taxonID	type	description	language	source
4438245CFFF3094887AEFF55FDB3A528.taxon	type_taxon	Type species: Tarsopolipus lagenaeformis Berlese, 1911, by original designation.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFF3094887AEFF55FDB3A528.taxon	diagnosis	Diagnosis. See Seeman (2019 a).	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFFC094687AEFBD9FA38A696.taxon	description	The key to species of the piratica subgroup was also published in Seeman (2019 b); E. piraticus is included although its host is Trichosternus. The key is primarily to adult females although larvae are helpful for separation of some species.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFFD094687AEFD73FA38A7BF.taxon	description	Key to adult females	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFFD095E87AEFB14FB43A339.taxon	description	(Figs 1 – 8, 10)	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFFD095E87AEFB14FB43A339.taxon	diagnosis	Diagnosis. Adult female: setae sc 1, c 1, c 2, d, f with thickened bases, finely tapering (rarely asymmetrically not tapering); anterior margin of prodorsal shield eroded, concave; setae v 1 9 – 11. Larviform male: sc 1 9. Larval female: Setae c 1 16 – 17, d 11 – 12, f 10 – 11.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFFD095E87AEFB14FB43A339.taxon	materials_examined	Material examined. Holotype female ex. Castelnaudia setosiceps. 9.5 km N of Ravenshoe, 17 ° 31 ′ 31 ″ S 145 ° 29 ′ 21 ″ E, 9 Feb- 15 Apr 1999, G. & S. Monteith, pitfall trap, 1060 masl, HR # T 78934 (QMS 113562). Paratypes. All ex. C. setosiceps. 10 females, 1 male, 3 larvae, as follows: 2 females, same data as holotype; 6 females, 1 male, 1 larva, The Crater NP, Atherton Tablelands, 17 ° 25 ′ 43 ″ S 145 ° 29 ′ 13 ″ E, 5 Dec 1988, G. Monteith & G. Thompson, 1000 masl, HR # T 19427 (QMS 113565 – 71, 1 female ANIC 52 - 003949); 2 females, 2 larvae, Millaa Millaa Lookout, 17 ° 31 ′ 23 ″ S 145 ° 34 ′ 05 ″ E, 1 Dec 1993 - 25 Feb 1994, J. Hasenpusch, pitfall trap, 1000 masl, HR # T 19429 (QMS 113572 – 74, ZMH-A 0015181). 1 female in each of ANIC, ZMH. Note: one female specimen from the Crater was too poor to include in the type series (QMS 113575). One pharate female from the Crater (QMS 113576).	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFFD095E87AEFB14FB43A339.taxon	description	Description. FEMALE (Figs 1 - 3, 10; n = 11). Gnathosoma. Length 60 (58 – 64), width 55 (52 – 62). Cheliceral stylets 73 (68 – 75), setae ch 21 (19 – 24), su 14 (11 – 13). Idiosoma (Figs 1 – 2, 10). Length 340 (340 – 400), width 265 (255 – 350). Prodorsal plate eroded mid-anteriorly and anterolaterally; setae v 1 11 (9 – 11), v 2 alveolar remnant, sc 1 20 (15 – 19), sc 2 66 (65 – 74); setae sc 1 thickened but not bulbous (Fig. 10). Distance between setae v 1 – v 1 46 (46 – 53), sc 1 – sc 1 92 (100 – 105), v 1 – sc 1 31 (28 – 35), v 2 – v 2 44 (50 – 52), sc 2 – sc 2 135 (135 – 140), sc 1 – sc 2 35 (31 – 35). Opisthosomal setae c 1, c 2, d, f thickened, sometimes appearing stout by lacking fine tips, especially setae c 2 (Figs 1, 10). Plate C setae c 1 17 (11 – 21), c 2 8 (8 – 18), distance between setae c 1 – c 1 96 (92 – 100). Plate D setae d 14 (13 – 20), d – d 145 (145 – 165), cupuli ia laterad d. Plate EF setae f 14 (14 – 18), f – f 130 (115 – 155), cupuli im laterad f. Plate H almost completely divided into two small plates, often ventral, seta h 7 (5 – 6). Venter (Fig. 2): coxisterna 1 and 2 well defined, coxisterna 3 weakly defined; coxal setae slender, 1 a 5 (4 – 5), 2 a 8 (7 – 8), 3 a 6 (7 – 8), 3 b 8 (9 – 11). Alveoli of 1 b, 2 b present. Legs (Fig. 3). Setal counts legs I – III, femur-tarsus: 3 - 2 - 6 (+ φ) - 8 (+ ω), 0 - 1 - 4 - 6, 0 - 1 - 4 - 6. Leg I. Femur I, d minute, l’ 3 (2 – 3), v ″ 12 (10 – 12); genu I, l′ 3 (3), l ″ 3 (3); tibia I, d 75 (60 – 75), l’ 7 (6 – 7), l’ ’ 8 (7 – 10), v’ 4 (4 – 5), v’ ’ 11 (10 – 12), k 5 (5 – 6), φ 10 (9 – 10); tarsus I, tc’ 15 (15 – 18), tc’ ’ 16 (16 – 19), pl’ 11 (9 – 11), pl’ ’ 17 (14 – 18), pv′ 3 (2 – 3), pv’ ’ 4 (4 – 5), ω 3 (3 – 4), s 8 (7 – 8), u′ ′ 2 (m – 2). Leg II. Genu II, l′ 4 (3 – 4); tibia II, d 56 (45 – 50), l’ 7 (6 – 7), v’ 17 (12 – 13), v’ ’ 15 (15 – 20); tarsus II, tc′ 2 (2), tc ″ 6 (5 – 6), pl ″ 51 (31 – 49), pv’ ’ 4 (4 – 5), u’ 7 (7 – 8), u′ ′ minute. Leg III. Genu III, l′ 3 (3 – 4); tibia III, d 38 (35 – 45), l’ 7 (6 – 7), v’ 13 (11 – 15), v’ ’ 16 (16 – 22); tarsus III, tc’ 2 (2 – 3), tc ″ 5 (5 – 6), pl ″ 45 (41 – 50), pv’ ’ 3 (3 – 4), u’ 7 (7), u′ ′ minute. LARVIFORM ADULT MALE (Figs 4 – 5; n = 1). Gnathosoma. Length 33, width 30. Cheliceral stylets 25, setae ch 5, su 6. Idiosoma (Figs 4 – 5). Length 165, width 130. Prodorsal plate with setae v 1 5, v 2 alveolar remnant, sc 1 9, sc 2 85. Distance between setae v 1 – v 1 22, sc 1 – sc 1 48, v 1 – sc 1 18, v 2 – v 2 28, sc 2 – sc 2 60, sc 1 – sc 2 21. Plate CD setae c 1 6, c 2 5, d 6, distance between setae c 1 – c 1 40, c 1 – c 2 30, d – d 41; cupuli ia laterad setae d (asymmetrical expression). Plate EF concave, eroded posteriorly; with setae f 3, distance f – f 28; cupuli im anterolaterad setae f. Genital capsule length 29, width 34, setae h minute. Venter (Fig. 4): coxal setae 1 a 3, 2 a 4, 3 a 5, 3 b 5. Legs. Setal counts legs I – III, femur-tarsus: 3 - 2 - 6 (+ φ) - 8 (+ ω), 0 - 1 - 4 - 6, 0 - 1 - 4 - 6. Leg I. Femur I, d minute, l′ minute, v ″ 6; genu I l′ 2, l ″ 2; tibia I, d 45, l’ 2, l’ ’ 3, v ’ 3, v ’’ 8, k 4, φ 9; tarsus I, tc’ 13, tc’ ’ 15, pl’ 8, pl’ ’ 14, pv ′ 2, pv’ ’ 2, ω 3, s 5, u ′′ minute. Leg II. Genu II, l′ 2; tibia II, d 21, l’ 5, v’ 11, v’ ’ 15; tarsus II, tc′ 2, tc ″ 5, pl ″ 27, pv’ ’ 2, u’ 5, u ″ 2. Leg III. Tibia III, d 22, l’ 4, v’ 10, vʺ 13; tarsus III, tc′ 2, tc ″ 5, pl ″ 28, pv’ ’ 3, u ’ 5, u ″ minute. LARVAL FEMALE (Figs 6 – 8; n = 3). Gnathosoma. Length 50 – 51, width 46 – 50. Palp with minute dorsal femoral seta. Cheliceral stylets 62 – 65, setae ch 20 – 27, su 10 – 11. Idiosoma (Fig. 6). Length 225 – 240, width 175 – 185. Prodorsal plate overlapping folded membranous cuticle (outline shown in figure), with setae v 1 5 – 6, v 2 alveolar remnant, sc 1 9 – 10, sc 2 100 – 120. Distance between setae v 1 – v 1 33 – 37, sc 1 – sc 1 68 – 70, v 1 – sc 1 26 – 28, v 2 – v 2 36 – 39, sc 2 – sc 2 85 – 88, sc 1 – sc 2 34 – 37. Plate C setae c 1 16 – 17, c 2 10 – 11, distance between setae c 1 – c 1 49 – 52. Plate D setae d 11 – 12, d – d 61 – 62, cupuli ia very close to setae d, anteriad to anterolaterad d. Plate EF setae f 10 – 11, f – f 52 – 57, cupuli im close to setae f, anterolaterad f. Plate H narrow, 16 – 19 wide, ca. 25 – 30 long bearing setae h 1 ca. 180, h 2 11 – 12. Venter (Fig. 7): coxisterna 1 - 2 well defined, coxisterna 3 poorly defined, membranous; coxal setae slender, 1 a 2 – 3, 2 a 4 – 5, 3 a 6 – 7, 3 b 5. Alveoli of 1 b, 2 b present. Legs (Fig. 8). Setal counts legs I – III, femur-tarsus: 3 - 2 - 6 (+ φ) - 8 (+ ω), 0 - 1 - 4 - 6, 0 - 1 - 4 - 6. Leg I. Femur I, d minute, l′ minute, v ″ 6 – 9; genu I, l′ 2, l ″ 2; tibia I, d 53 – 60, l’ 8 – 9, l’ ’ 11 – 13, v’ 2 – 3, v’ ’ 12 – 13, k 6 – 7, φ 10 – 11; tarsus I, tc’ 15 – 16, tc’ ’ 16 – 18, pl’ 9 – 10, pl’ ’ 17 – 20, pv′ 2, pv’ ’ 3, ω 3, s 5, u′ ′ minute. Leg II. Genu II, l′ 2 – 3; tibia II, d 35 – 40, l’ 6 – 7, v’ 13 – 15, v’ ’ 14 – 20; tarsus II, tc′ 2 – 3, tc ″ 4 – 5, pl ″ 40 – 43, pv’ ’ 3, u’ 6, u′ ′ 2. Leg III. Genu III, l′ 2 – 3; tibia III, d 29 – 45, l’ 6 – 8, v’ 12 – 17, v’ ’ 15 – 17; tarsus III, tc’ 2 – 3, tc ″ 4 – 5, pl ″ 30 – 35, pv’ ’ 3, u’ 6 – 7, u′ ′ minute.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFFD095E87AEFB14FB43A339.taxon	diagnosis	Differential diagnosis. Female E. hadros sp. nov. has idiosomal setae that are less thickened than those of E. mixtus sp. nov. and E. umbonatus sp. nov., giving these setae a comparatively slender appearance. See Fig. 10 and the Differential diagnosis for E. mixtus for further discussion of this feature. The anterior margin of the prodorsal shield of female E. hadros is also deeply eroded medially; this erosion is weak in E. umbonatus and absent in E. mixtus. The male of E. hadros has a longer seta sc 1 (9 versus 3 – 6 in E. mixtus and E. umbonatus), but the single male specimen of E. hadros reduces confidence in this possible diagnostic feature. Larval female E. hadros are difficult to distinguish from E. mixtus, but the size of setae f (10 – 11 versus 14 – 19) is useful. Larvae of E. hadros are distinguished from E. umbonatus by having longer setae c 1 (16 – 19 versus 8 – 11).	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFFD095E87AEFB14FB43A339.taxon	etymology	Etymology. The specific name hadros (Greek, strong, stout) alludes to the strong dorsal setae.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFE6095B87AEFF5AFC7EA2F0.taxon	description	(Figs 9 – 10)	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFE6095B87AEFF5AFC7EA2F0.taxon	diagnosis	Diagnosis. Adult female: setae sc 1, c 1, c 2, d, f with strongly thickened bases, often blunt-tipped; anterior margin of prodorsal shield eroded, concave; setae v 1 10 – 13. Larviform male: sc 1 3 – 6. Larval female: Setae c 1 13 – 20, d 11 – 12, f 14 – 19.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFE6095B87AEFF5AFC7EA2F0.taxon	materials_examined	Material examined. Holotype female ex. Castelnaudia mixta. Broken River, Eungella, 21 ° 10 ′ 05 ″ S 148 ° 30 ′ 30 ″ E, 17 Nov 1992 - 15 Apr 1993, G. Monteith & D. Cook, flight intercept & pitfall trap, HR # T 82585 (QMS 113035). Paratypes. 12 females, 4 males, 4 larvae, as follows: 11 females, 3 males, 1 larva, same data as holotype (QMS 113036 - 43, 113045, 113047, ANIC 52 - 003950 – 51, ZMH-A 0015182 – 83); 1 male, 1 larva, same data except HR # T 82586 (QMS 113046, ZMH-A 0015184). 1 female, same data except HR # T 82582 (QMS 113044); 1 larva, same data except HR # T 82584 (ANIC 52 - 003952); 1 larva, Pearse’s Lookout, Eungella, 21 ° 06 ′ 53 ″ S 148 ° 31 ′ 06 ″ S, 17 Nov 1992 - 15 Apr 1993, G. Monteith & D. Cook, IN 5934, HR # T 82526 (QMS 113048). All in QM except 1 female, male, larva in ANIC and ZMH.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFE6095B87AEFF5AFC7EA2F0.taxon	description	Description. FEMALE (Figs 9 – 10; n = 13). Gnathosoma. Length 59 (57 – 65), width 55 (53 – 60). Cheliceral stylets 75 (67 – 78), setae ch 18 (16 – 25), su 17 (11 – 18). Idiosoma (Fig. 9). Length 395 (365 – 400), width 280 (270 – 330). Prodorsal plate eroded mid-anteriorly and anterolaterally; setae v 1 13 (10 – 13), v 2 alveolar remnant, sc 1 thickened, 11 / 16 (asymmetry) (11 – 17), sc 2 75 (70 – 95) (Fig. 10). Distance between setae v 1 – v 1 48 (41 – 52), sc 1 – sc 1 94 (82 – 100), v 1 – sc 1 33 (26 – 40), v 2 – v 2 43 (41 – 51), sc 2 – sc 2 130 (130 – 140), sc 1 – sc 2 35 (24 – 34). Opisthosomal setae c 1, c 2, d, f thickened, sometimes appearing stout by lacking fine tips, especially setae c 2. Plate C setae c 1 12 / 20 (asymmetry) (10 – 20), c 2 9 (10 – 15), distance between setae c 1 – c 1 100 (78 – 95). Plate D setae d 17 (12 – 21), d – d 145 (135 – 150), cupuli ia laterad d. Plate EF setae f 13 / 18 (asymmetry) (11 – 19), f – f 135 (110 – 125), cupuli im laterad f. Plate H divided into two small plates, or fused weakly, often ventral, seta h 5 (5 – 6). Venter: coxisterna 1 and 2 well defined, coxisterna 3 weakly defined; coxal setae slender, 1 a 4 (3 – 4), 2 a 8 (6 – 8), 3 a thickened, 5 (5 – 7), 3 b 9 (7 – 9). Alveoli of 1 b, 2 b present. Legs (holotype only, except if holotype different from E. hadros). Setal counts legs I – III, femur-tarsus: 3 - 2 - 6 (+ φ) - 8 (+ ω), 0 - 1 - 4 - 6, 0 - 1 - 4 - 6. Leg I. Femur I, d minute, l’ 4 (3 – 5), v ″ 11; genu I, l′ 2, l ″ 3; tibia I, d 65, l’ 8, l’ ’ 10, v’ 5, v’ ’ 10, k 5, φ 7 (7 – 9); tarsus I, tc’ 16, tc’ ’ 15, pl’ 9, pl’ ’ 13, pv′ 3, pv’ ’ 3, ω 3, s 7, u′ ′ minute. Leg II. Genu II, l′ 3; tibia II, d 57, l’ 7, v’ 13, v’ ’ 19; tarsus II, tc′ 2, tc ″ 6, pl ″ 60, pv’ ’ 4, u’ 7, u′ ′ minute. Leg III. Genu III, l′ 3; tibia III, d 42, l’ 7, v’ 11, v’ ’ 15; tarsus III, tc’ 2, tc ″ 5, pl ″ 39, pv’ ’ 3, u’ 6, u′ ′ minute. LARVIFORM ADULT MALE (n = 4). Gnathosoma. Length 32 – 35, width 30 – 34. Cheliceral stylets 23 – 27, setae ch 5 – 7, su 5 – 10. Idiosoma. Length 160 – 205, width 125 – 150. Prodorsal plate with setae v 1 4 – 5, v 2 alveolar remnant, sc 1 3 – 6, sc 2 60 – 70. Distance between setae v 1 – v 1 21 – 26, sc 1 – sc 1 40 – 52, v 1 – sc 1 20 – 25, v 2 – v 2 22 – 27, sc 2 – sc 2 57 – 77, sc 1 – sc 2 16 – 22. Plate CD setae c 1 4 – 6, c 2 4 – 5, d 4 – 5, distance between setae c 1 – c 1 46 – 59, c 1 – c 2 22 – 28, d – d 36 – 52; cupuli ia anterolaterad setae d. Plate EF eroded medially, almost divided; with setae f 2 – 3, distance f – f 24 – 30; cupuli im anterolaterad to laterad setae f. Genital capsule length 28 – 30, width 34 – 38, setae h minute. Venter: coxal setae 1 a 2 – 3, 2 a 4 – 5, 3 a 3 – 4, 3 b 5 – 6. Legs. Setal counts legs I – III, femur-tarsus: 3 - 2 - 6 (+ φ) - 8 (+ ω), 0 - 1 - 4 - 6, 0 - 1 - 4 - 6. Leg I. Femur I, d minute, l′ 2, v ″ 7; genu I l′ 2, l ″ 2; tibia I, d 38, l’ 3, l’ ’ 2, v’ 3, v’ ’ 10, k 4, φ 6 – 8; tarsus I, tc’ 11 – 14, tc’ ’ 11 – 14, pl’ 7, pl’ ’ 11, pv ′ 2, pv’ ’ 3, ω 3, s 4, u ′′ minute. Leg II. Genu II, l′ 2; tibia II, d 16, l’ 5, v’ 10, v’ ’ 13; tarsus II, tc′ 2, tc ″ 4, pl ″ 27, pv’ ’ 3, u’ 5, u ″ 2. Leg III. Tibia III, d 16, l’ 4, v’ 12, v ″ 12; tarsus III, tc′ 2, tc ″ 3 – 5, pl ″ 20, pv’ ’ 3, u’ 5, u ″ present. LARVAL FEMALE (n = 4). Gnathosoma. Length 46 – 51, width 43 – 49. Cheliceral stylets 61 – 64, setae ch 23 – 27, su 7 – 10. Idiosoma. Length 205 – 260, width 165 – 215. Prodorsal plate overlapping folded membranous cuticle, with setae v 1 6 – 8, v 2 alveolar remnant, sc 1 7 – 10, sc 2 120 – 135. Distance between setae v 1 – v 1 31 – 35, sc 1 – sc 1 59 – 63, v 1 – sc 1 23 – 25, v 2 – v 2 30 – 34, sc 2 – sc 2 72 – 83, sc 1 – sc 2 31 – 33. Plate C setae c 1 13 – 20, c 2 8 – 11, distance between setae c 1 – c 1 42 – 48. Plate D setae d 11 – 12, d – d 53 – 61, cupuli ia very close to setae d, anteriad to anterolaterad d. Plate EF setae f 14 – 19, f – f 46 – 52, cupuli im close to setae f, anterolaterad f. Plate H narrow, 15 – 20 wide, ca. 25 – 30 long bearing setae h 1 ca. 150, h 2 13 – 17. Venter: coxisterna 1 - 2 well defined, coxisterna 3 poorly defined, membranous; coxal setae slender, 1 a 3, 2 a 4 – 5, 3 a 4 – 5, 3 b 4 – 5. Alveoli of 1 b, 2 b present. Legs. Setal counts legs I – III, femur-tarsus: 3 - 2 - 6 (+ φ) - 8 (+ ω), 0 - 1 - 4 - 6, 0 - 1 - 4 - 6. Leg I. Femur I, d minute, l′ 2, v ″ 7; genu I, l′ 2, l ″ 3; tibia I, d 54, l’ 9, l’ ’ 10, v’ 4, v’ ’ 8 – 12, k 6, φ 10; tarsus I, tc’ 13, tc’ ’ 15, pl’ 8, pl’ ’ 16, pv′ 2, pv’ ’ 3, ω 3, s 5, u′ ′ minute. Leg II. Genu II, l′ 4; tibia II, d 40, l’ 7, v’ 14, v’ ’ 16; tarsus II, tc′ 2, tc ″ 4, pl ″ 35, pv’ ’ 3, u’ 5, u′ ′ 2. Leg III. Genu III, l′ 4; tibia III, d 35, l’ 7, v’ 13, v’ ’ 14; tarsus III, tc’ 2, tc ″ 4, pl ″ 21 – 40, pv’ ’ 3, u’ 6, u′ ′ minute.