taxonID	type	description	language	source
038487E62F24243EFF12B7D2FBBDF8C7.taxon	description	Figs 1 A, 2	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F24243EFF12B7D2FBBDF8C7.taxon	materials_examined	Material examined. Holotype: Thailand, Surat Thani Province, Ang Thong Islands, Wua Talap Island, ascent to Buaboke Cave, 9 ° 37.993 ’ N, 99 ° 40.369 ’ E, ca. 50 m a. s. l. (locality code: Th 1), Sep. 2007, A. Reischütz leg., at base of limestone rocks, NHMW 113484 (D: 2.03 mm, H: 1.23 mm). Paratype: 1 shell, same data as holotype, RE.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F24243EFF12B7D2FBBDF8C7.taxon	etymology	Etymology. The specific epithet (noun used in apposition) derives from the name of the cave where the new species was collected.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F24243EFF12B7D2FBBDF8C7.taxon	diagnosis	Diagnosis. A tiny Chamalycaeus species with finely ribbed R 1, elevated, narrow swelling on R 3, and great distance between inner and outer peristomes.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F24243EFF12B7D2FBBDF8C7.taxon	description	Description. Shell off-white, probably somewhat translucent when fresh; shell outline oval in dorsal view; spire slightly elevated, low conical; body whorl rounded; protoconch low, finely granular, spiral striae lacking, 1.25 – 1.5 whorls; R 1 of 1.25 – 1.5 whorls, with strong, elevated, slim, densely, regularly arranged ribs, fine spiral striation between ribs; boundary between R 1 and R 2 distinct due to change of rib density; R 2 ribs slightly curved towards aperture, spaces between ribs approximately as wide or slightly wider than ribs; ca. 26 ribs present; R 2 + R 3 less than quarter whorl (ca. 80 ˚), R 2 somewhat shorter than R 3; boundary between R 2 and R 3 distinct due to deep constriction and ribless posterior portion of R 3; R 3 with short, narrow, ribbed central swelling; anterior slope of swelling steeper than posterior slope; aperture oblique to shell axis, rounded with small basal incision; boundary between inner and outer peristomes conspicuous, inner peristome strongly thickened, protruding, and slightly expanded; outer peristome thin, strongly expanded, distinctly reflected portion over umbilicus; distance between inner and outer peristomes significant, swelling of R 3 and edge of outer peristome closely situated; umbilicus relatively narrow, ca. one fourth of shell width. Measurements. D: 2.03 – 2.29 mm, H: 1.23 – 1.33 mm. Operculum. Unknown.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F24243EFF12B7D2FBBDF8C7.taxon	diagnosis	Differential diagnosis. The most similar species is Chamalycaeus canaliculatus (Möllendorff, 1894) (examined material: Golf von Siam: Koh-Samui, coll. Möllendorff, SMF 109468, lectotype, Fig. 1 B), which was described from the nearby island, Koh Samui. The new species from Wua Talap Island is slightly smaller (C. canaliculatus D: 2.4 mm), has a more elevated R 3, and much greater distance between the inner and outer peristomes. The new species differs from Dicharax ovatus n. sp. by the slightly smaller size, lower spire, very rounded body whorl and aperture, blunt swelling on R 3, and the shorter distance between the inner and outer peristomes.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F24243EFF12B7D2FBBDF8C7.taxon	distribution	Distribution. This new species is known only from the type locality (Fig. 3).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F24243EFF12B471FAA0FDA3.taxon	type_taxon	Type species. Alycaeus (Chamalycaeus) fruhstorferi Möllendorff, 1897, by monotypy.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F24243EFF12B471FAA0FDA3.taxon	diagnosis	Diagnosis. Genus characterized by the combination of a spirally striated teleoconch, protoconch without spiral striae, and wide umbilicus (see Páll-Gergely et al. (2020) for extended diagnosis and taxonomic remarks).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F272432FF12B3D9FBBDFCCD.taxon	description	Fig. 4	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F272432FF12B3D9FBBDFCCD.taxon	materials_examined	Material examined. Holotype: Thailand, Kanchanaburi Province, Erawan National Park, Erawan Falls Trail, 14 ° 22.310 ’ N, 99 ° 8.699 ’ E, 90 m a. s. l. (locality code: 2015 / 28), 17 Feb. 2015, A. Hunyadi leg., HNHM 104869 (D: 2.45 mm, H: 1.53 mm). Paratypes: 4 shells, same data as for holotype, HA; 1 shell, Thailand, Kanchanaburi Province, 4.3 km SW Kanchanaburi, Taoist Buddhist temple, 13 ° 59 ’ N, 99 ° 31 ’ E, 40 m a. s. l., 13 May 1988, K. Auffenberg leg., litter sample, UF 345935.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F272432FF12B3D9FBBDFCCD.taxon	etymology	Etymology. The specific epithet refers to the type locality (Erawan National Park).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F272432FF12B3D9FBBDFCCD.taxon	diagnosis	Diagnosis. A small Chamalycaeus species with bluntly angled body whorl, short R 2, rounded aperture, and strongly expanded outer peristome.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F272432FF12B3D9FBBDFCCD.taxon	description	Description. Shell off-white to light grey, rather glossy, very slightly translucent; shell outline slightly oval in dorsal view; spire slightly elevated, conical-domed; body whorl rather bluntly angled, depressed inward basolaterally in apertural view; protoconch rather low, glossy, spiral striae lacking, 1.25 – 1.5 whorls; R 1 of 1.5 – 1.75 whorls, with irregular, narrow, but rather low ribs and some weak spiral striation; boundary between R 1 and R 2 not distinct, but visible due to change in rib density; R 2 + R 3 approximately a quarter whorl; R 2 short, about one third of R 3 length, only ca. 14 – 16 ribs; R 2 ribs low, dorsally rounded; boundary between R 2 and R 3 distinct due to moderately deep constriction and almost smooth early R 3; R 3 with elongated, almost smooth (only a few low riblets / growth lines) central swelling; aperture oblique to shell axis, round; distance between inner and outer peristomes conspicuous, inner peristome protruding, thin, but robust, and slightly expanded; outer peristome strongly expanded, but not reflected, reduced in umbilical area; umbilicus narrow, less than one third shell width. Measurements. D: 2.25 – 2.45 mm, H: 1.49 – 1.56 mm. Operculum. Unknown.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F272432FF12B3D9FBBDFCCD.taxon	diagnosis	Differential diagnosis. This new species differs from Chamalycaeus irmatallus n. sp. in the larger size, longer R 3, and more expanded peristome.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F272432FF12B3D9FBBDFCCD.taxon	distribution	Distribution. This species is known only from two nearby localities (Fig. 5).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F2A2431FF12B589FB7EFEC5.taxon	description	Figs 6 – 8	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F2A2431FF12B589FB7EFEC5.taxon	materials_examined	Material examined. Holotype: Thailand, Krabi Province, N of Krabi, Wat Tham Suea (Tiger Cave Temple), along steps to the cave, 8 ° 7.603 ’ N, 98 ° 55.466 ’ E, ca. 90 m a. s. l. (locality code: Th 11), Sep. 2007, A. Reischütz leg., at base of limestone rocks, NHMW 113485 (D: 2.42 mm, H: 1.53 mm). Paratypes: 12 shells, same data as holotype, RE; 42 shells, Thailand, Krabi Province, Railay Beach West, Viewpoint Hill, 8 ° 0.511 ’ N, 98 ° 50.248 ’ E, ca. 30 m a. s. l. (locality code: Th 14), Sep. 2007, A. Reischütz leg., at base of limestone rocks, RE; 2 shells, same data as preceding, CUMZ 5268; 1 shell (Fig. 6 C), Thailand, Krabi Province, E of Krabi, Khao Khanab Nam limestone hill, 8 ° 4.707 ’ N, 98 ° 55.263 ’ E, ca. 2 m a. s. l. (locality code: Th 15), Sep. 2007, A. Reischütz leg., at base of limestone rocks just above tidal influence, NHMW 113486; 17 shells, same data as preceding, RE; 3 shells, limestone hill at river Khanab Nam (Krabi Yai) at the northeastern city limits of Krabi, by the road Thanon Lan Gu (Mueang Kao), ca. 400 m E of crossing with road Thanon Thong Chai, S of road, 8 ° 5.817 ’ N, 98 ° 55.068 ’ E, ca. 10 m a. s. l. (locality code: Th 17), Mar. 2010, A. Reischütz leg., at base of limestone rocks, RE; 2 shells, Thailand, Krabi Province, Wat Tham Sua, 8 ° 7.567 ’ N, 98 ° 55.496 ’ E, 40 m a. s. l. (locality code: 2015 / 34), 21 Feb. 2015, A. Hunyadi leg., HA. Other material: 7 juvenile / broken shells; same data as holotype, RE; 6 juvenile / broken shells, Thailand, Krabi Province, Railay Beach West, Viewpoint Hill; 8 ° 0.511 ’ N, 98 ° 50.248 ’ E; ca. 30 m a. s. l. (locality code: Th 14), Sep. 2007, A. Reischütz leg., at base of limestone rocks, RE; 4 juvenile / broken shells, Thailand, Krabi Province, E of Krabi, Khao Khanab Nam limestone hill, 8 ° 4.707 ’ N, 98 ° 55.263 ’ E, ca. 2 m a. s. l. (locality code: Th 15), Sep. 2007, A. Reischütz leg., at base of limestone rocks just above tidal influence, RE; 2 juvenile / broken shells, Thailand, Krabi Province, limestone hill at river Khanab Nam (Krabi Yai) at northeastern city limits of Krabi, by road Thanon Lan Gu (Mueang Kao), ca. 400 m E of crossing with road Thanon Thong Chai, S of road, 8 ° 5.817 ’ N, 98 ° 55.068 ’ E, ca. 10 m a. s. l. (locality code: Th 17), Mar. 2010, A. Reischütz leg., at base of limestone rocks, RE; 3 juvenile / broken shells, Thailand, Krabi Province, Wat Tham Sua, 8 ° 7.567 ’ N, 98 ° 55.496 ’ E, 40 m a. s. l. (locality code: 2015 / 34), 21 Feb. 2015, A. Hunyadi leg., HA; 2 broken shells, Thailand, Krabi Province, granite rock in evergreen forest at Tone Nga Chang Waterfall, Cha Lung, Hat Yai, Songkhla, 6 ° 56.810 ’ N, 100 ° 13.895 ’ E (locality code: AC 21), C. Sutcharit et al. leg., CUMZ 5271 (Fig. 8, identification questionable, not shown on Fig. 3).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F2A2431FF12B589FB7EFEC5.taxon	etymology	Etymology. The specific epithet refers to the R 2 rib morphology, which, with the numerous tooth-like projections, is reminiscent of a harrow, an agricultural tool consisting of a framework with teeth used to break up plowed land (hirpex in Latin).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F2A2431FF12B589FB7EFEC5.taxon	diagnosis	Diagnosis. A small Chamalycaeus species with R 2 + R 3 shorter than a quarter whorl and R 2 with serrulate ribs.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F2A2431FF12B589FB7EFEC5.taxon	description	Description (based on the type series). Shell off-white, light grey to slightly yellowish; shell outline rounded in dorsal view, R 2 conspicuously bulging / angled in dorsal view, but variable in some populations; spire rather depressed, but variable among populations, low conical; body whorl rounded or rather parabolic; protoconch elevated, finely granular, spiral striation lacking, 1.5 whorls (Figs 7 C – D); R 1 ca. 1.5 whorls, first ca. 0.75 whorl dominated by fine ribbing that gradually increases strength toward the R 1 termination; spiral striation as strong as radial ribs at end of R 1, while due to domination of ribs at end of R 1, relative strength of spiral striae decreases; R 1 and R 2 boundary conspicuous due to denser R 2 ribs; R 2 + R 3 less than a quarter whorl (ca. 70 – 80 ˚); R 2 bulging, forming a blunt angle in dorsal view, and rounded in apertural view; R 2 ribs lamella-like, rather low, characteristically serrate (i. e. a series of tiny spikes pointing anteriorly, see Figs 7 A – B), ca. 24 – 26 R 2 ribs present; R 2 and R 3 of comparable lengths or R 2 slightly shorter; R 3 with spiral striation and weaker ribs than R 1; boundary between R 2 and R 3 distinct due to lower R 3 ribs, although constriction is shallow; aperture very strongly oblique to shell axis; rounded, with very weak upper (parieto-palatal) incision, a weak basal incision occasionally also indicated; boundary between inner and outer peristomes distinct; inner peristome protruding, rather sharp and relatively thin; outer peristome expanded except in umbilical area; umbilicus relatively wide, over one third shell width. A cross-sectional view of R 2 was examined in one specimen (Figs 7 E – H): anterior crust forms an elevated rib, which forms a series of hollow spikes (visible in cross-sectional image as widened tip) toward aperture, posterior crust low, adnate to anterior crust, which slightly folds over posterior crust; cross-sectional view of microtunnels nearly rounded, drop shaped, or somewhat triangular. Measurements. D: 2.00 – 2.61 mm, H: 1.32 – 1.67 mm. Operculum. Strongly concave, outer surface with no structures, covered with thick mud layer; inner side with elevated central nipple (Fig. 6 B).