identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
4438245CFFF3094887AEFF55FDB3A528.text	4438245CFFF3094887AEFF55FDB3A528.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eutarsopolipus Berlese 1913	<div><p>Genus Eutarsopolipus Berlese, 1913</p> <p>Type species: Tarsopolipus lagenaeformis Berlese, 1911, by original designation.</p> <p>Diagnosis. See Seeman (2019a).</p></div> 	http://treatment.plazi.org/id/4438245CFFF3094887AEFF55FDB3A528	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Seeman, Owen D.	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFFC094687AEFBD9FA38A696.text	4438245CFFFC094687AEFBD9FA38A696.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eutarsopolipus Berlese 1913	<div><p>Key to species of Eutarsopolipus from Castelnaudia</p> <p>The key to species of the piratica subgroup was also published in Seeman (2019b); E. piraticus is included although its host is Trichosternus. The key is primarily to adult females although larvae are helpful for separation of some species.</p> <p>1. Claw on leg I absent, unguinal setae on tarsi II-III massive, hook-like. (piratica subgroup)........................... 2</p> <p>- Claw on leg I present, unguinal setae on tarsi II-III enlarged, bifid, never hook like................................. 4</p> <p>2. Tarsi II-III with seta pl″ long, 75–95. Hosts: Castelnaudia eungella and C. wilsoni........... E. verberatus Seeman, 2019b</p> <p>- Tarsi II-III with seta pl″ shorter, 35–53.................................................................... 3</p> <p>3. Setae v1 far apart (63–94); setae tc″ on tarsi II– III comparatively large and obviously thorn-like, length 4–5… Host: Castelnaudia obscuripennis.............................................................. E. uncatus Seeman, 2019b</p> <p>- Setae v1 closer together (47–58); setae tc″ on tarsi II– III comparatively smaller and merely thickened, length 2–3… Hosts: Trichosternus frater and T. mutatus................................................. E. piraticus Seeman, 2019b</p> <p>4. At least some idiosomal setae significantly thickened, sometimes bulbous. (hadros subgroup)......................... 5</p> <p>- All idiosomal setae slender.............................................................................. 7</p> <p>5. Setae sc1 sometimes thickened but not bulbous (Fig. 10); anterior margin of prodorsal shield deeply concave (Fig. 1)… Host: Castelnaudia setosiceps.................................................................. E. hadros sp. nov.</p> <p>- Setae sc1 strongly thickened to bulbous (Fig. 10); anterior margin of prodorsal shield weakly concave or straight......... 6</p> <p>6. Larvae with shorter setae c1 8–11, d 7–10, f 7–10; female generally with shorter dorsal setae: v1 7-10, c1 6–12, d 10–15, f 7–12 (Fig. 11)… Host: Castelnaudia cordata................................................... E. umbonatus sp. nov.</p> <p>- Larvae with longer setae c1 13–20, d 11–12, f 14–19; female generally with longer dorsal setae: v1 10–13, c1 10–20, d 12–21, f 11–19 (Fig. 9)… Host: Castelnaudia mixta.................................................. E. mixtus sp. nov.</p> <p>7. Anterior tip of gnathosoma rounded....................................................................... 8</p> <p>- Anterior tip of gnathosoma indented, giving a lip-like appearance. (basiatus subgroup)............................. 12</p> <p>8. Setae h small but developed. (nahmani subgroup)..........................................................… 9</p> <p>- Setae h minute (not larger than alveolus) or absent. (despoticus subgroup)....................................... 11</p> <p>9. Gnathosoma large, 67–70 x 69–72; distance d-d 200–230; genua II-III setae l′ 5–7, tibia I seta l′ 9–11 (Fig. 12)… Host: Castelnaudia septemcostata................................................................... E. nahmani sp. nov.</p> <p>- Gnathosoma smaller, 57–61 x 48–54; distance d-d 115–140; genua II-III setae l′ 2–4, tibia I seta l′ 4–7................ 10</p> <p>10. Setae h small, 2–4; tarsus III with small setae tc′ 2, pv′ 2–3 (Figs. 17, 18)… Host: Castelnaudia septemcostata. E. raveni sp. nov.</p> <p>- Setae h longer, 7–10; tarsus III with larger setae tc′ 3–4, pv′ 4 (Figs. 21, 22)… Host: Castelnaudia porphyriaca. E. teuceri sp. nov.</p> <p>11. Gnathosoma large, 61–65 x 60–67, cheliceral stylets 75–84, tarsi II-III setae tcʹʹ 3–4 (Figs 25–26)… Host: Castelnaudia cordata................................................................................ E. hebronae sp. nov.</p> <p>- Gnathosoma smaller, 45–51 x 47–51, cheliceral stylets 50–55, tarsi II-III setae tcʹʹ 2–3 (Figs 24–26)… Host: Castelnaudia porphyriaca........................................................................ E. despoticus sp. nov.</p> <p>12. Setae sc1 &lt;10; distance d -d 90–105, f-f 75–92............................................................. 13</p> <p>- Setae sc1&gt; 15; distance d -d 135–150, f-f 100–120.......................................................... 14</p> <p>13. Setae c1 15–17, seta d 16–22; distance v2-v2 38–40; sc1-sc1 61–65; v1-sc1 16–20; tibia I seta v′ 2; genua II-III setae l′ minute (Figs 37–38). Host: Castelnaudia cordata................................................... E. osculum sp. nov.</p> <p>- Setae c1 6–9, d 7–9; distance v2-v2 48–62; sc1-sc1 82–100; v1-sc1 36–46; tibia I seta v′ 5–6; genua II-III setae l′ 3–4 (Figs 28–29). Host: Castelnaudia porphyriaca.................................................... E. basiatus sp. nov.</p> <p>14. Cheliceral seta 24–39 (Fig. 39); larva with short setae c1 6–9, c2 9–12, d 7–10. Host: Castelnaudia eungella... E. savatus sp. nov.</p> <p>- Cheliceral seta 17-21 (Fig. 32); larva with longer setae c1 15–18, c2 19–24, d 17–19 (Fig. 35). Host: Castelnaudia marginifera.................................................................................................................................................................................. E. labiatus sp. nov.</p></div> 	http://treatment.plazi.org/id/4438245CFFFC094687AEFBD9FA38A696	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Seeman, Owen D.	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFFD094687AEFD73FA38A7BF.text	4438245CFFFD094687AEFD73FA38A7BF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eutarsopolipus Berlese 1913	<div><p>Key to species of Eutarsopolipus from Notonomus</p> <p>Key to adult females</p> <p>1. Leg I with one claw; genu I with one seta (l″ absent)........................ E. lambkinae Constantine &amp; Seeman, 2014</p> <p>- Leg I with paired claws; genu I with two setae (l″ present)..................................................... 2</p> <p>2. Setae sc2 short, 15–20; tarsi II-III with setae tc′ short (3–7) and tc″ small (5–6).................... E. biuncatus sp. nov.</p> <p>- Setae sc2 longer, 36–40; tarsi II-III with setae tc′ longer (8–11) and tc″ larger (8–9).................... E. janus sp. nov.</p></div> 	http://treatment.plazi.org/id/4438245CFFFD094687AEFD73FA38A7BF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Seeman, Owen D.	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFFD095E87AEFB14FB43A339.text	4438245CFFFD095E87AEFB14FB43A339.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eutarsopolipus hadros Seeman 2021	<div><p>Eutarsopolipus hadros sp. nov.