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFE6095B87AEFF5AFC7EA2F0.taxon	diagnosis	Differential diagnosis. Eutarsopolipus mixtus sp. nov. is similar to E. hadros sp. nov. but is most easily distinguished by having thicker dorsal setae. This feature is useful for species separation, but is complicated by intraspecific variation in setal form, which is often expressed asymmetrically. However, setae sc 1 are always thicker in E. mixtus, even if they taper. Likewise, other dorsal idiosomal setae tend to be thicker in E. mixtus, exemplified by setae f (Fig. 10). The prodorsal shield of E. umbonatus is weakly eroded anteriorly, in contrast to the deep erosion of E. hadros and lack of erosion in E. umbonatus. However, the intermediate state of this feature reduces its effectiveness. In larvae, E. mixtus have longer setae f (14 – 19 versus 10 – 11 in E. hadros and 7 – 10 in E. umbonatus).	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFE6095B87AEFF5AFC7EA2F0.taxon	etymology	Etymology. The species name mixtus (Latin, mixed) refers both to the host species C. mixta but also the variation in form of the dorsal setae.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFE6095B87AEFF5AFC7EA2F0.taxon	discussion	Remarks. Three paratype females have fully developed larvae within their bodies, with no egg, suggestive of vivipary. Two additional specimens of C. mixta from Finch Hatton Gorge (HR # T 82535, T 82529) had a male and a larva (one per host) of an undermined species of Eutarsopolipus (Table 1).	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFE0095887AEF8E9FC3FA6E5.taxon	description	(Fig. 11)	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFE0095887AEF8E9FC3FA6E5.taxon	diagnosis	Diagnosis. Adult female: setae sc 1, c 1, c 2, d, f with strongly thickened bases, often blunt-tipped; anterior margin of prodorsal shield not or only slightly eroded, straight or weakly concave; setae v 1 7 – 10. Larviform male: sc 1 5 – 6. Larval female: Setae c 1 8 – 11, d 7 – 10, f 7 – 10.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFE0095887AEF8E9FC3FA6E5.taxon	materials_examined	Material examined. Holotype female ex. Castelnaudia cordata, Mt Mee SF, 6 km NNW barracks, 27 ° 2 ʹ 53 ʺ S 152 ° 40 ʹ 49 ʺ E, 8 Jan- 3 Mar 1992, D. Cook, flight intercept trap, IN 5148, HR # T 68607 (QMS 113481). Paratypes. 16 females, 7 males, 15 larvae, as follows: all ex. C. cordata. 15 females, 6 males, 13 larvae, same data as holotype (QMS 113482 – 92, ANIC 52 - 003953 – 57, ZMH-A 0015185 – 89); 1 female, 1 larva, McIntosh Rd, Unumbar SF, 28 ° 25 ʹ 19 ʺ S 152 ° 42 ʹ 14 ʺ E, 25 Nov 1972, G. Monteith, IN 6467, HR # T 64329 (QMS 113493 – 94); 1 male, 1 larva, “ Glen Witheren ”, shelf scrub, 28 ° 2 ʹ 58 ʺ S 153 ° 6 ʹ 58 ʺ E, 30 Dec 1991 - 24 May 1992, G. Monteith, pitfall trap, IN 5682, rainforest, HR # 68606 (QMS 113495); 1 larva, Lower Coomera, 28 ° 12 ʹ 20 ʺ S 153 ° 10 ʹ 46 ʺ E, 9 Jan- 6 Apr 1995, G. Monteith, pitfall trap, IN 5998, HR # T 68610. All in QM except 3 females, 1 male, 1 larva in each of ANIC and ZMH. Other material examined. 5 females, 2 larvae, on same slide as specimens of E. hebronae, same data as holotype. Not included as paratypes.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFE0095887AEF8E9FC3FA6E5.taxon	description	Description. FEMALE (Fig. 11; n = 17). Gnathosoma. Length 62 (56 – 61), width 57 (55 – 58). Cheliceral stylets 68 (68 – 70), setae ch 22 (25 – 30), su 10 (10 – 18). Idiosoma (Fig. 11). Length 330 (320 – 400), width 290 (250 – 325). Prodorsal plate slightly or not eroded midanteriorly, more substantially eroded anterolaterally; setae v 1 7 (7 – 10), v 2 alveolar remnant, sc 1 usually thickened, thorn-like 7 (7 – 12) [14 / 16 in specimen # 5, 12 / 15 in specimen # 8, 9 / 15 in specimen # 11], sc 2 50 – 80. Distance between setae v 1 – v 1 47 (41 – 49), sc 1 – sc 1 105 (94 – 105), v 1 – sc 1 38 (32 – 36), v 2 – v 2 45 (41 – 50), sc 2 – sc 2 130 (125 – 130), sc 1 – sc 2 27 (25 – 32). Opisthosomal setae c 1, c 2, d, f usually stout but sometimes with fine tips. Plate C setae c 1 7 (6 – 12; 15 in specimen # 8), c 2 10 (10 – 15), distance between setae c 1 – c 1 84 (82 – 100). Plate D setae d 10 (10 – 15), d – d 155 (160 – 185), cupuli ia laterad d. Plate EF setae f 7 / 10 (asymmetry) (7 – 12; 10 / 13 in specimen # 8; 15 in specimen # 15), f – f 115 (130 – 150), cupuli im laterad f. Plate H divided or fused weakly, seta h 7 (5 – 7). Venter: coxisterna 1 well defined, coxisterna 2 well defined but sometimes with folds, coxisterna 3 weakly defined; coxal setae slender, 1 a 4 (3 – 4), 2 a 6 (5 – 6), 3 a thickened, 3 – 6, 3 b 4 – 6. Alveoli of 1 b, 2 b present. Legs (holotype only, except if holotype different from E. hadros / mixtus). Setal counts legs I – III, femur-tarsus: 3 - 2 - 6 (+ φ) - 8 (+ ω), 0 - 1 - 4 - 6, 0 - 1 - 4 - 6. Leg I. Femur I, d minute, l’ 2, v ″ 14; genu I, l′ 3, l ″ 3; tibia I, d 60, l’ 7, l’ ’ 10, v’ 5, v’ ’ 8, k 5, φ 8 (7 – 9); tarsus I, tc’ 14, tc’ ’ 15, pl’ 7, pl’ ’ 13, pv′ 3, pv’ ’ 4, ω 4, s 7, u′ ′ minute. Leg II. Genu II, l′ 4; tibia II, d 40, l’ 7, v’ 12, v’ ’ 23; tarsus II, tc′ 2, tc ″ 6, pl ″ 55, pv’ ’ 4, u’ 7, u′ ′ minute. Leg III. Genu III, l′ 3; tibia III, d 29, l’ 8, v’ 11, v’ ’ 21; tarsus III, tc’ 2, tc ″ 6, pl ″ 52, pv’ ’ 4, u’ 8, u′ ′ minute. LARVIFORM ADULT MALE (n = 7). Gnathosoma. Length 33 – 35, width 30 – 32. Cheliceral stylets 25 – 28, setae ch 6 – 7, su 5 – 7. Idiosoma. Length 170 – 185, width 120 – 130. Prodorsal plate with setae v 1 3 – 5, v 2 alveolar remnant, sc 1 5 – 6, sc 2 70 – 80. Distance between setae v 1 – v 1 19 – 21, sc 1 – sc 1 46 – 50, v 1 – sc 1 22 – 26, v 2 – v 2 19 – 22, sc 2 – sc 2 57 – 63, sc 1 – sc 2 20 – 21. Plate CD setae c 1 5 – 7, c 2 4 – 6, d 5 – 6, distance between setae c 1 – c 1 42 – 45, c 1 – c 2 27 – 31, d – d 40 – 43; cupuli ia anterolaterad setae d. Plate EF eroded medially, sometimes almost divided; with setae f 2 – 4, distance f – f 27 – 28; cupuli im anterolaterad to laterad setae f. Genital capsule length 28 – 31, width 31 – 32, setae h minute. Venter: coxal setae 1 a 2 – 3, 2 a 4, 3 a 2 – 3, 3 b 4 – 5. Legs. Setal counts legs I – III, femur-tarsus: 3 - 2 - 6 (+ φ) - 8 (+ ω), 0 - 1 - 4 - 6, 0 - 1 - 4 - 6. Leg I. Femur I, d minute, l’ minute, v ″ 7; genu I l′ 2, l ″ 2; tibia I, d 44, l’ 2, l’ ’ 3, v ’ 3, v ’’ 7, k 4, φ 8; tarsus I, tc’ 12, tc’ ’ 12, pl’ 7, pl’ ’ 12, pv ′ 2, pv’ ’ 2, ω 3, s 5, u ′′ 2. Leg II. Genu II, l′ 2; tibia II, d 18, l’ 5, v’ 10, v’ ’ 17; tarsus II, tc′ 3, tc ″ 4, pl ″ 28, pv’ ’ 3, u’ 5, u ″ 2. Leg III. Tibia III, d 14, l’ 5, v’ 11, vʺ 11; tarsus III, tc′ 3, tc ″ 4, pl ″ 22, pv’ ’ 3, u’ 5, u ″ minute. LARVAL FEMALE (n = 15). Gnathosoma. Length 48 – 52, width 46 – 54. Cheliceral stylets 63 – 70, setae ch 27 – 29, su 7 – 9. Idiosoma. Length 195 – 280, width 135 – 215. Prodorsal plate with setae v 1 4 – 7, v 2 alveolar remnant, sc 1 5 – 8, sc 2 100 – 120. Distance between setae v 1 – v 1 28 – 37, sc 1 – sc 1 65 – 75, v 1 – sc 1 23 – 26, v 2 – v 2 32 – 37, sc 2 – sc 2 71 – 85, sc 1 – sc 2 32 – 37. Plate C setae c 1 8 – 11, c 2 5 – 10, distance between setae c 1 – c 1 40 – 48. Plate D setae d 7 – 10, d – d 49 – 63, cupuli ia very close to setae d, anteriad to anterolaterad d. Plate EF setae f 7 – 10, f – f 48 – 50, cupuli im close to setae f, anteriad f. Plate H narrow, 13 – 15 wide, ca. 25 – 35 long bearing setae h 1 ca. 170, h 2 6 – 8 (Mt Mee specimens 12 – 15). Venter: coxisterna 1 - 2 well defined, coxisterna 3 poorly defined, membranous; coxal setae 1 a 2 – 3, slender, 2 a 4 – 5, slender, 3 a 3 – 5 sometimes slightly thickened, 3 b 4 – 5, slender. Alveoli of 1 b, 2 b present. Legs. Setal counts legs I – III, femur-tarsus: 3 - 2 - 6 (+ φ) - 8 (+ ω), 0 - 1 - 4 - 6, 0 - 1 - 4 - 6. Leg I. Femur I, d minute, l′ 2, v ″ 8; genu I, l′ 3, l ″ 2; tibia I, d 62, l’ 9, l’ ’ 10, v’ 3, v’ ’ 11, k 5, φ 10; tarsus I, tc’ 12, tc’ ’ 13, pl’ 10, pl’ ’ 16, pv′ 2, pv’ ’ 3, ω 3, s 5, u′ ′ 2. Leg II. Genu II, l′ 3; tibia II, d 35, l’ 5, v’ 15, v’ ’ 15; tarsus II, tc′ 2, tc ″ 4, pl ″ 37, pv’ ’ 3, u’ 6, u′ ′ minute. Leg III. Genu III, l′ 3; tibia III, d 33, l’ 6, v’ 13, v’ ’ 15; tarsus III, tc’ 2, tc ″ 4, pl ″ 32, pv’ ’ 3, u’ 6, u′ ′ minute.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFE0095887AEF8E9FC3FA6E5.taxon	diagnosis	Differential diagnosis. Eutarsopolipus umbonatus sp. nov. is similar to E. mixtus sp. nov. by having much stronger dorsal setae compared with E. hadros. However, like all species and particularly E. mixtus, the form of these setae differed intraspecifically: although the thicker nature of these setae are useful for separating E. umbonatus from E. hadros, they are too variable to separate E. umbonatus from E. mixtus. Larvae are more useful for distinguishing the latter two species. Larva of E. umbonatus have smaller idiosomal setae: setae c 1 (8 – 11 versus 13 – 20), d (7 – 10 versus 11 – 12) and f (7 – 10 versus 14 – 19) are slightly to obviously shorter in E. umbonatus.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFE0095887AEF8E9FC3FA6E5.taxon	etymology	Etymology. The specific name umbonatus (knob-like) refers to the short, more knob-like setae in females of this species.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFE0095887AEF8E9FC3FA6E5.taxon	discussion	Remarks. In larvae, setae h 2 were longer in the type series from Mt Mee (12 – 15) compared with the three specimens from other sites (6 – 8). No other features distinguished these populations. However, Mt Mee is part of a mountain range (the D’Aguilar Range) to the northwest of Brisbane, while all other collections were from the Border Range to the south to southwest of Brisbane. The geographical separation of these populations, combined with the difference in length of seta h, suggests that these may be separate species on isolated populations of Castelnaudia cordata. Furthermore, these differences also suggest that the host species deserves closer attention. The D’Aguilar Range has several endemic taxa, such as a unique assemblage of anamid spiders (Rix et al. 2020). Likewise, the beetle specimens from the D’Aguilar Range may be an endemic species closely related to C. cordata. The above three species, E. hadros, E. mixtus and E. umbonatus, are very closely related and it is tempting to consider all three as one species. However, each species does exhibit subtle differences, they occur on different host species, and are highly isolated geographically with the hosts C. setosiceps, C. mixta and C. cordata in north-east Queensland, middle-east Queensland and south-east Queensland, respectively. One paratype female has a fully developed larva within its body, with no egg, indicating vivipary. Two larvae of another species close to Eutarsopolipus echinatus Seeman, 2019 a (i. e. same thorn-like setae, but clearly a different species) were also collected from C. cordata: one from the type host (on a separate slide) and one from Lower Coomera and on the same slide as the larva of E. umbonatus (QMS 113496). This species was found co-infesting its host with E. hebronae sp. nov. and E. osculum sp. nov. (Tables 1, 2). The ochoai species group - nahmani subgroup Diagnosis. Female with dorsal idiosomal setae fine, slender; setae h small but developed. All life stages with gnathosomal apex evenly rounded.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFE0095887AEF8E9FC3FA6E5.taxon	discussion	Remarks. In larvae, setae h 2 were longer in the type series from Mt Mee (12 – 15) compared with the three specimens from other sites (6 – 8). No other features distinguished these populations. However, Mt Mee is part of a mountain range (the D’Aguilar Range) to the northwest of Brisbane, while all other collections were from the Border Range to the south to southwest of Brisbane. The geographical separation of these populations, combined with the difference in length of seta h, suggests that these may be separate species on isolated populations of Castelnaudia cordata. Furthermore, these differences also suggest that the host species deserves closer attention. The D’Aguilar Range has several endemic taxa, such as a unique assemblage of anamid spiders (Rix et al. 2020). Likewise, the beetle specimens from the D’Aguilar Range may be an endemic species closely related to C. cordata. The above three species, E. hadros, E. mixtus and E. umbonatus, are very closely related and it is tempting to consider all three as one species. However, each species does exhibit subtle differences, they occur on different host species, and are highly isolated geographically with the hosts C. setosiceps, C. mixta and C. cordata in north-east Queensland, middle-east Queensland and south-east Queensland, respectively. One paratype female has a fully developed larva within its body, with no egg, indicating vivipary. Two larvae of another species close to Eutarsopolipus echinatus Seeman, 2019 a (i. e. same thorn-like setae, but clearly a different species) were also collected from C. cordata: one from the type host (on a separate slide) and one from Lower Coomera and on the same slide as the larva of E. umbonatus (QMS 113496). This species was found co-infesting its host with E. hebronae sp. nov. and E. osculum sp. nov. (Tables 1, 2).	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFE0095887AEF8E9FC3FA6E5.taxon	diagnosis	Diagnosis. Female with dorsal idiosomal setae fine, slender; setae h small but developed. All life stages with gnathosomal apex evenly rounded.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFE3095887AEFC81FE52A7DD.taxon	diagnosis	Diagnosis. Female with dorsal idiosomal setae fine, slender; setae h small but developed. All life stages with gnathosomal apex evenly rounded.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFE3095387AEFBD9FE6EA29E.taxon	description	(Figs 12 – 16)	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFE3095387AEFBD9FE6EA29E.taxon	diagnosis	Diagnosis. Adult female: Gnathosomal length 67 – 70, width 69 – 72; anterior margin of prodorsal shield not eroded medially; v 1 6 – 7, sc 1 10 – 12, c 1 17 – 19, c 2 14 – 19, d 15 – 19, f 15 – 18, h 3 – 6, v 1 - v 1 54 – 64, d-d 200 – 230, f-f 150 – 170; genua II-III setae l′ 5 – 7, tibia I seta l′ 9 – 11, tarsi II-III setae tc′ ′ 5 – 6, pv′ ′ 2 – 3. Larval female: Gnathosomal length 57 – 59, width 57 – 60; idiosomal length 335 – 400; c 1 (10 – 11) and c 2 (12) subequal, v 1 – v 1 47 – 59, sc 1 – sc 1 100 – 125, c 1 – c 1 105 – 120, d – d 130 – 135; genua II-III setae l′ 3 – 5, tibia I seta l′ 12 – 13, tarsi II-III setae tc′ 3 – 4, tc′ ′ 5 – 6, pv′ ′ 2 – 3.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFE3095387AEFBD9FE6EA29E.taxon	materials_examined	Material examined. Holotype female, ex. Castelnaudia septemcostata. O’Reilly’s Guesthouse, 28 ° 14 ʹ 08 ʺ S 152 ° 08 ʹ 12 ʺ E, 1 - 31 Dec 1909, H. Hacker, IN 7977, HR # T 88472 (QMS 113011). In QM. Paratypes. All ex. C. septemcostata. 