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F2A2431FF12B589FB7EFEC5.taxon	diagnosis	Differential diagnosis. The reticulated R 2 sculpture distinguishes this species from all congeners except for C. serratus n. sp. See that species for comparisons.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F2A2431FF12B589FB7EFEC5.taxon	distribution	Distribution. This new species is known from four populations occurring in Krabi Province, Thailand (Fig. 3).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F2A2431FF12B589FB7EFEC5.taxon	discussion	Remarks. The examined populations show a degree of morphological variability, see Table 1.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F2D2434FF12B7EAFD6BFF0D.taxon	description	Figs 9, 10 A	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F2D2434FF12B7EAFD6BFF0D.taxon	materials_examined	Material examined. Holotype: Thailand, Surat Thani Province, limestone mtn., 6 km S of Na San, 8 ° 39.583 ’ N, 99 ° 23.883 ’ E, 100 m a. s. l., 3 Jun. 1987, F. G. Thompson leg., UF 279508 (D: 2.02 mm, H: 1.32 mm). Paratypes: 11 shells, same data as holotype, UF 551216. Other material: 6 shells, Myanmar, Thayet-myo, Pegu, coll. Blanford, NHMUK 1906.4.4.71; 1 shell, Thailand, Phang Nga Province, Phang Nga, Tham Nam Phud, 8 ° 27.829 ’ N, 98 ° 32.617 ’ E, 40 m a. s. l. (locality code: 2015 / 33), 20 Feb. 2015, A. Hunyadi leg., around the cave, HA.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F2D2434FF12B7EAFD6BFF0D.taxon	etymology	Etymology. The specific epithet is an anagram for armillatus, a species with which this new species has been confused (see Differential diagnosis).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F2D2434FF12B7EAFD6BFF0D.taxon	diagnosis	Diagnosis. A tiny Chamalycaeus species with glossy shell, irregularly, finely ribbed R 1, very short R 2 with blunt, low ribs, and somewhat longer R 3 (the two latter regions are shorter than a quarter whorl combined).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F2D2434FF12B7EAFD6BFF0D.taxon	description	Description. Shell off-white, rather glossy, although available shells are somewhat eroded; shell outline slightly oval in dorsal view; spire slightly elevated, conical-domed; body whorl somewhat “ parabolic ” in apertural view; protoconch rather low, glossy, spiral striae lacking, 1.25 whorls; R 1 of 1.5 – 1.75 whorls, with very indistinct, irregular, low and rounded, relatively wide ribs, and some weak spiral striation; boundary between R 1 and R 2 distinct due to change in intensity and strength of ribbing; R 2 short, only ca. 16 – 20 ribs; R 2 + R 3 less than quarter whorl (ca. 70 – 80 ˚); R 3 up to twice as long as R 2; boundary between R 2 and R 3 clear due to change in rib morphology, although constriction very shallow; R 3 with very fine, rounded ribs, and low, convex, elongated central swelling; aperture round, slightly oblique to shell axis; boundary between inner and outer peristomes distinct, inner peristome strongly protruding, thickened, slightly expanded; outer peristome slightly weaker or as strong, also expanded, only slightly reflected toward umbilicus; umbilicus relatively narrow, ca. one third of shell width. Measurements. D: 1.75 – 2.02 mm, H: 1.15 – 1.32 mm. Operculum. Unknown.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F2D2434FF12B7EAFD6BFF0D.taxon	diagnosis	Differential diagnosis. Chamalycaeus armillatus (Benson, 1856) (material examined: Thayet-Mio, UMZC 102995, holotype, Fig. 10 B) has a more rounded body whorl, longer R 2 and R 3, and a thicker peristome with greater distance between the edges of the inner and outer peristomes.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F2D2434FF12B7EAFD6BFF0D.taxon	distribution	Distribution. The type locality is in Thailand’s Surat Thani Province. A single shell was collected in the neighbouring Phang Nga Province (locality code: 2015 / 33, coll. HA). Shells collected more than 1,000 km north in Thayet, Myanmar are so similar to the Thai populations that they could not be separated (see below) (Fig. 3).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F2D2434FF12B7EAFD6BFF0D.taxon	discussion	Remarks. Chamalycaeus armillatus was described from an eroded specimen. Some spiral striations are visible on the holotype, but they might be the part of the lower shell layer. Thus, this species was provisionally assigned to Chamalycaeus in Páll-Gergely et al. (2020). Specimens from Thayet (NHMUK 1906.4.4.71, labelled as armillatus) have slightly less densely arranged R 2 ribs than typical C. irmatallus n. sp. A single, eroded shell from Tham Nam Phud (Thailand) is somewhat larger than the holotype of C. irmatallus n. sp., and has a longer R 2. Since all other shell characters are identical, we provisionally identify the Tham Nam Phud and Thayet specimens as C. cf. irmatallus n. sp.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F30242AFF12B589FBDAF939.taxon	description	Fig. 11	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F30242AFF12B589FBDAF939.taxon	materials_examined	Material examined. Holotype: Thailand, Krabi Province, Phi-Phi Islands, Phi Phi Don Island, climbing rock at W end of beach in Ton Sai Bay, 7 ° 44.011 ’ N, 98 ° 45.936 ’ E, ca. 70 m a. s. l. (locality code: Th 22), Mar. 2010, A. Reischütz leg., at base of limestone rocks in the forest, NHMW 113487 (D: 3.61 mm, H: 2.62 mm). Paratypes: 1 shell, same data as holotype; 1 shell, Thailand, Krabi Province, Phi-Phi Islands, Phi Phi Don Island, ascent to climbing rock at W end of beach in Ton Sai Bay, 7 ° 44.049 ’ N, 98 ° 45.966 ’ E, ca. 20 m a. s. l. (locality code: Th 21), Mar. 2010, A. Reischütz leg., at base of limestone rocks in forest, NHMW 113488 (photographed paratype: Fig. 11 B, D: 3.73 mm, H: 2.37 mm); 9 shells, same data as preceding, RE; 2 shells, same data as preceding, CUMZ 5269; 7 shells, Thailand, Krabi Province, Phi-Phi Islands, Phi Phi Don Island, limestone rocks at Monkey Beach, 7 ° 43.386 ’ N, 98 ° 46.135 ’ E, ca. 10 m a. s. l. (locality code: Th 23), Mar. 2010, A. Reischütz leg., RE. Other material: 4 juvenile / broken shells, Thailand, Krabi Province, Phi-Phi Islands, Phi Phi Don Island, ascent to climbing rock at W end of beach in Ton Sai Bay, 7 ° 44.049 ’ N, 98 ° 45.966 ’ E, ca. 20 m a. s. l. (locality code: Th 21), Mar. 2010, A. Reischütz leg., at base of limestone rocks in forest, RE; 5 juvenile / broken shells, Thailand, Krabi Province, Phi-Phi Islands, Phi Phi Don Island, limestone rocks at Monkey Beach, 7 ° 43.386 ’ N, 98 ° 46.135 ’ E, ca. 10 m a. s. l. (locality code: Th 23), Mar. 2010, A. Reischütz leg., RE; 7 shells, 1.5 km SE of Ao Luek Tai, 8 ° 22.386 ’ N, 98 ° 44.130 ’ E, 40 – 45 m a. s. l., 16 – 19 Mar. 2020, S. Aiken leg., SA; 4 shells (one photographed here: Fig. 11 C), 1.7 km SE of Ao Luek Tai, 8 ° 22.020 ’ N, 98 ° 44.004 ’ E, 25 m a. s. l., 19 Mar. 2020, S. Aiken leg., in dry debris on rock, at base of limestone cliff, SA.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F30242AFF12B589FBDAF939.taxon	etymology	Etymology. This new species is named for the Thai province in which it occurs.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F30242AFF12B589FBDAF939.taxon	diagnosis	Diagnosis. A medium-sized Chamalycaeus species with R 2 and R 3 of almost a quarter whorl each, blunt, elongated swelling on R 3, protruding inner peristome, and strongly expanded outer peristome.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F30242AFF12B589FBDAF939.taxon	description	Description. Shell off-white to light grey; shell outline slightly oval in dorsal view; spire elevated, conical; body whorl rounded or parabolic; protoconch somewhat elevated, finely granular, rather glossy, spiral striae lacking, 1.5 whorls; R 1 of 1.75 – 2.5 whorls, with strong, narrow, elevated, regularly arranged ribs, spaces between ribs relatively wide, fine spiral striation between ribs; boundary between R 1 and R 2 distinct due to change in rib density; R 2 ribs more elevated than those of R 1, narrow, lamella-like; ca. 26 – 28 ribs present; R 2 and R 3 of comparable lengths, covering approximately a half whorl or slightly less; boundary between R 2 and R 3 distinct due to rather deep constriction and change in sculpture; R 3 with elongated, convex, central swelling, posterior portion almost smooth, anterior bears ribs of similar density as on R 1; aperture strongly oblique to shell axis, rounded, with a very small upper (parieto-palatal) incision; boundary between inner and outer peristomes conspicuous, inner peristome strongly protruding (extreme in some specimens) and slightly expanded; outer peristome slim, strongly expanded except in umbilical region; umbilicus narrow, ca. one fourth of shell width. Measurements. D: 3.41 – 3.81 mm, H: 2.34 – 2.62 mm (type samples). Operculum. Unknown.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F30242AFF12B589FBDAF939.taxon	diagnosis	Differential diagnosis. Chamalycaeus reinhardti (Mörch, 1872) (examined material: Great Nicobar, coll. Godwin-Austen, NHMUK 1903.7.1.2711, 1 shell) and Chamalycaeus reinhardti sabangensis (Rensch, 1933) (examined material: Sumatra: Wald b. Sabang, Pulu Weh., exp. Rensch, 1927, SMF 6241, 1 paratype) both have a narrower swelling on R 3 and a more elevated spire.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F30242AFF12B589FBDAF939.taxon	distribution	Distribution. This species is known from the Phi-Phi Islands and the vicinity of Ao Luek Tai town, Krabi Province, Thailand (Fig. 3).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F30242AFF12B589FBDAF939.taxon	discussion	Remarks. The shells from the two sites near Ao Luek Tai are slightly smaller (D: 2.8 – 3.3 mm), more depressed and conical from above, have overall denser ribs, lower R 3 swelling, and the outer peristome is expanded in a smaller portion of the lip. However, although these differences occur, we consider them conspecific with those from Phi-Phi Islands due to the overall similar sculpture and proportions of shell regions.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F322429FF12B589FBBDFDE5.taxon	description	Fig. 12	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F322429FF12B589FBBDFDE5.taxon	materials_examined	Material examined. Holotype: Thailand, Nakhon Si Thammarat Province, Khao Luang Park, 15.2 km S junction Highways 4186 and 4188, 1.3 km S Park Headquarters, 8 ° 44 ’ N, 99 ° 43 ’ E, 200 m a. s. l., 16 Apr. 1988, K. Auffenberg leg., UF 345204 (D: 2.35 mm, H: 1.67 mm).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F322429FF12B589FBBDFDE5.taxon	etymology	Etymology. Named for the serrated R 2 ribs (Latin derivative serratus, notched like a saw).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F322429FF12B589FBBDFDE5.taxon	diagnosis	Diagnosis. A small Chamalycaeus species with R 2 + R 3 being longer than a quarter whorl, R 3 twice as long as R 2, which bears characteristically serrulate R 2 ribs.