</p> <p>(Figs 1–8, 10)</p> <p>Diagnosis. Adult female: setae sc1, c1, c2, d, f with thickened bases, finely tapering (rarely asymmetrically not tapering); anterior margin of prodorsal shield eroded, concave; setae v1 9–11. Larviform male: sc1 9. Larval female: Setae c1 16–17, d 11–12, f 10–11.</p> <p>Material examined. Holotype female ex. Castelnaudia setosiceps. 9.5 km N of <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=145.48917&amp;materialsCitation.latitude=-17.525278" title="Search Plazi for locations around (long 145.48917/lat -17.525278)">Ravenshoe</a>, 17° 31′ 31″ S 145° 29′ 21″E, 9 Feb-15 Apr 1999, G. &amp; S. Monteith, pitfall trap, 1060 masl, HR# T 78934 (QMS 113562). Paratypes. All ex. C. setosiceps. 10 females, 1 male, 3 larvae, as follows: 2 females, same data as holotype; 6 females, 1 male, 1 larva, The Crater NP, Atherton Tablelands, 17° 25′ 43″ S 145° 29′ 13″E, 5 Dec 1988, G. Monteith &amp; G. Thompson, 1000 masl, HR# T 19427 (QMS 113565–71, 1 female ANIC 52-003949); 2 females, 2 larvae, Millaa Millaa Lookout, 17° 31′ 23″ S 145° 34′ 05″E, 1 Dec 1993 - 25 Feb 1994, J. Hasenpusch, pitfall trap, 1000 masl, HR# T 19429 (QMS 113572–74, ZMH-A0015181). 1 female in each of ANIC, ZMH.</p> <p>Note: one female specimen from the Crater was too poor to include in the type series (QMS 113575). One pharate female from the Crater (QMS 113576).</p> <p>Description. FEMALE (Figs 1-3, 10; n = 11).</p> <p>Gnathosoma. Length 60 (58–64), width 55 (52–62). Cheliceral stylets 73 (68–75), setae ch 21 (19–24), su 14 (11–13).</p> <p>Idiosoma (Figs 1–2, 10). Length 340 (340–400), width 265 (255–350). Prodorsal plate eroded mid-anteriorly and anterolaterally; setae v1 11 (9–11), v2 alveolar remnant, sc1 20 (15–19), sc2 66 (65–74); setae sc1 thickened but not bulbous (Fig. 10). Distance between setae v1–v1 46 (46–53), sc1–sc1 92 (100–105), v1–sc1 31 (28–35), v2–v2 44 (50–52), sc2–sc2 135 (135–140), sc1–sc2 35 (31–35). Opisthosomal setae c1, c2, d, f thickened, sometimes appearing stout by lacking fine tips, especially setae c2 (Figs 1, 10). Plate C setae c1 17 (11–21), c2 8 (8–18), distance between setae c1–c1 96 (92–100). Plate D setae d 14 (13–20), d–d 145 (145–165), cupuli ia laterad d. Plate EF setae f 14 (14–18), f–f 130 (115–155), cupuli im laterad f. Plate H almost completely divided into two small plates, often ventral, seta h 7 (5–6). Venter (Fig. 2): coxisterna 1 and 2 well defined, coxisterna 3 weakly defined; coxal setae slender, 1a 5 (4–5), 2a 8 (7–8), 3a 6 (7–8), 3b 8 (9–11). Alveoli of 1b, 2b present.</p> <p>Legs (Fig. 3). Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Leg I. Femur I, d minute, l’ 3 (2–3), v″ 12 (10–12); genu I, l′ 3 (3), l″ 3 (3); tibia I, d 75 (60–75), l’ 7 (6–7), l’’ 8 (7–10), v’ 4 (4–5), v’’ 11 (10–12), k 5 (5–6), φ 10 (9–10); tarsus I, tc’ 15 (15–18), tc’’ 16 (16–19), pl’ 11 (9–11), pl’’ 17 (14–18), pv′ 3 (2–3), pv’’ 4 (4–5), ω 3 (3–4), s 8 (7–8), u′′ 2 (m–2). Leg II. Genu II, l′ 4 (3–4); tibia II, d 56 (45–50), l’ 7 (6–7), v’ 17 (12–13), v’’ 15 (15–20); tarsus II, tc′ 2 (2), tc ″ 6 (5–6), pl ″ 51 (31–49), pv’’ 4 (4–5), u’ 7 (7–8), u′′ minute. Leg III. Genu III, l′ 3 (3–4); tibia III, d 38 (35–45), l’ 7 (6–7), v’ 13 (11–15), v’’ 16 (16–22); tarsus III, tc’ 2 (2–3), tc ″ 5 (5–6), pl ″ 45 (41–50), pv’’ 3 (3–4), u’ 7 (7), u′′ minute.</p> <p>LARVIFORM ADULT MALE (Figs 4–5; n = 1).</p> <p>Gnathosoma. Length 33, width 30. Cheliceral stylets 25, setae ch 5, su 6.</p> <p>Idiosoma (Figs 4–5). Length 165, width 130. Prodorsal plate with setae v1 5, v2 alveolar remnant, sc1 9, sc2 85. Distance between setae v1–v1 22, sc1–sc1 48, v1–sc1 18, v2–v2 28, sc2–sc2 60, sc1–sc2 21. Plate CD setae c1 6, c2 5, d 6, distance between setae c1–c1 40, c1–c2 30, d–d 41; cupuli ia laterad setae d (asymmetrical expression). Plate EF concave, eroded posteriorly; with setae f 3, distance f–f 28; cupuli im anterolaterad setae f. Genital capsule length 29, width 34, setae h minute. Venter (Fig. 4): coxal setae 1a 3, 2a 4, 3a 5, 3b 5.</p> <p>Legs. Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Leg I. Femur I, d minute, l′ minute, v″ 6; genu I l′ 2, l″ 2; tibia I, d 45, l’ 2, l’’ 3, v ’ 3, v ’’ 8, k 4, φ 9; tarsus I, tc’ 13, tc’’ 15, pl’ 8, pl’’ 14, pv ′ 2, pv’’ 2, ω 3, s 5, u ′′ minute. Leg II. Genu II, l′ 2; tibia II, d 21, l’ 5, v’ 11, v’’ 15; tarsus II, tc′ 2, tc ″ 5, pl ″ 27, pv’’ 2, u’ 5, u″ 2. Leg III. Tibia III, d 22, l’ 4, v’ 10, vʺ 13; tarsus III, tc′ 2, tc ″ 5, pl ″ 28, pv’’ 3, u ’ 5, u″ minute.</p> <p>LARVAL FEMALE (Figs 6–8; n = 3).</p> <p>Gnathosoma. Length 50–51, width 46–50. Palp with minute dorsal femoral seta. Cheliceral stylets 62–65, setae ch 20–27, su 10–11.</p> <p>Idiosoma (Fig. 6). Length 225–240, width 175–185. Prodorsal plate overlapping folded membranous cuticle (outline shown in figure), with setae v1 5–6, v2 alveolar remnant, sc1 9–10, sc2 100–120. Distance between setae v1–v1 33–37, sc1–sc1 68–70, v1–sc1 26–28, v2–v2 36–39, sc2–sc2 85–88, sc1–sc2 34–37. Plate C setae c1 16–17, c2 10–11, distance between setae c1–c1 49–52. Plate D setae d 11–12, d–d 61–62, cupuli ia very close to setae d, anteriad to anterolaterad d. Plate EF setae f 10–11, f–f 52–57, cupuli im close to setae f, anterolaterad f. Plate H narrow, 16–19 wide, ca. 25–30 long bearing setae h1 ca. 180, h2 11–12. Venter (Fig. 7): coxisterna 1-2 well defined, coxisterna 3 poorly defined, membranous; coxal setae slender, 1a 2–3, 2a 4–5, 3a 6–7, 3b 5. Alveoli of 1b, 2b present.</p> <p>Legs (Fig. 8). Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Leg I. Femur I, d minute, l′ minute, v″ 6–9; genu I, l′ 2, l″ 2; tibia I, d 53–60, l’ 8–9, l’’ 11–13, v’ 2–3, v’’ 12–13, k 6–7, φ 10–11; tarsus I, tc’ 15–16, tc’’ 16–18, pl’ 9–10, pl’’ 17–20, pv′ 2, pv’’ 3, ω 3, s 5, u′′ minute. Leg II. Genu II, l′ 2–3; tibia II, d 35–40, l’ 6–7, v’ 13–15, v’’ 14–20; tarsus II, tc′ 2–3, tc ″ 4–5, pl ″ 40–43, pv’’ 3, u’ 6, u′′ 2. Leg III. Genu III, l′ 2–3; tibia III, d 29–45, l’ 6–8, v’ 12–17, v’’ 15–17; tarsus III, tc’ 2–3, tc ″ 4–5, pl ″ 30–35, pv’’ 3, u’ 6–7, u′′ minute.</p> <p>Differential diagnosis. Female E. hadros sp. nov. has idiosomal setae that are less thickened than those of E. mixtus sp. nov. and E. umbonatus sp. nov., giving these setae a comparatively slender appearance. See Fig. 10 and the Differential diagnosis for E. mixtus for further discussion of this feature. The anterior margin of the prodorsal shield of female E. hadros is also deeply eroded medially; this erosion is weak in E. umbonatus and absent in E. mixtus. The male of E. hadros has a longer seta sc1 (9 versus 3–6 in E. mixtus and E. umbonatus), but the single male specimen of E. hadros reduces confidence in this possible diagnostic feature. Larval female E. hadros are difficult to distinguish from E. mixtus, but the size of setae f (10–11 versus 14–19) is useful. Larvae of E. hadros are distinguished from E. umbonatus by having longer setae c1 (16–19 versus 8–11).</p> <p>Etymology. The specific name hadros (Greek, strong, stout) alludes to the strong dorsal setae.</p> </div>	http://treatment.plazi.org/id/4438245CFFFD095E87AEFB14FB43A339	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Seeman, Owen D.	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFE6095B87AEFF5AFC7EA2F0.text	4438245CFFE6095B87AEFF5AFC7EA2F0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eutarsopolipus mixtus Seeman 2021	<div><p>Eutarsopolipus mixtus sp. nov.</p> <p>(Figs 9–10)</p> <p>Diagnosis. Adult female: setae sc1, c1, c2, d, f with strongly thickened bases, often blunt-tipped; anterior margin of prodorsal shield eroded, concave; setae v1 10–13. Larviform male: sc1 3–6. Larval female: Setae c1 13–20, d 11–12, f 14–19.