5 females, 2 males, 2 larvae, as follows: 1 larva, same data as holotype (QMS 113017); 3 females, Mt Cougal, shelf, 28 ° 14 ʹ 10 ʺ S 152 ° 19 ʹ 34 ʺ E, 19 Nov 1987 - 17 Mar 1988, D. Cook & G. Monteith, pitfall trap, rainforest, IN 6062, HR # T 64265 (QMS 113012, ANIC 52 - 003958, ZMH-A 0015190); 1 female, 1 male, Bar Mountain, Wiangaree, 28 ° 27 ʹ 37 ʺ S 153 ° 8 ʹ 2 ʺ E, 11 Oct 1977 - 2 Oct 1978, G. & S. R. Monteith, pitfall trap, rainforest, IN 5624, HR # T 57344 (QMS 113013, 113016); 1 female, 1 male, Tweed Lookout, Wiangaree, 28 ° 23 ʹ 14 ʺ S 153 ° 6 ʹ 33 ʺ E, 16 Nov- 27 Dec 1974, G. & S. R. Monteith, pitfall trap, rainforest, IN 5349, HR # T 57316 (QMS 113014 - 15); 1 larva, Upper Tallebudgera Creek, 28 ° 14 ʹ 10 ʺ S 152 ° 19 ʹ 34 ʺ E, 19 Nov 1987 - 17 Mar 1988, D. Cook & G. Monteith, pitfall trap, rainforest, IN 6062, HR # T 64272 (QMS 113018). All in QM except 1 female in ANIC, 1 female in ZMH.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFE3095387AEFBD9FE6EA29E.taxon	description	Description. FEMALE (Figs 12 – 13, n = 6). Gnathosoma. Length 69 (67 – 70), width 70 (69 – 72). Cheliceral stylets 72 (73 – 82), setae ch 16 (15 – 24), su 16 (13 – 19). Idiosoma (Fig. 12). Length 520 (380 – 600), width 405 (295 – 470). Prodorsal plate not eroded anteriorly; setae slender, v 1 7 (6 – 7), v 2 alveolar remnant, sc 1 10 (10 – 12), sc 2 60 (60 – 70). Distance between setae v 1 – v 1 68 (54 – 64), sc 1 – sc 1 135 (135 – 140), v 1 – sc 1 47 (46 – 53), v 2 – v 2 74 (70 – 79), sc 2 – sc 2 205 (200 – 225), sc 1 – sc 2 55 (52 – 59). Opisthosomal setae slender. Plate C setae c 1 16 (17 – 19), c 2 13 (14 – 19), distance between setae c 1 – c 1 150 (155 – 165). Plate D setae d 17 (15 – 19), d – d 215 (200 – 230), cupuli ia anteriad to anterolaterad d. Plate EF setae f 17 (15 – 18), f – f 170 (150 – 165), cupuli im anterolaterad f. Plate H divided or fused weakly, seta h 3 (3 – 6). Venter: coxisterna 1 well defined, coxisterna 2 - 3 well defined but sometimes with some folds; coxal setae slender, 1 a 4 (3 – 4), 2 a 5 (5 – 6), 3 a 8 (8 – 10), 3 b 6 (5 – 6). Alveoli of 1 b, 2 b present. Legs (Fig. 13). Setal counts legs I – III, femur-tarsus: 3 - 2 - 6 (+ φ) - 8 (+ ω), 0 - 1 - 4 - 6, 0 - 1 - 4 - 6. Leg I. Femur I, d minute, l’ 3 (2 – 3), v ″ 13 (10 – 15); genu I, l′ 3 (3 – 4), l ″ 3 (3 – 4); tibia I, d 63 (60 – 68), l’ 10 (9 – 11), l’ ’ 14 (11 – 15), v’ 7 (6 – 8), v’ ’ 10 (10 – 15), k 7 (6 – 8), φ 13 (12 – 13); tarsus I, tc’ 18 (18 – 19), tc’ ’ 18 (18 – 19), pl’ 13 (12 – 16), pl’ ’ 19 (17 – 20), pv′ 3 (2 – 3), pv’ ’ 2 (2 – 3), ω 4 (3 – 4), s 7 (6 – 8), u′ ′ 2 (2). Leg II. Genu II, l′ 7 (5 – 7); tibia II, d 50 (40 – 45), l’ 12 (8 – 12), v’ 11 (11 – 15), v’ ’ 27 (20 – 26); tarsus II, tc′ 4 (3 – 4), tc ″ 6 (5 – 6), pl ″ 62 (42 – 60), pv’ ’ 2 (2 – 3), u’ 8 (7 – 8), u′ ′ 2 (2). Leg III. Genu III, l′ 6 (5 – 6); tibia III, d 35 (28 – 35), l’ 7 (7), v’ 15 (15 – 17), v’ ’ 26 (20 – 22); tarsus III, tc’ 3 (3), tc ″ 5 (4 – 5), pl ″ 60 (50 – 70), pv’ ’ 2 (2), u’ 8 (7 – 8), u′ ′ 2 (m – 2). LARVIFORM ADULT MALE (Fig. 14; n = 2) Gnathosoma. Length 37 – 38, width 36. Cheliceral stylets 24 – 27, setae ch 6 – 7, su 6. Idiosoma (Fig. 14). Length 225 – 235, width 150 – 170. Prodorsal plate with setae v 1 6 – 7, v 2 alveolar remnant, sc 1 5, sc 2 50 – 55. Distance between setae v 1 – v 1 23 – 24, sc 1 – sc 1 49 – 50, v 1 – sc 1 24 – 26, v 2 – v 2 28 – 33, sc 2 – sc 2 67 – 70, sc 1 – sc 2 34 – 36. Plate CD setae c 1 6 – 8, c 2 7 – 8, d 7, distance between setae c 1 – c 1 62 – 65, c 1 – c 2 33 – 37, d – d 56 – 63; cupuli ia anterolaterad setae d. Plate EF slightly eroded medially; with setae f 6 – 7, distance f – f 39 – 45; cupuli im anteriad to anterolaterad setae f. Genital capsule length 36 – 39, width 36, setae h m – 2. Venter: coxal setae 1 a 2, 2 a 2 – 3, 3 a 5, 3 b 4. Legs. Setal counts legs I – III, femur-tarsus: 3 - 2 - 6 (+ φ) - 8 (+ ω), 0 - 1 - 4 - 6, 0 - 1 - 4 - 6. Leg I. Femur I, d minute, l′ minute, v ″ 5 – 6; genu I l′ 2, l ″ 2; tibia I, d 26 – 30, l’ 4 – 5, l’ ’ 6 – 7, v ’ 5 – 7, v ’’ 7 – 8, k 3 – 4, φ 7 – 8; tarsus I, tc’ 15, tc’ ’ 15, pl’ 6 – 7, pl’ ’ 12 – 15, pv ′ 2, pv’ ’ 2, ω 2 – 3, s 4 – 5, u ′′ m. Leg II. Genu II, l′ 2 – 3; tibia II, d 15 – 23, l’ 5 – 6, v’ 9 – 11, v’ ’ 15 – 19; tarsus II, tc′ m – 2, tc ″ 3 – 4, pl ″ 23 – 30, pv’ ’ 2, u ’ 5 – 6, u ″ minute. Leg III. Genu III, l′ 2 – 3; tibia III, d 13 – 15, l’ 5 – 7, v’ 9 – 11, vʺ 12 – 16; tarsus III, tc′ 2, tc ″ 3 – 4, pl ″ 30 – 35, pv’ ’ 2, u’ 5 – 6, u ″ minute. LARVAL FEMALE (Figs 15 – 16). Gnathosoma. Length 57 – 59, width 57 – 60. Cheliceral stylets 60 – 62, setae ch 16 – 25, su 10 – 11. Idiosoma (Fig. 15). Length 335 – 400, width 270 – 350. Prodorsal plate with setae v 1 4 – 5, v 2 alveolar remnant, sc 1 6 – 7, sc 2 90 – 105. Distance between setae v 1 – v 1 47 – 59, sc 1 – sc 1 100 – 125, v 1 – sc 1 40 – 41, v 2 – v 2 61 – 78, sc 2 – sc 2 140 – 155, sc 1 – sc 2 52 – 53. Plate C setae c 1 10 – 11, c 2 12, distance between setae c 1 – c 1 105 – 120. Plate D setae d 9 – 10, d – d 130 – 135, cupuli ia anterolaterad setae d. Plate EF setae f 10, f – f 83, cupuli im anterolaterad f. Plate H narrow, 17 – 20 wide, 23 – 30 long, bearing setae h 1 ca. 130, h 2 9 – 10. Venter: coxisterna 1 - 2 well defined, coxisterna 3 with some folds; coxal setae slender, 1 a 3, 2 a 4, 3 a 7 – 8, 3 b 5. Alveoli of 1 b, 2 b present. Legs (Fig. 16). Setal counts legs I – III, femur-tarsus: 3 - 2 - 6 (+ φ) - 8 (+ ω), 0 - 1 - 4 - 6, 0 - 1 - 4 - 6. Leg I. Femur I, d minute, l′ 2, v ″ 7 – 8; genu I, l′ 2, l ″ 2; tibia I, d 50 – 65, l’ 12 – 13, l’ ’ 11 – 13, v’ 6 – 8, v’ ’ 10 – 11, k 5, φ 12; tarsus I, tc’ 19 – 22, tc’ ’ 18 – 21, pl’ 12 – 14, pl’ ’ 14 – 20, pv′ 2, pv’ ’ 2, ω 2, s 5 – 6, u′ ′ m. Leg II. Genu II, l′ 4 – 5; tibia II, d 25 – 42, l’ 10 – 11, v’ 11 – 17, v’ ’ 20 – 22; tarsus II, tc′ 2 – 3, tc ″ 3 – 4, pl ″ 44 – 47, pv’ ’ 2, u’ 7 – 8, u′ ′ minute. Leg III. Genu III, l′ 3 – 5; tibia III, d 32 – 35, l’ 10 – 11, v’ 13 – 16, v’ ’ 21 – 22; tarsus III, tc’ 2 – 3, tc ″ 3 – 4, pl ″ 44 – 50, pv’ ’ 2 – 3, u’ 6, u′ ′ minute.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFE3095387AEFBD9FE6EA29E.taxon	diagnosis	Differential diagnosis. The nahmani subgroup also includes Eutarsopolipus burwelli Seeman, 2019 a (from Nurus medius) and Eutarsopolipus rutherfordae Constantine & Seeman, 2014 (from Trichosternus subvirens). Eutarsopolipus burwelli has a large gnathosoma and in this manner is closer to E. nahmani than the other species of this subgroup. Considering their presence on different genera, the species are surprisingly similar. Females differ by: setae v 1 smaller in E. nahmani (6 – 7 versus 9 – 13), as are setae sc 2 (60 – 70 versus 95 – 120) and the gnathosomal cheliceral setae (15 – 24 versus 30 – 37); plate H also seems more rudimentary in E. burwelli, although this plate is often distorted or obscured, making comparisons difficult. Larvae also have smaller setae: v 1 (4 – 5 versus 8 – 11), sc 2 (90 – 105 versus 130 – 140) and ch (16 – 25 versus 30 – 38).	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFE3095387AEFBD9FE6EA29E.taxon	etymology	Etymology. It is with great pleasure that I name this species for my son Harry Nahman, who was great company during school- and work-from-home periods in 2020, when he also helped by entering measurement data for several species.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFE3095387AEFBD9FE6EA29E.taxon	discussion	Remarks. The paratype females from Bar Mountain and Tweed Lookout have fully developed larvae in their body. No larvae seem to have egg shells, suggestive of vivipary. These larvae are not included as paratypes. One male (unregistered) from Tweed Lookout is an unknown species of Eutarsopolipus. Its most distinctive features are much larger sc 1 - sc 1 (74 vs 49 – 50), the reduction of tc ″ on tarsi II-III to a minute seta (ca. 1 µm, versus 4 – 5), and the tiny setae pv ″ on tarsi II-III are not spur-like, instead being slender. A larva (unregistered) from Bar Mountain is the same species (setae tcʺ are similarly reduced). These specimens are tentatively considered closest to the despoticus subgroup.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFE8096E87AEF91EFDE1A124.taxon	description	(Figs 17 – 20)	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFE8096E87AEF91EFDE1A124.taxon	diagnosis	Diagnosis. Adult female: Gnathosomal length 58 – 60, width 48 – 50; anterior margin of prodorsal shield eroded medially; v 1 7 – 10, sc 1 13 – 16, c 1 15 – 20, c 2 19 – 23, d 17 – 23, f 14 – 20, h 2 – 4, v 1 - v 1 40 – 46, d-d 115 – 125, f-f 115 – 125; genua II-III setae l′ 2 – 3, tibia I seta l′ 4 – 5, tarsi II-III setae tc′ ′ 5 – 6, pv′ ′ 3 – 4. Larval female: Gnathosomal length 45 – 50, width 38 – 41; idiosomal length 195 – 230; c 1 7 – 11 and c 2 7 – 10 subequal, v 1 – v 1 30 – 33, sc 1 – sc 1 57 – 62, c 1 – c 1 44 – 54, d – d 55 – 67; genua II-III setae l′ m – 2, tibia I seta l′ 6 – 9, tarsi II-III setae tc′ 2 – 3, tc′ ′ 4 – 5, pv′ ′ 2 – 3.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFE8096E87AEF91EFDE1A124.taxon	materials_examined	Material examined. Holotype, female, ex. Castelnaudia septemcostata. Mt Cougal, shelf, 28 ° 14 ʹ 10 ʺ S 152 ° 19 ʹ 34 ʺ E, 19 Nov 1987 - 17 Mar 1988, D. Cook & G. Monteith, pitfall trap, rainforest, IN 6062, HR # T 64265 (QMS 113019). In QM. Paratypes. All ex. C. septemcostata. 4 females, 6 males, 14 larvae, as follows. 1 female, 2 larvae, same data as holotype (QMS 113027 - 28, ZMH-A 0015191); 1 female, O’Reilly’s Guesthouse, 28 ° 14 ʹ 08 ʺ S 152 ° 08 ʹ 12 ʺ E, 1 - 31 Dec 1909, H. Hacker, IN 7977, HR # T 88472 (QMS 113020); 1 female, 1 male, 3 larvae, Tweed Lookout, Wiangaree, 28 ° 23 ʹ 14 ʺ S 153 ° 6 ʹ 33 ʺ E, 16 Nov- 27 Dec 1974, G. & S. R. Monteith, pitfall trap, rainforest, IN 5349, HR # T 57316 (QMS 113024, 113029 - 30, ANIC 52 - 003959); 1 female, 5 males, 6 larvae, Rotary Park, Lismore, 28 ° 48 ʹ 37 ʺ S 153 ° 17 ʹ 43 ʺ E, 16 Nov- 26 Dec 1974, G. & S. R. Monteith, pitfall trap, rainforest, IN 5415, HR # T 57330 (QMS 113021 - 23, 25 - 26, 31 - 32, ANIC 52 - 003960 – 61, ZMH-A 0015192); 1 larva, same data as holotype except 18 Jan- 27 Apr 1987, IN 6056, HR # T 64261; 1 larva, Upper Tallebudgera Creek, 28 ° 14 ʹ 10 ʺ S 152 ° 19 ʹ 34 ʺ E, 19 Nov 1987 - 17 Mar 1988, D. Cook & G. Monteith, pitfall trap, rainforest, IN 6062, HR # T 64272 (QMS 113034); 1 larva, Bar Mountain, Wiangaree, 28 ° 27 ʹ 37 ʺ S 153 ° 8 ʹ 2 ʺ E, 11 Oct 1977 - 2 Oct 1978, IN 5624, G & S. R. Monteith, pitfall trap, rainforest, IN 5624, HR # T 57344 (QMS 113033). All in QM except 1 female, 1 male, 1 larva in ANIC; 1 male, 1 larva in ZMH.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFE8096E87AEF91EFDE1A124.taxon	description	Description. FEMALE (Figs 17 – 18; n = 5). Gnathosoma. Length 58 (58 – 60), width 48 (46 – 50). Cheliceral stylets 68 (68 – 72), setae ch 22 (24 – 27), su 13 (10 – 11). Idiosoma (Fig. 17). Length 335 (325 – 480), width 240 (255 – 320). Prodorsal plate eroded anteriorly; setae slender, v 1 9 (7 – 10), v 2 alveolar remnant, sc 1 16 (13 – 16), sc 2 85 (85 – 90). Distance between setae v 1 – v 1 46 (40 – 51), sc 1 – sc 1 79 (78 – 96), v 1 – sc 1 25 (23 – 27), v 2 – v 2 48 (44 – 58), sc 2 – sc 2 105 (110 – 125), sc 1 – sc 2 25 (23 – 27). Opisthosomal setae slender. Plate C setae c 1 16 (15 – 20), c 2 23 (19 – 23), distance between setae c 1 – c 1 71 (73 – 92). Plate D setae d 21 (17 – 23), d – d 125 (115 – 125), cupuli ia laterad to anterolaterad d. Plate EF setae f 16 (14 – 20), f – f 125 (115 – 125), cupuli im anterolaterad f. Plate H fused weakly, seta h 2 (3 – 4). Venter: coxisterna 1 - 2 well defined, coxisterna 3 weakly defined; coxal setae slender, 1 a 3 (3 – 4), 2 a 5 (5 – 7), 3 a 6 (4 – 7), 3 b 6 (6 – 7). Alveoli of 1 b, 2 b present. Legs (Fig. 18). Setal counts legs I – III, femur-tarsus: 3 - 2 - 6 (+ φ) - 8 (+ ω), 0 - 1 - 4 - 6, 0 - 1 - 4 - 6. Leg I. Femur I, d minute, l’ 2 (2 – 3), v ″ 8 (10 – 16); genu I, l′ 2 (2), l ″ 2 (2); tibia I, d 70 (60 – 65), l’ 5 (4 – 5), l’ ’ 8 (6 – 9), v’ 4 (3 – 4), v’ ’ 10 (8 – 10), k 5 (4), φ 9 (8 – 10); tarsus I, tc’ 14 (15 – 17), tc’ ’ 14 (13 – 17), pl’ 11 (11 – 14), pl’ ’ 19 (14 – 17), pv′ 2 (2), pv’ ’ 2 (2), ω 3 (3 – 4), s 6 (5 – 6), u′ ′ m (m – 2). Leg II. Genu II, l′ 2 (2); tibia II, d 35 (30 – 40), l’ 5 (4 – 5), v’ 11 (8 – 12), v’ ’ 20 (14 – 17); tarsus II, tc′ 3 (2 – 3), tc ″ 5 (5), pl ″ 58 (45 – 46), pv’ ’ 3 (3 – 4), u’ 7 (7), u′ ′ 2 (m – 2). Leg III. Genu III, l′ 2 (2 – 3); tibia III, d 40 (33 – 39), l’ 6 (5), v’ 12 (10 – 12), v’ ’ 24 (15 – 17); tarsus III, tc’ 2 (2), tc ″ 5 (5 – 6), pl ″ 55 (45 – 50), pv’ ’ 2 (2 – 3), u’ 7 (6 – 7), u′ ′ m (m – 2). LARVIFORM ADULT MALE (Fig. 19; n = 6). Gnathosoma. Length 31 – 35, width 29 – 29. Cheliceral stylets 21 – 25, setae ch 7 – 9, su 6 – 7. Idiosoma (Fig. 19). Length 145 – 180, width 105 – 130. Prodorsal plate with setae v 1 4, v 2 alveolar remnant, sc 1 4 – 6, sc 2 45 – 50. Distance between setae v 1 – v 1 18 – 21, sc 1 – sc 1 41 – 45, v 1 – sc 1 14 – 19, v 2 – v 2 19 – 23, sc 2 – sc 2 51 – 60, sc 1 – sc 2 18 – 27. Plate CD setae c 1 5 – 6, c 2 5 – 6, d 5 – 6, distance between setae c 1 – c 1 45 – 53, c 1 – c 2 25 – 33, d – d 37 – 48; cupuli ia not seen or absent. Plate EF eroded posteromedially; with setae f 3 – 4, distance f – f 24 – 30; cupuli im anterolaterad to laterad setae f. Genital capsule length 25 – 30, width 28 – 30, setae h minute. Venter: coxal setae 1 a 2 – 3, 2 a 3 – 4, 3 a 3 – 4, 3 b 4 – 5. Legs. Setal counts legs I – III, femur-tarsus: 3 - 2 - 6 (+ φ) - 8 (+ ω), 0 - 1 - 4 - 6, 0 - 1 - 4 - 6. Leg I. Femur I, d minute, l′ minute, v ″ 5 – 6 (asymmetrical absence on drawn specimen); genu I l′ m – 2, l ″ m – 2; tibia I, d 36 – 42, l’ 2, l’ ’ 3 – 4, v’ 3 – 4, v’ ’ 6 – 7, k 3, φ 7 – 8; tarsus I, tc’ 12 – 14, tc’ ’ 13 – 14, pl’ 6 – 7, pl’ ’ 11 – 13, pv ′ m – 2, pv’ ’ 2, ω 2, s 4 – 5, u ′′ m. Leg II. Genu II, l′ m – 2; tibia II, d 23 – 29, l’ 4, v’ 7 – 10, v’ ’ 14 – 16; tarsus II, tc′ m – 2, tc ″ 4 – 5, pl ″ 24 – 34, pv’ ’ 3, u’ 5 – 6, u ″ minute. Leg III. Genu III, l′ 2; tibia III, d 19 – 24, l’ 4 – 5, v’ 7 – 12, v’ ’ 10 – 15; tarsus III, tc′ minute, tc ″ 4 – 5, pl ″ 23 – 29, pv’ ’ 2, u’ 5 – 6, u ″ minute. LARVAL FEMALE (Fig. 20; n = 14). Gnathosoma. Length 45 – 50, width 38 – 41. Cheliceral stylets 57 – 63, setae ch 22 – 33, su 5 – 9. Idiosoma (Fig. 20). Length 195 – 230, width 140 – 190. Prodorsal plate with setae v 1 5 – 6, v 2 alveolar remnant, sc 1 6 – 7, sc 2 80 – 95. Distance between setae v 1 – v 1 30 – 33, sc 1 – sc 1 57 – 62, v 1 – sc 1 19 – 22, v 2 – v 2 31 – 36, sc 2 – sc 2 68 – 79, sc 1 – sc 2 27 – 33. Plate C setae c 1 7 – 11, c 2 7 – 10, distance between setae c 1 – c 1 44 – 54. Plate D setae d 9 – 12, d – d 55 – 67, cupuli ia very close to setae d, anteriad to anterolaterad d. Plate EF setae f 7 – 13, f – f 44 – 51, cupuli im close to setae f, anteriad to anterolaterad f. Plate H 20 – 22 wide, 23 – 26 long, bearing setae h 1 ca. 120, h 2 7 – 9. Venter: coxisterna 1 - 2 well defined, coxisterna 3 with some folds; coxal setae slender, 1 a 2 – 3, 2 a 3 – 4, 3 a 4 – 5, 3 b 4 – 5. Alveoli of 1 b, 2 b present. Legs. Setal counts legs I – III, femur-tarsus: 3 - 2 - 6 (+ φ) - 8 (+ ω), 0 - 1 - 4 - 6, 0 - 1 - 4 - 6. Leg I. Femur I, d minute, l′ minute, v ″ 6 – 9; genu I, l′ minute, l ″ m – 2; tibia I, d 45 – 61, l’ 6 – 9, l’ ’ 8 – 10, v’ 3 – 4, v’ ’ 5 – 8, k 4 – 5, φ 8 – 10; tarsus I, tc’ 13 – 15, tc’ ’ 14 – 15, pl’ 6 – 7, pl’ ’ 14 – 20, pv′ m – 2, pv’ ’ 2, ω 2 – 3, s 5, u′ ′ minute. Leg II. Genu II, l′ m – 2; tibia II, d 29 – 39, l’ 5 – 6, v’ 8 – 12, v’ ’ 17 – 22; tarsus II, tc′ 2, tc ″ 4, pl ″ 31 – 38, pv’ ’ 2 – 3, u’ 5 – 6, u′ ′ m – 2. Leg III. Genu III, l′ m – 2; tibia III, d 25 – 36, l’ 4 – 5, v’ 8 – 11, v’ ’ 15 – 18; tarsus III, tc’ m – 2, tc ″ 4 – 5, pl ″ 27 – 37, pv’ ’ 2 – 3, u’ 5 – 6, u′ ′ m – 2.