>Ichoronyssus pollerae.— Wen, 1975: 351.</p> <p>Ichoronyssus leprosus.—Strandtmann &amp; Warton, 1958: 91.</p> <p>Ichoronyssus granulosus.— Vshivkov, 1963: 324; Costa, 1967: 107; Dubovchenko, 1968: 8; Pinchuk, 1970: 73.</p> <p>Macronyssus granulosus.— Radovsky, 1966: 94; 1967: 132; Beron, 1968: 159; 1974: 65; Uchikawa, 1976: 841; Ogadzhanyan &amp; Arutyunyan, 1974: 79; Cicolani &amp; Manilla, 1980: 35; Rybin, 1983: 356; Rybin et al., 1989: 425; Whitaker et al., 1990: 54; Estrada-Peña &amp; Serra-Cobo, 1991: 345; Lourenço &amp; Palmeirim, 2007: 163; Radovsky, 2010: 56.</p> <p>Type locality: Banat (Romania) (Kolenati, 1856).</p> <p>Type host: Miniopterus schreibersi (Kolenati, 1856).</p> <p>Distribution in Russian Federation: Crimea (as Ichoronyssus granulosus — Vshivkov, 1963; Stanyukovich, 1997; Orlova &amp; Orlov, 2018), Voronezh Province (Stanyukovich, 1997), Krasnoyarsk Region (Stanyukovich, 1997), Altai Republic (Orlova &amp; Orlov, 2015; Orlova et al., 2015c), Primorskiy Region (Medvedev et al., 1991; Tiunov et al., 2021), Sakhalin Province (Kunashir island) (Orlova &amp; Zhigalin, 2015a; Orlova et al., 2015b), [Far East] (Stanyukovich, 1997).</p> <p>Distribution outside Russian Federation: Eurasia (Radovsky, 1967); Africa (Radovsky, 1967).</p> <p>Hosts. Myotis blythii (Vshivkov, 1963; Stanyukovich, 1997; Orlova &amp; Orlov, 2018), M. myotis (Radovsky, 1967), M. macrodactylus (Stanyukovich, 1997; present paper), M. petax (as M. daubentonii — Medvedev et al., 1991), M. sibiricus (as M. brandtii — Medvedev et al., 1991) M. nattereri (Stanyukovich, 1997), M. dasycneme (Stanyukovich, 1997), M. daubentonii (Stanyukovich, 1997), M. brandtii (Stanyukovich, 1997), M. mystacinus (Stanyukovich, 1997), M. emarginatus E. Geoffroy, 1806 (Vshivkov, 1963), M. cappaccinii (Stanyukovich, 1997), M. tricolor (Radovsky, 1967), Barbastella barbastellus (Stanyukovich, 1997), Pipistrellus javanicus (Beron, 2014), Plecotus auritus (Stanyukovich, 1997), Murina leucogaster (Stanyukovich, 1997), Nyctalus leisleri (Stanyukovich, 1997), Miniopterus fuliginosus (as M. schreibersi — Medvedev et al., 1991), Mi. schreibersi (Dusbábek, 1964), Mi. fraterculus Thomas &amp; Schwann, 1906 (Radovsky, 1967), Rhinolophus clivosus Cretzschmar, 1828 (Radovsky, 1967), Rh. euryale (Radovsky, 1967), Rosettus lanosus Thomas, 1906 (Radovsky, 1967). Ro. aegyptiacus (E. Geoffroy, 1810) (Beron, 2014).</p> </div>	http://treatment.plazi.org/id/E35487F5352AFFB00F93F91DFE6FD724	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Orlova, M. V.;Klimov, P. B.;Orlov, O. L.;Smirnov, D. G.;Zhigalin, A. V.;Budaeva, I. V.;Emelyanova, A. A.;Anisimov, N. V.	Orlova, M. V., Klimov, P. B., Orlov, O. L., Smirnov, D. G., Zhigalin, A. V., Budaeva, I. V., Emelyanova, A. A., Anisimov, N. V. (2021): A checklist of bat-associated macronyssid mites (Acari: Gamasina: Macronyssidae) of Russia, with new host and geographical records. Zootaxa 4974 (3): 537-564, DOI: https://doi.org/10.11646/zootaxa.4974.3.4
E35487F5352BFFB00F93FB35FB5AD40E.text	E35487F5352BFFB00F93FB35FB5AD40E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macronyssus heteromorphus Dusbabek & Radovsky 1972	<div><p>Macronyssus heteromorphus Dusbábek &amp; Radovsky, 1972</p> <p>Macronyssus heteromorphus Dusbábek &amp; Radovsky, 1972: 134.</p> <p>Macronyssus heteromorphus.— Uchikawa, 1979b: 255; Medvedev et al., 1991; Radovsky, 2010: 56.</p> <p>Type locality: Shikotan Island (Kuril Islands) (Dusbábek &amp; Radovsky, 1972).</p> <p>Type host: [Rattus norvegicus (Berkenhout, 1769)] (Dusbábek &amp; Radovsky, 1972).</p> <p>Material. F and M ex Myotis davidii from Tes-Khem river, 23 July 2016, leg. A. V. Zhigalin.</p> <p>Distribution in Russian Federation: Russian Federation: Khanty-Mansi Autonomous Region (Orlova et al., 2019), Altai Region (Orlova et al., 2017b), Novosibirsk Province (Orlova et al., 2017a), Khakassia Republic (Orlova et al., 2017a), Tuva Republic (Orlova et al., 2015a, 2015d; present paper), Krasnoyarsk Region (Stanyukovich, 1997; Orlova et al., 2017a), Kamchatka Region (Medvedev et al., 1991), Primorskiy Region (Medvedev et al., 1991), Sakhalin Province (Shikotan Island) (Dusbábek &amp; Radovsky, 1972), [Far East] (Stanyukovich, 1997).</p> <p>Distribution outside Russian Federation: Unknown.</p> <p>Hosts. [Rattus norvegicus] (Dusbábek &amp; Radovsky, 1972), Myotis petax (as M. daubentonii — Stanyukovich, 1997), M. sibiricus (as Myotis brandtii and M. mystacinus — Stanyukovich, 1997), M. brandtii (Orlova et al., 2019), M. ikonnikovi (Beron, 2014), Myotis davidii (present paper), Eptesicus nilssonii (Medvedev et al., 1991; Stanyukovich, 1997), Plecotus ognevi (as Plecotus auritus — Stanyukovich, 1997), Murina hilgendorfi (as Murina leucogaster — Medvedev et al., 1991; Stanyukovich, 1997), Miniopterus schreibersi (Beron, 2014).</p> </div>	http://treatment.plazi.org/id/E35487F5352BFFB00F93FB35FB5AD40E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Orlova, M. V.;Klimov, P. B.;Orlov, O. L.;Smirnov, D. G.;Zhigalin, A. V.;Budaeva, I. V.;Emelyanova, A. A.;Anisimov, N. V.	Orlova, M. V., Klimov, P. B., Orlov, O. L., Smirnov, D. G., Zhigalin, A. V., Budaeva, I. V., Emelyanova, A. A., Anisimov, N. V. (2021): A checklist of bat-associated macronyssid mites (Acari: Gamasina: Macronyssidae) of Russia, with new host and geographical records. Zootaxa 4974 (3): 537-564, DOI: https://doi.org/10.11646/zootaxa.4974.3.4