</p> <p>Material examined. Holotype female ex. Castelnaudia mixta. <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=148.50833&amp;materialsCitation.latitude=-21.168055" title="Search Plazi for locations around (long 148.50833/lat -21.168055)">Broken River</a>, Eungella, 21° 10′ 05″ S 148° 30′ 30″E, 17 Nov 1992 - 15 Apr 1993, G. Monteith &amp; D. Cook, flight intercept &amp; pitfall trap, HR# T 82585 (QMS 113035). Paratypes. 12 females, 4 males, 4 larvae, as follows: 11 females, 3 males, 1 larva, same data as holotype (QMS 113036-43, 113045, 113047, ANIC 52-003950 – 51, ZMH-A0015182–83); 1 male, 1 larva, same data except HR# T 82586 (QMS 113046, ZMH-A0015184). 1 female, same data except HR# T 82582 (QMS 113044); 1 larva, same data except HR# T 82584 (ANIC 52-003952); 1 larva, Pearse’s Lookout, Eungella, 21° 06′ 53″ S 148° 31′ 06″S, 17 Nov 1992 - 15 Apr 1993, G. Monteith &amp; D. Cook, IN5934, HR# T 82526 (QMS 113048). All in QM except 1 female, male, larva in ANIC and ZMH.</p> <p>Description. FEMALE (Figs 9–10; n = 13).</p> <p>Gnathosoma. Length 59 (57–65), width 55 (53–60). Cheliceral stylets 75 (67–78), setae ch 18 (16–25), su 17 (11–18).</p> <p>Idiosoma (Fig. 9). Length 395 (365–400), width 280 (270–330). Prodorsal plate eroded mid-anteriorly and anterolaterally; setae v1 13 (10–13), v2 alveolar remnant, sc1 thickened, 11/16 (asymmetry) (11–17), sc2 75 (70–95) (Fig. 10). Distance between setae v1–v1 48 (41–52), sc1–sc1 94 (82–100), v1–sc1 33 (26–40), v2–v2 43 (41–51), sc2–sc2 130 (130–140), sc1–sc2 35 (24–34). Opisthosomal setae c1, c2, d, f thickened, sometimes appearing stout by lacking fine tips, especially setae c2. Plate C setae c1 12/20 (asymmetry) (10–20), c2 9 (10–15), distance between setae c1–c1 100 (78–95). Plate D setae d 17 (12–21), d–d 145 (135–150), cupuli ia laterad d. Plate EF setae f 13/18 (asymmetry) (11–19), f–f 135 (110–125), cupuli im laterad f. Plate H divided into two small plates, or fused weakly, often ventral, seta h 5 (5–6). Venter: coxisterna 1 and 2 well defined, coxisterna 3 weakly defined; coxal setae slender, 1a 4 (3–4), 2a 8 (6–8), 3a thickened, 5 (5–7), 3b 9 (7–9). Alveoli of 1b, 2b present.</p> <p>Legs (holotype only, except if holotype different from E. hadros). Setal counts legs I–III, femur-tarsus: 3-2- 6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Leg I. Femur I, d minute, l’ 4 (3–5), v″ 11; genu I, l′ 2, l″ 3; tibia I, d 65, l’ 8, l’’ 10, v’ 5, v’’ 10, k 5, φ 7 (7–9); tarsus I, tc’ 16, tc’’ 15, pl’ 9, pl’’ 13, pv′ 3, pv’’ 3, ω 3, s 7, u′′ minute. Leg II. Genu II, l′ 3; tibia II, d 57, l’ 7, v’ 13, v’’ 19; tarsus II, tc′ 2, tc ″ 6, pl ″ 60, pv’’ 4, u’ 7, u′′ minute. Leg III. Genu III, l′ 3; tibia III, d 42, l’ 7, v’ 11, v’’ 15; tarsus III, tc’ 2, tc ″ 5, pl ″ 39, pv’’ 3, u’ 6, u′′ minute.</p> <p>LARVIFORM ADULT MALE (n = 4).</p> <p>Gnathosoma. Length 32–35, width 30–34. Cheliceral stylets 23–27, setae ch 5–7, su 5–10.</p> <p>Idiosoma. Length 160–205, width 125–150. Prodorsal plate with setae v1 4–5, v2 alveolar remnant, sc1 3–6, sc2 60–70. Distance between setae v1–v1 21–26, sc1–sc1 40–52, v1–sc1 20–25, v2–v2 22–27, sc2–sc2 57–77, sc1–sc2 16–22. Plate CD setae c1 4–6, c2 4–5, d 4–5, distance between setae c1–c1 46–59, c1–c2 22–28, d–d 36–52; cupuli ia anterolaterad setae d. Plate EF eroded medially, almost divided; with setae f 2–3, distance f–f 24–30; cupuli im anterolaterad to laterad setae f. Genital capsule length 28–30, width 34–38, setae h minute. Venter: coxal setae 1a 2–3, 2a 4–5, 3a 3–4, 3b 5–6.</p> <p>Legs. Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Leg I. Femur I, d minute, l′ 2, v″ 7; genu I l′ 2, l″ 2; tibia I, d 38, l’ 3, l’’ 2, v’ 3, v’’ 10, k 4, φ 6–8; tarsus I, tc’ 11–14, tc’’ 11–14, pl’ 7, pl’’ 11, pv ′ 2, pv’’ 3, ω 3, s 4, u ′′ minute. Leg II. Genu II, l′ 2; tibia II, d 16, l’ 5, v’ 10, v’’ 13; tarsus II, tc′ 2, tc ″ 4, pl ″ 27, pv’’ 3, u’ 5, u″ 2. Leg III. Tibia III, d 16, l’ 4, v’ 12, v″ 12; tarsus III, tc′ 2, tc ″ 3–5, pl ″ 20, pv’’ 3, u’ 5, u″ present.</p> <p>LARVAL FEMALE (n = 4).</p> <p>Gnathosoma. Length 46–51, width 43–49. Cheliceral stylets 61–64, setae ch 23–27, su 7–10.</p> <p>Idiosoma. Length 205–260, width 165–215. Prodorsal plate overlapping folded membranous cuticle, with setae v1 6–8, v2 alveolar remnant, sc1 7–10, sc2 120–135. Distance between setae v1–v1 31–35, sc1–sc1 59–63, v1–sc1 23–25, v2–v2 30–34, sc2–sc2 72–83, sc1–sc2 31–33. Plate C setae c1 13–20, c2 8–11, distance between setae c1–c1 42–48. Plate D setae d 11–12, d–d 53–61, cupuli ia very close to setae d, anteriad to anterolaterad d. Plate EF setae f 14–19, f–f 46–52, cupuli im close to setae f, anterolaterad f. Plate H narrow, 15–20 wide, ca. 25–30 long bearing setae h1 ca. 150, h2 13–17. Venter: coxisterna 1-2 well defined, coxisterna 3 poorly defined, membranous; coxal setae slender, 1a 3, 2a 4–5, 3a 4–5, 3b 4–5. Alveoli of 1b, 2b present.</p> <p>Legs. Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Leg I. Femur I, d minute, l′ 2, v″ 7; genu I, l′ 2, l″ 3; tibia I, d 54, l’ 9, l’’ 10, v’ 4, v’’ 8–12, k 6, φ 10; tarsus I, tc’ 13, tc’’ 15, pl’ 8, pl’’ 16, pv′ 2, pv’’ 3, ω 3, s 5, u′′ minute. Leg II. Genu II, l′ 4; tibia II, d 40, l’ 7, v’ 14, v’’ 16; tarsus II, tc′ 2, tc ″ 4, pl ″ 35, pv’’ 3, u’ 5, u′′ 2. Leg III. Genu III, l′ 4; tibia III, d 35, l’ 7, v’ 13, v’’ 14; tarsus III, tc’ 2, tc ″ 4, pl ″ 21–40, pv’’ 3, u’ 6, u′′ minute.</p> <p>Differential diagnosis. Eutarsopolipus mixtus sp. nov. is similar to E. hadros sp. nov. but is most easily distinguished by having thicker dorsal setae. This feature is useful for species separation, but is complicated by intraspecific variation in setal form, which is often expressed asymmetrically. However, setae sc1 are always thicker in E. mixtus, even if they taper. Likewise, other dorsal idiosomal setae tend to be thicker in E. mixtus, exemplified by setae f (Fig. 10).</p> <p>The prodorsal shield of E. umbonatus is weakly eroded anteriorly, in contrast to the deep erosion of E. hadros and lack of erosion in E. umbonatus. However, the intermediate state of this feature reduces its effectiveness. In larvae, E. mixtus have longer setae f (14–19 versus 10–11 in E. hadros and 7–10 in E. umbonatus).</p> <p>Etymology. The species name mixtus (Latin, mixed) refers both to the host species C. mixta but also the variation in form of the dorsal setae.</p> <p>Remarks. Three paratype females have fully developed larvae within their bodies, with no egg, suggestive of vivipary. Two additional specimens of C. mixta from Finch Hatton Gorge (HR# T82535, T82529) had a male and a larva (one per host) of an undermined species of Eutarsopolipus (Table 1).</p> </div>	http://treatment.plazi.org/id/4438245CFFE6095B87AEFF5AFC7EA2F0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Seeman, Owen D.	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFE0095887AEF8E9FC3FA6E5.text	4438245CFFE0095887AEF8E9FC3FA6E5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eutarsopolipus umbonatus Seeman 2021	<div><p>Eutarsopolipus umbonatus sp. nov.</p> <p>(Fig. 11)</p> <p>Diagnosis. Adult female: setae sc1, c1, c2, d, f with strongly thickened bases, often blunt-tipped; anterior margin of prodorsal shield not or only slightly eroded, straight or weakly concave; setae v1 7–10. Larviform male: sc1 5–6. Larval female: Setae c1 8–11, d 7–10, f 7–10.