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFE8096E87AEF91EFDE1A124.taxon	diagnosis	Differential diagnosis. Eutarsopolipus raveni sp. nov. shares the same host as E. nahmani but is a much smaller species and is instead closer to E. teuceri. See the differential diagnosis of E. teuceri for differences.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFE8096E87AEF91EFDE1A124.taxon	etymology	Etymology. It is with great pleasure that I name this species for Robert Raven, Curator of Arachnida at the Queensland Museum, and a friend and superb arachnologist, who has given great support to the development of acarology at the Queensland Museum.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFD5096B87AEFA41FEE8A1D5.taxon	description	(Figs 21 – 23)	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFD5096B87AEFA41FEE8A1D5.taxon	diagnosis	Diagnosis. Adult female: Gnathosomal length 58 – 61, width 51 – 54; anterior margin of prodorsal shield slightly eroded medially; v 1 8 – 10, sc 1 18 – 20, c 1 17 – 22, c 2 15 – 16, d 18 – 19, f 15 – 18, h 7 – 10, v 1 - v 1 42 – 48, d-d 130 – 140, f-f 125 – 130; genua II-III setae l′ 2 – 4, tibia I seta l′ 6 – 7, tarsi II-III setae tc′ 3 – 4, tc′ ′ 5 – 6, pv′ ′ large, 4. Larval female: Unknown.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFD5096B87AEFA41FEE8A1D5.taxon	materials_examined	Material examined. Holotype female, ex Castelnaudia porphyriaca. 1.5 km from Mt Cabinet, via Jimna, Sunday Ck, 26 ° 42 ʹ 39 ʺ S 152 ° 33 ʹ 49 ʺ, 9 Sep- 30 Nov 1974, G. & S. Monteith, pitfall trap, IN 5370, HR # T 57070 (QMS 113506). Paratypes. 2 females, 3 males, ex C. porphyriaca, Conondale Range, 27 Nov. 1974, G. Monteith, unregistered specimen (QMS 113507, 113510 - 12, ANIC 52 - 003962). All in QM except one female in ANIC.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFD5096B87AEFA41FEE8A1D5.taxon	description	Description. FEMALE (Figs 21 – 22; n = 3). Gnathosoma. Length 58 (60 – 61), width 54 (51 – 54). Cheliceral stylets 77 (75 – 82), setae ch 26 (20 – 27), su 8 (10). Idiosoma (Fig. 21). Length 360 (310 – 325), width 250 (225 – 250). Prodorsal plate not eroded anteriorly; setae slender, v 1 10 (8 – 9), v 2 alveolar remnant, sc 1 20 (18 – 20), sc 2 broken (85 – 105). Distance between setae v 1 – v 1 48 (42 – 44), sc 1 – sc 1 92 (80 – 86), v 1 – sc 1 31 (26 – 30), v 2 – v 2 48 (41 – 43), sc 2 – sc 2 105 (100 – 105), sc 1 – sc 2 28 (23 – 29). Opisthosomal setae slender. Plate C setae c 1 17 (17 – 22), c 2 17 (15 – 16), distance between setae c 1 – c 1 80 (70 – 75). Plate D setae d 19 (18 – 19), d – d 140 (130 – 135), cupuli ia anterolaterad d. Plate EF setae f 18 (15 – 16), f – f 130 (125), cupuli im anterolaterad f. Plate H divided or fused weakly, seta h 7 (7 – 10). Venter: coxisterna 1 - 3 moderately well defined, with few folds, coxisterna 3 weaker than 1 - 2; coxal setae slender, 1 a 4 (3 – 4), 2 a 5 (5 – 6), 3 a 4 (5), 3 b 7 (5 – 6). Alveoli of 1 b, 2 b present. Legs (Fig. 22). Setal counts legs I – III, femur-tarsus: 3 - 2 - 6 (+ φ) - 8 (+ ω), 0 - 1 - 4 - 6, 0 - 1 - 4 - 6. Leg I. Femur I, d minute, l’ m (2), v ″ 8 (7 - 9); genu I, l′ 2 (2 – 3), l ″ 2 (2 – 3); tibia I, d broken (60 – 68), l’ 7 (5 – 6), l’ ’ 6 (8), v’ 4 (4 – 5), v’ ’ 9 (8 – 10), k 4 (3 – 4), φ 9 (8); tarsus I, tc’ 15 (13 – 14), tc’ ’ 14 (15), pl’ 8 (11 – 12), pl’ ’ broken (9 – 15), pv′ 3 (3 – 4), pv’ ’ 3 (4), ω 2 (3), s 6 (6 – 7), u′ ′ m (m). Leg II. Genu II, l′ 2 (3 – 4); tibia II, d 40 (35 – 43), l’ 6 (7), v’ 10 (10 – 14), v’ ’ 17 (18 – 20); tarsus II, tc′ 3 (3 – 4), tc ″ 5 (5 – 6), pl ″ 50 (41 – 43), pv’ ’ 4 (4), u’ 6 (7), u′ ′ m (m). Leg III. Genu III, l′ 2 (3); tibia III, d obscured (32 – 35), l’ 6 (5 – 6), v’ obscured (11 – 13), v’ ’ obscured (14 – 15); tarsus III, tc’ 4 (3), tc ″ 5 (5), pl ″ broken (38 – 42), pv’ ’ 4 (4), u’ 7 (6 – 7), u′ ′ m (m). LARVIFORM ADULT MALE (Fig. 23; n = 3). Gnathosoma. Length 32 – 34, width 21 – 25. Cheliceral stylets 21 – 25, setae ch 5 – 6, su 5 – 6. Idiosoma. Length 155 – 165, width 110 – 125. Prodorsal plate with setae v 1 3 – 4, v 2 alveolar remnant, sc 1 5 – 6, sc 2 62 – 70. Distance between setae v 1 – v 1 16 – 19, sc 1 – sc 1 41 – 42, v 1 – sc 1 15 – 19, v 2 – v 2 18 – 21, sc 2 – sc 2 49 – 50, sc 1 – sc 2 21 – 25. Plate CD setae c 1 6 – 7, c 2 5 – 6, d 6 – 7, distance between setae c 1 – c 1 42 – 45, c 1 – c 2 24 – 29, d – d 35 – 37; cupuli ia anterolaterad setae d. Plate EF slightly eroded medially; with setae f 2 – 4 (thickened in one specimen), distance f – f 22 – 24; cupuli im anterolaterad setae f. Genital capsule length 26 – 29, width 30 – 32, setae h minute. Venter: coxal setae 1 a 2 – 3, 2 a 3 – 4, 3 a 3 – 4, 3 b 4 – 5. Legs. Setal counts legs I – III, femur-tarsus: 3 - 2 - 6 (+ φ) - 8 (+ ω), 0 - 1 - 4 - 6, 0 - 1 - 4 - 6. Leg I. Femur I, d minute, l′ minute, v ″ 3 – 4; genu I l′ 2, l ″ m – 2; tibia I, d 32 – 37, l’ 3 – 4, l’ ’ 3 – 4, v ’ 2 – 3, v ’’ 6 – 10, k 2 – 3, φ 6 – 7; tarsus I, tc’ 13, tc’ ’ 13 – 14, pl’ 6 – 7, pl’ ’ 9 – 12, pv ′ m, pv’ ’ 2, ω 2, s 4 – 5, u ′′ m. Leg II. Genu II, l′ 2; tibia II, d 16 – 17, l’ 5 – 6, v’ 9 – 12, v’ ’ 10 – 15; tarsus II, tc′ 2, tc ″ 4, pl ″ 27 – 33, pv ″ 2 – 3, u’ 4 – 5, u ″ minute. Leg III. Genu III, l′ 2 – 3; tibia III, d 12 – 16, l’ 4, v’ 8 – 10, v ″ 12 – 15; tarsus III, tc′ 2, tc ″ 4 – 5, pl ″ 26 – 32, pv ″ 3, u ’ 5 – 6, u ″ minute. LARVAL FEMALE. Unknown.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFD5096B87AEFA41FEE8A1D5.taxon	diagnosis	Differential diagnosis. Eutarsopolipus teuceri sp. nov. is close to E. raveni. Females are most easily separated by the length of seta h, which is short (2 – 4) in E. raveni but longer (7 – 10) in E. teuceri. Other small differences are the size of setae tc′ (3 – 4 versus 2) and pv′ (4 versus 2 – 3) on tarsus III, which are larger in E. teuceri, and the longer cheliceral stylets (75 – 82 versus 68 – 72). Males are almost identical, differing only by the length of setae sc 2 (45 – 50 vs 62 – 70 in E. raveni), although the smaller tc′ is also apparent (2 versus minute in E. raveni). Eutarsopolipus raveni and E. teuceri both share a small gnathosoma with E. rutherfordae, which is also a member of the nahmani subgroup but its host is Trichosternus subvirens. These two species are most easily distinguished from E. rutherfordae by the strong, stiff setae d and f in E. rutherfordae (very fine, not stiff in E. raveni and E. teuceri). Eutarsopolipus raveni also has small setae h (2 – 4 compared to 7 – 10 in E. rutherfordae and E. teuceri), while E. teuceri has longer cheliceral stylets (75 – 82 versus 60), shorter cheliceral setae (20 – 27 versus 42), shorter seta v′ ′ on femur II (7 – 9 versus 16) and much larger setae pv′ ′ on tarsi II-III (4 versus 2).	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFD5096B87AEFA41FEE8A1D5.taxon	etymology	Etymology. The specific name teuceri refers to the legendry archer Teucer of the Odyssey, who in battle hid behind the glittering shield of Ajax, alluding to the spectacular metallic purple sheen of the elytra of the host beetle E. porphyriaca.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFD5096B87AEFA41FEE8A1D5.taxon	discussion	Remarks. Castelnaudia porphyriaca hosted three species described herein, but also had one larva of another species collected at Conondale. The larva belongs to the pterostichi species group and has an enlarged tibia I seta v′, the only femoral seta is d, and the only setae on all genua is on leg I (seta l′). The ochoai species group - despoticus subgroup	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFD5096B87AEFA41FEE8A1D5.taxon	diagnosis	Diagnosis. Female with dorsal idiosomal setae fine, slender; setae h minute. All life stages with gnathosomal apex evenly rounded.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFD0096987AEF9D1FD11A04D.taxon	description	(Figs 24 – 27)	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFD0096987AEF9D1FD11A04D.taxon	diagnosis	Diagnosis. Adult female: Gnathosomal length 45 – 51, width 47 – 51, cheliceral stylets 50 – 55; tarsi II-III setae tc′ ′ 2 – 3. Larval female: Gnathosomal length 42 – 45, width 43 – 45, cheliceral stylets 45 – 52; tarsi II-III setae tc′ ′ m – 2.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFD0096987AEF9D1FD11A04D.taxon	materials_examined	Material examined. Holotype female, ex Castelnaudia porphyriaca. Conondale Range, 27 Nov. 1974, G. Monteith, unregistered specimen (QMS 113508). Paratypes. All ex C. porphyriaca. 6 females, 2 larvae, as follows: 3 females, 2 larvae, same data as holotype (QMS 113509 – 10, 13 – 4, ANIC 52 - 003963); 3 females, 1.5 km from Mt Cabinet, via Jimna, Sunday Ck, 26 ° 42 ʹ 39 ʺ S 152 ° 33 ʹ 49 ʺE, 9 Sep- 30 Nov 1974, G. & S. Monteith, pitfall trap, HR # T 57070 (QMS 113515 – 16, ZMH-A 0015193). All in QM except 1 female in ANIC and ZMH. Note: two females from Mt Cabinet have fully developed larvae in their body; measurements from these larvae are included where features could be measured.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFD0096987AEF9D1FD11A04D.taxon	description	Description. FEMALE (Figs 24 – 26; n = 4). Gnathosoma (Fig. 25 A). Length 46 (45 – 51), width 50 (47 – 51). Cheliceral stylets 53 (50 – 55), setae ch 27 (28 – 34), su 12 (9 – 16). Idiosoma (Fig. 24). Length 285 (245 – 325), width 255 (230 – 300). Prodorsal plate not eroded anteriorly; setae slender, v 1 11 (9 – 12), v 2 alveolar remnant, sc 1 10 (9 – 10), sc 2 85 (65 – 80). Distance between setae v 1 – v 1 60 (55 – 59), sc 1 – sc 1 98 (87 – 97), v 1 – sc 1 35 (32 – 37; asymmetrically 27 in one specimen), v 2 – v 2 67 (61 – 68), sc 2 – sc 2 125 (125 – 130), sc 1 – sc 2 32 (31 – 35). Opisthosomal setae slender. Plate C setae c 1 9 (7 – 9), c 2 15 (12 – 14), distance between setae c 1 – c 1 100 (105 – 120). Plate D setae d 11 (9 – 11), d – d 130 (125 – 140), cupuli ia close to and anterolaterad d. Plate EF setae f 12 (10 – 11), f – f 74 (70 – 77), cupuli im distant from and anterolaterad f. Plate H entire, seta h minute. Venter: coxisterna 1 - 2 moderately well defined, coxisterna 3 somewhat less defined; coxal setae slender, 1 a 3 (2 – 3), 2 a 5 (4 – 5), 3 a 9 (7 – 9), 3 b 6 (5 – 6). Alveoli of 1 b, 2 b present. Legs (Figs 26 A-C). Setal counts legs I – III, femur-tarsus: 3 - 2 - 6 (+ φ) - 8 (+ ω), 0 - 1 - 4 - 6, 0 - 1 - 4 - 6. Leg I. Femur I, d minute, l’ 2 (2 – 4), v ″ 6 (5 – 7); genu I, l′ 2 (2 – 3), l ″ 3 (2 – 3); tibia I, d 50 (50 – 56), lʹ 11 (8 – 10), lʺ 12 (11 – 12), v’ 6 (6 – 7), v’ ’ 18 (13 – 20), k 5 (4), φ 10 (9 - 10); tarsus I, tc’ 18 (16 – 18), tcʺ 17 (16 – 17), pl’ 15 (12 – 15), plʺ 19 (16 – 17), pv′ minute, pvʺ 2 (2), ω 4 (4 – 5), s 5 (5 – 6), u′ ′ m (m). Leg II. Genu II, l′ 4 (3 – 4); tibia II, d 35 (31 – 37), l’ 9 (8 – 10), v’ 20 (16 – 25), vʺ 26 (22 – 29); tarsus II, tc′ 8 (5 – 9), tc ″ 3 (2 – 3), pl ″ 50 (43 – 52), pvʺ 2 (2), u’ 5 (5 – 6), u′ ′ m (m). Leg III. Genu III, l′ 3 (2 – 3); tibia III, d 30 (28 – 37), l’ 7 (7 – 9), v’ 23 (18 – 25), vʺ 26 (18 – 28); tarsus III, tc’ 4 (4 – 5), tc ″ 3 (2 – 3), pl ″ 50 (36 – 50), pvʺ 2 (2), u’ 5 (5 – 6), u′ ′ m (m). LARVIFORM MALE. Unknown. LARVAL FEMALE (Fig. 27; n = 4). Gnathosoma. Length 42 – 45, width 43 – 45. Cheliceral stylets 45 – 52, setae ch 24 – 27, su 9 – 10. Idiosoma (Fig. 27). Length 195 – 205, width 145 – 165. Prodorsal plate with setae v 1 7 – 9, v 2 alveolar remnant, sc 1 6 – 8, sc 2 90 – 100. Distance between setae v 1 – v 1 34 – 36, sc 1 – sc 1 65 – 70, v 1 – sc 1 26 – 27, v 2 – v 2 45 – 48, sc 2 – sc 2 74 – 81, sc 1 – sc 2 29 – 35. Plate C setae c 1 4 – 8, c 2 10, distance between setae c 1 – c 1 62 – 71. Plate D setae d 8 – 10, d – d 58 – 61, cupuli ia distant from setae d, anterolaterad d. Plate EF setae f 9 – 10, f – f 42 – 45, cupuli im distant to setae f, anteriad f (not seen in drawn specimen). Plate H broad, 25 – 31 wide, ca. 34 – 35 long bearing setae h 1 ca. 180, h 2 6 – 7. Venter: coxisterna 1 - 2 well defined, coxisterna 3 with some membranous folds; coxal setae slender, 1 a 3, 2 a 4 – 5, 3 a 6 – 9, 3 b 5 – 6. Alveoli of 1 b, 2 b present. Legs. Setal counts legs I – III, femur-tarsus: 3 - 2 - 6 (+ φ) - 8 (+ ω), 0 - 1 - 4 - 6, 0 - 1 - 4 - 6. Leg I. Femur I, d minute, l′ 2, v ″ 3 – 4; genu I, l′ 2, l ″ 2; tibia I, d 60 – 65, l’ 11 – 12, l’ ’ 10 – 12, v’ 4 – 5, v’ ’ 14 – 17, k 3 – 4, φ 10; tarsus I, tc’ 16 – 18, tc’ ’ 18, pl’ 14 – 17, pl’ ’ 19 – 25, pv′ minute, pv’ ’ m – 2, ω 4 – 5, s 5, u′ ′ minute. Leg II. Genu II, l′ 2 – 3; tibia II, d 40 – 45, l’ 8 – 9, v’ 17 – 18, v’ ’ 22; tarsus II, tc′ 3 – 4, tc ″ m – 2, pl ″ 32 – 38, pv’ ’ m – 2, u’ 4 – 5, u′ ′ m – 2. Leg III. Genu III, l′ 2; tibia III, d 35 – 39, l’ 6 – 7, v’ 20 – 22, v’ ’ 23 – 29; tarsus III, tc’ 3 – 5, tc ″ m – 2, pl ″ 35 – 43, pv’ ’ m – 2, u’ 4 – 5, u′ ′ minute.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFD0096987AEF9D1FD11A04D.taxon	diagnosis	Differential diagnosis. Eutarsopolipus despoticus sp. nov. is most similar to E. hebronae sp. nov., but differs by having a smaller gnathosoma, cheliceral stylets and setae tc ″ on tarsi II-III. See below the Differential diagnosis of E. hebronae sp. nov. for further details.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFD0096987AEF9D1FD11A04D.taxon	etymology	Etymology. The host’s specific name porphyriaca refers to its purple wings, and as the colour purple was associated with rulers, the name despoticus (a despot, a tyrant) alludes to this species living beneath the purple wings of its host while engaged in a parasitic life style.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFD2096787AEFB69FC50A2D8.taxon	description	(Figs 25 – 26)	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFD2096787AEFB69FC50A2D8.taxon	diagnosis	Diagnosis. Adult female: Gnathosomal length 61 – 65, width 60 – 67, cheliceral stylets 75 – 84; tarsi II-III setae tc′ ′ 3 – 4. Larval female: Gnathosomal length 50 – 55, width 50 – 53, cheliceral stylets 64 – 66; tarsi II-III setae tc′ ′ 2 – 3.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFD2096787AEFB69FC50A2D8.taxon	materials_examined	Material examined. Holotype female, ex Castelnaudia cordata. Mt Mee SF, 6 km NNW barracks, 27 ° 2 ʹ 53 ʺ S 152 ° 40 ʹ 49 ʺ E, 8 Jan- 3 Mar 1992, D. Cook, flight intercept trap, IN 5148, HR # T 68607 (QMS 113463). Paratypes. All ex C. cordata. 