E35487F5352CFFB70F93FF64FACAD03C.text	E35487F5352CFFB70F93FF64FACAD03C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macronyssus hosonoi Uchikawa 1979	<div><p>Macronyssus hosonoi Uchikawa, 1979</p> <p>Macronyssus sp. 5 Uchikawa, 1979a: 35.</p> <p>Macronyssus hosonoi Uchikawa, 1979b: 252.</p> <p>Macronyssus hosonoi.— Radovsky, 2010: 56.</p> <p>Macronyssus evansi Stanyukovich, 1990b: 71 (synonymy by Stanyukovich, 1997).</p> <p>Type locality: Hokujo Village, Nagano Pref (Japan) (Uchikawa, 1979b).</p> <p>Type host: Myotis sibiricus (as Myotis hosonoi) (Uchikawa, 1979b).</p> <p>Material. 3 FF (all with intrauterine larvae), M ex Myotis davidii from Bayan settlement (6 July 2016), leg. D. V. Kazakov; 5 FF (including 4 with intrauterine larvae), M, N1 ex Myotis ikonnikovi from park “Tunkinskiy” (6 August 2016), leg. D. V. Kazakov; 5 MM, 4 N1 ex Murina hilgendorfi from park “Tunkinskiy” (8 August 2016), leg. D. V. Kazakov.</p> <p>Distribution in Russian Federation: Altai Region (Orlova et al., 2014), Khakassia Republic (Orlova et al., 2017), Krasnoyarsk Region (as Macronyssus evansi — Stanyukovich, 1990b, 1997), Irkutsk Region (Orlova et al., 2017), Buryatiya Republic (Orlova et al., 2017; present paper), Kamchatka Territory (Stanyukovich, 1997), Sakhalin Province (Kunashir island) (Orlova et al., 2015d), [Far East] (as Macronyssus evansi — Stanyukovich, 1997).</p> <p>Distribution outside Russian Federation: Japan (Uchikawa, 1979); Mongolia (Scheffler et al., 2012).</p> <p>Hosts. Myotis sibiricus (as Myotis brandtii, M. hosonoi and M. mystacinus — Stanyukovich, 1997; as M. mystacinus — Uchikawa, 1979), M. ikonnikovi (Stanyukovich, 1997; present paper), M. davidii (present paper), Barbastella darjelingensis (Stanyukovich, 1997), Plecotus ognevi (as Plecotus auritus — Stanyukovich, 1997).</p> </div>	http://treatment.plazi.org/id/E35487F5352CFFB70F93FF64FACAD03C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Orlova, M. V.;Klimov, P. B.;Orlov, O. L.;Smirnov, D. G.;Zhigalin, A. V.;Budaeva, I. V.;Emelyanova, A. A.;Anisimov, N. V.	Orlova, M. V., Klimov, P. B., Orlov, O. L., Smirnov, D. G., Zhigalin, A. V., Budaeva, I. V., Emelyanova, A. A., Anisimov, N. V. (2021): A checklist of bat-associated macronyssid mites (Acari: Gamasina: Macronyssidae) of Russia, with new host and geographical records. Zootaxa 4974 (3): 537-564, DOI: https://doi.org/10.11646/zootaxa.4974.3.4
E35487F5352CFFB70F93FC4DFA90D443.text	E35487F5352CFFB70F93FC4DFA90D443.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macronyssus kolenatii (Oudemans 1902)	<div><p>Macronyssus kolenatii (Oudemans, 1902)</p> <p>Liponyssus kolenatii Oudemans, 1902: 48.</p> <p>Liponyssus kolenatii.—Oudemans, 1904: 25.</p> <p>Hirstesia britteni Radford, 1953: 239 (synonymy by Radovsky, 1966).</p> <p>Macronyssus britteni.— Evans et al., 1961: 140.</p> <p>Ichoronyssus kolenatii.— Pinchuk, 1970: 85.</p> <p>Macronyssus kolenatii.— Radovsky, 1966: 94; 1967: 136; 2010: 56; Mehl, 1979: 3; Baker &amp; Beccaloni, 2006: 173. Macronyssus kolenati.— Nikitchenko, 1995: 88.</p> <p>Type locality: Osterholz near Bremen (Germany) (Oudemans, 1902).</p> <p>Type host: Pipistrellus pipistrellus (Oudemans, 1902).</p> <p>Material. N1 ex soprano pipistrelle Pipistrellus pygmaeus from biological station “Venevitinovo”, 4 June 2019, leg. O.L. Orlov, I. V. Budaeva.</p> <p>Distribution in Russian Federation: Crimea (Stanyukovich, 1997), Pskov Province (Medvedev et al., 2000), Penza Region (Stanyukovich, 1997), Voronezh Region (Stanyukovich, 1997; present paper), Volgograd Region (Stanyukovich, 1997), Rostov Region (Zabashta et al., 2019), Sverdlovsk Region (Stanyukovich, 1997), [Pre-Urals, Urals— Orlova et al., 2015d].</p> <p>Distribution outside Russian Federation: Europe (Radovsky, 1967; Haitlinger &amp; Walter, 1997), Africa (Egypt— Radovsky, 1967).</p> <p>Hosts. Pipistrellus pipistrellus (Stanyukovich, 1997), P. nathusii (Stanyukovich, 1997), P. pygmaeus (present paper), P. kuhlii (Stanyukovich, 1997; Zabashta et al., 2019), Myotis dasycneme (Stanyukovich, 1997), M. daubentonii (Beron, 2014), M. blythii (Beron, 2014), M. brandtii (Stanyukovich, 1997), M. mystacinus (Stanyukovich, 1997), M. bechsteini (Haitlinger &amp; Walter, 1997), Nyctalus noctula (Haitlinger &amp; Walter, 1997), Pipistrellus maderensis (Beron, 2014), Vespertilio murinus (Stanyukovich, 1997), Eptesicus nilssonii (Stanyukovich, 1997).