</p> <p>Material examined. Holotype female ex. Castelnaudia cordata, Mt Mee SF, 6 km NNW barracks, 27° 2ʹ 53ʺ S 152° 40ʹ 49ʺ E, 8 Jan-3 Mar 1992, D. Cook, flight intercept trap, IN5148, HR# T 68607 (QMS113481). Paratypes. 16 females, 7 males, 15 larvae, as follows: all ex. C. cordata. 15 females, 6 males, 13 larvae, same data as holotype (QMS 113482–92, ANIC 52-003953 – 57, ZMH-A0015185–89); 1 female, 1 larva, McIntosh Rd, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=152.70389&amp;materialsCitation.latitude=-28.421944" title="Search Plazi for locations around (long 152.70389/lat -28.421944)">Unumbar</a> SF, 28° 25ʹ 19ʺ S 152° 42ʹ 14ʺ E, 25 Nov 1972, G. Monteith, IN6467, HR# T 64329 (QMS 113493–94); 1 male, 1 larva, “ <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=153.11612&amp;materialsCitation.latitude=-28.049444" title="Search Plazi for locations around (long 153.11612/lat -28.049444)">Glen Witheren</a> ”, shelf scrub, 28° 2ʹ 58ʺ S 153° 6ʹ 58ʺ E, 30 Dec 1991 - 24 May 1992, G. Monteith, pitfall trap, IN5682, rainforest, HR#68606 (QMS 113495); 1 larva, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=153.17944&amp;materialsCitation.latitude=-28.205557" title="Search Plazi for locations around (long 153.17944/lat -28.205557)">Lower Coomera</a>, 28° 12ʹ 20ʺ S 153° 10ʹ 46ʺ E, 9 Jan-6 Apr 1995, G. Monteith, pitfall trap, IN5998, HR# T 68610. All in QM except 3 females, 1 male, 1 larva in each of ANIC and ZMH.</p> <p>Other material examined. 5 females, 2 larvae, on same slide as specimens of E. hebronae, same data as holotype. Not included as paratypes.</p> <p>Description. FEMALE (Fig. 11; n = 17).</p> <p>Gnathosoma. Length 62 (56–61), width 57 (55–58). Cheliceral stylets 68 (68–70), setae ch 22 (25–30), su 10 (10–18).</p> <p>Idiosoma (Fig. 11). Length 330 (320–400), width 290 (250–325). Prodorsal plate slightly or not eroded midanteriorly, more substantially eroded anterolaterally; setae v1 7 (7–10), v2 alveolar remnant, sc1 usually thickened, thorn-like 7 (7–12) [14/ 16 in specimen #5, 12/ 15 in specimen #8, 9/ 15 in specimen #11], sc2 50–80. Distance between setae v1–v1 47 (41–49), sc1–sc1 105 (94–105), v1–sc1 38 (32–36), v2–v2 45 (41–50), sc2–sc2 130 (125– 130), sc1–sc2 27 (25–32). Opisthosomal setae c1, c2, d, f usually stout but sometimes with fine tips. Plate C setae c1 7 (6–12; 15 in specimen #8), c2 10 (10–15), distance between setae c1–c1 84 (82–100). Plate D setae d 10 (10–15), d–d 155 (160–185), cupuli ia laterad d. Plate EF setae f 7/10 (asymmetry) (7–12; 10/ 13 in specimen #8; 15 in specimen #15), f–f 115 (130–150), cupuli im laterad f. Plate H divided or fused weakly, seta h 7 (5–7). Venter: coxisterna 1 well defined, coxisterna 2 well defined but sometimes with folds, coxisterna 3 weakly defined; coxal setae slender, 1a 4 (3–4), 2a 6 (5–6), 3a thickened, 3–6, 3b 4–6. Alveoli of 1b, 2b present.</p> <p>Legs (holotype only, except if holotype different from E. hadros / mixtus). Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Leg I. Femur I, d minute, l’ 2, v″ 14; genu I, l′ 3, l″ 3; tibia I, d 60, l’ 7, l’’ 10, v’ 5, v’’ 8, k 5, φ 8 (7–9); tarsus I, tc’ 14, tc’’ 15, pl’ 7, pl’’ 13, pv′ 3, pv’’ 4, ω 4, s 7, u′′ minute. Leg II. Genu II, l′ 4; tibia II, d 40, l’ 7, v’ 12, v’’ 23; tarsus II, tc′ 2, tc ″ 6, pl ″ 55, pv’’ 4, u’ 7, u′′ minute. Leg III. Genu III, l′ 3; tibia III, d 29, l’ 8, v’ 11, v’’ 21; tarsus III, tc’ 2, tc ″ 6, pl ″ 52, pv’’ 4, u’ 8, u′′ minute.</p> <p>LARVIFORM ADULT MALE (n = 7).</p> <p>Gnathosoma. Length 33–35, width 30–32. Cheliceral stylets 25–28, setae ch 6–7, su 5–7.</p> <p>Idiosoma. Length 170–185, width 120–130. Prodorsal plate with setae v1 3–5, v2 alveolar remnant, sc1 5–6, sc2 70–80. Distance between setae v1–v1 19–21, sc1–sc1 46–50, v1–sc1 22–26, v2–v2 19–22, sc2–sc2 57–63, sc1–sc2 20–21. Plate CD setae c1 5–7, c2 4–6, d 5–6, distance between setae c1–c1 42–45, c1–c2 27–31, d–d 40–43; cupuli ia anterolaterad setae d. Plate EF eroded medially, sometimes almost divided; with setae f 2–4, distance f–f 27–28; cupuli im anterolaterad to laterad setae f. Genital capsule length 28–31, width 31–32, setae h minute. Venter: coxal setae 1a 2–3, 2a 4, 3a 2–3, 3b 4–5.</p> <p>Legs. Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Leg I. Femur I, d minute, l’ minute, v″ 7; genu I l′ 2, l″ 2; tibia I, d 44, l’ 2, l’’ 3, v ’ 3, v ’’ 7, k 4, φ 8; tarsus I, tc’ 12, tc’’ 12, pl’ 7, pl’’ 12, pv ′ 2, pv’’ 2, ω 3, s 5, u ′′ 2. Leg II. Genu II, l′ 2; tibia II, d 18, l’ 5, v’ 10, v’’ 17; tarsus II, tc′ 3, tc ″ 4, pl ″ 28, pv’’ 3, u’ 5, u″ 2. Leg III. Tibia III, d 14, l’ 5, v’ 11, vʺ 11; tarsus III, tc′ 3, tc ″ 4, pl ″ 22, pv’’ 3, u’ 5, u″ minute.</p> <p>LARVAL FEMALE (n = 15).</p> <p>Gnathosoma. Length 48–52, width 46–54. Cheliceral stylets 63–70, setae ch 27–29, su 7–9.</p> <p>Idiosoma. Length 195–280, width 135–215. Prodorsal plate with setae v1 4–7, v2 alveolar remnant, sc1 5–8, sc2 100–120. Distance between setae v1–v1 28–37, sc1–sc1 65–75, v1–sc1 23–26, v2–v2 32–37, sc2–sc2 71–85, sc1–sc2 32–37. Plate C setae c1 8–11, c2 5–10, distance between setae c1–c1 40–48. Plate D setae d 7–10, d–d 49–63, cupuli ia very close to setae d, anteriad to anterolaterad d. Plate EF setae f 7–10, f–f 48–50, cupuli im close to setae f, anteriad f. Plate H narrow, 13–15 wide, ca. 25–35 long bearing setae h1 ca. 170, h2 6–8 (Mt Mee specimens 12–15). Venter: coxisterna 1-2 well defined, coxisterna 3 poorly defined, membranous; coxal setae 1a 2–3, slender, 2a 4–5, slender, 3a 3–5 sometimes slightly thickened, 3b 4–5, slender. Alveoli of 1b, 2b present.</p> <p>Legs. Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Leg I. Femur I, d minute, l′ 2, v″ 8; genu I, l′ 3, l″ 2; tibia I, d 62, l’ 9, l’’ 10, v’ 3, v’’ 11, k 5, φ 10; tarsus I, tc’ 12, tc’’ 13, pl’ 10, pl’’ 16, pv′ 2, pv’’ 3, ω 3, s 5, u′′ 2. Leg II. Genu II, l′ 3; tibia II, d 35, l’ 5, v’ 15, v’’ 15; tarsus II, tc′ 2, tc ″ 4, pl ″ 37, pv’’ 3, u’ 6, u′′ minute. Leg III. Genu III, l′ 3; tibia III, d 33, l’ 6, v’ 13, v’’ 15; tarsus III, tc’ 2, tc ″ 4, pl ″ 32, pv’’ 3, u’ 6, u′′ minute.</p> <p>Differential diagnosis. Eutarsopolipus umbonatus sp. nov. is similar to E. mixtus sp. nov. by having much stronger dorsal setae compared with E. hadros. However, like all species and particularly E. mixtus, the form of these setae differed intraspecifically: although the thicker nature of these setae are useful for separating E. umbonatus from E. hadros, they are too variable to separate E. umbonatus from E. mixtus. Larvae are more useful for distinguishing the latter two species. Larva of E. umbonatus have smaller idiosomal setae: setae c1 (8–11 versus 13–20), d (7–10 versus 11–12) and f (7–10 versus 14–19) are slightly to obviously shorter in E. umbonatus.</p> <p>Etymology. The specific name umbonatus (knob-like) refers to the short, more knob-like setae in females of this species.</p> <p>Remarks. In larvae, setae h2 were longer in the type series from Mt Mee (12–15) compared with the three specimens from other sites (6–8). No other features distinguished these populations. However, Mt Mee is part of a mountain range (the D’Aguilar Range) to the northwest of Brisbane, while all other collections were from the Border Range to the south to southwest of Brisbane. The geographical separation of these populations, combined with the difference in length of seta h, suggests that these may be separate species on isolated populations of Castelnaudia cordata. Furthermore, these differences also suggest that the host species deserves closer attention. The D’Aguilar Range has several endemic taxa, such as a unique assemblage of anamid spiders (Rix et al. 2020). Likewise, the beetle specimens from the D’Aguilar Range may be an endemic species closely related to C. cordata.