6 females, 3 larvae, as follows: all ex. C. cordata. 3 females, 1 larva, same data as holotype (QMS 113464 – 66, ANIC 52 - 003964); 1 larva, McIntosh Rd, Unumbar SF, 28 ° 25 ʹ 19 ʺ S 152 ° 42 ʹ 14 ʺ E, 25 Nov 1972, G. Monteith, IN 6467, HR # T 64329 (ANIC 52 - 003965); 1 female, Lower Coomera, 28 ° 12 ʹ 20 ʺ S 153 ° 10 ʹ 46 ʺ E, 9 Jan- 6 Apr 1995, G. Monteith, pitfall trap, IN 5998, HR # T 68610 (QMS 113467); 2 females, 1 larva, Lever’s Plateau, 28 ° 18 ʹ 57 ʺ S 152 ° 50 ʹ 53 ʺ E, 22 Feb- 8 May 1976, G. & S. R. Monteith, pitfall trap, rainforest, IN 5485, HR # T 68619 (QMS 113468 – 69, ZMH-A 0015194). 1 female, 1 larva in ANIC; 1 female in ZMH. Other material examined. 4 females from type locality, mounted with specimens of E. umbonatus. Not included as paratypes. Notes: 1 female from each of Mt Mee and Lever’s Plateau are E. hebronae but too poor to be considered paratypes. There is also a pharate female from McIntosh Rd.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFD2096787AEFB69FC50A2D8.taxon	description	Description. FEMALE (Figs 25 – 26; n = 6). Gnathosoma (Fig. 25 B). Length 61 (61 – 65), width 60 (60 – 67). Cheliceral stylets 78 (75 – 84), setae ch 35 (29 – 38), su 17 (13 – 15). Idiosoma. Length 420 (345 – 430), width 290 (255 – 340). Prodorsal plate eroded mid-anteriorly; setae slender; setae v 1 6 (5 – 7), v 2 alveolar remnant, sc 1 10 (6 – 10), sc 2 95 (85 – 105). Distance between setae v 1 – v 1 56 (54 – 66), sc 1 – sc 1 105 (93 – 105), v 1 – sc 1 33 (21 – 30), v 2 – v 2 64 (67 – 69), sc 2 – sc 2 140 (135 – 150), sc 1 – sc 2 40 (33 – 43). Opisthosomal setae slender. Plate C setae c 1 9 (7 – 9), c 2 12 (11 – 13), distance between setae c 1 – c 1 82 (69 – 85). Plate D setae d 10 (7 – 10), d – d 145 (130 – 150), cupuli ia well apart from d, anterolaterad d. Plate EF setae f 10 (9 – 13), f – f 86 (79 – 95), cupuli im well apart from f, anterolaterad f. Plate H divided or fused weakly, seta h minute. Venter: coxisterna 1 well defined, coxisterna 2 with some folds, coxisterna 3 weakly defined; coxal setae slender, 1 a 3 (2 – 3), 2 a 5 (4 – 5), 3 a 7 (6 – 7), 3 b 5 (5). Alveoli of 1 b, 2 b present. Legs (Figs 26 D-E). Setal counts legs I – III, femur-tarsus: 3 - 2 - 6 (+ φ) - 8 (+ ω), 0 - 1 - 4 - 6, 0 - 1 - 4 - 6. Leg I. Femur I, d minute, l’ 3 (3), vʺ 8 (6 – 7); genu I, l′ 3 (3), l ″ 3 (3 – 4); tibia I, d 50 (50 – 65), l’ 11 (11 – 13), lʺ 3 (3 – 4), v’ 8 (5 – 7), vʺ 12 (10 – 15), k 5 (5 – 7), φ 10 (10 – 12); tarsus I, tc’ 22 (19 – 20), tcʺ 18 (16 – 20), pl’ 15 (12 – 16), pl’ ’ 23 (20 – 23), pv′ 2 (2), pvʺ 2 (2), ω 5 (4 – 5), s 5 (5 – 6), u′ ′ 2 (m – 2). Leg II. Genu II, l′ 4 (4 – 6); tibia II, d 45 (45 – 50), l’ 11 (9 – 12), v’ 19 (17 – 19), vʺ 21 (22 – 29); tarsus II, tc′ 4 (3 – 4), tc ″ 4 (3 – 4), pl ″ 52 (38 – 50), pvʺ 2 (2), u’ 6 (6 – 7), u′ ′ 2 (m – 2). Leg III. Genu III, l′ 5 (4 – 5); tibia III, d 40 (35 – 50), l’ 8 (8 – 10), v’ 15 (14 – 18), vʺ 30 (26 – 30); tarsus III, tc’ 3 (2 – 3), tc ″ 4 (3 – 4), pl ″ 40 (40 – 50), pvʺ 2 (2), u’ 6 (6), u′ ′ m (m). LARVIFORM MALE. Unknown. LARVAL FEMALE (n = 3). Gnathosoma. Length 50 – 55, width 50 – 53. Cheliceral stylets 64 – 66, setae ch 35 – 36, su 8 – 10. Idiosoma. Length 210 – 270, width 165 – 210. Prodorsal plate with setae v 1 6 – 7, v 2 alveolar remnant, sc 1 7 – 9, sc 2 90 – 120. Distance between setae v 1 – v 1 33 – 40, sc 1 – sc 1 74 – 86, v 1 – sc 1 29 – 34, v 2 – v 2 51 – 60, sc 2 – sc 2 80 – 90, sc 1 – sc 2 25 – 30. Plate C setae c 1 7 – 9, c 2 10 – 13, distance between setae c 1 – c 1 50 – 59. Plate D setae d 8 – 10, d – d 61 – 74, cupuli ia anterolaterad d. Plate EF setae f 9 – 11, f – f 43 – 52, cupuli im anteriad to anterolaterad f. Plate H long and narrow, 16 – 20 wide, ca. 30 – 40 long bearing setae h 1 ca. 150, h 2 8 – 10. Venter: coxisterna 1 - 2 moderately well defined, with few folds, coxisterna 3 poorly defined, membranous; coxal setae slender, 1 a 3, 2 a 5, 3 a 5, 3 b 5 – 6. Alveoli of 1 b, 2 b present. Legs. Setal counts legs I – III, femur-tarsus: 3 - 2 - 6 (+ φ) - 8 (+ ω), 0 - 1 - 4 - 6, 0 - 1 - 4 - 6. Leg I. Femur I, d minute, l′ minute, v ″ 5 – 7; genu I, l′ 3, l ″ 2 – 3; tibia I, d 75 – 80, l’ 3, l’ ’ 2 – 3, v’ 6 – 8, v’ ’ 9 – 12, k 3 – 5, φ 11 – 12; tarsus I, tc’ 15 – 16, tc’ ’ 16, pl’ 9 – 13, pl’ ’ 18 – 20, pv′ m – 2, pv’ ’ 2, ω 3 – 4, s 5 – 6, u′ ′ minute. Leg II. Genu II, l′ 3 – 4; tibia II, d 45 – 50, l’ 9 – 10, v’ 13 – 18, v’ ’ 23 – 26; tarsus II, tc′ 2 – 3, tc ″ 2 – 3, pl ″ 45 – 50, pv’ ’ 2, u’ 5 – 6, u′ ′ minute. Leg III. Genu III, l′ 3 – 4; tibia III, d 40 – 42, l’ 9 – 13, v’ 12 – 15, v’ ’ 20 – 26; tarsus III, tc’ 2, tc ″ 2 – 3, pl ″ 40 – 50, pv’ ’ 2, u’ 5 – 6, u′ ′ m – 2.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFD2096787AEFB69FC50A2D8.taxon	diagnosis	Differential diagnosis. Eutarsopolipus hebronae sp. nov. differs from E. despoticus sp. nov. by having a much larger gnathosoma (61 – 65 × 60 – 67 versus 45 – 51 × 47 – 51) and cheliceral stylets (75 – 84 versus 50 – 55). The species also differ in the size of setae tc ″ on tarsi II-III, being much larger in E. hebronae (3 – 4) than E. despoticus (2 – 3). Even though these setae are only about 1 μm longer, they are obviously larger, being around twice as massive in E. hebronae. These differences in gnathosomal size and tarsal setae are present in the adult and larval female.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFD2096787AEFB69FC50A2D8.taxon	etymology	Etymology. It is with great pleasure that I name this species for Wendy Hebron, with whom I have shared the role of collection manager for 14 years. Not only has her welcome and proficient assistance in the Queensland Museum arachnology collection been greatly appreciated, but her fine efforts have allowed me to dedicate more time to acarology.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFD2096787AEFB69FC50A2D8.taxon	discussion	Remarks. This species was found co-infesting its host species with E. umbonatus sp. nov. and an undescribed species of Eutarsopolipus close to E. echinatus Seeman, 2019 a (Tables 1, 2). The ochoai species group - basiatus subgroup Diagnosis. Female with dorsal idiosomal setae fine, slender; setae h minute or alveolar vestiges. All life stages with gnathosomal apex with small paired lobes.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFD2096787AEFB69FC50A2D8.taxon	discussion	Remarks. This species was found co-infesting its host species with E. umbonatus sp. nov. and an undescribed species of Eutarsopolipus close to E. echinatus Seeman, 2019 a (Tables 1, 2).	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFD2096787AEFB69FC50A2D8.taxon	diagnosis	Diagnosis. Female with dorsal idiosomal setae fine, slender; setae h minute or alveolar vestiges. All life stages with gnathosomal apex with small paired lobes.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFDC096787AEF8DDFDD1A3B0.taxon	diagnosis	Diagnosis. Female with dorsal idiosomal setae fine, slender; setae h minute or alveolar vestiges. All life stages with gnathosomal apex with small paired lobes.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFDD096087AEFF5AFE69A749.taxon	description	(Figs 28 – 31)	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFDD096087AEFF5AFE69A749.taxon	diagnosis	Diagnosis. Adult female: sc 1 5 – 7, c 1 6 – 9, c 2 9 – 14, d 7 – 9, f 6 – 7, v 2 - v 2 48 – 62, sc 1 - sc 1 36 - 46, d-d 91 – 115, f-f 76 – 92; genua II-III setae l′ 3 – 4, tibia I setae l′ 4 – 6, v′ 5 – 6, tarsi II-III setae tc′ ′ 2 – 3. Adult male: leg I with two claws; c 1 3 – 4, c 2 4 – 5, v 2 - v 2 26 – 33; tarsi II-III setae tc ″ m – 2. Larval female: c 1 6 – 8, c 2 8 – 10, d 6 – 10, f 7 – 9, sc 2 – sc 2 64 – 71, v 1 - sc 1 27 – 32; tarsi II-III setae tc′ ′ minute.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFDD096087AEFF5AFE69A749.taxon	materials_examined	Material examined. Holotype female, ex Castelnaudia porphyriaca. Conondale Range, 27 Nov. 1974, G. Monteith, unregistered specimen. Paratypes. All ex C. porphyriaca. 4 females, 4 males, 5 larvae, as follows: 1 female, 2 males, 2 larvae, same data as holotype (QMS 113518 – 20, ANIC 52 - 003966 – 67); 3 females, 2 males, 2 larvae, 1.5 km from Mt Cabinet, via Jimna, Sunday Ck, 26 ° 42 ʹ 39 ʺ S 152 ° 33 ʹ 49 ʺE, 9 Sep- 30 Nov 1974, G. & S. Monteith, pitfall trap, IN 5370, HR # T 57070 (QMS 113521 – 25, ANIC 52 - 003968, ZMH-A 0015195); 1 female, Conondale Range, site 1, 26 ° 42 ʹ 46 ʺ S 152 ° 37 ʹ 08 ʺE, 20 Apr – 2 May 1995, B. Nomann & G. Thompson, pitfall trap, IN 7291, HR # T 68573 (QMS 113526). All in QM except 1 female, 1 male, 1 larva in ANIC and 1 female in ZMH. Note: 1 female from Conondale has a fully developed larva in its body; measurements from this larva are included where features could be measured.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFDD096087AEFF5AFE69A749.taxon	description	Description. FEMALE (Figs 28 – 29; n = 5). Gnathosoma (Fig. 29 D). Length 45 (45 – 46), width 44 (42 – 45), apex indented, terminating in small but distinct lobes. Cheliceral stylets 40 (38 – 44), setae ch 20 (16 – 24), su 9 (8 – 12). Idiosoma (Fig. 28). Length 315 (275 – 325) width 405 (295 – 470). Prodorsal plate not eroded anteriorly; setae slender, v 1 6 (5 – 7), v 2 alveolar remnant, sc 1 6 (5 – 7), sc 2 76 (69 – 82). Distance between setae v 1 – v 1 41 (34 – 38), sc 1 – sc 1 100 (82 – 94), v 1 – sc 1 42 (36 – 37; asymmetry in one specimen: 36 / 46), v 2 – v 2 62 (48 – 58), sc 2 – sc 2 110 (90 – 105), sc 1 – sc 2 26 (27 – 32). Opisthosomal setae slender. Plate C setae c 1 9 (6 – 8), c 2 13 (9 – 14), distance between setae c 1 – c 1 83 (69 – 77). Plate D setae d 9 (7 – 9), d – d 115 (91 – 110), cupuli ia anterolaterad d. Plate EF setae f 7 (6 – 7), f – f 85 (76 – 92), cupuli im anteriad to anterolaterad f. Plate H divided, seta h minute. Venter: coxisterna 1 - 2 moderately well defined, with few folds, coxisterna 3 membranous; coxal setae slender, 1 a 3 (2 – 3), 2 a 4 (4 – 5), 3 a 5 (4 – 6), 3 b 6 (5 – 6). Alveoli of 1 b, 2 b present. Legs (Figs 29 A-C). Setal counts legs I – III, femur-tarsus: 3 - 2 - 6 (+ φ) - 8 (+ ω), 0 - 1 - 4 - 6, 0 - 1 - 4 - 6. Leg I. Femur I, d minute, l’ m (m – 2), v ″ 4 (4 – 5); genu I, l′ 3 (2 – 3), l ″ 2 (2 – 3); tibia I, d 60 (45 – 55), l’ 4 (5 – 6), l’ ’ 7 (7 – 8), v’ 6 (5 – 6), v’ ’ 18 (14 – 17), k 5 (5 – 6), φ 10 (9 – 10); tarsus I, tc’ 17 (14 – 17), tc’ ’ 16 (15 – 16), pl’ 10 (8 – 9), pl’ ’ 20 (15 – 21), pv′ m (m), pv’ ’ 2 (2), ω 5 (4 – 5), s 5 (5), u′ ′ m (m). Leg II. Genu II, l′ 4 (3 – 4); tibia II, d 35 (30 – 42), l’ 5 (6 – 8), v’ 21 (19 – 21), v’ ’ 28 (21 – 28); tarsus II, tc′ 2 (2 – 4), tc ″ 2 (2), pl ″ 45 (37 – 46), pv’ ’ 2 (2), u’ 5 (5 – 6), u′ ′ m (2). Leg III. Genu III, l′ 4 (3 – 4); tibia III, d 35 (32 – 43), l’ 7 (5 – 8), v’ 19 (15 – 21), v’ ’ 24 (19 – 25); tarsus III, tc’ 3 (2 – 4), tc ″ 2 (2 – 3), pl ″ 35 (30 – 37), pv’ ’ 2 (2), u’ 5 (4 – 5), u′ ′ m (m). LARVIFORM ADULT MALE (Fig. 30; n = 4). Gnathosoma. Length 28 – 30, width 29 – 31, apex indented, terminating in small but distinct lobes. Cheliceral stylets 17 – 22, setae ch 5 – 7, su 4 – 7. Idiosoma (Fig. 30). Length 140 – 175, width 105 – 120. Prodorsal plate with setae v 1 4 – 5, v 2 alveolar remnant, sc 1 3 – 5, sc 2 55 – 65. Distance between setae v 1 – v 1 19 – 24, sc 1 – sc 1 42 – 46, v 1 – sc 1 23 – 26, v 2 – v 2 26 – 32, sc 2 – sc 2 53 – 58, sc 1 – sc 2 16 – 17 (one asymmetrical specimen 14 / 19). Plate CD setae c 1 3 – 4, c 2 4 – 5, d 4 – 5, distance between setae c 1 – c 1 43 – 51, c 1 – c 2 23 – 26, d – d 37 – 42; cupuli ia laterad setae d. Plate EF slightly eroded medially; with setae f 4 – 5, distance f – f 27 – 28; cupuli im anteriad setae f. Genital capsule length 24 – 27, width 37 – 41, setae h m – 2. Venter: coxal setae 1 a 3 – 4, 2 a 3 – 4, 3 a 3 – 5, 3 b 4 – 6. Legs. Leg I with two claws. Setal counts legs I – III, femur-tarsus: 3 - 2 - 6 (+ φ) - 8 (+ ω), 0 - 1 - 4 - 6, 0 - 1 - 4 - 6. Leg I. Femur I, d minute, v ″ 3 – 4; genu I l′ 2 – 3, l ″ 5 – 6; tibia I, d 29 – 45, l’ 5, l’ ’ 2, v ’ 3 – 4, v ’’ 8 – 13, k 3 – 4, φ 8 – 9; tarsus I, tc’ 14 – 15, tc’ ’ 13 – 15, pl’ 7 – 10, pl’ ’ 11 – 15, pv ′ minute, pv’ ’ m – 2, ω 4 – 5, s 4, u ′′ minute. Leg II. Genu II, l′ 2 – 3; tibia II, d 29 – 38, l’ 4 – 6, v’ 10 – 14, v’ ’ 15 – 19; tarsus II, tc′ 2 – 3, tc ″ m – 2, pl ″ 26 – 32, pv’ ’ minute, u’ 4 – 5, u ″ minute. Leg III. Genu III, l′ 2; tibia III, d 18 – 26, l’ 3 – 5, v’ 10 – 14, v’ ’ 15 – 22; tarsus III, tc′ 2 – 3, tc ″ m – 2, pl ″ 24 – 30, pv’ ’ minute, u’ 4, u ″ minute. LARVAL FEMALE (Fig. 31; n = 6). Gnathosoma. Length 36 – 41, width 37 – 39, apex indented, terminating in small but distinct lobes. Cheliceral stylets 34 – 37, setae ch 34 – 37, su 7 – 10. Idiosoma (Fig. 31). Length 220 – 245, width 150 – 175. Prodorsal plate with setae v 1 4 – 8, v 2 alveolar remnant, sc 1 3 – 6, sc 2 70 – 85. Distance between setae v 1 – v 1 29 – 32, sc 1 – sc 1 63 – 73, v 1 – sc 1 31 – 32 (asymmetrical in one specimen, 33 / 27), v 2 – v 2 42 – 45, sc 2 – sc 2 64 – 71, sc 1 – sc 2 25 – 26 (asymmetrical in one specimen, 27 / 23). Plate C setae c 1 6 – 8, c 2 8 – 10, distance between setae c 1 – c 1 51 – 57. Plate D setae d 6 – 10, d – d 48 – 54, cupuli ia anterolaterad setae d. Plate EF setae f 7 – 9 (one specimen expressing two f setae on left, possibly in place of cupule im), f – f 40 – 51, cupuli im anteriad f. Plate H narrow, large, 23 – 29 wide, 35 – 38 long, bearing setae h 1 ca. 170, h 2 6 – 8. Venter: coxisterna 1 - 2 moderately well defined, with some folds, coxisterna 3 membranous; coxal setae slender, 1 a 3, 2 a 3 – 5, 3 a 5 – 7, 3 b 4 – 5. Alveoli of 1 b, 2 b present. Legs. Setal counts legs I – III, femur-tarsus: 3 - 2 - 6 (+ φ) - 8 (+ ω), 0 - 1 - 4 - 6, 0 - 1 - 4 - 6. Leg I. Femur I, d minute, l′ minute, v ″ 3 – 4; genu I, l′ 2, l ″ 2; tibia I, d 50 – 55, l’ 5 – 8, l’ ’ 8 – 10, v’ 5 – 6, v’ ’ 11 – 14, k 4 – 5, φ 10 – 11; tarsus I, tc’ 15 – 16, tc’ ’ 14 – 15, pl’ 10 – 13, pl’ ’ 20 – 23, pv′ minute, pv’ ’ minute, ω 5, s 4 – 5, u′ ′ minute. Leg II. Genu II, l′ 2 – 3; tibia II, d 32 – 40, l’ 6 – 7, v’ 15 – 18, v’ ’ 17 – 27; tarsus II, tc′ m – 2, tc ″ minute, pl ″ 26 – 38, pv’ ’ m – 2, u’ 4 – 5, u′ ′ minute. Leg III. Genu III, l′ 2; tibia III, d 31 – 40, l’ 5 – 7, v’ 16 – 17, v’ ’ 21 – 23; tarsus III, tc’ 2, tc ″ minute, pl ″ 31 – 38, pv’ ’ m – 2, u’ 4 – 5, u′ ′ minute.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFDD096087AEFF5AFE69A749.taxon	diagnosis	Differential diagnosis. The most obvious difference between E. basiatus sp. nov. and all other species on Castelnaudia is the retention of two claws on leg I of the male. This species is also distinctive within the basiatus subgroup by having: tiny setae tc ″ on tarsi II-III (minute- 2 versus distinctly larger (3 – 5) in all other species, being at least twice as massive); and shorter idiosomal setae (e. g., females with short d (7 – 9) versus 12 – 23 in the other species).	