</p> </div>	http://treatment.plazi.org/id/E35487F5352CFFB70F93FC4DFA90D443	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Orlova, M. V.;Klimov, P. B.;Orlov, O. L.;Smirnov, D. G.;Zhigalin, A. V.;Budaeva, I. V.;Emelyanova, A. A.;Anisimov, N. V.	Orlova, M. V., Klimov, P. B., Orlov, O. L., Smirnov, D. G., Zhigalin, A. V., Budaeva, I. V., Emelyanova, A. A., Anisimov, N. V. (2021): A checklist of bat-associated macronyssid mites (Acari: Gamasina: Macronyssidae) of Russia, with new host and geographical records. Zootaxa 4974 (3): 537-564, DOI: https://doi.org/10.11646/zootaxa.4974.3.4
E35487F5352CFFB60F93F896FC5AD211.text	E35487F5352CFFB60F93F896FC5AD211.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macronyssus leislerianus Fain 2003	<div><p>Macronyssus leislerianus Fain et al., 2003</p> <p>Macronyssus leislerianus Fain, Walter &amp; Heddergott, 2003: 57.</p> <p>Macronyssus leislerianus.— Radovsky, 2010: 56.</p> <p>Type locality: Leinefelde, Breites Holz (Germany) (Fain et al., 2003).</p> <p>Type host: Nyctalus leisleri (Fain et al., 2003).</p> <p>Distribution in Russian Federation: Dagestan Republic (Orlova et al., 2020a).</p> <p>Distribution outside Russian Federation: Europe (Germany) (Fain et al., 2003).</p> <p>Host. Nyctalus leisleri (Fain et al., 2003; Orlova et al., 2020a).</p> </div>	http://treatment.plazi.org/id/E35487F5352CFFB60F93F896FC5AD211	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Orlova, M. V.;Klimov, P. B.;Orlov, O. L.;Smirnov, D. G.;Zhigalin, A. V.;Budaeva, I. V.;Emelyanova, A. A.;Anisimov, N. V.	Orlova, M. V., Klimov, P. B., Orlov, O. L., Smirnov, D. G., Zhigalin, A. V., Budaeva, I. V., Emelyanova, A. A., Anisimov, N. V. (2021): A checklist of bat-associated macronyssid mites (Acari: Gamasina: Macronyssidae) of Russia, with new host and geographical records. Zootaxa 4974 (3): 537-564, DOI: https://doi.org/10.11646/zootaxa.4974.3.4
E35487F5352DFFB60F93FE60FAC3D0EF.text	E35487F5352DFFB60F93FE60FAC3D0EF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macronyssus sibiricus Orlova & Zhigalin 2015	<div><p>Macronyssus sibiricus Orlova &amp; Zhigalin, 2015</p> <p>Macronyssus sibiricus Orlova &amp; Zhigalin, 2015b: 314.</p> <p>Macronyssus sibiricus.— Orlova et al., 2015d: 75; Orlova et al., 2017a: 359.</p> <p>Type locality: Barsukovskaya cave (Novosibirsk province, Russia) (Orlova &amp; Zhigalin, 2015b).</p> <p>Type host: Myotis petax (Orlova &amp; Zhigalin, 2015b).</p> <p>Material. 9 FF, M ex Myotis petax from Barsukovskaya cave, 3 January 2014, leg. O.L.Orlov; N1 ex Murina hilgendorfi from Barsukovskaya cave, 26 March 2014, leg. O.L.Orlov; F (with intrauterine egg) ex Plecotus ognevi from Tigirekskiy State Natural Reserve, 2 March 2016, leg. O.L. Orlov.</p> <p>Distribution in Russian Federation: Altai Region (present paper), Novosibirsk Province (Orlova &amp; Zhigalin, 2015b; Orlova et al., 2015d), Khakassia Republic (Orlova et al., 2017a), Krasnoyarsk Region (Orlova et al., 2017a), Irkutsk Province (Orlova et al., 2017a), Buryatya Republic (Orlova et al., 2017a).</p> <p>Distribution outside Russian Federation: Unknown.</p> <p>Hosts. Myotis sibiricus (Orlova &amp; Kazakov, 2017; Orlova et al., 2017a), M. dasycneme (Orlova &amp; Zhigalin, 2015b), M. petax (present paper), Murina hilgendorfi (present paper), Plecotus ognevi (present paper).</p> </div>	http://treatment.plazi.org/id/E35487F5352DFFB60F93FE60FAC3D0EF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Orlova, M. V.;Klimov, P. B.;Orlov, O. L.;Smirnov, D. G.;Zhigalin, A. V.;Budaeva, I. V.;Emelyanova, A. A.;Anisimov, N. V.	Orlova, M. V., Klimov, P. B., Orlov, O. L., Smirnov, D. G., Zhigalin, A. V., Budaeva, I. V., Emelyanova, A. A., Anisimov, N. V. (2021): A checklist of bat-associated macronyssid mites (Acari: Gamasina: Macronyssidae) of Russia, with new host and geographical records. Zootaxa 4974 (3): 537-564, DOI: https://doi.org/10.11646/zootaxa.4974.3.4