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F322429FF12B589FBBDFDE5.taxon	description	Description. Shell off-white, although dead-collected; shell outline rounded / slightly oval in dorsal view; spire elevated, conical; body whorl rounded with some depression inwardly in dorso-basal region; protoconch 1.5 whorls, elevated, finely granular, without spiral striation; R 1 slightly more than 1.5 whorls, first ca. 0.75 whorl dominated by fine ribs, gradually becoming stronger toward R 1 termination; spiral striation as strong as radial ribs at R 1 termination; boundary between R 1 and R 2 conspicuous because of denser R 2 ribs; R 2 + R 3 more than quarter whorl (approximately 100 ˚); R 2 ribs lamella-like, rather low, characteristically serrate with series of tiny spikes pointing anteriorly, ca. 28 R 2 ribs present; R 3 ca. twice as long as R 2; R 3 with rather strong spiral striation and weaker ribs than R 1; boundary between R 2 and R 3 clear due to less pronounced R 3 ribs, although constriction is shallow; aperture strongly oblique to shell axis, rounded; boundary between inner and outer peristomes distinct; inner peristome protruding, narrow, and relatively thin; outer peristome expanded, especially in basal and upper palatal directions; umbilicus narrow, ca. one fourth of shell width. Measurements. D: 2.35 mm, H: 1.67 mm. Operculum. Unknown.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F322429FF12B589FBBDFDE5.taxon	diagnosis	Differential diagnosis. This new species is very similar to Chamalycaeus hirpex n. sp. in terms of R 2 morphology. However, it differs from that species in the long R 3, which has an elongated, blunt swelling, and a more expanded aperture.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F322429FF12B589FBBDFDE5.taxon	distribution	Distribution. This new species is known only from the type locality (Fig. 3).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F34242EFF12B589FBBDFAD6.taxon	description	Fig. 13	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F34242EFF12B589FBBDFAD6.taxon	materials_examined	Material examined. Holotype: Thailand, Rayong Province, limestone range Khao San Ta, 2.0 km W of Ban Syaek Batan, 12 ° 58.75 ’ N, 101 ° 40.00 ’ E, 150 m a. s. l., 25 Apr. 1987, F. G. Thompson leg., UF 279157 (D: 1.8 mm, H: 1.16 mm). Paratypes: 5 shells, same data as holotype, UF 551217.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F34242EFF12B589FBBDFAD6.taxon	etymology	Etymology. This new species is named for its conspicuous spiral striae.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F34242EFF12B589FBBDFAD6.taxon	diagnosis	Diagnosis. A tiny Chamalycaeus species with strong, conspicuous spiral striations on entire teleoconch, R 2 + R 3 slightly shorter than a quarter whorl, and distance between the edges of the inner and outer peristomes is great, outer peristome not reflected.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F34242EFF12B589FBBDFAD6.taxon	description	Description. Shell off-white to yellowish-corneous, probably somewhat translucent when fresh; shell outline rounded in dorsal view; spire slightly elevated, low conical; body whorl rounded; protoconch somewhat elevated, glossy, finely granular, spiral striae lacking, 1.5 whorls; R 1 of 1.25 – 1.5 whorls, with very strong, broad spiral striae on last 0.75 – 1.0 whorl, preceding area rather smooth; last ca. half whorl of R 1 with strong but low ribs, similar strength to spiral striae, widely and irregularly spaced; boundary between R 1 and R 2 distinct due to change in rib density; R 2 ribs curve toward aperture, spaces between ribs approximately as wide as ribs; ca. 18 – 20 ribs present; R 2 + R 3 less than quarter whorl (ca. 70 – 80 ˚), comparable in length; boundary between R 2 and R 3 distinct due to change in rib morphology, constriction lacking; R 3 practically without swelling, with similar spiral striae to that of R 1, ribs virtually absent, replaced by very fine, weak growth lines; aperture slightly oblique to shell axis, round; boundary between inner and outer peristomes conspicuous, inner peristome strongly protruding and slightly expanded; edge of outer peristome thin, expanded weakly toward umbilicus, not reflected; umbilicus relatively narrow, approximately one third of shell width. Measurements. 1.72 – 1.93 mm, H: 1.07 – 1.16 mm. Operculum. Unknown.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F34242EFF12B589FBBDFAD6.taxon	diagnosis	Differential diagnosis. The small size, prominent, thin outer peristome, strong ribs and spiral striae distinguish this species from all other congeners. The most similar species in size and sculpture are Chamalycaeus microconus (Möllendorff, 1887) (examined material: Malakka: Bukit Pondong (Perak), SMF 109493, lectotype, and SMF 109494, 2 paralectotypes) and Chamalycaeus mixtus Zilch, 1957 (examined material: Malakka: Bukit Pondong (Perak), SMF 109510, holotype, and SMF 109511, 4 paratypes) from the Malay Peninsula (Bukit Pondong), but they have higher spires, comparatively larger apertures, and shorter R 3 s. For comparisons, see photos in Páll-Gergely et al. (2020).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F34242EFF12B589FBBDFAD6.taxon	distribution	Distribution. This new species is known only from the type locality (Fig. 5). Genus Dicharax Kobelt & Möllendorff, 1900 Charax Benson, 1859: 177. Dicharax Kobelt & Möllendorff, 1900: 186. (replacement name for Charax Benson, 1859, non Charax Scopoli, 1777 [Pisces]). Type species. Alycaeus hebes Benson, 1857, by subsequent designation (Gude 1921). Diagnosis. Typical Dicharax species possess R 2 ribs curved toward aperture. Genus generally characterized by complete absence of spiral striation. Only two of 164 species assigned to genus by Páll-Gergely et al. (2020), have indication of spiral striation. A few other species reported to have striations by Jirapatrasilp et al. (2021). Despite occurrence of spiral striations in relatively few species, the absence of striations is generally a valuable character by which to recognize Dicharax species, although examination of additional characters is also critical.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F34242EFF12B589FBBDFAD6.taxon	distribution	Distribution. This new species is known only from the type locality (Fig. 5).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F34242EFF12B589FBBDFAD6.taxon	type_taxon	Type species. Alycaeus hebes Benson, 1857, by subsequent designation (Gude 1921).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F34242EFF12B589FBBDFAD6.taxon	diagnosis	Diagnosis. Typical Dicharax species possess R 2 ribs curved toward aperture. Genus generally characterized by complete absence of spiral striation. Only two of 164 species assigned to genus by Páll-Gergely et al. (2020), have indication of spiral striation. A few other species reported to have striations by Jirapatrasilp et al. (2021). Despite occurrence of spiral striations in relatively few species, the absence of striations is generally a valuable character by which to recognize Dicharax species, although examination of additional characters is also critical.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F34242CFF12B232FE18F822.taxon	description	Figs 14 – 15	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F34242CFF12B232FE18F822.taxon	materials_examined	Material examined. Holotype: Thailand, Loei Province, 9.2 km NW Loei, Buddhist temple, up and N of main cave entrance, 17 ° 35 ’ N, 101 ° 44 ’ E, 250 m a. s. l., 28 Apr. 1988, K. Auffenberg leg., base of cliff under Georissa zone, UF 345861 (D: 2.9 mm, H: 1.9 mm). Paratype: 1 shell, same data as holotype, UF 551218.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F34242CFF12B232FE18F822.taxon	etymology	Etymology. The specific epithet admirandus (Latin for astonishing, remarkable, extraordinary) refers to several notable shell characters of this new species, namely the strong keel, strongly fringed aperture, and the spiral striation.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F34242CFF12B232FE18F822.taxon	diagnosis	Diagnosis. A medium-sized Dicharax species with strong peripheral keel, strongly ribbed and spirally striated R 1, and strongly fringed aperture.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F34242CFF12B232FE18F822.taxon	description	Description. Shell off-white, although both available shells were somewhat eroded; shell outline slightly oval in dorsal view; spire slightly elevated, low conical, body whorl with conspicuous peripheral keel; protoconch somewhat elevated, finely granular, spiral striation lacking, 1.75 whorls; R 1 of 1.5 whorls, first half whorl practically smooth, gradually changing to widely, regularly-spaced, strong ribs (ca. 30 ribs on last half whorl of R 1), present on entire R 1 except area between peripheral keel and outer edge of periumbilical keel, which has fine spiral striation; some spiral striation visible on dorsum of R 1, and between ribs; boundary between R 1 and R 2 distinct due to dense R 2 ribs; R 2 ribs curved toward aperture, ca. 34 – 36 R 2 ribs present; spaces between R 2 ribs narrower than ribs; R 2 + R 3 more than quarter whorl (ca. 110 ˚); R 2 considerably shorter than R 3, slightly more half its length; boundary between R 2 and R 3 distinct due to change in sculpture and deep constriction; R 3 with rounded, elongated swelling situated closer to constriction than to peristome; aperture strongly oblique to shell axis, rounded with basal incision; boundary between inner and outer peristomes not conspicuous; inner peristome strong, somewhat protruding, expanded, basal and palatal portions strongly fringed, outer peristome weaker, expanded, but reflected only in direction of umbilicus; umbilicus funnel-shaped, relatively narrow, less than one third of shell width. Measurements. D: 2.9 – 3 mm, H: 1.9 mm Operculum. Unknown.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F34242CFF12B232FE18F822.taxon	diagnosis	Differential diagnosis. The angled body whorl, the funnel-shaped umbilicus, the flattened dorsal side, and the strongly convex R 3 swelling distinguish this new species from the most similar D. fimbriatus (Bavay & Dautzenberg, 1912), which is known from Laos and Vietnam (Páll-Gergely et al. 2017). The R 1 ribs of this species are stronger than those of most D. fimbriatus populations, and the new species has a spirally striated R 1.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F34242CFF12B232FE18F822.taxon	materials_examined	Dicharax sylheticus (Godwin-Austen, 1914) (Examined material: S. Sylhet Hills, W. Channel leg., NHMUK 1903.7.1.55, holotype, Fig. 16) is similar, but has a more elevated spire, a less keeled body whorl, a longer R 2 with less dense ribs, and a rounded aperture.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F34242CFF12B232FE18F822.taxon	distribution	Distribution. This new species is known only from the type locality (Fig. 17).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F34242CFF12B232FE18F822.taxon	discussion	Remarks. Similarly to some populations of D. omissus (Godwin-Austen, 1914), this species also possesses spiral striation on R 1, which is very rare in Dicharax. Spiral striation on the teleoconch is characteristic for Chamalycaeus, but this species is classified in Dicharax due to the curved R 2 ribs characterizing that genus and its overall shell shape. Possible homology of the spiral striation of these few Dicharax species and other alycaeid genera needs further investigation.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F382423FF12B589FE5BFE74.taxon	description	Fig. 18	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F382423FF12B589FE5BFE74.taxon	materials_examined	Material examined. Holotype: Myanmar, Ataran, Burma, ex Dr. F. Stoliczka, NZSI M. 8073 (holotype [single specimen mentioned in original description]). Other material: 1 shell, Myanmar, Kayin State, Mudon centre ESE ca. 51 km, Htimahto NE, NE point of Kwooprai Hill, 16 ° 2.396 ’ N, 97 ° 58.200 ’ E, 30 m a. s. l. (locality code: 2018 / 47), 10 Oct. 2018, A. Hunyadi, K. Okubo & J. U. Otani leg., HA. Type locality. “ 1 Ataran ”.