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFDD096087AEFF5AFE69A749.taxon	etymology	Etymology. The specific name basiatus (a kisser) refers to the distinctive lip-like protrusion of the gnathosoma found in this species group.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFDB097B87AEFC6DFEB7A3EE.taxon	description	(Figs 29, 32 – 36)	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFDB097B87AEFC6DFEB7A3EE.taxon	diagnosis	Diagnosis. Adult female: sc 1 15 – 19, c 1 20 – 22, c 2 21 – 24, d 20 – 24, f 10 – 16, v 2 - v 2 57 – 64, sc 1 - sc 1 84 - 92, d-d 135 – 150, f-f 100 – 120; genua II-III setae l′ 2 – 3, tibia I setae l′ 7 – 8, v′ 3 – 4, tarsi II-III setae tc′ ′ 4 – 5. Adult male: leg I with one claw; c 1 7 – 12, c 2 8 – 10, v 2 - v 2 34 – 40; tarsi II-III setae tc ″ 3 – 4. Larval female: c 1 15 – 18, c 2 19 – 24, d 17 – 19, f 11 – 15, sc 2 – sc 2 83 – 100, v 1 - sc 1 22 – 28; tarsi II-III setae tc′ ′ 3 – 4.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFDB097B87AEFC6DFEB7A3EE.taxon	materials_examined	Material examined. Holotype female, ex. Castelnaudia marginifera. Mt Mistake Plateau, via Goomburra, 27 ° 58 ʹ 37 ʺ S 152 ° 22 ʹ 58 ʺ E, 17 - 22 Nov 1987, G. Monteith, rainforest, IN 6919, HR # T 88508 (QMS 113432). Paratypes. All ex. C. marginifera. 27 females, 10 males, 11 larvae, as follows: 6 females, 2 males, 2 larvae, same data as holotype (QMS 113433 – 39); 8 females, 4 males, Bald Mtn area, via Emu Vale, 28 ° 13 ʹ 57 ʺ S 152 ° 24 ʹ 54 ʺ E, 26 Jan- 30 Jan 1973, G. Monteith, IN 9131, HR # T 88513 (QMS 113440 – 49, ANIC 52 - 003969); 13 females, 4 males, 9 larvae, Lever’s Plateau, 28 ° 18 ʹ 57 ʺ S 152 ° 50 ʹ 53 ʺ E, 29 Feb- 11 Apr 1992, D. Cook, pitfall trap, rainforest, IN 5139, HR # T 64339 (QMS 113450 – 62, ANIC 52 - 003970 – 71, ZMH-A 0015196 – 98). All in QM except 1 female, 1 male, 1 larva in ANIC and ZMH. Other material examined. 1 pharate female, same data as holotype.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFDB097B87AEFC6DFEB7A3EE.taxon	description	Description. FEMALE (Figs 29, 32 – 33; n = 27, 7 measured). Gnathosoma (Fig. 29 E). Length 49 (47 – 51), width 42 (41 – 43), apex indented, terminating in small but distinct lobes. Cheliceral stylets 41 (40 – 45), setae ch 17 (17 – 21), su 10 (7 – 12). Idiosoma (Figs 32 – 33). Length 305 (300 – 370) width 235 (265 – 300). Prodorsal plate not eroded anteriorly; setae slender, v 1 7 (7 – 10), v 2 alveolar remnant, sc 1 asymmetrical lengths, 17 and 12 (15 – 19, not asymmetrical), sc 2 80 (75 – 84). Distance between setae v 1 – v 1 45 (43 – 48), sc 1 – sc 1 88 (85 – 92), v 1 – sc 1 24 (27 – 33), v 2 – v 2 57 (58 – 64), sc 2 – sc 2 150 (140 – 155), sc 1 – sc 2 48 (38 – 48). Opisthosomal setae slender. Plate C setae c 1 21 (20 – 22), c 2 22 (21 – 24), distance between setae c 1 – c 1 83 (72 – 97). Plate D setae d 20 (21 – 24), d – d 140 (135 – 150), cupuli ia anterolaterad d. Plate EF setae f 11 (10 – 16), f – f 110 (100 – 120), cupuli im anteriad to anterolaterad f. Plate H divided, seta h minute. Venter: coxisterna 1 - 2 moderately well defined, with few folds, coxisterna 3 defined but membranous; coxal setae slender, 1 a 3 (3), 2 a 5 (4 – 5), 3 a 6 (6 – 7), 3 b 6 (6 – 7). Alveoli of 1 b, 2 b present. Legs (Figs 33 B-D). Setal counts legs I – III, femur-tarsus: 3 - 2 - 6 (+ φ) - 8 (+ ω), 0 - 1 - 4 - 6, 0 - 1 - 4 - 6. Ranges given only when holotype measurement different from E. basiatus. Leg I. Femur I, d minute, l’ 2, v ″ 6 (5 – 8); genu I, l′ 2, l ″ 2; tibia I, d 45 (40 – 50), l’ 8 (7 – 8), l’ ’ 14 (10 – 12), v’ 4 (3 – 4), v’ ’ 16, k 4 (4 – 5), φ 11 (10 – 11); tarsus I, tc’ 14, tc’ ’ 15, pl’ 10, pl’ ’ 19, pv′ 2 (m – 2), pv’ ’ 2, ω 4, s 5, u′ ′ m (m). Leg II. Genu II, l′ 3; tibia II, d 40, l’ 7, v’ 13 (13 – 15), v’ ’ 40 (35 – 40); tarsus II, tc′ 2, tc ″ 5 (4 – 5), pl ″ 40, pv’ ’ 2, u’ 7 (6 – 7), u′ ′ minute. Leg III. Genu III, l′ 2 (2); tibia III, d 35, l’ 5, v’ 18, v’ ’ 21; tarsus III, tc’ 2 (2), tc ″ 5 (4 – 5), pl ″ 30, pv’ ’ 2, u’ 7 (6 – 7), u′ ′ minute. LARVIFORM ADULT MALE (Fig. 34; n = 10, 5 measured). Gnathosoma. Length 31 – 33, width 29 – 30, apex indented, terminating in small but distinct lobes; pharynx distorted in drawn specimen. Cheliceral stylets 20 – 23, ch 5 – 6, su 4 – 5. Idiosoma. Length 180 – 200, width 145 – 175. Prodorsal plate with setae v 1 3 – 5, v 2 alveolar remnant, sc 1 5 – 7, sc 2 50 – 65. Distance between setae v 1 – v 1 28 – 32, sc 1 – sc 1 47 – 53, v 1 – sc 1 17 – 22 (asymmetry in one specimen, 11 / 19), v 2 – v 2 34 – 40, sc 2 – sc 2 69 – 78, sc 1 – sc 2 16 – 25 (one asymmetrical specimen 32 / 25). Plate CD setae c 1 7 – 12, c 2 8 – 10, d 6 – 10 (asymmetrical in one specimen, 7 / 13), distance between setae c 1 – c 1 48 – 57, c 1 – c 2 34 – 42 (asymmetrical in one specimen, 25 / 39), d – d 44 – 53; cupuli ia anterolaterad setae d. Plate EF slightly eroded medially; with setae f 4 – 5, distance f – f 26 – 32; cupuli im anterolaterad setae f. Genital capsule length 28 – 29, width 38 – 42, setae h minute. Venter: coxal setae 1 a 3, 2 a 4 – 5, 3 a 2 – 3, 3 b 4 – 5. Legs. Setal counts legs I – III, femur-tarsus: 3 - 2 - 6 (+ φ) - 8 (+ ω), 0 - 1 - 4 - 6, 0 - 1 - 4 - 6. Ranges given only when first measurement different from E. basiatus. Leg I. Femur I, d minute, l′ minute, v ″ 4 – 6; genu I l′ m – 2, l ″ m – 2; tibia I, d 45, l’ m – 2, l’ ’ m – 2, v ’ 4 – 5, v ’’ 9 – 14, k 3, φ 8; tarsus I, tc’ 13 – 14, tc’ ’ 13 – 14, pl’ 10 – 11, pl’ ’ 12, pv ′ m – 2, pv’ ’ m – 2, ω 2 – 3, s 4 – 5, u ′′ minute. Leg II. Genu II, l′ 2; tibia II, d 21 – 30, l’ 6 – 7, v’ 14, v’ ’ 16; tarsus II, tc′ 2, tc ″ 3 – 4, pl ″ 26, pv’ ’ 2, u’ 5, u ″ minute. Leg III. Genu III, l′ minute; tibia III, d 19, l’ 5 – 6, v’ 14, v’ ’ 14 – 18; tarsus III, tc′ 2, tc ″ 3 – 4, pl ″ 26, pv’ ’ 2, u’ 5, u′ ′ minute. LARVAL FEMALE (Figs 35 – 36; n = 11, 5 measured). Gnathosoma. Length 42 – 44, width 37 – 40, apex indented, terminating in small but distinct lobes. Cheliceral stylets 36 – 43, setae ch 20 – 24, su 5 – 9. Idiosoma (Fig. 35). Length 220 – 265, width 155 – 205. Prodorsal plate with setae v 1 5 – 7, v 2 alveolar remnant, sc 1 6 – 8, sc 2 75 – 100. Distance between setae v 1 – v 1 36 – 46, sc 1 – sc 1 66 – 81, v 1 – sc 1 22 – 28 (asymmetrical in one specimen: v 2 and sc 1 reversed in position), v 2 – v 2 48 – 64, sc 2 – sc 2 83 – 100, sc 1 – sc 2 28 – 32. Plate C setae c 1 15 – 18, c 2 19 – 24, distance between setae c 1 – c 1 49 – 60. Plate D setae d 17 – 19, d – d 60 – 80, cupuli ia closely anteriad to anterolaterad setae d. Plate EF setae f 11 – 15, f – f 46 – 64, cupuli im anteriad to anterolaterad f. Plate H narrow, large, 20 – 24 wide, 34 – 42 long, bearing setae h 1 ca. 170, h 2 10 – 17. Venter: coxisterna 1 - 2 moderately well defined, with some folds, coxisterna 3 membranous; coxal setae slender, 1 a 2 – 3, 2 a 4 – 5, 3 a 4 – 5, 3 b 5 – 6. Alveoli of 1 b, 2 b present. Legs (Fig. 36). Setal counts legs I – III, femur-tarsus: 3 - 2 - 6 (+ φ) - 8 (+ ω), 0 - 1 - 4 - 6, 0 - 1 - 4 - 6. Ranges given only when first measurement different from E. basiatus. Leg I. Femur I, d minute, l′ minute, v ″ 4 – 6; genu I, l′ m – 2, l ″ m – 2; tibia I, d 50 – 60, l’ 8 – 12, l’ ’ 7 – 12, v’ 6 – 7, v’ ’ 11 – 15, k 4, φ 10; tarsus I, tc’ 12 – 14, tc’ ’ 13 – 14, pl’ 10, pl’ ’ 15 – 19, pv′ minute, pv’ ’ m – 2, ω 3 – 4, s 5, u′ ′ minute. Leg II. Genu II, l′ m – 2; tibia II, d 35, l’ 7, v’ 12 – 17, v’ ’ 19; tarsus II, tc′ 2 – 3, tc ″ 3 – 4, pl ″ 30, pv’ ’ 2, u’ 5, u′ ′ minute. Leg III. Genu III, l′ minute; tibia III, d 28 – 34, l’ 7, v’ 16, v’ ’ 18 – 25; tarsus III, tc’ 2, tc ″ 3 – 4, pl ″ 27 – 36, pv’ ’ 2, u’ 5 – 6, u′ ′ minute.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFDB097B87AEFC6DFEB7A3EE.taxon	diagnosis	Differential diagnosis. Eutarsopolipus labiatus sp. nov. is most similar to E. savatus sp. nov. by sharing long setae sc 1 (exceeding 10) and greater distance d-d (135 – 150) and f-f (100 – 120). Females are very similar and differ by the shorter cheliceral seta (17 – 21 versus 24 – 39) and slightly longer idiosomal setae. The idiosomal setal lengths are however more distinctive in the larvae: c 1 15 – 18, c 2 19 – 24, d 17 – 19 in E. labiatus versus c 1 6 – 9, c 2 9 – 12, d 7 – 10 in E. savatus.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFDB097B87AEFC6DFEB7A3EE.taxon	etymology	Etymology. The specific name labiatus (lipped) refers to the protruding lip-like gnathosoma found in this species group.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFC1097887AEFF5AFD30A067.taxon	description	(Figs 37 – 38)	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFC1097887AEFF5AFD30A067.taxon	diagnosis	Diagnosis. Adult female: sc 1 7 – 8, c 1 15 – 17, c 2 16 – 18, d 16 – 22, f 6 – 11, v 2 - v 2 35 – 40, sc 1 - sc 1 60 – 65, d-d 95 – 98, f-f 67 – 80; genua II-III setae l′ m – 2, tibia I setae l′ 7 – 8, v′ 2 – 3, tarsi II-III setae tc′ ′ 3 – 4. Adult male: leg I with one claw; c 1 5, c 2 5, v 2 - v 2 27; tarsi II-III setae tc ″ 2 – 3. Larval female: c 1 11, c 2 9, d 10, f 7, sc 2 – sc 2 71, v 1 - sc 1 22; tarsi II-III setae tc′ ′ 3.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFC1097887AEFF5AFD30A067.taxon	materials_examined	Material examined. Holotype female, ex. Castelnaudia cordata. “ Glen Witheren ”, shelf scrub, 28 ° 2 ʹ 58 ʺ S 153 ° 6 ʹ 58 ʺ E, 30 Dec 1991 - 24 May 1992, G. Monteith, pitfall trap, IN 5682, rainforest, HR # T 68606 (QMS 113471). In QM. Paratypes. All ex. C. cordata. 2 females, 1 male, 1 larva, as follows: 1 female, 1 male, 1 larva, same data as holotype (QMS 113472 - 74); 1 female, Mt Clunie summit, 28 ° 18 ʹ 31 ʺ S 152 ° 31 ʹ 33 ʺ E, 16 Dec 1972, G. Monteith, rainforest, IN 6361, HR # T 64324 (ANIC 52 - 003972). All in QM except 1 female in ANIC. Note: The holotype and paratype females from “ Glen Witherin ” are on one slide.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFC1097887AEFF5AFD30A067.taxon	description	Description. FEMALE (Figs 37 – 38; n = 3). Gnathosoma. Length 42 (44), width 40 (36 – 38), apex indented, terminating in small but distinct lobes. Cheliceral stylets 21 (23 – 25), setae ch 36 (35 – 41), su 8 (12 – 13). Idiosoma (Fig. 37). Length 275 (290 – 310) width 185 (180 – 200). Prodorsal plate not eroded anteriorly; setae slender, v 1 6 (6 – 7), v 2 alveolar remnant, sc 1 7 (7 – 8), sc 2 90 (80 – 85). Distance between setae v 1 – v 1 33 (30 – 36), sc 1 – sc 1 65 (60 – 61), v 1 – sc 1 20 (16 – 22), v 2 – v 2 38 (35 – 40), sc 2 – sc 2 105 (93 – 105), sc 1 – sc 2 35 (31 – 37). Opisthosomal setae slender. Plate C setae c 1 17 (15 – 17), c 2 18 (16), distance between setae c 1 – c 1 75 (59 – 72). Plate D setae d 22 (16 – 20), d – d 98 (95 – 98), cupuli ia anterolaterad d. Plate EF setae f 10 (6 – 11), f – f 75 (67 – 80), cupuli im anterolaterad f. Plate H divided, seta h minute. Venter: coxisterna 1 - 2 moderately well defined, with few folds, coxisterna 3 membranous; coxal setae slender, 1 a 3 (2 – 3), 2 a 4 (4 – 5), 3 a 5 (4 – 6), 3 b 5 (5 – 6). Alveoli of 1 b, 2 b present. Legs (Fig. 38). Setal counts legs I – III, femur-tarsus: 3 - 2 - 6 (+ φ) - 8 (+ ω), 0 - 1 - 4 - 6, 0 - 1 - 4 - 6. Ranges given only when holotype measurement different from E. basiatus. Leg I. Femur I, d minute, l’ minute, v ″ 5 (5 – 8); genu I, l′ 2 (2), l ″ 2; tibia I, d 50, l’ 7 (7 – 8), l’ ’ 8, v’ 2 (2 – 3), v’ ’ 8 (8 – 10), k 5, φ 10; tarsus I, tc’ 15, tc’ ’ 13 (13 – 14), pl’ 8, pl’ ’ 21, pv′ 2 (2), pv’ ’ 2, ω 3 (3), s 5, u′ ′ m. Leg II. Genu II, l′ m (m); tibia II, d 36, l’ 6, v’ 15 (18), v’ ’ 22; tarsus II, tc′ 2 (m – 2), tc ″ 4 (3 – 4), pl ″ 45, pv’ ’ 2, u’ 6, u′ ′ m. Leg III. Genu III, l′ m (m – 2); tibia III, d 33, l’ 7, v’ 10 (10 – 17), v’ ’ 25; tarsus III, tc’ 2 (2), tc ″ 4 (3 – 4), pl ″ 45 (42 – 45), pv’ ’ 2, u’ 7 (6), u′ ′ m. LARVIFORM ADULT MALE (n = 1). Gnathosoma. Length 28, width 26, apex indented, terminating in small but distinct lobes. Cheliceral stylets 20, setae ch 6, su 5. Idiosoma. Length 140, width 110. Prodorsal plate with setae v 1 4 (absent one side), v 2 alveolar remnant, sc 1 3, sc 2 55. Distance between setae v 1 – v 1 not measureable, sc 1 – sc 1 41, v 1 – sc 1 17, v 2 – v 2 27, sc 2 – sc 2 54, sc 1 – sc 2 16. Plate CD setae c 1 5, c 2 5, d 6, distance between setae c 1 – c 1 42, c 1 – c 2 27, d – d 39; cupuli ia anteriad setae d. Plate EF slightly eroded medially; with setae f 4, distance f – f 25; cupuli im anteriad setae f. Genital capsule length 25, width 31, setae h minute. Venter: coxal setae 1 a 2, 2 a 3, 3 a 3, 3 b 3. Legs. Setal counts legs I – III, femur-tarsus: 3 - 2 - 6 (+ φ) - 8 (+ ω), 0 - 1 - 4 - 6, 0 - 1 - 4 - 6. Leg I. Femur I, d minute, l′ minute, v ″ 3; genu I, l′ minute, l ″ minute; tibia I, d 38, l’ 4, l’ ’ 4, v ’ 3, v ’’ not measurable, k 3, φ 8; tarsus I, tc’ 13, tc’ ’ 10, pl’ 5, pl’ ’ 15, pv ′ minute, pv’ ’ minute, ω 3, s 4, u ′′ minute. Leg II. Genu II, l′ 2; tibia II, d 19, l’ 4, v’ 11, v’ ’ 16; tarsus II, tc′ minute, tc ″ 2, pl ″ 31, pv’ ’ minute, u’ 4, u ″ minute. Leg III. Genu III, l′ minute; tibia III, d 19, l’ 5, v’ 9, v’ ’ 13; tarsus III, tc′ minute, tc ″ 3, pl ″ 25, pv’ ’ minute, u’ 5, u ″ minute. LARVAL FEMALE (n = 1). Gnathosoma. Length 40, width 35, apex indented, terminating in small but distinct lobes. Cheliceral stylets 35, setae ch 28, su 7. Idiosoma. Length 195, width 140. Prodorsal plate with setae v 1 5, v 2 alveolar remnant, sc 1 6, sc 2 95. Distance between setae v 1 – v 1 28, sc 1 – sc 1 58, v 1 – sc 1 22, v 2 – v 2 33, sc 2 – sc 2 71, sc 1 – sc 2 22. Plate C setae c 1 11, c 2 9, distance between setae c 1 – c 1 46. Plate D setae d 10, d – d 51, cupuli ia anterolaterad setae d. Plate EF setae f 7, f – f 38, cupuli im anterolaterad f. Plate H narrow, large, 20 wide, 33 long, bearing setae h 1, broken, h 2 7. Venter: coxisterna 1 - 2 moderately well defined, with some folds, coxisterna 3 membranous; coxal setae slender, 1 a 4, 2 a 4, 3 a 4, 3 b 5. Alveoli of 1 b, 2 b present. Legs. Setal counts legs I – III, femur-tarsus: 3 - 2 - 6 (+ φ) - 8 (+ ω), 0 - 1 - 4 - 6, 0 - 1 - 4 - 6. Leg I. Femur I, d minute, l′ minute, v ″ 5; genu I, l′ minute, l ″ minute; tibia I, d 55, l’ 8, l’ ’ 8, v’ 3, v’ ’ 13, k 5, φ 10; tarsus I, tc’ 13, tc’ ’ 14, pl’ 6, pl’ ’ 20, pv′ minute, pv’ ’ 2, ω 3, s 5, u′ ′ minute. Leg II. Genu II, l′ minute; tibia II, d 27, l’ 6, v’ 12, v’ ’ 17; tarsus II, tc′ 2, tc ″ 3, pl ″ 35, pv’ ’ 2, u’ 5, u′ ′ minute. Leg III. Genu III, l′ 2; tibia III, d 34, l’ 7, v’ 13, v’ ’ 15; tarsus III, tc’ minute, tc ″ 3, pl ″ 34, pv’ ’ minute, u’ 6, u′ ′ minute.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFC1097887AEFF5AFD30A067.taxon	diagnosis	Differential diagnosis. Eutarsopolipus osculum sp. nov. is most similar to E. basiatus sp. nov. in having short setae sc 1 (<10) and a shorter distance between setae d-d (95 – 115) and f-f (67 – 92). In addition to the features noted in the Differential diagnosis of E. basiatus, E. osculum has a shorter seta c 1 (15 – 17 versus 6 – 9).	