E35487F5352DFFB60F93FBE3FCECD444.text	E35487F5352DFFB60F93FBE3FCECD444.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macronyssus stanyukovichi Orlova & Zhigalin 2015	<div><p>Macronyssus stanyukovichi Orlova &amp; Zhigalin, 2015</p> <p>Macronyssus stanyukovichi Orlova &amp; Zhigalin, 2015b: 315.</p> <p>Macronyssus stanyukovichi.— Orlova et al., 2015d: 76; Orlova &amp; Kazakov, 2017: 165; Orlova et al., 2017a: 359.</p> <p>Type locality: Strashnaya cave (Altai Region, Russia) (Orlova &amp; Zhigalin, 2015b).</p> <p>Type host: Murina hilgendorfi (Orlova &amp; Zhigalin, 2015b).</p> <p>Material. M, 3 N1 ex Murina hilgendorfi from Arkheologicheskaya cave, 13 June 2016, leg. L.B. Kravchenko; F ex Mu. hilgendorfi from Argaley-3 cave, 18 September 2015, leg. D. V. Kazakov; 11 N1 ex Mu. hilgendorfi from Okhotnichiya cave, 4 September 2015, leg. D. V. Kazakov; 4 N1 ex Mu. hilgendorfi from Okhotnichiya cave, 12 September 2016, leg. D. V. Kazakov; 34 FF (including 3 with intrauterine larvae); 9 N1 ex Mu. hilgendorfi from Dolganskaya Yama cave, 18 August 2016, leg. D. V. Kazakov; 3 FF, 2 MM, 11 N1 ex Mu. hilgendorfi from Dolganskaya Yama cave, 20 August 2016, leg. D. V. Kazakov; 33 FF, 11 MM, 721 N1 ex Mu. hilgendorfi from Dolganskaya Yama cave, 11 September 2017, leg. D. V. Kazakov; 8 FF, 4 MM, 71 N1 ex Mu. hilgendorfi from Dolganskaya Yama cave, 12 September 2017, leg. D. V. Kazakov; 7 FF, 3 MM, 133 N1 ex Mu. hilgendorfi from Dolganskaya Yama cave, 13 September 2017, leg. D. V. Kazakov.</p> <p>Distribution in Russian Federation: Altai Region (Orlova &amp; Zhigalin, 2015b; Orlova et al., 2017a), Novosibirsk Province (Orlova &amp; Zhigalin, 2015b; Orlova et al., 2015d), Krasnoyarsk Region (Orlova &amp; Kazakov, 2017), Irkutsk Province (present paper), Buryatya Republic (present paper), Primorskiy Region (Tiunov et al., 2021).</p> <p>Distribution outside Russian Federation: Unknown.</p> <p>Hosts. Murina hilgendorfi (Orlova &amp; Zhigalin, 2015b; Orlova et al., 2015d; Orlova &amp; Kazakov, 2017), Myotis sibiricus (Orlova et al., 2017), M. bombinus (present paper).</p> </div>	http://treatment.plazi.org/id/E35487F5352DFFB60F93FBE3FCECD444	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Orlova, M. V.;Klimov, P. B.;Orlov, O. L.;Smirnov, D. G.;Zhigalin, A. V.;Budaeva, I. V.;Emelyanova, A. A.;Anisimov, N. V.	Orlova, M. V., Klimov, P. B., Orlov, O. L., Smirnov, D. G., Zhigalin, A. V., Budaeva, I. V., Emelyanova, A. A., Anisimov, N. V. (2021): A checklist of bat-associated macronyssid mites (Acari: Gamasina: Macronyssidae) of Russia, with new host and geographical records. Zootaxa 4974 (3): 537-564, DOI: https://doi.org/10.11646/zootaxa.4974.3.4
E35487F5352DFFB50F93F895FCE4D2A5.text	E35487F5352DFFB50F93F895FCE4D2A5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macronyssus tigirecus Orlova & Zhigalin 2015	<div><p>Macronyssus tigirecus Orlova &amp; Zhigalin, 2015</p> <p>Macronyssus tigirecus Orlova &amp; Zhigalin, 2015b: 318.</p> <p>Macronyssus tigirecus.— Orlova et al., 2015d: 76.</p> <p>Type locality: Yashchur cave (Altai Region, Russia) (Orlova &amp; Zhigalin, 2015b).</p> <p>Type host: Murina hilgendorfi (Orlova &amp; Zhigalin, 2015b).</p> <p>Material. 12 FF, 9 MM, 25 N1 ex Murina hilgendorfi from Okhotnichiya cave, 4 September 2015, leg. D. V. Kazakov; 12 FF, 3 MM, 59 N1 ex Murina hilgendorfi from Okhotnichiya cave, 12 September 2016, leg. D. V. Kazakov; 5 FF, 2 MM, 13 N1 ex Murina hilgendorfi from Okhotnichiya cave, 12 September 2016, leg. D. V. Kazakov; F, 3 MM, 7 N1 ex Murina hilgendorfi from Argaley-3 cave, 18 September 2015, leg. D. V. Kazakov.</p> <p>Distribution in Russian Federation: Altai Region (Orlova &amp; Zhigalin, 2015b; Orlova et al., 2015d), Irkutsk Province (present paper), Buryatya Republic (present paper).</p> <p>Distribution outside Russian Federation: Unknown.</p> <p>Host. Murina hilgendorfi (Orlova &amp; Zhigalin, 2015b).</p> </div>	http://treatment.plazi.org/id/E35487F5352DFFB50F93F895FCE4D2A5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Orlova, M. V.;Klimov, P. B.;Orlov, O. L.;Smirnov, D. G.;Zhigalin, A. V.;Budaeva, I. V.;Emelyanova, A. A.;Anisimov, N. V.	Orlova, M. V., Klimov, P. B., Orlov, O. L., Smirnov, D. G., Zhigalin, A. V., Budaeva, I. V., Emelyanova, A. A., Anisimov, N. V. (2021): A checklist of bat-associated macronyssid mites (Acari: Gamasina: Macronyssidae) of Russia, with new host and geographical records. Zootaxa 4974 (3): 537-564, DOI: https://doi.org/10.11646/zootaxa.4974.3.4
E35487F5352EFFB50F93FC7CFC9FD7B2.text	E35487F5352EFFB50F93FC7CFC9FD7B2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Steatonyssus aglaiae Stanjukovich 1991	<div><p>Steatonyssus aglaiae Stanjukovich, 1991</p> <p>Steatonyssus aglaiae Stanjukovich, 1991: 270.</p> <p>Steatonyssus aglaiae.— Radovsky, 2010: 101.</p> <p>Type locality: Karadagskiy Reserve (Crimea).</p> <p>Type host: Rhinolophus ferrumequinum (Stanyukovich, 1991).</p> <p>Distribution in Russian Federation: Crimea (Stanyukovich, 1991).</p> <p>Distribution outside Russian Federation: Unknown.</p> <p>Host. Rhinolophus ferrumequinum (Stanyukovich, 1991).</p> </div>	http://treatment.plazi.org/id/E35487F5352EFFB50F93FC7CFC9FD7B2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Orlova, M. V.;Klimov, P. B.;Orlov, O. L.;Smirnov, D. G.;Zhigalin, A. V.;Budaeva, I. V.;Emelyanova, A. A.;Anisimov, N. V.	Orlova, M. V., Klimov, P. B., Orlov, O. L., Smirnov, D. G., Zhigalin, A. V., Budaeva, I. V., Emelyanova, A. A., Anisimov, N. V. (2021): A checklist of bat-associated macronyssid mites (Acari: Gamasina: Macronyssidae) of Russia, with new host and geographical records. Zootaxa 4974 (3): 537-564, DOI: https://doi.org/10.11646/zootaxa.4974.3.4