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F382423FF12B589FE5BFE74.taxon	diagnosis	Diagnosis (emended based on a newly collected specimen). Protoconch relatively low, glossy, without spiral striation; R 1 glossy, widely-spaced ribs near the suture and inside the umbilicus, edge of R 1 smooth; R 2 + R 3 over a quarter whorl long, R 2 ca. half as long as R 3; R 2 ribs typical for Dicharax (curved towards the aperture); R 3 smooth, with blunt, but large, conspicuous, elevated swelling.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F382423FF12B589FE5BFE74.taxon	discussion	Remarks. Until recently the eroded holotype was the only known specimen of the species making the assignment to Dicharax questionable. The newly collected fresh shell allowed examination of shell sculpture which confirmed the assignment.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F392421FF12B4E0FBBDFC7D.taxon	description	Fig. 19	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F392421FF12B4E0FBBDFC7D.taxon	materials_examined	Material examined. Holotype: Thailand, Nakhon Ratchasima Province, 3.4 km W of Ban Mu Si, limestone hill, 14 ° 32.016 ’ N, 101 ° 22.783 ’ E, 380 m a. s. l., 5 May 1987, F. G. Thompson leg., UF 346812 (D: 2.54 mm, H: 1.54 mm). Paratypes: 18 shells, same data as holotype, UF 551219. Other material: 2 juvenile shells, same data as holotype, UF 551220.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F392421FF12B4E0FBBDFC7D.taxon	etymology	Etymology. This species is named for the very round aperture (Greek cyclo-, ring and stoma, mouth, opening, noun in apposition).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F392421FF12B4E0FBBDFC7D.taxon	diagnosis	Diagnosis. A small, depressed Dicharax species with glossy R 1, protruding inner peristome, and round aperture; R 2 + R 3 shorter than a quarter whorl, R 2 half as long as R 3.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F392421FF12B4E0FBBDFC7D.taxon	description	Description. Shell white, some specimens with very faint brownish or yellowish colouration; shell outline rounded or very slightly oval in dorsal view, shell depressed, spire only slightly elevated, body whorl rounded; protoconch low, glossy, 1.25 – 1.5 whorls; R 1 of 1.25 – 1.75 whorls, glossy without sculptural elements; boundary between R 1 and R 2 conspicuous due to R 2 ribs; R 2 with ribs curved towards aperture, ca. 24 – 26 ribs present; spaces between R 2 ribs narrower than ribs; R 2 + R 3 less than quarter whorl (ca. 80 ˚); R 2 half or less length of R 3; R 3 glossy, smooth; boundary between R 2 and R 3 distinct due to change in sculpture and moderately deep constriction; middle of R 3 with rounded swelling; aperture strongly oblique to shell axis, round; boundary between inner and outer peristomes not conspicuous; inner peristome somewhat protruding, rather slim, outer peristome multi-layered, thickened and expanded, particularly toward umbilicus, but not reflected; umbilicus wide, slightly more than a third of shell width. Measurements. D: 2.22 – 2.68 mm, H: 1.31 – 1.56 mm. Operculum. Unknown.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F392421FF12B4E0FBBDFC7D.taxon	diagnosis	Differential diagnosis. Dicharax politus (W. T. Blanford, 1865) (examined material: Phungdo, Arakan, coll. Blanford, NHMUK 1906.4.4.178, 3 probable syntypes, Fig. 20 C) is larger, has a more elevated spire, and longer R 3. Dicharax (?) immaculatus Páll-Gergely, 2017 from Gansu Province, China is similar with low spire and smooth R 1, but with shorter R 2 without elevated ribs, and its peristome much thinner than that of new species. Some populations of D. cristatus (Möllendorff, 1886) possess a smooth R 1 and a low spire, but it has a longer R 2, and is always larger than the new species (Páll-Gergely et al. 2017).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F392421FF12B4E0FBBDFC7D.taxon	distribution	Distribution. This new species is known only from the type locality (Fig. 5).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F3B2426FF12B6E8FBAEFE99.taxon	description	Fig. 21	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F3B2426FF12B6E8FBAEFE99.taxon	materials_examined	Material examined. Holotype: Thailand, Mae Hong Son Province, 42 km NW of Pai, Road 1095, 19 ° 32 ’ N, 98 ° 32 ’ E, 750 m a. s. l., 20 Mar. 1988, K. Auffenberg leg., UF 345577 (D: 2.89 mm, H: 1.97 mm). Paratypes: 22 shells, same data as holotype, UF 551221. Other material: 9 juvenile shells, same data as holotype, UF 551222.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F3B2426FF12B6E8FBAEFE99.taxon	etymology	Etymology. The dorsal view of the shell of this new species looks like a face of a drunken (ebrius in Latin) man due to glossy R 3 swelling.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F3B2426FF12B6E8FBAEFE99.taxon	diagnosis	Diagnosis. A small Dicharax species with a finely ribbed R 1, glossy swelling on R 3 situated far from peristome, thickened outer peristome and slightly fringed inner peristome.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F3B2426FF12B6E8FBAEFE99.taxon	description	Description. Shell off-white to yellowish corneous, slightly translucent; shell outline slightly oval in dorsal view; spire slightly elevated, low conical, body whorl rounded; protoconch low, glossy, 1.25 – 1.5 whorls; R 1 of 1.75 – 2 whorls, finely, regularly ribbed from beginning, ribs gradually become more widely-spaced toward R 1 termination; boundary between R 1 and R 2 conspicuous due to dense R 2 ribs; R 2 ribs curved toward aperture, ca. 44 – 46 ribs present; R 2 ribs almost adnate; R 2 + R 3 slightly more than quarter whorl (ca. 100 ˚); R 2 approximately as long as R 3 or slightly shorter; boundary between R 2 and R 3 distinct due to change in sculpture and deep constriction; R 3 with prominent, but rounded, glossy swelling situated close to constriction; between swelling and peristome with few fine ribs, in some specimens this area slightly swollen, seemingly forming secondary swelling; aperture strongly oblique to shell axis, rounded with basal incision; boundary between inner and outer peristomes indistinct; inner peristome strong, protruding, expanded, basal and palatal portions slightly fringed, outer peristome expanded, and reflected toward umbilicus; peristome edges merge to form a multi-layered, thickened apertural lip; umbilicus narrow, approximately one fourth of shell width. Measurements. D: 2.65 – 2.89 mm, H: 1.85 – 1.97 mm. Operculum. Outer surface concave, yellowish corneous, with some traces of a deciduous lamina; inner surface with a small, pointed, low central nipple.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F3B2426FF12B6E8FBAEFE99.taxon	diagnosis	Differential diagnosis. The small shell size, blunt, but prominent, glossy swelling on R 3 makes this new species easy to recognize. Dicharax notus (Godwin-Austen, 1914) is similar in overall shell shape, but has blunter, less glossy R 3 swelling situated closer to peristome and a rounded aperture. Due to the weak traces of lower, secondary R 3 swelling and overall shell shape, this new species is similar to D. birugosus (Godwin-Austen, 1893) (syn. Alycaeus canaliculus Godwin-Austen, 1914, see Páll-Gergely et al. 2020) from the Khasi Hills, India. However, the Indian species has shorter R 2 and R 3, the R 2 ribs are more widely-spaced, and prominent secondary swelling.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F3B2426FF12B6E8FBAEFE99.taxon	distribution	Distribution. This new species is known only from the type locality (Fig. 17).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F3C2424FF12B3EDFE8FFC7D.taxon	description	Fig. 22	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F3C2424FF12B3EDFE8FFC7D.taxon	materials_examined	Material examined. Holotype: Thailand, Phrae Province, 4 km N of Ban Nim Rin, valley with limestone ridges, 18 ° 10.133 ’ N, 99 ° 56.233 ’ E, 200 m a. s. l., 16 May 1988, F. G. Thompson leg., leaf litter at base of boulders, UF 380401 (D: 1.65 mm, H: 1.15 mm).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F3C2424FF12B3EDFE8FFC7D.taxon	etymology	Etymology. The specific epithet gnomus (Medieval Latin for gnome) refers to the tiny shell that has unusual shell area ratios compared to most other species (to be used as a noun in apposition).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F3C2424FF12B3EDFE8FFC7D.taxon	diagnosis	Diagnosis. Tiny Dicharax species with wide, flat R 1 ribs and fine spiral striation, short R 2 with flat ribs, strongly reduced R 3, and circular aperture.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F3C2424FF12B3EDFE8FFC7D.taxon	description	Description. Shell light grey, slightly semi-translucent; shell outline rounded in dorsal view; spire slightly elevated, body whorl rounded; protoconch low, finely granular, glossy, 1.5 whorls; R 1 of 1.25 whorls, beginning with low ribs that appear curved anteriorly, developing into wide, flattened ribs separated by grooves, forming a smoothish, glossy surface; first whorl of R 1 with some fine spiral striation; boundary between R 1 and R 2 not conspicuous due to similar rib morphology, although ribs become denser, ca. 30 R 2 ribs present; R 2 + R 3 less than quarter whorl (ca. 70 – 80 ˚); R 3 considerably shorter than R 2; boundary between R 2 and R 3 distinct due to sculpture change and rather deep constriction; R 3 strongly reduced, represented only as a slight swelling just behind peristome; aperture almost parallel with shell axis, circular, round; peristome weak, very slightly thickened; boundary between inner and outer peristomes barely discernible; inner peristome weak, somewhat protruding and very slightly expanded, outer peristome very weakly developed, only indicated; umbilicus relatively narrow, less than one third of shell width. Measurements. D: 1.65 mm, H: 1.15 mm. Operculum. Unknown.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F3C2424FF12B3EDFE8FFC7D.taxon	diagnosis	Differential diagnosis. The R 3 region practically absent in this species. No other low-spired alycaeid species has this trait in Thailand or Myanmar. The Dicharax species described as Awalycaeus Kuroda, 1951 in Japan also have a strongly reduced R 3, but we consider this is a case of convergent evolution.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F3C2424FF12B3EDFE8FFC7D.taxon	distribution	Distribution. This new species is known only from the type locality (Fig. 17).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F3C2424FF12B3EDFE8FFC7D.taxon	discussion	Remarks. This species has unusual conchological characters, and therefore, its generic placement is unclear. The spiral striation on R 1 and the lack of spiral striation on the protoconch suggests a relationship to Chamalycaeus, whereas the glossy shell surface, the flattened R 2 ribs, and the low protoconch indicate a position in Dicharax. This study revealed other Dicharax species / populations that possess some spiral striae on R 1 (see under Dicharax admirandus, n. sp. and D. omissus and Páll-Gergely et al. 2017). Thus, the placement of this species in Dicharax is justified.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F3E2424FF12B6E8FBAEF84D.taxon	description	Fig. 23	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F3E2424FF12B6E8FBAEF84D.taxon	materials_examined	Material examined. Holotype: Thailand, Tak Province, Knoll 7 km E of Ban Huei Hin Fond, 16 ° 46.350 ’ N, 98 ° 40.383 ’ E, 610 m a. s. l., 02 May 1987, F. G. Thompson leg., UF 347421 (D: 3.13 mm, H: 2.08 mm).