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFC1097887AEFF5AFD30A067.taxon	etymology	Etymology. The name osculum (little mouth or little kiss) alludes to both the lip-like protrusion of the gnathosoma but also the smaller size of this species.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFC3097587AEFB07FE1AA7F9.taxon	description	(Figs 39 – 40)	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFC3097587AEFB07FE1AA7F9.taxon	diagnosis	Diagnosis. Adult female: sc 1 11 – 19, c 1 11 – 19, c 2 14 – 20, d 12 – 19, f 8 – 12, v 2 - v 2 48 – 58, sc 1 - sc 1 76 – 88, d-d 120 – 160, f-f 95 – 120; genua II-III setae l′ 2 – 4, tibia I setae l′ 8 – 10, v′ 4 – 5, tarsi II-III setae tc′ ′ 4 – 5. Adult male: leg I with one claw; c 1 4 – 5, c 2 5 – 6, v 2 - v 2 34 – 41; tarsi II-III setae tc ″ 3 – 4. Larval female: c 1 6 – 9, c 2 9 – 12, d 7 – 10, f 7, sc 2 - sc 2 78 – 100, v 1 - sc 1 22 – 24; tarsi II-III setae tc′ ′ 3.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFC3097587AEFB07FE1AA7F9.taxon	materials_examined	Material examined. Holotype female, ex. Castelnaudia eungella. Pease’s Lookout, Eungella, 21 ° 6 ʹ 53 ʺ S 148 ° 31 ʹ 6 ʺ E, 17 Nov 1992 - 15 Apr 1993, D. Cook, G. Monteith, flight intercept & pitfall trap, IN 5934, HR # T 82502 (QMS 113497). Paratypes. All ex. C. eungella. 9 females, 4 males, 11 larvae, as follows: 1 female, 4 larvae same data as holotype (QMS 113498, 113500 – 503); 2 larvae, same data except another beetle, HR # T 82503 (QMS 113504, 113578); 2 females, 1 male, 2 larvae, same data except another beetle, HR # T 82506 (QMS 113577, 11379 – 80, ANIC 52 - 003973, ZMH-A 0015199); 2 females, 1 male, 1 larva, Mt Dalrymple, 21 ° 1 ʹ 37 ʺ S 148 ° 38 ʹ 2 ʺ E, 21 Dec 1992 – 10 Jan 1993, ANZSES, HR # T 82501 (QMS 113581 – 83, ZMH-A 0015200); 1 female, 2 larvae, Eungella National Park, Mt Dalrymple track, 1200 m site, 21.026 ° S 148.627 ° E, 23 Nov 2013, C. Burwell, rainforest, sampcode 25933, beetle CB 6 (QMS 113584 – 85, ANIC 52 - 003974); 1 female, 1 male, Mt William, 21 ° 1 ʹ 5 ʺ S 148 ° 35 ʹ 57 ʺ E, 21 Dec 1992 – 10 Jan 1993, ANZSES, HR # T 82489 (QMS 113586 – 87); 2 females, 1 male, Upper Cattle Creek, Eungella, 21 ° 1 ʹ 41 ʺ S 148 ° 36 ʹ 11 ʺ E, 17 Nov 1992, G. Monteith, G. Thompson, H. Janetzki, HR # T 82513 (QMS 113588 – 90). All in QM except 1 female and 1 larva in each of ANIC and ZMH. Other material examined. 1 pharate female ex HR # T 82503 (QMS 113505).	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFC3097587AEFB07FE1AA7F9.taxon	description	Description. FEMALE (Figs 39 – 40; n = 10). Gnathosoma. Length 43 (43 – 48), width 41 (40 – 45), apex indented, terminating in small but distinct lobes. Cheliceral stylets 44 (38 – 40), setae ch 31 (24 – 39), su 14 (13 – 16). Idiosoma (Fig. 39). Length 360 (305 – 385), width 405 (250 – 350). Prodorsal plate not eroded anteriorly; setae slender, v 1 9 (6 – 9), v 2 alveolar remnant, sc 1 16 (11 – 19; one specimen asymmetrical 10 / 15), sc 2 80 (80 – 110). Distance between setae v 1 – v 1 45 (37 – 45), sc 1 – sc 1 82 (73 – 88), v 1 – sc 1 26 (often asymmetrical, 23 – 32), v 2 – v 2 58 (48 – 58), sc 2 – sc 2 130 (125 – 135), sc 1 – sc 2 38 (often asymmetrical, 31 – 46). Opisthosomal setae slender. Plate C setae c 1 19 (9 – 16), c 2 19 (14 – 20), distance between setae c 1 – c 1 85 (80 – 115). Plate D setae d 19 (12 – 19), d – d 140 (120 – 160), cupuli ia anterolaterad d. Plate EF setae f 10 (8 – 12), f – f 115 (95 – 120), cupuli im anteriad to anterolaterad f. Plate H divided, seta h minute. Venter: coxisterna 1 - 2 moderately well defined, with few folds, coxisterna 3 membranous; coxal setae slender, 1 a 4 (3 – 4), 2 a 5 (4 – 5), 3 a 6 (5 – 6), 3 b 6 (6 – 7). Alveoli of 1 b, 2 b present. Legs (Fig. 40). Setal counts legs I – III, femur-tarsus: 3 - 2 - 6 (+ φ) - 8 (+ ω), 0 - 1 - 4 - 6, 0 - 1 - 4 - 6. Paratype measurement given only when holotype measurement different from E. basiatus. Leg I. Femur I, d minute, l’ 3, v ″ 8 (7 – 10); genu I, l′ 3, l ″ 3; tibia I, d 60, l’ 10 (8 – 10), l’ ’ 9 (8 – 9), v’ 5 (4 – 5), v’ ’ 11 (10 – 17), k 5, φ 9; tarsus I, tc’ 15, tc’ ’ 13 (13 – 15), pl’ 9, pl’ ’ 16 (15 – 18), pv′ 2 (2), pv’ ’ 2, ω 4, s 6 (5), u′ ′ m. Leg II. Genu II, l′ 4; tibia II, d 48 (48 – 55), l’ 7, v’ 17 (14 – 15), v’ ’ 17 (20 – 27); tarsus II, tc′ 2, tc ″ 4 (5), pl ″ 59 (50), pv’ ’ 2, u’ 6, u′ ′ minute. Leg III. Genu III, l′ 3; tibia III, d 55 (40 – 49), l’ 7, v’ 16, v’ ’ 17 (22 – 29); tarsus III, tc’ 2, tc ″ 5 (4 – 5), pl ″ 39 (37 – 40), pv’ ’ 2, u’ 7 (6 – 7), u′ ′ minute. LARVIFORM ADULT MALE (n = 4). Gnathosoma. Length 31 – 39, width 28 – 36, apex indented, terminating in small but distinct lobes. Cheliceral stylets 20 – 22, setae ch 6 – 7, su 6 – 7. Idiosoma. Length 150 – 180, width 125 – 150. Prodorsal plate with setae v 1 2 – 3, v 2 alveolar remnant, sc 1 3 – 5, sc 2 60 – 70. Distance between setae v 1 – v 1 30 – 36, sc 1 – sc 1 43 – 50, v 1 – sc 1 15 – 16, v 2 – v 2 34 – 41, sc 2 – sc 2 63 – 77, sc 1 – sc 2 19 – 25. Plate CD setae c 1 4 – 5, c 2 5 – 6, d 5 – 6, distance between setae c 1 – c 1 46 – 54, c 1 – c 2 32 – 38, d – d 40 – 46; cupuli ia anterolaterad setae d. Plate EF eroded medially; with setae f 4 – 5, distance f – f 25 – 29; cupuli im anterolaterad setae f. Genital capsule length 27 – 28, width 38 – 40, setae h minute. Venter: coxal setae 1 a 2 – 3, 2 a 3 – 4, 3 a 2 – 3, 3 b 4 – 5. Legs. Setal counts legs I – III, femur-tarsus: 3 - 2 - 6 (+ φ) - 8 (+ ω), 0 - 1 - 4 - 6, 0 - 1 - 4 - 6. Leg I. Femur I, d minute, l′ minute, v ″ 4 – 6; genu I, l′ minute, l ″ minute; tibia I, d 42 – 52, l’ 2 – 3, l’ ’ 3 – 4, v ’ 3, v ’’ 10, k 3, φ 8; tarsus I, tc’ 12 – 14, tc’ ’ 12 – 14, pl’ 9, pl’ ’ 10 – 14, pv ′ m – 2, pv’ ’ m – 2, ω 3 – 4, s 4 – 5, u ′′ minute. Leg II. Genu II, l′ minute; tibia II, d 26 – 30, l’ 5, v’ 11, v’ ’ 18 – 21; tarsus II, tc′ 2, tc ″ 3 – 4, pl ″ 25 – 28, pv’ ’ minute, u’ 4, u ″ minute. Leg III. Genu III, l′ minute; tibia III, d 26 – 28, l’ 4 – 6, v’ 8 – 12, v’ ’ 16; tarsus III, tc′ 2, tc ″ 4, pl ″ 29 – 31, pv’ ’ 2, u’ 4, u ″ minute. LARVAL FEMALE (n = 11). Gnathosoma. Length 38 – 41, width 33 – 36, apex indented, terminating in small but distinct lobes. Cheliceral stylets 30 – 35, setae ch 22 – 27, su 6 – 7. Idiosoma. Length 185 – 260, width 140 – 220. Prodorsal plate with setae v 1 4 – 5, v 2 alveolar remnant, sc 1 5 – 6, sc 2 105 – 110. Distance between setae v 1 – v 1 32 – 35, sc 1 – sc 1 59 – 65, v 1 – sc 1 22 – 24, v 2 – v 2 44 – 46, sc 2 – sc 2 78 – 100, sc 1 – sc 2 22 – 34. Plate C setae c 1 6 – 9, c 2 9 – 10, distance between setae c 1 – c 1 50 – 71. Plate D setae d 7 – 10, d – d 54 – 72, cupuli ia anteriad to anteromesad d. Plate EF setae f 7 – 10, f – f 47 – 59, cupuli im anteriad to anterolaterad f. Plate H narrow, large, 18 – 21 wide, 31 – 36 long, bearing setae h 1 ca. 160, h 2 6 – 8. Venter: coxisterna 1 - 2 moderately well defined, with some folds, coxisterna 3 membranous; coxal setae slender, 1 a 3, 2 a 4, 3 a 2 – 4, 3 b 5 (one specimen lacking setae 3 a and 3 b expressed as alveoli). Alveoli of 1 b, 2 b present. Legs. Setal counts legs I – III, femur-tarsus: 3 - 2 - 6 (+ φ) - 8 (+ ω), 0 - 1 - 4 - 6, 0 - 1 - 4 - 6. Ranges given only when first measurement different from E. basiatus. Leg I. Femur I, d minute, l′ minute, v ″ 4 – 6; genu I, l′ m – 2, l ″ 2; tibia I, d 55 – 65, l’ 7 – 11, l’ ’ 8 – 11, v’ 4 – 5, v’ ’ 8 – 10, k 4, φ 10; tarsus I, tc’ 15, tc’ ’ 15, pl’ 6 – 7, pl’ ’ 14 – 17, pv′ minute, pv’ ’ minute, ω 4, s 5, u′ ′ minute. Leg II. Genu II, l′ m – 2; tibia II, d 40, l’ 6, v’ 10 – 14, v’ ’ 21; tarsus II, tc′ minute, tc ″ 3, pl ″ 28, pv’ ’ minute, u’ 5, u′ ′ minute. Leg III. Genu III, l′ 3; tibia III, d 36, l’ 7, v’ 10 – 15, v’ ’ 21; tarsus III, tc’ m – 2, tc ″ 3, pl ″ 38 – 43, pv’ ’ minute, u’ 5, u′ ′ minute.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFC3097587AEFB07FE1AA7F9.taxon	diagnosis	Differential diagnosis. Eutarsopolipus savatus sp. nov. is most similar to E. labiatus and the distinguishing features are discussed under the Differential Diagnosis for the latter species.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFC3097587AEFB07FE1AA7F9.taxon	etymology	Etymology. The name savatus (a tender kiss) alludes to both the lip-like protrusion of the gnathosoma but also alludes to the larger size of this species.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFC3097587AEFB07FE1AA7F9.taxon	discussion	Remarks. One larva similar to E. echinatus Seeman, 2019 a was found on HR # T 82502; one male closest to the nahmani subgroup group was found on HR # T 82502 (QMS 113499) (Tables 1, 2). The leytei species group	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFCE097187AEFBD9FA39A3AD.taxon	description	(Figs 41 – 43)	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFCE097187AEFBD9FA39A3AD.taxon	diagnosis	Diagnosis. Adult female: leg I with two claws; v 1 8 – 10, sc 2 15 – 20; genu I with two setae (l ″ present); tarsus II setae tc ″ 5 – 6, u′ 6 – 7. Adult male: setae sc 2 2 – 3. Larval female: sc 2 51 – 55.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFCE097187AEFBD9FA39A3AD.taxon	materials_examined	Material examined. Holotype female, ex. Notonomus transitus. Blue Mt, 0.6 km SE, 21 ° 36 ʹ 19 ʺ S 148 ° 58 ʹ 21 ʺ E, 3 – 4 Oct 1999, G. Monteith, D. Cook, C. Burwell, S. Evans, rainforest, IN 7801, HR # T 89396 (QMS 113608). Paratypes. All ex. Notonomus transitus. 5 females, 5 males, 14 larvae, as follows: 2 females, 2 males, 10 larvae, same data as holotype (QMS 113609 – 18, ANIC 52 - 003975 – 77, ZMH-A 0015201); 1 female, 1 male, 1 larva, 50 mi. W of Mackay, 21 ° 08 ʹ 12 ʺ S 148 ° 29 ʹ 30 ʺ E, 6 Jan 1973, J. Hammond, IN 7864, HR # T 81753 (QMS 113621 – 23); 2 females, 2 males, 1 larva, Broken River, Eungella, 21 ° 10 ʹ 5 ʺ S 148 ° 30 ʹ 30 ʺ E, 4 Dec 1992 – 30 Apr 1993, P. Lawless, M. Shaw, R. Raven, IN 9781, HR # T 110084 (QMS 113624 – 26, ZMH-A 0015202 – 03); 2 larvae, Dalrymple Rd, Eungella NP, 21 ° 1 ʹ 44 ʺ S 148 ° 35 ʹ 7 ʺ E, 9 Nov 1991 – 29 Jul 1992, P. Lawless, R. Raven, M. Shaw, rainforest, IN 7768, HR # T 81768 (QMS 113627 – 28). All in QM except 1 female, 1 male, 1 larva in ANIC and ZMH. Other material examined. 2 larvae, same data as holotype, in poor condition (QMS 113619 – 20).	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFCE097187AEFBD9FA39A3AD.taxon	description	Description. FEMALE (Fig. 41; n = 6). Gnathosoma. Length 45 (44 – 46), width 39 (34 – 37). Palp with alveolar remnant of femoral seta. Cheliceral stylets 38 (35 – 36), setae ch 22 (14 – 16), su 11 (8 – 12). Idiosoma (Fig. 41 A). Length ca. 500 (270 – 430), width ca. 300 (215 – 310), bigger specimens swollen, distorted. Stigmata apparent, large, in soft cuticle anteriad setae v 1 to posterolaterad gnathosoma (varies according to mount), respiratory system weakly developed, tracheae not extending past small atrium, tracheae and atrium sometimes not visible. Prodorsal plate with setae v 1 10 (8 – 9), v 2 alveolar remnant, sc 1 9 (7 – 8), sc 2 20 (15 – 17). Distance between setae v 1 – v 1 44 (39 – 51), sc 1 – sc 1 69 (64 – 72), v 1 – sc 1 22 (23 – 25), v 2 – v 2 52 (53 – 57), sc 2 – sc 2 84 (83 – 110), sc 1 – sc 2 34 (24 – 32). Plate C appearing divided, each plate bearing its slender setae at their margins, c 1 6 (5 – 7), c 2 6 (4 – 5). Plate D setae d 5 (4 – 5), d – d 92 (85 – 95), cupuli ia anteriad d. Plate EF setae f 4 (4 – 5), f – f broken (49 – 60), cupuli im anterolaterad f. Plate H absent. Venter: coxisterna 1 - 3 well defined; coxal setae slender, 1 a 5 (5 – 6), 2 a 5 (4 – 5), 3 a 6 (5 – 6), 3 b 5 (4 – 5). Alveoli of 1 b, 2 b present. Legs (Figs 41 B-C). Setal counts legs I – III, femur-tarsus: 3 - 2 - 6 (+ φ) - 9 (+ ω), 0 - 1 - 4 - 7 (+ ω), 0 - 1 - 4 - 7. Leg I. Femur I, d 1 (1 – 3), l’ 5 (4 – 5), v ″ 8 (9 – 10); genu I, l’ 3 (2 – 3), l ″ 2 (2 – 3); tibia I, d 16 (14 – 19), l’ 3 (2 – 3), l’ ’ 5 (5 – 6), v’ 3 (3 – 5), v’ ’ 9 (8 – 14), k 3 (3 – 4), φ 6 (6 – 7); tarsus I, tc’ 9 (9 – 10), tc’ ’ 9 (9 – 10), pl’ 8 (8 – 10), pl’ ’ 10 (8 – 12), pv′ 2 (2 – 3), pv’ ’ 2 (2 – 3), ω 3 (3 – 4), s 6 (5 – 6), u′ ′ minute (m – 2), p′ m (m – 2). Leg II. Genu II, l’ 3 (2 – 3); tibia II, d 9 (7 – 9), l’ 5 (3 – 5), v’ 8 (8 – 11), v’ ’ 14 (13 – 16); tarsus II, tc′ 8 (6 – 8), tc ″ 5 (5 – 6), pl ″ 10 (7 – 9), pv′ 3 (3 – 4), pv’ ’ 4 (4 – 5), ω 3 (2 – 3), u’ 7 (6 – 7), u ″ 2 (2). Leg III. Genu III, l’ 3 (2 – 3); tibia III, d 4 (5 – 6), l’ 4 (3 – 4), v’ 8 (6 – 9), v’ ’ 11 (13 – 15); tarsus III, tc’ 6 (6 – 7), tc ″ 5 (5 – 6), pl ″ 8 (9 – 10), pv′ 3 (2 – 3), pv’ ’ 2 (2 – 3), u’ 6 (6 – 7), u ″ 2 (2). Claws on legs I-III well developed and paired. LARVIFORM ADULT MALE (Fig. 42; n = 5). Gnathosoma. Length 28 – 31, width 28 – 29. Palp with alveolar remnant of femoral seta. Cheliceral stylets 17 – 18, setae ch 1 – 2, su 4 – 6. Idiosoma. Length 155 – 165, width 110 – 120. Prodorsal plate with setae v 1 m – 2, v 2 alveolar remnant, sc 1 minute, sc 2 2 – 3, thickened. Distance between setae v 1 – v 1 19 – 22, sc 1 – sc 1 45 – 48, v 1 – sc 1 19 – 21, v 2 – v 2 34 – 36, sc 2 – sc 2 54 – 56, sc 1 – sc 2 22 – 23. Plate CD incised posterolaterally, setae c 1, c 2, d minute, distance between setae c 1 – c 1 44 – 46, c 1 – c 2 26 – 35, d – d 31 – 32; cupuli ia laterad to anterolaterad setae d. Plate EF with setae f minute, distance f – f 21 – 23; cupuli im anterolaterad setae f. Genital capsule length 22 – 25, width 34 – 36, setae h minute. Venter: coxal setae slender, 1 a 2 – 3, 2 a 2 – 3, 3 a 2, 3 b 2 – 3. Legs. Setal counts legs I – III, femur-tarsus: 2 - 0 - 6 (+ φ) - 8 (+ ω), 0 - 0 - 4 - 7 (+ ω), 0 - 0 - 4 - 7. Compared to female: Femur I seta v ″ absent, genu I l′ absent, genua I-III l ″ absent; tarsus I p′ absent. Leg I. Femur I, d minute, l’ 2; genu I, l′ minute; tibia I, d 4 – 5, l’ minute, l’ ’ m – 2, v ’ 2, v ’’ 6 – 7, k minute, φ 4 – 5; tarsus I, tc’ 7 – 8, tc’ ’ 8 – 9, pl’ 4 – 6, pl’ ’ 6 – 9, pv ′ 2, pv’ ’ 2 – 3, ω 3, s 4 – 5, u ′′ 1 – 2. Leg II. Tibia II, d 3 – 5, l’ 2 – 3, v’ 6 – 7, v’ ’ 7 – 8; tarsus II, tc′ 3, tc ″ 4, pl ″ 5 – 6, pv ′ 2, pv’ ’ 2 – 3, ω 2 – 3, u’ 5 – 6, u ″ minute. Leg III. Tibia III, d 2, l’ 2, v’ 6 – 8, vʺ 7 – 9; tarsus III, tc′ 2 – 3, tc ″ 4, pl ″ 4 – 5, pv ′ 2, pv’ ’ 2, u ’ 5, u ″ minute. Claws on leg II-III present. Setal form similar to that in female, but with tibia I setae l′, l ″, v′ as tiny spines. LARVAL FEMALE (Fig. 43; n = 14). Gnathosoma. Length 30 – 34, width 27 – 30. Palp with alveolar remnant of femoral seta. Cheliceral stylets 25 – 27, setae ch 19 – 23, su 8 – 10. Idiosoma (Fig. 43). Length 165 – 225, width 105 – 185; larger specimens swollen laterally between prodorsal shield and plate CD. Prodorsal plate with setae v 1 12 – 14, v 2 alveolar remnant, sc 1 5 – 6, sc 2 51 – 55. Distance between setae v 1 – v 1 20 – 23, sc 1 – sc 1 48 – 51, v 1 – sc 1 25 – 28, v 2 – v 2 29 – 35, sc 2 – sc 2 49 – 54, sc 1 – sc 2 27 – 33. Plates C and D fused medially; line of fusion visible. Plate CD setae c 1 5 – 6, c 2 3 – 4, d 4 – 5, cupuli ia anteriad to anteromesad d, distance between setae c 1 – c 1 26 – 27, d – d 24 – 28. Plate EF setae f 4 – 5, f – f 23 – 26, cupuli im anteriad f. Plate H poorly expressed, caudal cone 22 – 30 wide, 8 – 13 long, bearing setae h 1 ca. 65 – 70, h 2 3 – 4. Venter: coxisterna 1 - 3 well-defined; coxal setae slender, 1 a 2 – 3, 2 a 3 – 4, 3 a 4 – 5, 3 b 3 – 4. Alveoli of 1 b, 2 b present. Legs. Setal counts legs I – III, femur-tarsus: 3 - 2 - 6 (+ φ) - 8 (+ ω), 0 - 1 - 4 - 6 (+ ω), 0 - 1 - 4 - 6 (setae pv′ absent on tarsi II- III). Leg I. Femur I, d minute, l’ 2 – 3, v’ ’ 5 – 7; genu I, l’ 2 – 3, l ″ m – 2; tibia I, d 17 – 21, l’ 6 – 7, l’ ’ 5 – 7, v’ 3 – 4, v’ ’ 7 – 9, k 2 – 3, φ 6 – 7; tarsus I, tc’ 8 – 9, tc’ ’ 9 – 10, pl’ 5 – 6, pl’ ’ 9 – 10, pv′ 2, pv’ ’ 2, ω 3 – 4, s 4 – 5, u′ ′ minute. Leg II. Genu II, l’ 2 – 3; tibia II, d 10 – 12, l’ 4 – 5, v’ 7 – 9, v’ ’ 11 – 13; tarsus II, tc′ 3 – 4, tc ″ 4 – 5, pl ″ 7 – 9, pv’ ’ 3 – 4, ω 3 – 4, u’ 5 – 6, u′ ′ minute. Leg III. Genu III, l’ 2; tibia III, d 10 – 11, l’ 4 – 5, v’ 7 – 8, v’ ’ 11 – 12; tarsus III, tc’ 4 – 5, tc ″ 4 – 5, pl ″ 8 – 10, u’ 5 – 6, pv’ ’ 5 – 6, u′ ′ minute.