E35487F5352EFFB50F93FDB5FD0BD06D.text	E35487F5352EFFB50F93FDB5FD0BD06D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Steatonyssus Kolenati 1858	<div><p>Genus Steatonyssus Kolenati, 1858</p> <p>Steatonyssus Kolenati, 1858: 6.</p> <p>Steatonyssus.— Radovsky, 2010: 96.</p> <p>Type species: Steatonyssus periblepharus Kolenati, 1858.</p> <p>Ceratonyssus Ewing 1923: 6.</p> <p>Type species: Ceratonyssus ceratognathus Ewing, 1923.</p> </div>	http://treatment.plazi.org/id/E35487F5352EFFB50F93FDB5FD0BD06D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Orlova, M. V.;Klimov, P. B.;Orlov, O. L.;Smirnov, D. G.;Zhigalin, A. V.;Budaeva, I. V.;Emelyanova, A. A.;Anisimov, N. V.	Orlova, M. V., Klimov, P. B., Orlov, O. L., Smirnov, D. G., Zhigalin, A. V., Budaeva, I. V., Emelyanova, A. A., Anisimov, N. V. (2021): A checklist of bat-associated macronyssid mites (Acari: Gamasina: Macronyssidae) of Russia, with new host and geographical records. Zootaxa 4974 (3): 537-564, DOI: https://doi.org/10.11646/zootaxa.4974.3.4
E35487F5352EFFB50F93FAC7FE98D4F8.text	E35487F5352EFFB50F93FAC7FE98D4F8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Steatonyssus noctulus Rybin 1992	<div><p>Steatonyssus noctulus Rybin, 1992</p> <p>Steatonyssus noctulus Rybin, 1992: 157.</p> <p>Steatonyssus noctulus.—Rybin, 1995: 47; Rupp &amp; Ludwig, 2000: 275; Baker &amp; Craven, 2003: 6; Radovsky, 2010: 101.</p> <p>Type locality: Osh, Oshskaya province (Kirghizstan).</p> <p>Type host: Nyctalus noctula.</p> <p>Material. 2 FF, 21 N1 ex Eptesicus serotinus from Biological station “Venevitinovo”, 17 June 2019, leg. I. V. Budaeva; M ex Nyctalus lasiopterus from State Nature Reserve “Samarskaya Luka”, 7 July 2016, leg. D.G. Smirnov; N1 ex Nyctalus leisleri from SNR “Samarskaya Luka”, 8 July 2016, leg. D.G. Smirnov.</p> <p>Distribution in Russian Federation: Crimea (Stanyukovich, 1997), Pskov Province (Medvedev et al., 2000), Penza Province (Stanyukovich, 1997), Voronezh Province (present paper), Samara Province (present paper), Rostov Province (Zabashta et al., 2019), Dagestan Republic (Orlova et al., 2020a), [Russia] (Orlova et al., 2015d).</p> <p>Distribution outside Russian Federation: Europe (Stanyukovich, 1997; Rupp &amp; Ludwig 2000). Asia (Rybin 1992; Stanyukovich, 1997).</p> <p>Hosts. Nyctalus noctula (Stanyukovich, 1997; Medvedev et al., 2000; Zabashta et al., 2019), Ny. lasiopterus (present paper), Ny. leisleri (present paper), Miniopterus schreibersi (Stanyukovich, 1997); Eptesicus serotinus (present paper).</p> </div>	http://treatment.plazi.org/id/E35487F5352EFFB50F93FAC7FE98D4F8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Orlova, M. V.;Klimov, P. B.;Orlov, O. L.;Smirnov, D. G.;Zhigalin, A. V.;Budaeva, I. V.;Emelyanova, A. A.;Anisimov, N. V.	Orlova, M. V., Klimov, P. B., Orlov, O. L., Smirnov, D. G., Zhigalin, A. V., Budaeva, I. V., Emelyanova, A. A., Anisimov, N. V. (2021): A checklist of bat-associated macronyssid mites (Acari: Gamasina: Macronyssidae) of Russia, with new host and geographical records. Zootaxa 4974 (3): 537-564, DOI: https://doi.org/10.11646/zootaxa.4974.3.4
E35487F5352FFFB40F93FF64FD38D031.text	E35487F5352FFFB40F93FF64FD38D031.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Steatonyssus occidentalis (Ewing 1933)	<div><p>Steatonyssus occidentalis (Ewing, 1933)</p> <p>Ceratonyssus occidentalis Ewing, 1933: 10.</p> <p>Steatonyssus occidentalis.— Radford, 1942: 65; Fonseca, 1948: 319; Radovsky &amp; Furman, 1963: 272; Yunker &amp; Radovsky, 1966: 96; Miller et al., 1973: 609; Whitaker &amp; Easterla, 1975: 246; Saunders, 1975: 88; Rybin, 1983: 359; Whitaker &amp; Maser, 1985: 73; Rybin et al., 1989: 425; Durden et al., 1992: 507; Ritzi et al., 2001: tabl.; Rupp et al., 2004: 189.</p> <p>Steatonyssus (S.) occidentalis.— Till &amp; Evans, 1964: 559; Radovsky, 1966: 94; 1967: 206; 2010: 101; Pereira Lorenzo &amp; Quinteiro Alonso, 1993: 19.</p> <p>Type locality: Adel, Oregon (USA) (Ewing, 1933).</p> <p>Type host: “Common brown bat” (Ewing, 1933).</p> <p>Material. 14 FF (including 2 with intrauterine egg), 2 MM, 24 N1 ex Eptesicus serotinus from biological station “Venevitinovo”, 20 June 2019, leg. I. V. Budaeva.</p> <p>Distribution in Russian Federation: Voronezh Province (present paper), Volgograd Province (Stanyukovich, 1997), Stavropol Region (Stanyukovich, 1997).