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F3E2424FF12B6E8FBAEF84D.taxon	etymology	Etymology. The R 3 shape of this species is reminiscent of the large proboscis of elephant seal males (genus Mirounga) (see Galimberti et al. 2019).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F3E2424FF12B6E8FBAEF84D.taxon	diagnosis	Diagnosis. A medium-sized Dicharax species with glossy R 1 having some widely-spaced, typical Dicharax - ribs, long R 3 with an elongated swelling, strongly expanded outer peristome, and narrow umbilicus.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F3E2424FF12B6E8FBAEF84D.taxon	description	Description. Shell white, with slightly yellowish colouration; shell outline oval in dorsal view; spire elevated, conical, body whorl rounded; protoconch low, glossy, slightly more than 1.25 whorls; R 1 of 1.75 whorls, glossy, with widely-spaced ribs, strong only near suture and in umbilicus, represented as very fine axial lines on edge of peristome; boundary between R 1 and R 2 conspicuous due to dense R 2 ribs; R 2 ribs curved toward aperture, ca. 28 R 2 ribs present; spaces between R 2 ribs narrower than ribs; R 2 + R 3 more than quarter whorl (ca. 110 ˚); R 2 approximately half length of R 3; R 3 glossy, smooth, with very fine growth lines; boundary between R 2 and R 3 conspicuous due to change in sculpture, although constriction shallow; entire R 3 with an elongated, inflated swelling reminiscent of an elephant seal proboscis; aperture strongly oblique to shell axis, round; boundary between inner and outer peristomes distinct; inner peristome somewhat protruding, rather thin, outer peristome extremely expanded, forming uniformly wide rim around aperture, slightly reflected over umbilicus, notched at the parieto-palatal edge; umbilicus very narrow. Measurements. D. 3.13 mm, H: 2.08 mm. Operculum. Unknown.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F3E2424FF12B6E8FBAEF84D.taxon	diagnosis	Differential diagnosis. There are no other species recorded from Thailand, Myanmar, and neighbouring areas with such an expanded outer peristome. This trait, combined with the relatively small size, elevated spire, and smooth R 1 makes this species easily recognizable.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F3E2424FF12B6E8FBAEF84D.taxon	distribution	Distribution. This new species is known only from the type locality (Fig. 17).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F3F241EFF12B380FDF7FB31.taxon	description	Figs 24 – 27	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F3F241EFF12B380FDF7FB31.taxon	materials_examined	Material examined. Lectotype (hereby designated): Myanmar, Shan State, “ Fort Stedman, Burma, coll. Woodthorpe ”, NHMUK 1903.7. 1.3065.1. Paralectotypes: 14 shells, same data as for lectotype, NHMUK 1903.7. 1.3065. Other Material: 1 shell (Fig. 25 B), Myanmar, Mandalay State, Anesakhan, Dat Taw Gyaint Waterfall, 21 ° 58.760 ’ N, 96 ° 23.116 ’ E, 610 m a. s. l. (locality code: 2018 / 65), 18 Oct. 2018, A. Hunyadi leg., HNHM 104863 (D: 3.08 mm, H: 1.76 mm); 5 shells + 4 juvenile shells, same data as preceding, HA — Shan State; 15 shells + 6 strongly eroded shells, Myanmar, Mandalay State, Hopong, Sam Phu, Cave Ae- 5 at ridge above village Ho Hwe, 20 ° 41.103 ’ N, 97 ° 16.198 ’ E (locality code: JG 3), 02 Feb. 2019, J. Grego leg., JG; 21 shells + 1 juvenile shell, Myanmar, Mandalay State, Hsihseng centre E ca. 4.5 km, left side of rd. + 1.7 km on unpaved rd., limestone rocks, 20 ° 9.359 ’ N, 97 ° 17.883 ’ E, 1140 m a. s. l. (locality code: 2018 / 39), 07 Oct. 2018, A. Hunyadi, K. Okubo & J. U. Otani leg., HA; 1 shell (photographed), Myanmar, Mandalay State, Hsihseng centre E ca. 6 km, right side of rd. + 400 m on unpaved rd., limestone hill, 20 ° 7.983 ’ N, 97 ° 18.145 ’ E, 1010 m a. s. l. (locality code: 2018 / 40), 07 Oct. 2018, A. Hunyadi, K. Okubo & J. U. Otani leg., HNHM 104864; 6 shells, same data as preceding, HA; 14 shells, Myanmar, Mandalay State, Hsihseng centre E ca. 6 km, right side of rd. + 500 m on unpaved rd., limestone hill, 20 ° 8.002 ’ N, 97 ° 18.024 ’ E, 1000 m a. s. l. (locality code: 2018 / 41), 07 Oct. 2018, A. Hunyadi, K. Okubo & J. U. Otani leg., HA; 1 shell + 1 juvenile shell, Taunggyi, hill over Aye Say Tee Pagoda, Dragon Cave, 20 ° 47.489 ’ N, 97 ° 3.036 ’ E, 1380 m a. s. l. (locality code: 2018 / 44), 08 Oct. 2018, A. Hunyadi, K. Okubo & J. U. Otani leg., HA; 1 shell, Myanmar, Kayah State, Demoso, Demoso spring lake (locality code: JG 109), 16 May 2019, J. Grego leg., JG; 8 shells + 3 broken / eroded shells, Myanmar, Kayah State, Hpruso Distr., Maw Thi Do Village, Bo Pha Gu Cave sink, 19 ° 21.121 ’ N, 97 ° 3.759 ’ E (locality code: JG 10), 10 Feb. 2019, J. Grego leg. JG; 33 shells + 13 juvenile shells, Myanmar, Kayah State, Demoso Distr., Han Li village, Lokalay Loku Gu Cave (Myaug a San Cave), 19 ° 23.468 ’ N, 97 ° 01.372 ’ E (locality code: JG 11), 11 Feb. 2019, J. Grego leg., JG; 1 shell, Myanmar, Kayah State, Hpruso Distr., Maw Thi Do, road towards Han Li village, under bridge over Phruno River, 19 ° 22.966 ’ N, 97 ° 02.153 ’ E (locality code: JG 100), 12 Dec. 2019, J. Grego leg., HNHM 104865; 7 shells + 6 juvenile / broken shells, same data as preceding, JG; 26 shells + 4 juvenile / broken shells, Myanmar, Kayah State, Hpruso Distr., Maw Thi Do, road towards Han Li Village, rocks above bridge over Phruno River, 19 ° 23.011 ’ N, 97 ° 02.108 ’ E (locality code: JG 101), 12 Dec. 2019, J. Grego leg., JG; 50 shells + 17 juvenile / broken shells, Myanmar, Kayah State, Hpruso Distr., Maw Thi Do Village, Entrance of Phruno River Cave, 19 ° 22.744 ’ N, 97 ° 02.570 ’ E (locality code: JG 102), 12 Dec. 2019, J. Grego leg., JG; 7 shells, Thailand, Chiang Rai Province, Doi Tung, 50 m before Wat Phra That Doi Tung, around the car park, 20 ° 19.540 ’ N, 99 ° 49.987 ’ E, 1350 m a. s. l. (locality code: 2015 / 17), 12 Feb. 2015, A. Hunyadi leg., HA; 1 shell, Thailand, Chiang Rai Province, Doi Tung, 20 ° 20.533 ’ N, 99 ° 50.350 ’ E, 1320 m a. s. l., 08 May 1987, F. G. Thompson leg., UF 347139.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F3F241EFF12B380FDF7FB31.taxon	diagnosis	Diagnosis. A variable Dicharax species with glossy shell, regularly ribbed R 1, widely-spaced R 2 ribs, blunt to somewhat narrow central swelling on R 3, and strongly reflected outer peristome.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F3F241EFF12B380FDF7FB31.taxon	description	Description. Shell yellowish, corneous, occasionally with some reddish / pinkish colouration on initial whorls, sometimes semi-translucent; shell outline rounded in dorsal view, spire rather low, only slightly elevated, but somewhat variable, body whorl rounded; protoconch low, glossy, 1.25 – 1.5 whorls; R 1 approximately 1.75 whorls, sculpture variable, first 1 – 1.25 whorl with weak growth lines or finely ribbed, gradually becoming widely-spaced, rather prominent ribs; boundary between R 1 and R 2 not conspicuous but discernible because R 2 ribs denser than R 1; R 2 ribs curved toward aperture, ca. 20 – 22 R 2 ribs present; R 2 ribs relatively widely spaced or densely arranged; R 2 + R 3 ca. quarter whorl or more (90 – 110 ˚); R 2 and R 3 of comparable lengths, or R 2 slightly shorter; R 3 beginning smooth, glossy, anterior portion (between swelling and peristome) smooth or with few ribs similar to end of R 1; boundary between R 2 and R 3 conspicuous due to sculptural change and rather deep constriction; middle of R 3 with prominent, variably shaped swelling, high convex or low rounded; aperture strongly oblique to shell axis, rounded; boundary between inner and outer peristomes barely discernible, inner peristome not protruding; outer peristome strongly thickened and reflected in all directions, even to parietal shell wall; umbilicus relatively wide, slightly less than a third of shell width. A cross-sectional view of R 2 was examined in one specimen (Fig. 27): anterior crust forms significantly elevated, sometimes hollow rib, with anterior and posterior projections of comparable sizes resulting in a T-shaped cross-section; anterior crust folds over much weaker posterior crust; posterior crust thick; cross-sectional view of microtunnel is nearly round; Measurements. D: 2.98 – 4.61 mm, H: 1.62 – 2.56 mm. Operculum. The outer surface of the opercula of two live-collected specimens could be examined: it has a closely-coiled lamina, similar to that of Dicharax species known from Vietnam (Páll-Gergely et al. 2017), although the lamina is seemingly deciduous since only remnants were visible.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F3F241EFF12B380FDF7FB31.taxon	diagnosis	Differential diagnosis. Dicharax (?) bifrons (Theobald, 1870) (examined material: Shan States, NHMUK 1888.12.4.956 – 958, 3 syntypes, Fig. 20 A) is rather dome-shaped, possesses two swellings on R 3, and its outer peristome is not much reflected. Dicharax (?) woodthorpei (Godwin-Austen, 1914) (examined material: Fort Stedman, Burma, coll. Woodthorpe, NHMUK 1903.7.1.3064, 22 syntypes in two vials, Fig. 20 B) is also somewhat similar to this species in terms of general shell and aperture shape, but it has a higher spire, a less convex swelling on R 2, and R 2 bears no elevated ribs, but has light and dark alternating colour stripes. Dicharax politus (W. T. Blanford, 1865) (examined material: Phungdo, Arakan, coll. Blanford, NHMUK 1906.4.4.178, 3 probable syntypes, Fig. 20 C) is similar to this species in terms of general shell shape and sculpture, but differs in having a less thickened peristome, less expanded outer peristome, denser R 2 ribbing, and glossier R 1. See also under Dicharax omissus.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F3F241EFF12B380FDF7FB31.taxon	distribution	Distribution. Dicharax notus is known from the Mandalay, Shan, and Kayah states of Myanmar, and from Chiang Rai Province, Thailand (Fig. 28).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F3F241EFF12B380FDF7FB31.taxon	discussion	Remarks. This is a variable species in terms of shell size, R 1 sculpture, density of R 2 ribs, and the narrow width of the R 3 swelling. See Table 2. The Japanese Metalycaeus vinctus (Pilsbry, 1902) and M. minatoi Páll-Gergely, 2017 are probably sister species to each other with ribbed and smooth R 2 s, respectively (Páll-Gergely & Asami 2017). Dicharax notus and D. woodthorpei may present a similar situation, because they are very similar in shell shape and have geographically adjacent distributions, differing mostly in the fine morphology of the R 2 ribs, indicating that the character may be the subject of rapid evolutionary changes.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F05241DFF12B589FD88FDCE.taxon	description	Figs 29 – 30	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F05241DFF12B589FD88FDCE.taxon	materials_examined	Material examined. Lectotype (hereby designated): 1 shell, “ Siam & Shan boundary, coll. Woodthorpe ”, NHMUK 1903.7. 1.1228.1. Paralectotype: 1 shell, same data as for lectotype, NHMUK 1903.7. 1.1228. Other material: 1 shell (Fig. 29 B, D: 2.94 mm, H: 1.94 mm) + 10 shells, Myanmar, Shan State, Hopong centre 5.8 km — Namsang, left side of rd. # 4, near „ Hopong Spring Cave ”, 20 ° 49.028 ’ N, 97 ° 13.469 ’ E, 1110 m a. s. l. (locality code: 2018 / 35), 06 Oct. 2018, A. Hunyadi, K. Okubo & J. U. Otani leg., HA; 8 shells, Myanmar, Shan State, Taunggyi centre ca. 16 km — Hopong, rd. # 4 N 1.5 km, „ Shwe Pyi Aunchonda ” monastery, 20 ° 47.263 ’ N, 97 ° 8.239 ’ E, 1110 m a. s. l. (locality code: 2018 / 46), 08 Oct. 2018, A. Hunyadi, K. Okubo & J. U. Otani leg., HA; 1 shell (Fig. 30 A, D: 2.81 mm, H: 1.8 mm) + 7 shells, Thailand, Chiang Mai Province, 1.2 km N of Wat Tham Klaeb, 500 m side road from the junction, 19 ° 33.918 ’ N, 99 ° 3.690 ’ E, 650 m a. s. l. (locality code: 2015 / 08), 08 Feb. 2015, A. Hunyadi leg., HA; 1 shell (Fig. 30 B, D: 2.69 mm, H: 1.75 mm) + 2 shells, Thailand, Mae Hong Son Province, 800 m from Tham Nam Lod toward the Buddhist temple, 19 ° 33.928 ’ N, 98 ° 16.794 ’ E, 650 m a. s. l. (locality code: 2015 / 12), 09 Feb. 2015, A. Hunyadi leg., HA. Type locality. “ Siam and Shan boundary ”.