</p> <p>The above three species, E. hadros, E. mixtus and E. umbonatus, are very closely related and it is tempting to consider all three as one species. However, each species does exhibit subtle differences, they occur on different host species, and are highly isolated geographically with the hosts C. setosiceps, C. mixta and C. cordata in north-east Queensland, middle-east Queensland and south-east Queensland, respectively.</p> <p>One paratype female has a fully developed larva within its body, with no egg, indicating vivipary. Two larvae of another species close to Eutarsopolipus echinatus Seeman, 2019a (i.e. same thorn-like setae, but clearly a different species) were also collected from C. cordata: one from the type host (on a separate slide) and one from Lower Coomera and on the same slide as the larva of E. umbonatus (QMS 113496). This species was found co-infesting its host with E. hebronae sp. nov. and E. osculum sp. nov. (Tables 1, 2).</p> <p>The ochoai species group - nahmani subgroup</p> <p>Diagnosis. Female with dorsal idiosomal setae fine, slender; setae h small but developed. All life stages with gnathosomal apex evenly rounded.</p></div> 	http://treatment.plazi.org/id/4438245CFFE0095887AEF8E9FC3FA6E5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Seeman, Owen D.	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFE3095887AEFC81FE52A7DD.text	4438245CFFE3095887AEFC81FE52A7DD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eutarsopolipus nahmani Seeman 2021	<div><p>The ochoai species group - nahmani subgroup</p> <p>Diagnosis. Female with dorsal idiosomal setae fine, slender; setae h small but developed. All life stages with gnathosomal apex evenly rounded.</p></div> 	http://treatment.plazi.org/id/4438245CFFE3095887AEFC81FE52A7DD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Seeman, Owen D.	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFE3095387AEFBD9FE6EA29E.text	4438245CFFE3095387AEFBD9FE6EA29E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eutarsopolipus nahmani Seeman 2021	<div><p>Eutarsopolipus nahmani sp. nov.</p> <p>(Figs 12–16)</p> <p>Diagnosis. Adult female: Gnathosomal length 67–70, width 69–72; anterior margin of prodorsal shield not eroded medially; v1 6–7, sc1 10–12, c1 17–19, c2 14–19, d 15–19, f 15–18, h 3–6, v1-v1 54–64, d-d 200–230, f-f 150–170; genua II-III setae l′ 5–7, tibia I seta l′ 9–11, tarsi II-III setae tc′′ 5–6, pv′′ 2–3. Larval female: Gnathosomal length 57–59, width 57–60; idiosomal length 335–400; c1 (10–11) and c2 (12) subequal, v1–v1 47–59, sc1–sc1 100–125, c1–c1 105–120, d–d 130–135; genua II-III setae l′ 3–5, tibia I seta l′ 12–13, tarsi II-III setae tc′ 3–4, tc′′ 5–6, pv′′ 2–3.</p> <p>Material examined. Holotype female, ex. Castelnaudia septemcostata. O’Reilly’s Guesthouse, 28° 14ʹ 08ʺ S 152° 08ʹ 12ʺ E, 1-31 Dec 1909, H. Hacker, IN7977, HR# T 88472 (QMS 113011). <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=152.13666&amp;materialsCitation.latitude=-28.235556" title="Search Plazi for locations around (long 152.13666/lat -28.235556)">In</a> QM. Paratypes. All ex. C. septemcostata. 5 females, 2 males, 2 larvae, as follows: 1 larva, same data as holotype (QMS 113017); 3 females, Mt Cougal, shelf, 28° 14ʹ 10ʺ S 152° 19ʹ 34ʺ E, 19 Nov 1987 - 17 Mar 1988, D. Cook &amp; G. Monteith, pitfall trap, rainforest, IN6062, HR# T 64265 (QMS 113012, ANIC 52-003958, ZMH-A0015190); 1 female, 1 male, Bar Mountain, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=153.13388&amp;materialsCitation.latitude=-28.46028" title="Search Plazi for locations around (long 153.13388/lat -28.46028)">Wiangaree</a>, 28° 27ʹ 37ʺ S 153° 8ʹ 2ʺ E, 11 Oct 1977 - 2 Oct 1978, G. &amp; S. R. Monteith, pitfall trap, rainforest, IN5624, HR# T 57344 (QMS 113013, 113016); 1 female, 1 male, Tweed Lookout, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=153.10918&amp;materialsCitation.latitude=-28.387222" title="Search Plazi for locations around (long 153.10918/lat -28.387222)">Wiangaree</a>, 28° 23ʹ 14ʺ S 153° 6ʹ 33ʺ E, 16 Nov-27 Dec 1974, G. &amp; S. R. Monteith, pitfall trap, rainforest, IN5349, HR# T 57316 (QMS 113014-15); 1 larva, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=152.32611&amp;materialsCitation.latitude=-28.23611" title="Search Plazi for locations around (long 152.32611/lat -28.23611)">Upper Tallebudgera Creek</a>, 28° 14ʹ 10ʺ S 152° 19ʹ 34ʺ E, 19 Nov 1987 - 17 Mar 1988, D. Cook &amp; G. Monteith, pitfall trap, rainforest, IN6062, HR# T 64272 (QMS 113018). All in QM except 1 female in ANIC, 1 female in ZMH.</p> <p>Description. FEMALE (Figs 12–13, n = 6).</p> <p>Gnathosoma. Length 69 (67–70), width 70 (69–72). Cheliceral stylets 72 (73–82), setae ch 16 (15–24), su 16 (13–19).</p> <p>Idiosoma (Fig. 12). Length 520 (380–600), width 405 (295–470). Prodorsal plate not eroded anteriorly; setae slender, v1 7 (6–7), v2 alveolar remnant, sc1 10 (10–12), sc2 60 (60–70). Distance between setae v1–v1 68 (54–64), sc1–sc1 135 (135–140), v1–sc1 47 (46–53), v2–v2 74 (70–79), sc2–sc2 205 (200–225), sc1–sc2 55 (52–59). Opisthosomal setae slender. Plate C setae c1 16 (17–19), c2 13 (14–19), distance between setae c1–c1 150 (155–165). Plate D setae d 17 (15–19), d–d 215 (200–230), cupuli ia anteriad to anterolaterad d. Plate EF setae f 17 (15–18), f–f 170 (150–165), cupuli im anterolaterad f. Plate H divided or fused weakly, seta h 3 (3–6). Venter: coxisterna 1 well defined, coxisterna 2-3 well defined but sometimes with some folds; coxal setae slender, 1a 4 (3–4), 2a 5 (5–6), 3a 8 (8–10), 3b 6 (5–6). Alveoli of 1b, 2b present.</p> <p>Legs (Fig. 13). Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Leg I. Femur I, d minute, l’ 3 (2–3), v″ 13 (10–15); genu I, l′ 3 (3–4), l″ 3 (3–4); tibia I, d 63 (60–68), l’ 10 (9–11), l’’ 14 (11–15), v’ 7 (6–8), v’’ 10 (10–15), k 7 (6–8), φ 13 (12–13); tarsus I, tc’ 18 (18–19), tc’’ 18 (18–19), pl’ 13 (12–16), pl’’ 19 (17–20), pv′ 3 (2–3), pv’’ 2 (2–3), ω 4 (3–4), s 7 (6–8), u′′ 2 (2). Leg II. Genu II, l′ 7 (5–7); tibia II, d 50 (40–45), l’ 12 (8–12), v’ 11 (11–15), v’’ 27 (20–26); tarsus II, tc′ 4 (3–4), tc ″ 6 (5–6), pl ″ 62 (42–60), pv’’ 2 (2–3), u’ 8 (7–8), u′′ 2 (2). Leg III. Genu III, l′ 6 (5–6); tibia III, d 35 (28–35), l’ 7 (7), v’ 15 (15–17), v’’ 26 (20–22); tarsus III, tc’ 3 (3), tc ″ 5 (4–5), pl ″ 60 (50–70), pv’’ 2 (2), u’ 8 (7–8), u′′ 2 (m–2).</p> <p>LARVIFORM ADULT MALE (Fig. 14; n = 2)</p> <p>Gnathosoma. Length 37–38, width 36. Cheliceral stylets 24–27, setae ch 6–7, su 6.</p> <p>Idiosoma (Fig. 14). Length 225–235, width 150–170. Prodorsal plate with setae v1 6–7, v2 alveolar remnant, sc1 5, sc2 50–55. Distance between setae v1–v1 23–24, sc1–sc1 49–50, v1–sc1 24–26, v2–v2 28–33, sc2–sc2 67–70, sc1–sc2 34–36. Plate CD setae c1 6–8, c2 7–8, d 7, distance between setae c1–c1 62–65, c1–c2 33–37, d–d 56–63; cupuli ia anterolaterad setae d. Plate EF slightly eroded medially; with setae f 6–7, distance f–f 39–45; cupuli im anteriad to anterolaterad setae f. Genital capsule length 36–39, width 36, setae h m–2. Venter: coxal setae 1a 2, 2a 2–3, 3a 5, 3b 4.</p> <p>Legs. Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Leg I. Femur I, d minute, l′ minute, v″ 5–6; genu I l′ 2, l″ 2; tibia I, d 26–30, l’ 4–5, l’’ 6–7, v ’ 5–7, v ’’ 7–8, k 3–4, φ 7–8; tarsus I, tc’ 15, tc’’ 15, pl’ 6–7, pl’’ 12–15, pv ′ 2, pv’’ 2, ω 2–3, s 4–5, u ′′ m. Leg II. Genu II, l′ 2–3; tibia II, d 15–23, l’ 5–6, v’ 9–11, v’’ 15–19; tarsus II, tc′ m–2, tc ″ 3–4, pl ″ 23–30, pv’’ 2, u ’ 5–6, u″ minute. Leg III. Genu III, l′ 2–3; tibia III, d 13–15, l’ 5–7, v’ 9–11, vʺ 12–16; tarsus III, tc′ 2, tc ″ 3–4, pl ″ 30–35, pv’’ 2, u’ 5–6, u″ minute.</p> <p>LARVAL FEMALE (Figs 15–16).</p> <p>Gnathosoma. Length 57–59, width 57–60. Cheliceral stylets 60–62, setae ch 16–25, su 10–11.</p> <p>Idiosoma (Fig. 15). Length 335–400, width 270–350. Prodorsal plate with setae v1 4–5, v2 alveolar remnant, sc1 6–7, sc2 90–105. Distance between setae v1–v1 47–59, sc1–sc1 100–125, v1–sc1 40–41, v2–v2 61–78, sc2–sc2 140–155, sc1–sc2 52–53. Plate C setae c1 10–11, c2 12, distance between setae c1–c1 105–120. Plate D setae d 9–10, d–d 130–135, cupuli ia anterolaterad setae d. Plate EF setae f 10, f–f 83, cupuli im anterolaterad f. Plate H narrow, 17–20 wide, 23–30 long, bearing setae h1 ca. 130, h2 9–10. Venter: coxisterna 1-2 well defined, coxisterna 3 with some folds; coxal setae slender, 1a 3, 2a 4, 3a 7–8, 3b 5. Alveoli of 1b, 2b present.</p> <p>Legs (Fig. 16). Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Leg I. Femur I, d minute, l′ 2, v″ 7–8; genu I, l′ 2, l″ 2; tibia I, d 50–65, l’ 12–13, l’’ 11–13, v’ 6–8, v’’ 10–11, k 5, φ 12; tarsus I, tc’ 19–22, tc’’ 18–21, pl’ 12–14, pl’’ 14–20, pv′ 2, pv’’ 2, ω 2, s 5–6, u′′ m. Leg II. Genu II, l′ 4–5; tibia II, d 25–42, l’ 10–11, v’ 11–17, v’’ 20–22; tarsus II, tc′ 2–3, tc ″ 3–4, pl ″ 44–47, pv’’ 2, u’ 7–8, u′′ minute. Leg III. Genu III, l′ 3–5; tibia III, d 32–35, l’ 10–11, v’ 13–16, v’’ 21–22; tarsus III, tc’ 2–3, tc ″ 3–4, pl ″ 44–50, pv’’ 2–3, u’ 6, u′′ minute.</p> <p>Differential diagnosis. The nahmani subgroup also includes Eutarsopolipus burwelli Seeman, 2019a (from Nurus medius) and Eutarsopolipus rutherfordae Constantine &amp; Seeman, 2014 (from Trichosternus subvirens). Eutarsopolipus burwelli has a large gnathosoma and in this manner is closer to E. nahmani than the other species of this subgroup. Considering their presence on different genera, the species are surprisingly similar. Females differ by: setae v1 smaller in E. nahmani (6–7 versus 9–13), as are setae sc2 (60–70 versus 95–120) and the gnathosomal cheliceral setae (15–24 versus 30–37); plate H also seems more rudimentary in E. burwelli, although this plate is often distorted or obscured, making comparisons difficult. Larvae also have smaller setae: v1 (4–5 versus 8–11), sc2 (90–105 versus 130–140) and ch (16–25 versus 30–38).</p> <p>Etymology. It is with great pleasure that I name this species for my son Harry Nahman, who was great company during school- and work-from-home periods in 2020, when he also helped by entering measurement data for several species.</p> <p>Remarks. The paratype females from Bar Mountain and Tweed Lookout have fully developed larvae in their body. No larvae seem to have egg shells, suggestive of vivipary. These larvae are not included as paratypes.</p> <p>One male (unregistered) from Tweed Lookout is an unknown species of Eutarsopolipus. Its most distinctive features are much larger sc1-sc1 (74 vs 49–50), the reduction of tc″ on tarsi II-III to a minute seta (ca. 1 µm, versus 4–5), and the tiny setae pv″ on tarsi II-III are not spur-like, instead being slender. A larva (unregistered) from Bar Mountain is the same species (setae tcʺ are similarly reduced). These specimens are tentatively considered closest to the despoticus subgroup.</p> </div>	http://treatment.plazi.org/id/4438245CFFE3095387AEFBD9FE6EA29E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Seeman, Owen D.	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFE8096E87AEF91EFDE1A124.text	4438245CFFE8096E87AEF91EFDE1A124.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eutarsopolipus raveni Seeman 2021	<div><p>Eutarsopolipus raveni sp. nov.</p> <p>(Figs 17–20)</p> <p>Diagnosis. Adult female: Gnathosomal length 58–60, width 48–50; anterior margin of prodorsal shield eroded medially; v1 7–10, sc1 13–16, c1 15–20, c2 19–23, d 17–23, f 14–20, h 2–4, v1-v1 40–46, d-d 115–125, f-f 115–125; genua II-III setae l′ 2–3, tibia I seta l′ 4–5, tarsi II-III setae tc′′ 5–6, pv′′ 3–4. Larval female: Gnathosomal length 45–50, width 38–41; idiosomal length 195–230; c1 7–11 and c2 7–10 subequal, v1–v1 30–33, sc1–sc1 57–62, c1–c1 44–54, d–d 55–67; genua II-III setae l′ m–2, tibia I seta l′ 6–9, tarsi II-III setae tc′ 2–3, tc′′ 4–5, pv′′ 2–3.</p> <p>Material examined. Holotype, female, ex. Castelnaudia septemcostata. <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=152.32611&amp;materialsCitation.latitude=-28.23611" title="Search Plazi for locations around (long 152.32611/lat -28.23611)">Mt Cougal</a>, shelf, 28° 14ʹ 10ʺ S 152° 19ʹ 34ʺ E, 19 Nov 1987 - 17 Mar 1988, D. Cook &amp; G. Monteith, pitfall trap, rainforest, IN6062, HR# T 64265 (QMS 113019). In QM. Paratypes. All ex. C. septemcostata. 4 females, 6 males, 14 larvae, as follows. 1 female, 2 larvae, same data as holotype (QMS 113027-28, ZMH-A0015191); 1 female, O’Reilly’s Guesthouse, 28° 14ʹ 08ʺ S 152° 08ʹ 12ʺ E, 1-31 Dec 1909, H. Hacker, IN7977, HR# T 88472 (QMS 113020); 1 female, 1 male, 3 larvae, Tweed Lookout, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=152.13666&amp;materialsCitation.latitude=-28.235556" title="Search Plazi for locations around (long 152.13666/lat -28.235556)">Wiangaree</a>, 28° 23ʹ 14ʺ S 153° 6ʹ 33ʺ E, 16 Nov-27 Dec 1974, G. &amp; S. R. Monteith, pitfall trap, rainforest, IN5349, HR# T 57316 (QMS 113024, 113029-30, ANIC 52-003959); 1 female, 5 males, 6 larvae, Rotary Park, Lismore, 28° 48ʹ 37ʺ S 153° 17ʹ 43ʺ E, 16 Nov-26 Dec 1974, G. &amp; S. R. Monteith, pitfall trap, rainforest, IN5415, HR# T 57330 (QMS 113021-23, 25-26, 31-32, ANIC 52-003960 – 61, ZMH-A0015192); 1 larva, same data as holotype except 18 Jan-27 Apr 1987, IN6056, HR# T 64261; 1 larva, Upper Tallebudgera Creek, 28° 14ʹ 10ʺ S 152° 19ʹ 34ʺ E, 19 Nov 1987 - 17 Mar 1988, D. Cook &amp; G. Monteith, pitfall trap, rainforest, IN6062, HR# T 64272 (QMS113034); 1 larva, Bar Mountain, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=152.32611&amp;materialsCitation.latitude=-28.23611" title="Search Plazi for locations around (long 152.32611/lat -28.23611)">Wiangaree</a>, 28° 27ʹ 37ʺ S 153° 8ʹ 2ʺ E, 11 Oct 1977 - 2 Oct 1978, IN5624, G &amp; S. R. Monteith, pitfall trap, rainforest, IN5624, HR# T 57344 (QMS113033). All in QM except 1 female, 1 male, 1 larva in ANIC; 1 male, 1 larva in ZMH.</p> <p>Description. FEMALE (Figs 17–18; n = 5).</p> <p>Gnathosoma. Length 58 (58–60), width 48 (46–50). Cheliceral stylets 68 (68–72), setae ch 22 (24–27), su 13 (10–11).</p> <p>Idiosoma (Fig. 17). Length 335 (325–480), width 240 (255–320). Prodorsal plate eroded anteriorly; setae slender, v1 9 (7–10), v2 alveolar remnant, sc1 16 (13–16), sc2 85 (85–90). Distance between setae v1–v1 46 (40–51), sc1–sc1 79 (78–96), v1–sc1 25 (23–27), v2–v2 48 (44–58), sc2–sc2 105 (110–125), sc1–sc2 25 (23–27). Opisthosomal setae slender. Plate C setae c1 16 (15–20), c2 23 (19–23), distance between setae c1–c1 71 (73–92). Plate D setae d 21 (17–23), d–d 125 (115–125), cupuli ia laterad to anterolaterad d. Plate EF setae f 16 (14–20), f–f 125 (115–125), cupuli im anterolaterad f. Plate H fused weakly, seta h 2 (3–4). Venter: coxisterna 1-2 well defined, coxisterna 3 weakly defined; coxal setae slender, 1a 3 (3–4), 2a 5 (5–7), 3a 6 (4–7), 3b 6 (6–7). Alveoli of 1b, 2b present.