</p> <p>Distribution outside Russian Federation: Europe (Micherdzinski, 1980). Asia (Rybin 1983; Stanyukovich, 1997).</p> <p>Hosts. Eptesicus serotinus (Stanyukovich, 1997; present paper), Myotis mystacinus (Stanyukovich, 1997), M. daubentonii (Haitlinger &amp; Walter, 1997), M. blythii (Stanyukovich, 1997), Pipistrellus pipistrellus (as Vespertilio pipistrellus — Rybin, 1983), Plecotus auritus (Haitlinger &amp; Walter, 1997), Pl. austriacus (Haitlinger &amp; Walter, 1997), Nyctalus noctula (Haitlinger &amp; Walter, 1997).</p> </div>	http://treatment.plazi.org/id/E35487F5352FFFB40F93FF64FD38D031	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Orlova, M. V.;Klimov, P. B.;Orlov, O. L.;Smirnov, D. G.;Zhigalin, A. V.;Budaeva, I. V.;Emelyanova, A. A.;Anisimov, N. V.	Orlova, M. V., Klimov, P. B., Orlov, O. L., Smirnov, D. G., Zhigalin, A. V., Budaeva, I. V., Emelyanova, A. A., Anisimov, N. V. (2021): A checklist of bat-associated macronyssid mites (Acari: Gamasina: Macronyssidae) of Russia, with new host and geographical records. Zootaxa 4974 (3): 537-564, DOI: https://doi.org/10.11646/zootaxa.4974.3.4
E35487F5352FFFAB0F93FC40FEEAD1E0.text	E35487F5352FFFAB0F93FC40FEEAD1E0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Steatonyssus periblepharus Kolenati 1858	<div><p>Steatonyssus periblepharus Kolenati, 1858</p> <p>Steatonyssus periblepharus Kolenati, 1858: 6.</p> <p>Steatonyssus periblepharus.—Kolenati, 1859: 186; Oudemans, 1936: 288; Willmann, 1936: 153; Strandtmann &amp; Wharton, 1958: 113; Pinchuk, 1971: 114; Beron, 1968: 160; 1974: 66; Haitlinger &amp; Ruprecht, 1977: 333; Haitlinger, 1978: 714; Mehl, 1979: 33; Teng, 1980; Rybin, 1983: 358; Rybin et al., 1989: 425; Abul-hab &amp; Shihab, 1990; Stanjukovich, 1990: 197, 1997: 35; Cicek et al., 2007: 248.</p> <p>Liponyssus chiropteralis Hirst, 1922: 776 (synonymy by Radovsky, 1966).</p> <p>Steatonyssus murinus.— Turk, 1953: 13; Keegan, 1956: 221; Dusbábek, 1964b: 15; Turk &amp; Turk, 1952: 484; Willmann, 1955: 183; Costa, 1967: 107; Nikitchenko, 1995: 88.</p> <p>Steatonyssus (S.) periblepharus.— Till &amp; Evans, 1964: 517; Radovsky, 1967: 196; 2010: 101; Dusbábek, 1970: 67; Beron, 1968: 160; 1973: 125; Pereira Lorenzo &amp; Quinteiro Alonso, 1993: 19.</p> <p>Type locality: Unknown.</p> <p>Type host: Unknown.</p> <p>Material. F ex Hypsugo savii Bonaparte, 1837 from Shura-Ozen’ river, 17 May 2019, leg. D.G. Smirnov; 2 FF, 6 N1 ex Pipistrellus nathusii from village Elimbetovo, 20 August 2014, leg. A. V. Zhigalin; 8 N1 ex Myotis daubentonii from village Elimbetovo, 25 August 2014, leg. A. V. Zhigalin; M, 4 N1 ex P. nathusii from Mordovian State Natural Reserve, 21 June 2016, leg. D.G. Smirnov; N1 ex M. daubentonii from Mordovian State Natural Reserve, 22 June 2016, leg. D.G. Smirnov; F, 2 MM, 5 N1 ex M. dasycneme from Mordovian State Natural Reserve, 23 June 2016, leg. D.G. Smirnov; N1 ex Vespertilio murinus from State Nature Reserve “Samarskaya Luka”, 1-31 July 2006, leg. D.G. Smirnov; 10 FF, 8 N1 ex P. nathusii from village Sosnovka, 1-31 May 2006, leg. D.G. Smirnov; F ex V. murinus from Pervouralsk, 20 July 2015, leg. O.L. Orlov; F ex Eptesicus nilssonii from Serpievskaya cave, 27 July 2011, leg. O.L. Orlov; M ex M. dasycneme from Serpievskaya cave, 1 November 2012, leg. O.L. Orlov; F, M, 7 N1 ex P. nathusii from Natural Park “Nurgush”, 19 June 2006, leg. D.G. Smirnov; 3 N1 ex P. nathusii from Pogromnoye settlement, 26 June 2003, leg. D.G. Smirnov.</p> <p>Distribution in Russian Federation: Crimea (Bobkova 2004; Orlova &amp; Orlov, 2018), Pskov Province (Medvedev et al., 2000), Penza Province (Stanyukovich, 1997), Voronezh Province (Stanyukovich, 1997), Samara Province (Stanyukovich, 1997), Volgograd Province (Stanyukovich, 1997), North Osetiya Republic (Stanyukovich, 1997), Rostov Province (Zabashta et al., 2019; Orlova et al., 2020b), Dagestan Republic (Orlova et al., 2020a), Astrakhan Province (Zabashta et al., 2019; Orlova et al., 2020b), Mordovia Republic (present paper), Kirov Province (present paper), Bashkiria Republic (present paper), Sverdlovsk Province (present paper), Orenburg Province (present paper), Irkutsk Province (Stanyukovich, 1997), [Pre-Urals] (Orlova et al., 2015d).</p> <p>Distribution outside Russian Federation: Europe (Radovsky, 1967; Micherdzinski, 1980), Asia (Stanyukovich, 1997), Africa (Algeria, Egypt, Libya) (Benda et al., 2014).