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F05241DFF12B589FD88FDCE.taxon	diagnosis	Diagnosis. A variable Dicharax species with a relatively densely, regularly ribbed R 1, rarely with spiral striation, dense R 2 ribs, a rounded central swelling on R 3, and an expanded, but not reflected outer peristome.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F05241DFF12B589FD88FDCE.taxon	description	Measurements. D: 2.52 – 2.87 mm, H: 1.71 – 1.89 mm (2018 / 08 and 2015 / 12); D: 2.89 – 3.23 mm, H: 1.88 – 2.3 mm (2018 / 35 and 2015 / 46).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F05241DFF12B589FD88FDCE.taxon	diagnosis	Differential diagnosis. Typical Dicharax notus and D. omissus are very similar, and since only eroded specimens of the latter species were available, the distinction of these two species was somewhat questionable until newly collected material became available. These specimens reveal that D. notus is generally larger, R 1 bears denser ribs (although some populations of D. notus have less dense R 2 ribs), and its outer peristome is more reflected.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F05241DFF12B589FD88FDCE.taxon	distribution	Distribution. Dicharax omissus is known from the Shan States of Myanmar and neighbouring Thai provinces of Chiang Mai and Mae Hong Son (Jirapatrasilp et al. 2021; Fig. 28).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F05241DFF12B589FD88FDCE.taxon	discussion	Remarks. The syntypes and the newly collected 4 samples (possibly 3 populations) show some variability in terms of the sculpture of R 1 and R 3, and the presence or absence of the notch at the basal portion of the aperture. However, all of these traits are rather minor and do not justify the distinction of species. The conchological differences of these populations are summarized in Table 3. The shells of the two samples from Hopong have some slight spiral striation on the R 1, which is highly unusual in Dicharax. The spiral striae of this species might not be homologous with those of other alycaeid genera, although this requires further investigation.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F082412FF12B589FBBDFB1D.taxon	description	Fig. 31 A	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F082412FF12B589FBBDFB1D.taxon	materials_examined	Material examined. Holotype: Thailand, Surat Thani Province, Highway 401, 2.7 km W junction Highway 4142 and 401, evergreen forest on rocky hillside, 9 ° 10 ’ N, 99 ° 40 ’ E, 90 m a. s. l., 19 April 1988, K. Auffenberg leg., below cliff, base of cliff, UF 345330 (D: 1.86 mm, H: 0.99 mm). Paratypes: 5 shells, same data as holotype, UF 551223. Other Material: 3 broken shells, same data as holotype, UF 551224.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F082412FF12B589FBBDFB1D.taxon	etymology	Etymology. Named for the ovate (Latin derivation: ovatus) shell shape (in dorsal view).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F082412FF12B589FBBDFB1D.taxon	diagnosis	Diagnosis. A tiny Dicharax species with depressed shell, finely ribbed R 1, R 2 + R 3 less than a quarter whorl, R 2 with slightly curved ribs shorter than R 2.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F082412FF12B589FBBDFB1D.taxon	description	Description. Shell off-white, light yellowish; shell outline oval in dorsal view, spire depressed, nearly flat; body whorl rounded; protoconch low, rather glossy, 1.25 – 1.5 whorls; R 1 of 1.25 whorls; R 1 with fine, regular, dense ribs, gradually become more widely-spaced and elevated toward R 1 termination; boundary between R 1 and R 2 not conspicuous but discernible due to denser R 2 ribs than those of R 1; R 2 with low ribs that slightly curve toward aperture, ca. 24 – 26 R 2 ribs present; R 2 + R 3 less than quarter whorl (ca. 80 ˚); R 2 slightly shorter than R 3; R 3 with low ribs of similar density to those on R 1; boundary between R 2 and R 3 clear due to change in rib density and shallow constriction; very low swelling on R 3 only indicated; aperture slightly oblique to shell axis, rounded; boundary between inner and outer peristomes distinct; inner peristome thin, slightly protruding and expanded, rather narrow, outer peristome thicker than inner peristome, strongly expanded but not reflected; umbilicus wide, sigmoid, more than a third of shell width. Measurements. D: 1.81 – 1.95 mm, H: 0.93 – 1.1 mm. Operculum. unknown.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F082412FF12B589FBBDFB1D.taxon	diagnosis	Differential diagnosis. The most similar species regarding size and the dense, fine ribbing is the nearby occurring Chamalycaeus canaliculatus (examined material: Golf von Siam: Koh-Samui, coll. Möllendorff, SMF 109468, lectotype), which has a more pronounced constriction at border of R 2 and R 3, and a narrower swelling on R 3. Chamalycaeus armillatus (Benson, 1856) has a more elevated spire, the first 1 – 1.5 whorls of the teleoconch are smooth and glossy, and the last whorl of R 1 has spiral striation. Dicharax parvulus (Möllendorff, 1887) (examined material: Malakka: Bukit Pondong (Perak), coll. Möllendorff, SMF 109507, lectotype, Fig. 31 B) is smaller, has a comparatively larger aperture, a more thickened peristome, and a shorter R 2.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F082412FF12B589FBBDFB1D.taxon	distribution	Distribution. This new species is known only from the type locality (Fig. 3).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F082412FF12B009FF58F8D8.taxon	description	Fig. 32	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F082412FF12B009FF58F8D8.taxon	materials_examined	New material examined. 7 shells, Thailand, Chiang Rai Province, Doi Thung; 20 ° 20.533 ’ N, 99 ° 50.350 ’ E, 1320 m a. s. l., 08 May 1988, F. G. Thompson leg., UF 347150. Type locality. Thailand, Chiang Rai Province, Doi Tung, 50 m before Wat Phra That Doi Tung, around the car park, 20 ° 19.540 ’ N, 99 ° 49.987 ’ E, 1350 m a. s. l. Operculum. The operculum was not described in the original description, but a live-collected specimen from the Florida Museum allows description. The operculum is extremely thin, translucent, concave; outer surface glossy, without any notable structures, inner surface with a low but pointed, central nipple.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F082412FF12B009FF58F8D8.taxon	discussion	Remarks. The newly examined specimens were collected very near the type locality of D. stuparum (Fig. 17).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F082410FF12B24FFBBDFE09.taxon	description	Fig. 33	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F082410FF12B24FFBBDFE09.taxon	materials_examined	Material examined. Holotype: Thailand, Surat Thani Province, limestone hill right side off km. 105.2 rd. # 401 (Suratthani — Takuapa), 8 ° 53.622 ’ N, 98 ° 33.179 ’ E (locality code: Th. 119.00), Ch. & J. Hemmen leg., SMF 360000 (D: 2.79 mm, H: 1.83 mm). Paratype: 1 shell, same data as holotype, SMF 360001.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F082410FF12B24FFBBDFE09.taxon	etymology	Etymology. The specific epithet refers to the slight pinkish coloration of the available specimens.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F082410FF12B24FFBBDFE09.taxon	diagnosis	Diagnosis. A Dicharax species with a depressed shell, slight pinkish colouration, very widely spaced ribs on the last whorl of R 1, a short R 2 with straight, widely-spaced ribs, and a comparatively small, rounded aperture.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F082410FF12B24FFBBDFE09.taxon	description	Description. Shell whitish-yellowish with some pinkish colouration (holotype at the R 2 - R 3 area, paratype on initial whorls); shell outline rounded in dorsal view; spire only slightly elevated; body whorl rounded; protoconch low, rather glossy, 1.25 – 1.5 whorls; R 1 of 1.75 – 2 whorls; first whorl of R 1 with only weak growth lines, gradually changing to strongly ribbed, spaces between ribs increase width rapidly (last half whorl of R 1 with only 15 – 20 ribs); ribs straight, strong, narrow, but relatively low; boundary between R 1 and R 2 distinct due to denser R 2 ribs; R 2 with ca. 16, widely-spaced, low and rather narrow; R 2 + R 3 approximately a quarter whorl or slightly more; R 2 slightly shorter than R 3; boundary between R 2 and R 3 conspicuous due to smooth R 3 beginning and rather deep constriction; R 3 with widely-spaced ribs, only anterior to constriction similar to those on R 1, and blunt but prominent central swelling; aperture strongly oblique to shell axis, rounded, conspicuously small compared to shell size; boundary between inner and outer peristomes distinct; inner peristome prominent, protruding, relatively narrow; outer peristome as strong as inner peristome, strongly expanded, slightly reflected; umbilicus wide, approximately a third of shell width. Measurements. D: 2.74 – 2.79 mm, H: 1.7 – 1.83 mm. Operculum. The holotype possesses an operculum in the original position which appears to be separated into two layers. The outer layer, being partly broken, allows examination of the more deeply set layer. The outer surface is glossy without any surface structures. A central nipple is visible through the semi-translucent inner layer.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F082410FF12B24FFBBDFE09.taxon	diagnosis	Differential diagnosis. The combination of small size, widely-spaced R 1 ribs, and a relatively small aperture is unlike any other species in the region.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F082410FF12B24FFBBDFE09.taxon	distribution	Distribution. This new species is known only from the type locality (Fig. 3).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F0B2416FF12B589FD70FC11.taxon	description	Fig. 34	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F0B2416FF12B589FD70FC11.taxon	materials_examined	Material examined. Holotype: Myanmar, Shan State, Taunggyi, hill above Aye Say Tee Pagoda, Dragon Cave, 20 ° 47.489 ’ N, 97 ° 03.036 ’ E, 1380 m a. s. l. (locality code: 2018 / 44), 08 Oct. 2018, A. Hunyadi, K. Okubo & J. U. Otani leg., HNHM 104862 (holotype: D: 2.83 mm, H: 2.14 mm). Paratypes: 10 shells, same data as holotype, HA. Other material: 2 shells, Myanmar, Shan State, Hsihseng centre E ca. 6 km, right side of rd. + 400 m on unpaved rd., limestone hill, 20 ° 7.983 ’ N, 97 ° 18.145 ’ E, 1010 m a. s. l. (locality code: 2018 / 40), 07 Oct. 2018, A. Hunyadi, K. Okubo & J. U. Otani leg., HA.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F0B2416FF12B589FD70FC11.taxon	etymology	Etymology. The specific epithet verrucosus (Latin for excrescence / wart) refers to the small swelling on R 3, reminiscent of a wart.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F0B2416FF12B589FD70FC11.taxon	diagnosis	Diagnosis. A Dicharax species with a relatively high spire, glossy, smooth R 1, R 2 + R 3 about a quarter whorl (R 3 shorter than R 2), and blunt swelling on R 3.