</p> <p>Legs (Fig. 18). Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Leg I. Femur I, d minute, l’ 2 (2–3), v″ 8 (10–16); genu I, l′ 2 (2), l″ 2 (2); tibia I, d 70 (60–65), l’ 5 (4–5), l’’ 8 (6–9), v’ 4 (3–4), v’’ 10 (8–10), k 5 (4), φ 9 (8–10); tarsus I, tc’ 14 (15–17), tc’’ 14 (13–17), pl’ 11 (11–14), pl’’ 19 (14–17), pv′ 2 (2), pv’’ 2 (2), ω 3 (3–4), s 6 (5–6), u′′ m (m–2). Leg II. Genu II, l′ 2 (2); tibia II, d 35 (30–40), l’ 5 (4–5), v’ 11 (8–12), v’’ 20 (14–17); tarsus II, tc′ 3 (2–3), tc ″ 5 (5), pl ″ 58 (45–46), pv’’ 3 (3–4), u’ 7 (7), u′′ 2 (m–2). Leg III. Genu III, l′ 2 (2–3); tibia III, d 40 (33–39), l’ 6 (5), v’ 12 (10–12), v’’ 24 (15–17); tarsus III, tc’ 2 (2), tc ″ 5 (5–6), pl ″ 55 (45–50), pv’’ 2 (2–3), u’ 7 (6–7), u′′ m (m–2).</p> <p>LARVIFORM ADULT MALE (Fig. 19; n = 6).</p> <p>Gnathosoma. Length 31–35, width 29–29. Cheliceral stylets 21–25, setae ch 7–9, su 6–7.</p> <p>Idiosoma (Fig. 19). Length 145–180, width 105–130. Prodorsal plate with setae v1 4, v2 alveolar remnant, sc1 4–6, sc2 45–50. Distance between setae v1–v1 18–21, sc1–sc1 41–45, v1–sc1 14–19, v2–v2 19–23, sc2–sc2 51–60, sc1–sc2 18–27. Plate CD setae c1 5–6, c2 5–6, d 5–6, distance between setae c1–c1 45–53, c1–c2 25–33, d–d 37–48; cupuli ia not seen or absent. Plate EF eroded posteromedially; with setae f 3–4, distance f–f 24–30; cupuli im anterolaterad to laterad setae f. Genital capsule length 25–30, width 28–30, setae h minute. Venter: coxal setae 1a 2–3, 2a 3–4, 3a 3–4, 3b 4–5.</p> <p>Legs. Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Leg I. Femur I, d minute, l′ minute, v″ 5–6 (asymmetrical absence on drawn specimen); genu I l′ m–2, l″ m–2; tibia I, d 36–42, l’ 2, l’’ 3–4, v’ 3–4, v’’ 6–7, k 3, φ 7–8; tarsus I, tc’ 12–14, tc’’ 13–14, pl’ 6–7, pl’’ 11–13, pv ′ m–2, pv’’ 2, ω 2, s 4–5, u ′′ m. Leg II. Genu II, l′ m–2; tibia II, d 23–29, l’ 4, v’ 7–10, v’’ 14–16; tarsus II, tc′ m–2, tc ″ 4–5, pl ″ 24–34, pv’’ 3, u’ 5–6, u″ minute. Leg III. Genu III, l′ 2; tibia III, d 19–24, l’ 4–5, v’ 7–12, v’’ 10–15; tarsus III, tc′ minute, tc ″ 4–5, pl ″ 23–29, pv’’ 2, u’ 5–6, u″ minute.</p> <p>LARVAL FEMALE (Fig. 20; n = 14).</p> <p>Gnathosoma. Length 45–50, width 38–41. Cheliceral stylets 57–63, setae ch 22–33, su 5–9.</p> <p>Idiosoma (Fig. 20). Length 195–230, width 140–190. Prodorsal plate with setae v1 5–6, v2 alveolar remnant, sc1 6–7, sc2 80–95. Distance between setae v1–v1 30–33, sc1–sc1 57–62, v1–sc1 19–22, v2–v2 31–36, sc2–sc2 68–79, sc1–sc2 27–33. Plate C setae c1 7–11, c2 7–10, distance between setae c1–c1 44–54. Plate D setae d 9–12, d–d 55–67, cupuli ia very close to setae d, anteriad to anterolaterad d. Plate EF setae f 7–13, f–f 44–51, cupuli im close to setae f, anteriad to anterolaterad f. Plate H 20–22 wide, 23–26 long, bearing setae h1 ca. 120, h2 7–9. Venter: coxisterna 1-2 well defined, coxisterna 3 with some folds; coxal setae slender, 1a 2–3, 2a 3–4, 3a 4–5, 3b 4–5. Alveoli of 1b, 2b present.</p> <p>Legs. Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Leg I. Femur I, d minute, l′ minute, v″ 6–9; genu I, l′ minute, l″ m–2; tibia I, d 45–61, l’ 6–9, l’’ 8–10, v’ 3–4, v’’ 5–8, k 4–5, φ 8–10; tarsus I, tc’ 13–15, tc’’ 14–15, pl’ 6–7, pl’’ 14–20, pv′ m–2, pv’’ 2, ω 2–3, s 5, u′′ minute. Leg II. Genu II, l′ m–2; tibia II, d 29–39, l’ 5–6, v’ 8–12, v’’ 17–22; tarsus II, tc′ 2, tc ″ 4, pl ″ 31–38, pv’’ 2–3, u’ 5–6, u′′ m–2. Leg III. Genu III, l′ m–2; tibia III, d 25–36, l’ 4–5, v’ 8–11, v’’ 15–18; tarsus III, tc’ m–2, tc ″ 4–5, pl ″ 27–37, pv’’ 2–3, u’ 5–6, u′′ m–2.</p> <p>Differential diagnosis. Eutarsopolipus raveni sp. nov. shares the same host as E. nahmani but is a much smaller species and is instead closer to E. teuceri. See the differential diagnosis of E. teuceri for differences.</p> <p>Etymology. It is with great pleasure that I name this species for Robert Raven, Curator of Arachnida at the Queensland Museum, and a friend and superb arachnologist, who has given great support to the development of acarology at the Queensland Museum.</p> </div>	http://treatment.plazi.org/id/4438245CFFE8096E87AEF91EFDE1A124	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Seeman, Owen D.	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFD5096B87AEFA41FEE8A1D5.text	4438245CFFD5096B87AEFA41FEE8A1D5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eutarsopolipus teuceri Seeman 2021	<div><p>Eutarsopolipus teuceri sp. nov.</p> <p>(Figs 21–23)</p> <p>Diagnosis. Adult female: Gnathosomal length 58–61, width 51–54; anterior margin of prodorsal shield slightly eroded medially; v1 8–10, sc1 18–20, c1 17–22, c2 15–16, d 18–19, f 15–18, h 7–10, v1-v1 42–48, d-d 130–140, f-f 125–130; genua II-III setae l′ 2–4, tibia I seta l′ 6–7, tarsi II-III setae tc′ 3–4, tc′′ 5–6, pv′′ large, 4. Larval female: Unknown.</p> <p>Material examined. Holotype female, ex Castelnaudia porphyriaca. 1.5 km from Mt Cabinet, via Jimna, Sunday Ck, 26° 42ʹ 39ʺ S 152° 33ʹ 49ʺ, 9 Sep-30 Nov 1974, G. &amp; S. Monteith, pitfall trap, IN5370, HR# T 57070 (QMS 113506). Paratypes. 2 females, 3 males, ex C. porphyriaca, Conondale Range, 27 Nov. 1974, G. Monteith, unregistered specimen (QMS 113507, 113510-12, ANIC 52-003962). All in QM except one female in ANIC.</p> <p>Description. FEMALE (Figs 21–22; n = 3).</p> <p>Gnathosoma. Length 58 (60–61), width 54 (51–54). Cheliceral stylets 77 (75–82), setae ch 26 (20–27), su 8 (10).</p> <p>Idiosoma (Fig. 21). Length 360 (310–325), width 250 (225–250). Prodorsal plate not eroded anteriorly; setae slender, v1 10 (8–9), v2 alveolar remnant, sc1 20 (18–20), sc2 broken (85–105). Distance between setae v1–v1 48 (42–44), sc1–sc1 92 (80–86), v1–sc1 31 (26–30), v2–v2 48 (41–43), sc2–sc2 105 (100–105), sc1–sc2 28 (23–29). Opisthosomal setae slender. Plate C setae c1 17 (17–22), c2 17 (15–16), distance between setae c1–c1 80 (70–75). Plate D setae d 19 (18–19), d–d 140 (130–135), cupuli ia anterolaterad d. Plate EF setae f 18 (15–16), f–f 130 (125), cupuli im anterolaterad f. Plate H divided or fused weakly, seta h 7 (7–10). Venter: coxisterna 1-3 moderately well defined, with few folds, coxisterna 3 weaker than 1-2; coxal setae slender, 1a 4 (3–4), 2a 5 (5–6), 3a 4 (5), 3b 7 (5–6). Alveoli of 1b, 2b present.</p> <p>Legs (Fig. 22). Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Leg I. Femur I, d minute, l’ m (2), v″ 8 (7-9); genu I, l′ 2 (2–3), l″ 2 (2–3); tibia I, d broken (60–68), l’ 7 (5–6), l’’ 6 (8), v’ 4 (4–5), v’’ 9 (8–10), k 4 (3–4), φ 9 (8); tarsus I, tc’ 15 (13–14), tc’’ 14 (15), pl’ 8 (11–12), pl’’ broken (9–15), pv′ 3 (3–4), pv’’ 3 (4), ω 2 (3), s 6 (6–7), u′′ m (m). Leg II. Genu II, l′ 2 (3–4); tibia II, d 40 (35–43), l’ 6 (7), v’ 10 (10–14), v’’ 17 (18–20); tarsus II, tc′ 3 (3–4), tc ″ 5 (5–6), pl ″ 50 (41–43), pv’’ 4 (4), u’ 6 (7), u′′ m (m). Leg III. Genu III, l′ 2 (3); tibia III, d obscured (32–35), l’ 6 (5–6), v’ obscured (11–13), v’’ obscured (14–15); tarsus III, tc’ 4 (3), tc ″ 5 (5), pl ″ broken (38–42), pv’’ 4 (4), u’ 7 (6–7), u′′ m (m).</p> <p>LARVIFORM ADULT MALE (Fig. 23; n = 3).</p> <p>Gnathosoma. Length 32–34, width 21–25. Cheliceral stylets 21–25, setae ch 5–6, su 5–6.