</p> <p>Hosts. Pipistrellus nathusii (Medvedev et al., 2000; Zabashta et al., 2019; present paper), Pi. pipistrellus (Zabashta et al., 2019; Orlova &amp; Orlov, 2018), Pi. kuhlii (Zabashta et al., 2019; Orlova et al., 2020b), Pi. pygmaeus (Zabashta et al., 2019), Pi. coromandra Gray, 1838 (Stanyukovich, 1997), Myotis blythii (Stanyukovich, 1997), M. myotis (Stanyukovich, 1997), M. dasycneme (Stanyukovich, 1997; present paper), M. daubentonii (Stanyukovich, 1997; present paper), M. mystacinus (Stanyukovich, 1997), M. brandtii (Stanyukovich, 1997), M. cappaccinii (Stanyukovich, 1997), M. emarginatus (Stanyukovich, 1997), M. nattereri (Stanyukovich, 1997), Plecotus auritus (Stanyukovich, 1997), P. austriacus (Stanyukovich, 1997), Vespertilio murinus (Stanyukovich, 1997; present paper), Eptesicus nilssonii (Stanyukovich, 1997; present paper), E. serotinus (Stanyukovich, 1997), Barbastella barbastellus (Stanyukovich, 1997), Nyctalus noctula (Stanyukovich, 1997), Hypsugo savii (present paper), Rhinolophus ferrumequinum (Stanyukovich, 1997), Rh. euryale (Stanyukovich, 1997), Miniopterus schreibersi (Stanyukovich, 1997).</p> </div>	http://treatment.plazi.org/id/E35487F5352FFFAB0F93FC40FEEAD1E0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Orlova, M. V.;Klimov, P. B.;Orlov, O. L.;Smirnov, D. G.;Zhigalin, A. V.;Budaeva, I. V.;Emelyanova, A. A.;Anisimov, N. V.	Orlova, M. V., Klimov, P. B., Orlov, O. L., Smirnov, D. G., Zhigalin, A. V., Budaeva, I. V., Emelyanova, A. A., Anisimov, N. V. (2021): A checklist of bat-associated macronyssid mites (Acari: Gamasina: Macronyssidae) of Russia, with new host and geographical records. Zootaxa 4974 (3): 537-564, DOI: https://doi.org/10.11646/zootaxa.4974.3.4
E35487F53530FFAA0F93FCF1FBD9D24D.text	E35487F53530FFAA0F93FCF1FBD9D24D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Steatonyssus spinosus Willmann 1936	<div><p>Steatonyssus spinosus Willmann, 1936</p> <p>Steatonyssus spinosus Willmann, 1936: 152.</p> <p>Steatonyssus spinosus.— Fonseca, 1948: 319; Yunker &amp; Radovsky, 1966: 96; Beron, 1968: 160; 1969: 163; 1971: 100; Solomon, 1971: 313; Mahnert, 1971: 926; Pinchuk, 1971: 110; 1976: 129; Chow, 1973: 53; Rybin, 1983: 359; Uchikawa &amp; Kumada, 1977: 427; Haitlinger &amp; Ruprecht, 1977: 333; Haitlinger, 1978: 713; Ogadzhanyan &amp; Arutunyan, 1974: 80; Teng, 1980; Rybin, 1983: 359; Schmidt, 1987; Stanjukovich, 1990: 197; Haitlinger &amp; Lupicky, 2008: 124.</p> <p>Steatonyssus (S.) spinosus.— Till &amp; Evans, 1964: 567; Ah &amp; Radovsky, 1967: 425; Radovsky 1966: 94; 1967: 207; 2010: 101; Beron 1973: 124; Orlova et al., 2011: 136.</p> <p>Steatonyssus musculi.— Dusbábek, 1964b: 103.</p> <p>Type locality: Zoo of Hamburg (Germany) (Willmann, 1936).</p> <p>Type host: [Solenodon paradoxus Brandt, 1833] (from feces) (Willmann, 1936).</p> <p>Material. M ex Pipistrellus pygmaeus from biological station “Venevitinovo”, 4 June 2019, leg. I. V. Budaeva; F, 16 N1 ex Pipistrellus nathusii from State Nature Reserve “Samarskaya Luka”, 1-31 July 2006, leg. D.G. Smirnov; 2 N1 ex Vespertilio murinus from Biological station “Kolarovo”, 11 September 2015, leg. A. V. Zhigalin; N1 ex Myotis sibiricus from Tigirekskiy State Natural Reserve, 16 June 2016, leg. A. V. Zhigalin, O.L. Orlov; N1 ex Myotis sibiricus from State Natural Reserve “Barguzinskiy”, 2 July 2015, leg. D. V. Kazakov; N1 ex Myotis petax from Okhotnichiya cave, 4 September 2015, leg. D. V. Kazakov; F, M, N1 ex Eptesicus nilssonii from lake Gusinoe, 30 June 2016, leg. D. V. Kazakov; N1 ex Myotis sibiricus from Argaley-3 cave, 21 May 2016, leg. D. V. Kazakov; M ex Ep. nilssonii from Nizhniy Tsasuchey, 18 July 2015; M, 2 N1 ex Myotis davidii from Adun-Chelon, 24 July 2015, leg. D. V. Kazakov.</p> <p>Distribution in Russian Federation: Samara Province (present paper), Volgograd Province (Stanyukovich, 1997), Bashkiria Republic (present paper), Sverdlovsk Province (Orlova, 2013; Orlova et al., 2017c), Chelyabinsk Province (Orlova, 2013; Orlova et al., 2017c), Novosibirsk Province (Orlova et al., 2015d), Altai Region (present paper), Tomsk Province (Orlova et al., 2017c), Tuva Republic (Orlova et al., 2015a), Irkutsk Province (present paper), Buryatiya Republic (present paper), Zabaykalskiy Region (Orlova et al., 2017c), Primorskiy Region (Medvedev et al., 1991), [Ural] (Orlova et al., 2015d), [Southern Siberia] (Orlova et al., 2015c), [Far East] (Stanyukovich, 1997).</p> <p>Distribution outside Russian Federation: Europe (Radovsky, 1967). Asia (Radovsky, 1967).</p> <p>Hosts. [Solenodon paradoxus], Vespertilio murinus (Stanyukovich, 1997; Orlova et al., 2015d; present paper), V. sinensis (as V. superans — Till &amp; Evans, 1964; Medvedev et al., 1991), Myotis myotis (Stanyukovich, 1997), M. blythii (Stanyukovich, 1997), M. dasycneme (Orlova et al., 2015d), M. daubentonii (Stanyukovich, 1997), M. mystacinus (Stanyukovich, 1997), M. petax (Orlova et al., 2015c; present paper), M. davidii (present paper), M. sibiricus (present paper), Hypsugo alashanicus Bobrinskoy, 1926 (as Pipistrellus savii — Medvedev et al., 1991), Pipistrellus pygmaeus (present paper), Pi. pipistrellus (Stanyukovich, 1997), Pi. nathusii (Orlova et al., 2015d; present paper), Eptesicus nilssonii (Orlova et al., 2015d; present paper), Ep. serotinus (Stanyukovich, 1997), Nyctalus noctula (Stanyukovich, 1997), Ny. leisleri (Stanyukovich, 1997), Ny. aviator (Beron, 2014), Barbastella barbastellus (Stanyukovich, 1997), Plecotus auritus (Stanyukovich, 1997), Pl. ognevi (present paper), Rhinolophus ferrumequinum (Stanyukovich, 1997), Rh. hipposideros (Stanyukovich, 1997).