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F0B2416FF12B589FD70FC11.taxon	description	Description. Shell light grey, possibly translucent when fresh; shell outline rounded in dorsal view, spire elevated, body whorl rounded; protoconch low, rather glossy, 1.5 whorls; R 1 approximately 2.25 whorls, smooth, glossy, with occasional, weak growth lines; boundary between R 1 and R 2 conspicuous due to dense R 2 ribs; R 2 ribs curved toward aperture, ca. 36 – 38 R 2 ribs present; R 2 ribs situated close to each other, spaces between ribs much less than rib width; R 2 + R 3 covers ca. quarter whorl; R 3 very short; about half as long as R 2; sculpture of R 3 similar to R 1, boundary between R 2 and R 3 distinct, indicated by deep constriction; middle of R 3 with prominent, blunt swelling; aperture slightly oblique to shell axis; rounded, with slight upper incision in parieto-palatal region; boundary between inner and outer peristomes distinct, especially at columellar portion; inner peristome slim, slightly protruding, outer peristome strongly thickened, expanded and reflected, especially toward umbilicus (in sample 2018 / 40 more conspicuously than in 2018 / 44); umbilicus relatively narrow, less than a third of shell width. Measurements. D: 2.76 – 3.24 mm, H: 2.1 – 2.46 mm (2018 / 44); D: 3.21 – 3.55 mm, H: 2.46 – 2.5 mm (2018 / 40). Operculum. Unknown.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F0B2416FF12B589FD70FC11.taxon	diagnosis	Differential diagnosis. The combination of the elevated spire, smooth R 2, short R 3 with a prominent swelling distinguishes this species from all congeners.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F0B2416FF12B589FD70FC11.taxon	distribution	Distribution. Both known populations occur in the southeastern part of Shan States, Myanmar. The straightline distance between the two is ca. 77 km (Fig. 17).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F0B2416FF12B589FD70FC11.taxon	discussion	Remarks. The typical shells from Taunggyi are slightly smaller, and have less reflected peristomes than specimens from Hsihseng. However, since all other shell characters (shape, ratios of the shell regions, and shell sculpture) are identical, we treat them as the same species. Genus Metalycaeus Pilsbry, 1900 Alycaeus (Metalycaeus) Pilsbry, 1900: 382. Type species. Alycaeus (Metalycaeus) melanopoma Pilsbry, 1900 (synonym of Alycaeus nipponensis Reinhardt, 1877, see Minato 1988), by subsequent designation (Thiele 1929). Diagnosis. Metalycaeus is characterized by the presence of spiral striation on both the protoconch and teleoconch (Páll-Gergely et al. 2017, 2020). However, some shells of one Japanese species lacks spiral striation on the protoconch. Here we report a species (M. pygmachos n. sp.) lacking spiral striation on the entire shell, but based on other characters, is best assigned to this genus. Thus, examination of shell characters other than the spirally striated protoconch is also necessary.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F0B2416FF12B589FD70FC11.taxon	discussion	Remarks. The typical shells from Taunggyi are slightly smaller, and have less reflected peristomes than specimens from Hsihseng. However, since all other shell characters (shape, ratios of the shell regions, and shell sculpture) are identical, we treat them as the same species.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F0B2416FF12B589FD70FC11.taxon	type_taxon	Type species. Alycaeus (Metalycaeus) melanopoma Pilsbry, 1900 (synonym of Alycaeus nipponensis Reinhardt, 1877, see Minato 1988), by subsequent designation (Thiele 1929).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F0B2416FF12B589FD70FC11.taxon	diagnosis	Diagnosis. Metalycaeus is characterized by the presence of spiral striation on both the protoconch and teleoconch (Páll-Gergely et al. 2017, 2020). However, some shells of one Japanese species lacks spiral striation on the protoconch. Here we report a species (M. pygmachos n. sp.) lacking spiral striation on the entire shell, but based on other characters, is best assigned to this genus. Thus, examination of shell characters other than the spirally striated protoconch is also necessary.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F0C2415FF12B0F7FBAEF87D.taxon	description	Figs 35 – 36	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F0C2415FF12B0F7FBAEF87D.taxon	materials_examined	Material examined. Holotype: Thailand, Chiang Rai Province, Wat Phra That Tham Doi Khong Khao Meditation Centre, ca. 8 km W of Chiang Rai, cave clay at entrance of cave, 19 ° 54.771 ’ N, 99 ° 46.607 ’ E, ca. 420 m a. s. l. (locality code: Th 6), Sep. 2007, A. Reischütz leg., NHMW 113489 (D: 3.3 mm, H: 2.05 mm). Paratypes: 1 shell (fig. 35 B), same data as holotype, NHMW 113490; 7 shells, same data as holotype, RE; 2 shells, same data as holotype, CUMZ 5270. Other Material: 4 broken shells, same data as holotype, RE.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F0C2415FF12B0F7FBAEF87D.taxon	etymology	Etymology. The specific epithet (noun in apposition) derives from the shape of R 3 that resembles a duck’s beak (anas = duck in Latin).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F0C2415FF12B0F7FBAEF87D.taxon	diagnosis	Diagnosis. A medium-sized Metalycaeus species with a weakly sculptured R 1 (fine growth lines and spiral striae), almost smooth R 2, a R 3 with convex, central swelling and a rounded aperture.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F0C2415FF12B0F7FBAEF87D.taxon	description	Description. Shell light grey, probably somewhat translucent when fresh; shell outline slightly oval in dorsal view, spire somewhat elevated, low conical; body whorl rounded (rather “ parabolic ”); protoconch conspicuously large to shell size, somewhat elevated, last whorl spirally striated, 1.5 whorls; R 1 of 1.25 whorls, with very weak ribs (growth lines) occasionally moderately strong near suture, and fine spiral striations; boundary between R 1 and R 2 distinct due to R 2 axial sculpture; R 2 with narrow, somewhat elevated light-coloured stripes alternating with broader, darker stripes, ca. 24 – 32 R 2 stripes present; R 2 + R 3 approximately or slightly more than quarter whorl; R 2 slightly shorter than R 3; R 3 practically smooth, bearing only low growth lines; boundary between R 2 and R 3 distinct due to lack of R 3 light stripes (ribs) and rather deep constriction; R 3 with elevated, convex central swelling with anterior slope steeper than posterior; aperture strongly oblique to shell axis; round; boundary between inner and outer peristomes distinct; inner peristome protruding, expanded, rather slim; outer peristome of comparable strength, expanded, and slightly reflected toward umbilicus; umbilicus relatively narrow, approximately one fourth the shell width. A cross-sectional view of R 2 of one specimen was examined (Fig. 36 E – G): anterior crust very slightly elevated, forming very low, blunt rib; anterior crust and the slightly weaker posterior crust adnate, forming slender microtunnel in cross-sectional view. Measurements. D: 3.3 – 3.91 mm, H: 2.05 – 2.26 mm. Operculum. unknown.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F0C2415FF12B0F7FBAEF87D.taxon	diagnosis	Differential diagnosis. Metalycaeus rubinus (Godwin-Austen, 1893) (examined material: Ruby Mines District, Up. Burma, leg. Doherty, NHMUK 1903.7.1.2685, 2 syntypes, Fig. 37) is the only Metalycaeus species reported from Myanmar having atypical R 2 ribbing. It is far larger and has an elevated spire.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F0C2415FF12B0F7FBAEF87D.taxon	distribution	Distribution. This new species is known only from the type locality (Fig. 17).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F102409FF12B589FEE7FBEC.taxon	description	Figs 38, 39 A, 40 A – D	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F102409FF12B589FEE7FBEC.taxon	materials_examined	Material examined. Holotype: Thailand, Loei Province, limestone knoll 1 km E Ban Huai Muang, top of knoll, 17 ° 28.433 ’ N, 101 ° 53.333 ’ E, 375 m a. s. l., 21 May 1987, F. G. Thompson leg., in leaf litter, UF 343786 (D: 2.48 mm, H: 1.49 mm). Paratypes: 12 shells, same data as holotype, UF 551225. Other material: 1 shell (Fig. 38 B, D: 2.2 mm, H: 1.61 mm) + 5 shells + 4 juvenile shells, Thailand, Nakhon Sawan Province, limestone hill 4.0 km NW of Man Non San, 15 ° 56.517 ’ N, 99 ° 52.400 ’ E, 100 m a. s. l., 12 Jun. 1987, F. G. Thompson leg., UF 279699; 1 shell (Fig. 38 C, D: 2.04 mm, H: 1.25 mm) + 3 shells, Thailand, Phetchabun Province, 24.4 km S Phetchabun on Highway 2275 evergreen forest in back of outcrop, base of cliff, 16 ° 16 ’ N, 101 ° 10 ’ E, 120 m a. s. l., 02 May 1988, K. Auffenberg leg., UF 346105.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F102409FF12B589FEE7FBEC.taxon	etymology	Etymology. The shape of the shell of this new species is reminiscent of a coiled ram’s horn (aries in Latin). The specific epithet is used as a noun in apposition.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F102409FF12B589FEE7FBEC.taxon	diagnosis	Diagnosis. A tiny Metalycaeus species with strongly ribbed R 1, short combined R 2 + R 3, dense, curved R 2 ribs, and an angled R 2 visible from dorsal side.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F102409FF12B589FEE7FBEC.taxon	description	Description of the type series. Shell off-white, light grey, although all available shells dead collected; shell outline slightly oval in dorsal view, R 2 conspicuously swollen / angled; spire somewhat elevated, low conical; body whorl rounded with distinct lower, keel caused by angled ribs; protoconch of 1.5 whorls, elevated, first ca. 0.75 whorl finely granular and glossy, the last 0.75 whorl spirally striated; R 1 of 1.5 – 1.75 whorls, with strong, equidistant, narrow ribs and fine, dense spiral striae between ribs; boundary between R 1 and R 2 distinct due to densely-placed R 2 ribs; R 2 + R 3 less than a quarter whorl (ca. 70 ˚); R 2 swollen, forming blunt angle in dorsal view, and rounded in apertural view; R 2 ribs curved toward aperture, nearly adnate, ca. 24 – 26 R 2 ribs present; R 2 and R 3 of comparable length; R 3 with spiral striation and weaker ribs than R 1; boundary between R 2 and R 3 distinct due to lower R 3 ribs and descending portion of R 2 angle; aperture slightly oblique to shell axis, round; boundary between inner and outer peristomes distinct; inner peristome protruding and expanded, relatively thin; outer peristome thicker, wider, expanded, and very slightly reflected toward umbilicus; umbilicus moderately narrow, approximately one third of shell width. Measurements. 2.04 – 2.54 mm, H: 1.3 – 1.61 mm. Operculum. An operculum was present in one specimen (UF 279699). Outer surface multispiral without elevated lamina, originally covered with mud (a portion left on operculum for photography; Fig. 38 B), inner surface with very small, low, central nipple.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F102409FF12B589FEE7FBEC.taxon	diagnosis	Differential diagnosis. The only similar species is M. bicarinatus n. sp., which has two distinct keels in R 2, has slimmer, more elevated, and more widely-spaced ribs, and the protoconch lacks obvious spiral striae.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F102409FF12B589FEE7FBEC.taxon	distribution	Distribution. Three populations are known from the Thai provinces of Loei, Nakhon Sawan, and Phetchabun (Fig. 5).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F102409FF12B589FEE7FBEC.taxon	discussion	Remarks. The three populations classified as M. aries n. sp. show some conchological variability, which is compiled in Table 4.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F13240EFF12B069FB0DFD5E.taxon	description	Figs 39 B, 40 E – H	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F13240EFF12B069FB0DFD5E.taxon	materials_examined	Material examined. Holotype: Thailand, Phetchaburi Province, Khao Yoi, NE side of the hill, 13 ° 14.316 ’ N, 99 ° 49.493 ’ E, 30 m a. s. l. (locality code: 2015 / 39), 24 Feb. 2015, A. Hunyadi leg. HNHM 104868 (D: 2.33 mm, H: 1.63 mm). Paratype: 1 shell, same data as holotype, HA. Other Material: 4 juvenile / broken shells; same data as holotype, HA.