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFCE097187AEFBD9FA39A3AD.taxon	diagnosis	Differential diagnosis. Eutarsopolipus biuncatus sp. nov. and E. janus sp. nov. are the only members of the leytei species group to retain paired claws on tarsus I in females. The only other species of Eutarsopolipus with females retaining two claws on tarsus I is Eutarsopolipus diunculosus Eidelberg, 1994, a member of the pterostichi species group (lacking both a respiratory system and setae on genua II-III). In addition to the diagnostic features mentioned for E. janus, female specimens appear to have divided plate C. None of these specimens are of sufficient quality to be sure — they may have split when mounted — but the division of dorsal plates occurs elsewhere in Eutarsopolipus (e. g., Regenfuss 1968; Husband 2000).	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFCE097187AEFBD9FA39A3AD.taxon	etymology	Etymology. The specific name biuncatus (two-hooked) refers to the two claws on tarsus I of females.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFCE097187AEFBD9FA39A3AD.taxon	discussion	Remarks. All females had waste matter that obscured morphological features. This was partially removed by clearing some specimens in 5 % KOH, but no female was of high quality. The holotype was chosen on the basis of having the best quality prodorsum and legs I-II, where most diagnostic features occur. Both E. biuncatus sp. nov. and E. janus sp. nov. are unusual in retaining setae pv′ on tarsi II-III in the male, a seta usually absent in Tarsonemoidea (Lindquist 1986; Seeman 2019 a). Another specimen of N. transitus from the type locality (HR # T 95636) had an unusual male of another species of Eutarsopolipus (lacking all genual setae, large, blunt-tipped Y-shaped claws). Males are difficult to place in species groups as they always lack respiratory systems and sometimes lack genual setae that are present in the female.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFCB090A87AEFF5AFE85A5C5.taxon	description	(Figs 44 – 48)	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFCB090A87AEFF5AFE85A5C5.taxon	diagnosis	Diagnosis. Adult female: leg I with two claws; v 1 19 – 24, sc 2 36 – 40; genu I with two setae (l ″ present); tarsus II setae tc ″ 8 – 9, u′ 8 – 10. Adult male: setae sc 2 27 – 34. Larval female: sc 2 70 – 80.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFCB090A87AEFF5AFE85A5C5.taxon	materials_examined	Material examined. Holotype female, ex. Notonomus flos. Mt Lewis Rd, 29 km from Highway, Cow Bay, 16 ° 13 ʹ 51 ʺ S 145 ° 27 ʹ 43 ʺ E, 18 Nov 1997 – 9 Feb 1998, G. Monteith & D. Cook, pitfall trap, rainforest, IN 1702, HR # T 65790 (QMS 113527). Paratypes. All ex Notonomus flos. 7 females, 7 males, 9 larvae, as follows: 7 females, 7 males, 7 larvae, same data as holotype (QMS 113528 – 42, ANIC 52 - 003978 – 80, ZMH-A 0015204 – 05); 2 larvae, same data except different beetle, HR # T 65789 (QMS 113543, ZMH-A 0015206). All in QM except 1 female, 1 male, 1 larva in each of ANIC and ZMH. Other material examined. Ex N. aurifer: 7 females, 5 males, 7 larvae, as follows: 6 females, 2 males, 4 larvae, Lamb’s Head, 20 km SW Cairns, 17 ° 1 ʹ 23 ʺ S 145 ° 38 ʹ 33 ʺ E, 10 – 12 Dec 1989, G. Monteith, G. Thompson, H. Janetzki, HR # T 20079 (QMS 113544 – 54, 1 female ANIC); 2 males, 1 larva, same data except different beetle, HR # T 20078 (QMS 113555 – 57); 1 female, 1 male, 2 larvae, same data except different beetle, HR # T 20081 (QMS 113558 – 61). Ex N. spurgeoni: 5 females, 3 males, 4 larvae, as follows: 1 female, Mossman Bluff Track site 8, 16 ° 27 ʹ 36 ʺ S 145 ° 17 ʹ 12 ʺ E, 17 – 19 Dec 1988, G. Monteith, G. Thompson, ANZSES, HR # T 20799 (QMS 113591); 1 female, Mossman Bluff Track general, 16 ° 28 ʹ S 145 ° 18 ʹ E, 20 Dec 1989 – 15 Jan 1990, G. Monteith, G. Thompson, ANZSES, HR # T 20863 (QMS 113592); 1 male, 4 larvae, same data except different beetle, HR # T 20849 (QMS 113598 – 602); 3 females, 2 males, Mt Spurgeon, 7 km N, camp 2, 16 ° 22 ʹ 31 ʺ S 145 ° 12 ʹ 49 ʺ E, 17 – 19 Oct 1991, G. Monteith, H. Janetzki, D. Cook, L. Roberts, HR # T 20957 (QMS 113593 - 97). Ex N. dimorphicus: 1 female, 1 male, 3 larvae, as follows: 1 female, 1 larva, Mt Lewis, barracks, 16 ° 35 ʹ 39 ʺ S 145 ° 16 ʹ 28 ʺ E, no date, G. Wood, ex HR # T 20722 (QMS 113603); 1 male, 1 larva, Mt Lewis Rd, 22 km from Highway, ANZSES site 3, 16 ° 32 ʹ 48 ʺ S 145 ° 16 ʹ 53 ʺ E, 29 Nov 1997 – 9 Feb 1998, G. Monteith, D. Cook, rainforest, IN 1701, HR # T 65741 (QMS 113605 – 06); 1 larva, Mt Lewis, 2.5 km N, via Julatten, 16 ° 33 ʹ S 145 ° 15 ʹ E, 3 Nov 1983, D. Yeates, G. Thompson, HR # T 20721 (QMS 113607). Note: One male is fully developed within a broken egg and is included as a paratype. Description (excluding specimens from N. aurifer). FEMALE (Figs 44 – 46; n = 21). Gnathosoma. Length 55 (52 – 57), width 51 (45 – 51). Palp with alveolar remnant of femoral seta. Cheliceral stylets 45 (45 – 49), setae ch 25 (25 – 35), su 18 (15 – 20). Idiosoma (Figs 44 – 45). Length ca. 340 (290 – 600), width ca. 240 (230 – 450), bigger specimens swollen, distorted. Stigmata apparent, large, in soft cuticle anteriad setae v 1 to posterolaterad gnathosoma (varies according to mount), respiratory system not visible in most mounts but sometimes narrow trachea and small atrium apparent. Prodorsal plate with setae v 1 20 (19 – 24), v 2 alveolar remnant, sc 1 10 (9 – 11), sc 2 38 (36 – 40). Distance between setae v 1 – v 1 56 (53 – 60), sc 1 – sc 1 88 (87 – 96), v 1 – sc 1 31 (34 – 37), v 2 – v 2 69 (68 – 75), sc 2 – sc 2 95 (91 – 95), sc 1 – sc 2 37 (32 – 36). Plate C entire, setae c 1 8 (8 – 9), c 2 8 (8 – 10). Plate D setae d 9 (8 – 10), d – d 105 (110 – 125), cupuli ia anterolaterad d. Plate EF setae f 9 (7 – 8), f – f 71 (80 – 87), cupuli im anterolaterad to laterad f. Plate H present as small remnant. Venter: coxisterna 1 - 2 poorly defined, slightly broader than apodemes; coxisterna 3 well-defined; coxal setae slender, 1 a 6 (6 – 7), 2 a 6 (6 – 7), 3 a 6 (7 – 9), 3 b 6 (6 – 7). Alveoli of 1 b, 2 b present. Legs (Fig. 46). Setal counts legs I – III, femur-tarsus: 3 - 2 - 6 (+ φ) - 9 (+ ω), 0 - 1 - 4 - 7 (+ ω), 0 - 1 - 4 - 7. Leg I. Femur I, d 2 (2), l’ 13 (13 – 16), v ″ 14 (12 – 15); genu I, l’ 5 (5 – 6), l ″ 3 (3); tibia I, d 25 (27 – 30), l’ 10 (7 – 9), l’ ’ 6 (6 – 8), v’ 7 (7 – 8), v’ ’ 14 (11 – 12), k 4 (4 – 5), φ 6 (7 – 8); tarsus I, tc’ 13 (11 – 12), tc’ ’ 13 (13 – 15), pl’ 15 (14 – 17), pl’ ’ 18 (15 – 20), pv′ 3 (3 – 4), pv’ ’ 3 (2 – 3), ω 4 (4 – 5), s 6 (6 – 7), u′ ′ minute (m – 2), p′ 2 (m – 2). Leg II. Genu II, l’ 8 (5 – 6); tibia II, d 16 (15 – 20), l’ 9 (8 – 9), v’ 15 (14 – 16), v’ ’ 21 (20 – 23); tarsus II, tc′ 9 (9 – 11), tc ″ 8 (8 – 9), pl ″ 18 (16 – 18), pv′ 2 (2 – 3), pv’ ’ 4 (3 – 4), ω 4 (4 – 5), u’ 9 (9 – 10), u ″ m (m), difficult to discern next to large u′. Leg III. Genu III, l’ 5 (4 – 5); tibia III, d 14 (15 – 20), l’ 9 (7 – 9), v’ 12 (12 – 15), v’ ’ 17 (20 – 21); tarsus III, tc’ 8 (8 – 10), tc ″ 8 (8 – 9), pl ″ 20 (18 – 20), pv′ 4 (3 – 5), pv’ ’ 3 (3 – 5), u’ 8 (8 – 9), u ″ m (m), difficult to discern next to large u′. Claws on legs I-III well developed and paired. Tarsi I-III setae pvʹ, pvʺ slender. Large blunt spur-like setae: tarsus I s; tarsi II – III u’, tcʺ. Setae u′ pointed, not bifid. LARVIFORM ADULT MALE (Fig. 47; n = 16). Gnathosoma. Length 28 – 37, width 29 – 33. Cheliceral stylets 25 – 28, setae ch 3 – 4, su 8 – 9. Idiosoma (Fig. 47). Length 175 – 190, width 135 – 155. Prodorsal plate with setae v 1 4 – 9, v 2 alveolar remnant, sc 1 m – 2, sc 2 27 – 34. Distance between setae v 1 – v 1 24 – 30, sc 1 – sc 1 61 – 66, v 1 – sc 1 24 – 26, v 2 – v 2 42 – 49, sc 2 – sc 2 64 – 72, sc 1 – sc 2 24 – 28. Plate CD eroded posterolaterally, setae c 1 2 – 3, c 2 2, d m – 2, distance between setae c 1 – c 1 56 – 65, c 1 – c 2 32 – 35, d – d 42 – 47; cupuli ia laterad to anterolaterad setae d. Plate EF with setae f m – 2, distance f – f 22 – 25; cupuli im laterad to anterolaterad setae f. Genital capsule length 25 – 27, width 35 – 38, setae h minute. Venter: coxal setae slender, 1 a 4 – 5, 2 a 4 – 5, 3 a 5 – 6, 3 b 5 – 6. Legs. Setal counts legs I – III, femur-tarsus: 2 - 0 - 6 (+ φ) - 8 (+ ω), 0 - 0 - 4 - 7 (+ ω), 0 - 0 - 4 - 7. Compared to female: Femur I seta v ″ absent, genua I-III l′ and l ″ absent. Leg I. Femur I, d 2, l’ 2 – 4; tibia I, d 19 – 25, l’ 2, l’ ’ 2 – 3, v ’ 2 – 3, v ’’ 9 – 10, k 2 – 3, φ 6 – 7; tarsus I, tc’ 8 – 9, tc’ ’ 10 – 11, pl’ 8 – 11, pl’ ’ 10 – 12, pv ′ 2, pv’ ’ 2, ω 3 – 4, s 5, u ′′ minute. Leg II. Tibia II, d 6 – 11, l’ 3 – 4, v’ 9 – 12, v’ ’ 11 – 14; tarsus II, tc′ 6 – 8, tc ″ 5 – 6, pl ″ 11 – 12, pv ′ 2, pv’ ’ 3 – 4, ω 3 – 4, u’ 6, u ″ minute, difficult to discern. Leg III. Tibia III, d 2, l’ 3 – 5, v’ 8 – 11, vʺ 10 – 14; tarsus III, tc′ 3 – 4, tc ″ 6 – 7, pl ″ 9 – 11, pv ′ 2 – 3, pv’ ’ 3 – 5, u ’ 6 – 7, u ″ minute, difficult to discern. Claws on leg I single, claws on leg II-III paired. Setal form similar to that in female, but with tibia I setae l′, l ″ tiny spines and v′ slightly thickened. LARVAL FEMALE (Fig. 48). Gnathosoma. Length 38 – 46, width 31 – 34. Palp without alveolar remnant of femoral seta. Cheliceral stylets 36 – 42, setae ch 26 – 33, su 12 – 15. Idiosoma (Fig. 48). Length 180 – 225, width 135 – 240; larger specimens swollen evenly around body. Prodorsal plate with setae v 1 15 – 18, v 2 alveolar remnant, sc 1 8 – 10, sc 2 70 – 80. Distance between setae v 1 – v 1 29 – 31, sc 1 – sc 1 69 – 72, v 1 – sc 1 31 – 33, v 2 – v 2 50 – 54, sc 2 – sc 2 72 – 78, sc 1 – sc 2 33 – 37. Plates C and D fused medially; line of fusion visible. Plate C setae c 1 7 – 10, c 2 7 – 10, distance between setae c 1 – c 1 31 – 35. Plate D setae d 7 – 9, d – d 30 – 35, cupuli ia anterolaterad d. Plate EF setae f 6 – 8, f – f 29 – 33, cupuli im anteriad to anterolaterad f. Plate H small, 18 – 22 wide, 13 – 18 long, bearing setae h 1 ca. 100, h 2 4 – 6. Venter: coxisterna 1 - 3 well-defined; coxal setae slender, 1 a 4 – 5, 2 a 4 – 5, 3 a 7 – 8, 3 b 5 – 6. Alveoli of 1 b, 2 b present. Legs. Setal counts legs I – III, femur-tarsus: 3 - 2 - 6 (+ φ) - 8 (+ ω), 0 - 1 - 4 - 6 (+ ω), 0 - 1 - 4 - 6 (setae pv′ absent on tarsi II- III). Leg I. Femur I, d m – 2, l’ 2 – 3, v’ ’ 9 – 13; genu I, l’ 3, l ″ 2 – 3; tibia I, d 25 – 32, l’ 7 – 8, l’ ’ 7 – 9, v’ 4 – 5, v’ ’ 8 – 12, k 3 – 4, φ 7 – 8; tarsus I, tc’ 8 – 9, tc’ ’ 10 – 11, pl’ 8 – 10, pl’ ’ 13 – 15, pv′ 2 – 3, pv’ ’ 2 – 3, ω 4 – 5, s 5 – 6, u′ ′ 2. Leg II. Genu II, l’ 4 – 5; tibia II, d 13 – 17, l’ 8 – 9, v’ 9 – 12, v’ ’ 13 – 16; tarsus II, tc′ 7 – 8, tc ″ 5 – 6, pl ″ 13 – 17, pv’ ’ 4 – 5, ω 4 – 5, u’ 6 – 7, u′ ′ minute. Leg III. Genu III, l’ 3 – 4; tibia III, d 15 – 17, l’ 5 – 6, v’ 10 – 11, v’ ’ 16 – 17; tarsus III, tc’ 7 – 9, tc ″ 5 – 7, pl ″ 16 – 20, u’ 6 – 7, pv’ ’ 5 – 6, u′ ′ minute. Setal fprm similar to that in female. Note: one larva (# 2) has an unusual asymmetrical reversal of tcʺ and plʺ.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFCB090A87AEFF5AFE85A5C5.taxon	diagnosis	Differential diagnosis. Eutarsopolipus janus sp. nov. and E. biuncatus sp. nov. both have the unusual feature of paired claws on leg I in females. Female Eutarsopolipus janus differ by having longer setae sc 2 (36 – 40 versus 15 – 20) and tarsi II-III with longer setae tc′ (8 – 11 versus 3 – 7) and tc′ ′ (8 – 9 versus 5 – 6). Males are even more distinctive, with seta sc 2 being well-developed in E. janus (27 – 34) but greatly reduced in E. biuncatus (2 – 3).	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFCB090A87AEFF5AFE85A5C5.taxon	etymology	Etymology. The specific name janus refers to the god Janus, who symbolised beginnings and transitions, alluding to the questions regarding the evolution and speciation of this possible complex of cryptic species on several host species.	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFCB090A87AEFF5AFE85A5C5.taxon	discussion	Remarks. This species was found on four host species: the type host, N. flos, as well as N. aurifer, N. dimorphicus and N. spurgeoni. All these species are found in the Wet Tropics region of north Queensland and at least N. aurifer, N. dimorphicus and N. flos are closely related (Will 2015). Thus, these species may not have speciated when their hosts speciated, or they have done so cryptically. In support of the latter hypothesis, some minor differences were observed in N. aurifer. Females of this species differed by having: slightly longer cheliceral stylets (50 – 56 versus 43 – 50), slightly longer tibia I solenidion (6 – 8 versus 8 – 10), slightly longer seta tc′ on tarsus II (10 – 13 versus 8 – 10) and tarsus III (11 – 13 versus 8 – 10), slightly longer setae tcʺ on tarsus II (9 – 10 versus 8 – 9) and tarsus III (10 – 11 versus 8 – 9) and u′ on tarsus III (9 – 11 versus 8 – 9). Several of these differences are also apparent in the larvae. Larvae from N. aurifer have longer cheliceral stylets (43 – 49 versus 36 – 42), longer tarsal seta tc′ on tarsus III (10 – 12 versus 7 – 9) and tarsal seta tc ″ on tarsus II (6 – 7 versus 5 – 6). The species N. aurifer is also geographically separated from the other three species: the former is from a large fragment of the Wet Tropics south-west of Cairns, while the others are from a larger fragment southwest to west of Mossman comprising Mt Lewis (N. flos), Mt Spurgeon (N. spurgeoni) and Lamb’s Head (N. dimorphicus). These regions are separated by a biogeographical barrier, the Black Mountain Corridor, which separates radiations of other endemic rainforest animals (e. g., Krosch et al. 2009; Edwards & Melville 2010) and plants (e. g., Burke et al. 2013). Furthermore, N. dimorphicus and N. flos are more closely related to each other than they are to N. aurifer (Will 2015).	en	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