</p> </div>	http://treatment.plazi.org/id/E35487F53530FFAA0F93FCF1FBD9D24D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Orlova, M. V.;Klimov, P. B.;Orlov, O. L.;Smirnov, D. G.;Zhigalin, A. V.;Budaeva, I. V.;Emelyanova, A. A.;Anisimov, N. V.	Orlova, M. V., Klimov, P. B., Orlov, O. L., Smirnov, D. G., Zhigalin, A. V., Budaeva, I. V., Emelyanova, A. A., Anisimov, N. V. (2021): A checklist of bat-associated macronyssid mites (Acari: Gamasina: Macronyssidae) of Russia, with new host and geographical records. Zootaxa 4974 (3): 537-564, DOI: https://doi.org/10.11646/zootaxa.4974.3.4
E35487F53531FFAA0F93FE9CFD50D542.text	E35487F53531FFAA0F93FE9CFD50D542.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Steatonyssus superans Zemskaja 1951	<div><p>Steatonyssus superans Zemskaja, 1951</p> <p>Steatonyssus superans Zemskaja, 1951: 1.</p> <p>Steatonyssus superans.— Bregetova, 1953: 325; 1956: 162; Zhovtiy et al., 1962; Morozova et al., 1964: 162; Chow, 1973: 53; Uchikawa &amp; Kumada, 1977: 427; Teng, 1980; Micherdzinski, 1980: 182.</p> <p>Steatonyssus (S.) superans.—Evans &amp; Till, 1964: 624; Radovsky, 1966: 95; 1967: 207; 2010: 101; Ah &amp; Radovsky, 1967: 426.</p> <p>Type locality: Primorskiy Region (as Maritime Province).</p> <p>Type host: Vespertilio sinensis (as Vespertilio superans).</p> <p>Material. F (with intrauterine egg) ex Vespertilio murinus from biological station “Preduraliye”, 3 July 2012, leg. O.L. Orlov, M. V. Orlova; F ex V. murinus from State Nature Reserve “Samarskaya Luka”, 7 July 2016, leg. D.G. Smirnov; 4 FF, 8 MM, 6 N1 ex V. murinus from Novouralsk town, 18 July 2010, leg. O.L. Orlov; 2 FF, M, 3 N1 ex V. murinus from Ekaterinburg, 13 September 2016, leg. O.L. Orlov; 21 FF (including 16 with intrauterine egg), 3 MM, 14 N1 ex V. murinus from Pervouralsk town, 20 July 2015, leg. O.L. Orlov; 8 N1 ex Myotis petax from Tigirekskiy State Natural Reserve, 2 March 2016, leg. A. V. Zhigalin; 2 FF, 13 N1 ex V. murinus from biological station “Kolarovo”, 9 August 2014, leg. A. V. Zhigalin; 3 MM, 22 N1 ex V. murinus from Nizhnee Beloe lake, 4 July 2016, leg. D.G. Kazakov; 2 FF (both with intrauterine egg) ex Plecotus ognevi from Adun-Chelon, 26 July 2015, leg. D. V. Kazakov; 62 FF (including 57 with intrauterine egg), 16 MM, 243 N1 ex V. murinus from Adun-Chelon, 26 July 2015, leg. D. V. Kazakov; 3 FF (with intrauterine egg), 8 MM, 348 N1 ex Vespertilio sinensis from Adun-Chelon, 24 July 2015, leg. D. V. Kazakov.</p> <p>Distribution in Russian Federation: Samara Province (present paper), Volgograd Province (Stanyukovich, 1997), Perm Region (present paper), Sverdlovsk Province (present paper), Chelyabinsk Province (Orlova &amp; Orlov, 2013; Orlova et al., 2015d), Khanty-Mansi autonomous Region (Orlova &amp; Tomishina, 2019), Novosibirsk Province (Stanyukovich, 1997), Altai Region (present paper), Tomsk Province (present paper), Tuva Republic (Orlova et al., 2015a; Orlova et al., 2015d), Buryatia Republic (present paper), Zabaykal’skiy Region (Zhovtiy et al., 1962), Kamchatka Region (Medvedev et al., 1991), Primorskiy Region (Zemskaja 1951; Bregetova, 1956; Medvedev et al., 1991), [Far East] (Stanyukovich, 1997), [Ural, Western Siberia] (Orlova et al., 2015d).</p> <p>Distribution outside Russian Federation: Asia (Kazakhstan, Tajikistan— Stanyukovich, 1997; Korea—Ah &amp; Radovsky, 1967).</p> <p>Hosts. Vespertilio sinensis (Bregetova, 1956; as Vespertilio superans — Medvedev et al., 1991; present paper), V. murinus (Zemskaja 1951; Zhovtiy et al., 1962; Orlova et al., 2015d; present paper), Myotis petax (present paper), Eptesicus nilssonii (Medvedev et al., 1991), Plecotus ognevi (present paper). Eptesicus serotinus (Beron, 2014), Nyctalus lasiopterus aviator (Beron, 2014).</p> </div>	http://treatment.plazi.org/id/E35487F53531FFAA0F93FE9CFD50D542	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Orlova, M. V.;Klimov, P. B.;Orlov, O. L.;Smirnov, D. G.;Zhigalin, A. V.;Budaeva, I. V.;Emelyanova, A. A.;Anisimov, N. V.	Orlova, M. V., Klimov, P. B., Orlov, O. L., Smirnov, D. G., Zhigalin, A. V., Budaeva, I. V., Emelyanova, A. A., Anisimov, N. V. (2021): A checklist of bat-associated macronyssid mites (Acari: Gamasina: Macronyssidae) of Russia, with new host and geographical records. Zootaxa 4974 (3): 537-564, DOI: https://doi.org/10.11646/zootaxa.4974.3.4