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F13240EFF12B069FB0DFD5E.taxon	etymology	Etymology. This new species is named after the most remarkable conchological character, the double keel of R 2.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F13240EFF12B069FB0DFD5E.taxon	diagnosis	Diagnosis. A tiny Metalycaeus species with a strongly ribbed R 1, ribs narrow and elevated, short R 2 + R 3, dense, curved R 2 ribs, and an R 2 with double keel, visible from lateral and ventral sides. Differential diagnosis. Due to the peculiarly carinated R 2 no other alycaeid species is similar, except for M. aries n. sp. (see above).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F13240EFF12B069FB0DFD5E.taxon	description	Description of the type series. Shell off-white, light grey, although available shells were dead collected; shell rounded in dorsal view; spire elevated, low conical; R 1 body whorl rounded, R 2 has distinct upper and lower keels separated by flattened zone, resulting in rectangular-shaped R 2; protoconch consisting of 1.25 whorls, elevated, finely granular, last ca. quarter whorl with granules arranged into oblique striae; R 1 of 1.75 whorls, with strong, narrow, elevated, equidistant ribs, areas between ribs with fine, dense spiral striae; boundary between R 1 and R 2 distinct because of denser R 2 ribs; R 2 + R 3 less than a quarter whorl (ca. 80 ˚); R 2 very slightly swollen in dorsal view, and rectangular in lateral view due to upper and lower keels; R 2 ribs curved toward aperture, nearly adnate, ca. 26 R 2 ribs present; R 2 somewhat longer than R 3; R 3 strongly reduced, with remnants of spiral striation and rough growth lines; R 2 and R 3 boundary distinct due to change in rib morphology and moderate constriction; aperture slightly oblique to shell axis; round; boundary between inner and outer peristomes barely discernible; inner peristome slightly protruding and expanded, relatively thin; outer peristome also thin, slightly expanded, not reflected; umbilicus moderately narrow, approximately one third of shell width. Measurements. D: 2.33 – 2.4 mm, H: 1.63 – 1.87 mm. Operculum. Unknown.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F13240EFF12B069FB0DFD5E.taxon	distribution	Distribution. This new species is known only from the type locality (Fig. 5).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F13240EFF12B069FB0DFD5E.taxon	discussion	Remarks. This species is assigned to Metalycaeus even though no spiral striation has been found on the protoconch. Since all available material was dead collected, we believe that the fine shell sculpture that exists between the ribs at the end of the protoconch is probably remnants of spiral striation. Also, the protoconch of Metalycaeus aries n. sp., the conchologically most similar species is clearly spirally striated (see above).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F14240EFF12B7C8FED8F8EE.taxon	description	Fig. 41	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F14240EFF12B7C8FED8F8EE.taxon	materials_examined	Material examined. Holotype: Myanmar, Shan State, “ Kengtung, Siam Frontier, leg. Woodthorpe ”, NHMUK 1903.7. 1.3037 (holotype of A. kengtungensis). For the examined type specimens of other taxa see Páll-Gergely et al. (2017). Other material: 4 shells, Myanmar, Kayah State, Hpruso Distr., Maw Thi Do Village, Bo Pha Gu Cave sink, 19 ° 21.121 ’ N, 97 ° 3.759 ’ E (locality code: JG 10), 10 Feb. 2019, J. Grego leg., JG; 2 shells, Myanmar, Kayah State, Hpruso Distr., Maw Thi Do, road towards Han Li Village, under bridge over Phruno River, 19 ° 22.966 ’ N, 97 ° 02.153 ’ E (locality code: JG 100), 12 Dec. 2019, J. Grego leg., JG; 1 shell, Myanmar, Kayah State, Hpruso Distr., Maw Thi Do, road towards Han Li Village, rocks above bridge over Phruno River, 19 ° 23.011 ’ N, 97 ° 02.108 ’ E (locality code: JG 101), 12 Dec. 2019, J. Grego leg., JG; 1 shell, Myanmar, Kayah State, Hpruso Distr., Maw Thi Do Village, Entrance of Phruno River Cave, 19 ° 22.744 ’ N, 97 ° 02.570 ’ E (locality code: JG 102), 12 Dec. 2019, J. Grego leg., HNHM 104867 (photographed shell: 4.77 mm, H: 3.66 mm); 17 shells, same data as preceding, JG.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F14240EFF12B7C8FED8F8EE.taxon	diagnosis	Diagnosis of specimens from Myanmar. A medium-sized to large species with very long R 2 having typical Metalycaeus - type elevated, thin, widely spaced ribs, a narrow swelling on R 3, and a rounded aperture.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F14240EFF12B7C8FED8F8EE.taxon	description	Measurements. D: 4.65 – 5.05 mm, H: 3.66 – 3.71 mm (newly collected specimens).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F14240EFF12B7C8FED8F8EE.taxon	discussion	Remarks. Metalycaeus heudei is a widely distributed, variable species reported from China, Vietnam and Laos. Alycaeus kengtungensis (Fig. 41 B) from the eastern border of Shan State, Myanmar, and the newly collected shells from Kayah State (Fig. 5) fall within the known morphological variation of Metalycaeus heudei. Therefore, we consider A. kengtungensis a new synonym of M. heudei. With this, M. heudei is the most widely distributed alycaeid species known.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F16240DFF12B589FE92FC10.taxon	description	Fig. 42	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F16240DFF12B589FE92FC10.taxon	materials_examined	Material examined. Holotype: Myanmar, Mandalay State, near Maha Nandamu Peik Chin Myaung Cave & Waterfall, 22 ° 05.762 ’ N, 96 ° 37.087 ’ E, 740 m a. s. l. (locality code: 2018 / 66), 19 Oct. 2018, A. Hunyadi leg., HNHM 104866 (D: 4.8 mm, H: 2.91 mm). Paratype: 1 shell, same data as holotype, HA.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F16240DFF12B589FE92FC10.taxon	etymology	Etymology. The specific epithet pygmachos (πυγμαχος: Greek for boxer) refers to the peculiarly shaped R 3 resembling a boxer’s face after being punched.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F16240DFF12B589FE92FC10.taxon	diagnosis	Diagnosis. A large Metalycaeus species with a long, swollen R 2 having widely-spaced slim ribs, and a complex, conspicuous swelling on R 3.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F16240DFF12B589FE92FC10.taxon	description	Description. Shell off-white, although both available shells are eroded; shell outline oval in dorsal view due to conspicuously swollen R 2; spire slightly elevated, domed; body whorl rounded; protoconch low, finely granular, spiral striae lacking, 1.25 whorls; R 1 of 1.75 – 2 whorls, with strong, elevated, equidistant ribs and some faint, dense spiral striation; boundary between R 1 and R 2 not conspicuous because R 1 and R 2 ribs identical in shape and density; R 2 with ca. 30 – 32 R 2 ribs; R 2 + R 3 almost half whorl (ca. 150 – 160 ˚); R 2 slightly longer than R 3; boundary between R 2 and R 3 indicated with very deep constriction; R 3 characterized by very complex swelling strongly depressed dorsolaterally; this elongate depression parallel to constriction, corresponds with irregularly swollen ridge visible in oblique view in aperture; depression results in a U-shaped R 3 swelling; upper rim of U, visible in dorsal view, more swollen than narrower lower rim, visible in lateral view; additional small depression just anterior to upper swollen rim, i. e. between upper, swollen rim and peristome, corresponding to a separate, knob-like swelling inside aperture; aperture strongly oblique to shell axis, rounded with slight upper and lower incisions; boundary between inner and outer peristomes barely discernible, inner peristome slightly protruding and expanded, outer peristome much stronger, wider, expanded, and reflected toward umbilicus, wide distance between inner peristome and edge of outer peristome resulting in strongly thickened lip; umbilicus relatively narrow, approximately less than one third of shell width. Measurements. 4.8 – 5.1 mm, H: 2.91 – 2.98 mm. Operculum. Unknown.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F16240DFF12B589FE92FC10.taxon	diagnosis	Differential diagnosis. The peculiarly shaped R 3 swelling is a unique feature distinguishing this species from all Chamalycaeus and Metalycaeus species.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F16240DFF12B589FE92FC10.taxon	distribution	Distribution. This new species is known only from the type locality (Fig. 5).	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F16240DFF12B589FE92FC10.taxon	discussion	Remarks. This species is classified in the genus Metalycaeus due to its large size and the “ depressed swelling ” on R 3 as in other Metalycaeus species. The lack of spiral striation on the protoconch is considered a secondary reduction. Concluding remarks Nineteen new species are described here (Fig. 43) collected by a few field biologists at a few isolated localities in Myanmar and Thailand, suggesting that still many undescribed species await discovery in Southeast Asia’s limestone outcrops. Biogeography In addition to the description of the new species and new data on the morphology and distribution of a few known species, this paper presents the following conclusions: - The Metalycaeus species described here represent the first records of that genus in Thailand, and the southernmost localities for Metalycaeus (Fig. 44; Páll-Gergely et al. 2020). - The numerous Chamalycaeus species recorded from southern Thailand indicate a diversity centre for the genus. - The numerous Dicharax species from northern Thailand clearly show that this genus retains high diversity continuously from the southeastern Himalaya east into northern Thailand. In contrast, the genus is much less diverse in neighbouring northern Laos (Fig. 44). - The distribution of Metalycaeus heudei ranges from Kayah Province, Myanmar east and north to Hubei Province, China, covering a distance of more than 1800 km. Metalycaeus heudei has the largest distributional area of all alycaeids and one of the largest among all terrestrial caenogastropods. - For Dicharax diminutus (Heude, 1885) and D. dolichodeiros (Heude, 1890), populations were found several hundred kilometres apart from each other without material from intervening areas (see Páll-Gergely et al. 2017). A similar situation for Chamalycaeus irmatallus n. sp. is reported here. Four hypotheses were outlined by Páll-Gergely et al. (2017) to explain the previously reported distributional gaps: 1) the gaps between known populations are due to poor collecting effort and the lack of material; 2) those species may have once had wider distributions that have become restricted into smaller, relict populations; 3) long distance dispersal events; or 4) convergent evolution of shell characters resulting in misidentifications and thus, incorrect distributions. Additional material and genetic / anatomical data will hopefully clarify our understanding of these species. Predation Some shells we examined had scraped predation holes (Fig. 45), possibly caused by Atopos Simroth, 1891 slugs (see Liew & Schilthuizen 2014; M. Schilthuizen, pers. comm.). Samples of Dicharax notus were large enough to provide meaningful information (see Table 5). This indicates that the predation pressure is highly variable between sites (0 % for JG 10, 4 % in JG 102 and 42 % in JG 11); in some cases almost half of the shells show signs of predation.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
038487E62F16240DFF12B589FE92FC10.taxon	discussion	Remarks. This species is classified in the genus Metalycaeus due to its large size and the “ depressed swelling ” on R 3 as in other Metalycaeus species. The lack of spiral striation on the protoconch is considered a secondary reduction.	en	Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Auffenberg, Kurt (2021): Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 4973 (1): 1-61, DOI: https://doi.org/10.11646/zootaxa.4973.1.1