</p> <p>Idiosoma. Length 155–165, width 110–125. Prodorsal plate with setae v1 3–4, v2 alveolar remnant, sc1 5–6, sc2 62–70. Distance between setae v1–v1 16–19, sc1–sc1 41–42, v1–sc1 15–19, v2–v2 18–21, sc2–sc2 49–50, sc1–sc2 21–25. Plate CD setae c1 6–7, c2 5–6, d 6–7, distance between setae c1–c1 42–45, c1–c2 24–29, d–d 35–37; cupuli ia anterolaterad setae d. Plate EF slightly eroded medially; with setae f 2–4 (thickened in one specimen), distance f–f 22–24; cupuli im anterolaterad setae f. Genital capsule length 26–29, width 30–32, setae h minute. Venter: coxal setae 1a 2–3, 2a 3–4, 3a 3–4, 3b 4–5.</p> <p>Legs. Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Leg I. Femur I, d minute, l′ minute, v″ 3–4; genu I l′ 2, l″ m–2; tibia I, d 32–37, l’ 3–4, l’’ 3–4, v ’ 2–3, v ’’ 6–10, k 2–3, φ 6–7; tarsus I, tc’ 13, tc’’ 13–14, pl’ 6–7, pl’’ 9–12, pv ′ m, pv’’ 2, ω 2, s 4–5, u ′′ m. Leg II. Genu II, l′ 2; tibia II, d 16–17, l’ 5–6, v’ 9–12, v’’ 10–15; tarsus II, tc′ 2, tc ″ 4, pl ″ 27–33, pv″ 2–3, u’ 4–5, u″ minute. Leg III. Genu III, l′ 2–3; tibia III, d 12–16, l’ 4, v’ 8–10, v″ 12–15; tarsus III, tc′ 2, tc ″ 4–5, pl ″ 26–32, pv″ 3, u ’ 5–6, u″ minute.</p> <p>LARVAL FEMALE. Unknown.</p> <p>Differential diagnosis. Eutarsopolipus teuceri sp. nov. is close to E. raveni. Females are most easily separated by the length of seta h, which is short (2–4) in E. raveni but longer (7–10) in E. teuceri. Other small differences are the size of setae tc′ (3–4 versus 2) and pv′ (4 versus 2–3) on tarsus III, which are larger in E. teuceri, and the longer cheliceral stylets (75–82 versus 68–72). Males are almost identical, differing only by the length of setae sc2 (45–50 vs 62–70 in E. raveni), although the smaller tc′ is also apparent (2 versus minute in E. raveni).</p> <p>Eutarsopolipus raveni and E. teuceri both share a small gnathosoma with E. rutherfordae, which is also a member of the nahmani subgroup but its host is Trichosternus subvirens. These two species are most easily distinguished from E. rutherfordae by the strong, stiff setae d and f in E. rutherfordae (very fine, not stiff in E. raveni and E. teuceri). Eutarsopolipus raveni also has small setae h (2–4 compared to 7–10 in E. rutherfordae and E. teuceri), while E. teuceri has longer cheliceral stylets (75–82 versus 60), shorter cheliceral setae (20–27 versus 42), shorter seta v′′ on femur II (7–9 versus 16) and much larger setae pv′′ on tarsi II-III (4 versus 2).</p> <p>Etymology. The specific name teuceri refers to the legendry archer Teucer of the Odyssey, who in battle hid behind the glittering shield of Ajax, alluding to the spectacular metallic purple sheen of the elytra of the host beetle E. porphyriaca.</p> <p>Remarks. Castelnaudia porphyriaca hosted three species described herein, but also had one larva of another species collected at Conondale. The larva belongs to the pterostichi species group and has an enlarged tibia I seta v′, the only femoral seta is d, and the only setae on all genua is on leg I (seta l′).</p> <p>The ochoai species group - despoticus subgroup</p> <p>Diagnosis. Female with dorsal idiosomal setae fine, slender; setae h minute. All life stages with gnathosomal apex evenly rounded.</p></div> 	http://treatment.plazi.org/id/4438245CFFD5096B87AEFA41FEE8A1D5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Seeman, Owen D.	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFD0096987AEF9D1FD11A04D.text	4438245CFFD0096987AEF9D1FD11A04D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eutarsopolipus despoticus Seeman 2021	<div><p>Eutarsopolipus despoticus sp. nov.</p> <p>(Figs 24–27)</p> <p>Diagnosis. Adult female: Gnathosomal length 45–51, width 47–51, cheliceral stylets 50–55; tarsi II-III setae tc′′ 2–3. Larval female: Gnathosomal length 42–45, width 43–45, cheliceral stylets 45–52; tarsi II-III setae tc′′ m–2.</p> <p>Material examined. Holotype female, ex Castelnaudia porphyriaca. Conondale Range, 27 Nov. 1974, G. Monteith, unregistered specimen (QMS 113508). Paratypes. All ex C. porphyriaca. 6 females, 2 larvae, as follows: 3 females, 2 larvae, same data as holotype (QMS 113509–10, 13–4, ANIC 52-003963); 3 females, 1.5 km from Mt Cabinet, via <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=152.56361&amp;materialsCitation.latitude=-26.710835" title="Search Plazi for locations around (long 152.56361/lat -26.710835)">Jimna</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=152.56361&amp;materialsCitation.latitude=-26.710835" title="Search Plazi for locations around (long 152.56361/lat -26.710835)">Sunday Ck</a>, 26° 42ʹ 39ʺ S 152° 33ʹ 49ʺE, 9 Sep-30 Nov 1974, G. &amp; S. Monteith, pitfall trap, HR# T 57070 (QMS 113515–16, ZMH-A0015193). All in QM except 1 female in ANIC and ZMH.</p> <p>Note: two females from Mt Cabinet have fully developed larvae in their body; measurements from these larvae are included where features could be measured.</p> <p>Description. FEMALE (Figs 24–26; n = 4).</p> <p>Gnathosoma (Fig. 25A). Length 46 (45–51), width 50 (47–51). Cheliceral stylets 53 (50–55), setae ch 27 (28–34), su 12 (9–16).</p> <p>Idiosoma (Fig. 24). Length 285 (245–325), width 255 (230–300). Prodorsal plate not eroded anteriorly; setae slender, v1 11 (9–12), v2 alveolar remnant, sc1 10 (9–10), sc2 85 (65–80). Distance between setae v1–v1 60 (55–59), sc1–sc1 98 (87–97), v1–sc1 35 (32–37; asymmetrically 27 in one specimen), v2–v2 67 (61–68), sc2–sc2 125 (125– 130), sc1–sc2 32 (31–35). Opisthosomal setae slender. Plate C setae c1 9 (7–9), c2 15 (12–14), distance between setae c1–c1 100 (105–120). Plate D setae d 11 (9–11), d–d 130 (125–140), cupuli ia close to and anterolaterad d. Plate EF setae f 12 (10–11), f–f 74 (70–77), cupuli im distant from and anterolaterad f. Plate H entire, seta h minute. Venter: coxisterna 1-2 moderately well defined, coxisterna 3 somewhat less defined; coxal setae slender, 1a 3 (2–3), 2a 5 (4–5), 3a 9 (7–9), 3b 6 (5–6). Alveoli of 1b, 2b present.</p> <p>Legs (Figs 26 A-C). Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Leg I. Femur I, d minute, l’ 2 (2–4), v″ 6 (5–7); genu I, l′ 2 (2–3), l″ 3 (2–3); tibia I, d 50 (50–56), lʹ 11 (8–10), lʺ 12 (11–12), v’ 6 (6–7), v’’ 18 (13–20), k 5 (4), φ 10 (9-10); tarsus I, tc’ 18 (16–18), tcʺ 17 (16–17), pl’ 15 (12–15), plʺ 19 (16–17), pv′ minute, pvʺ 2 (2), ω 4 (4–5), s 5 (5–6), u′′ m (m). Leg II. Genu II, l′ 4 (3–4); tibia II, d 35 (31–37), l’ 9 (8–10), v’ 20 (16–25), vʺ 26 (22–29); tarsus II, tc′ 8 (5–9), tc″ 3 (2–3), pl″ 50 (43–52), pvʺ 2 (2), u’ 5 (5–6), u′′ m (m). Leg III. Genu III, l′ 3 (2–3); tibia III, d 30 (28–37), l’ 7 (7–9), v’ 23 (18–25), vʺ 26 (18–28); tarsus III, tc’ 4 (4–5), tc″ 3 (2–3), pl″ 50 (36–50), pvʺ 2 (2), u’ 5 (5–6), u′′ m (m).</p> <p>LARVIFORM MALE. Unknown.</p> <p>LARVAL FEMALE (Fig. 27; n = 4).</p> <p>Gnathosoma. Length 42–45, width 43–45. Cheliceral stylets 45–52, setae ch 24–27, su 9–10.</p> <p>Idiosoma (Fig. 27). Length 195–205, width 145–165. Prodorsal plate with setae v1 7–9, v2 alveolar remnant, sc1 6–8, sc2 90–100. Distance between setae v1–v1 34–36, sc1–sc1 65–70, v1–sc1 26–27, v2–v2 45–48, sc2–sc2 74–81, sc1–sc2 29–35. Plate C setae c1 4–8, c2 10, distance between setae c1–c1 62–71. Plate D setae d 8–10, d–d 58–61, cupuli ia distant from setae d, anterolaterad d. Plate EF setae f 9–10, f–f 42–45, cupuli im distant to setae f, anteriad f (not seen in drawn specimen). Plate H broad, 25–31 wide, ca. 34–35 long bearing setae h1 ca. 180, h2 6–7. Venter: coxisterna 1-2 well defined, coxisterna 3 with some membranous folds; coxal setae slender, 1a 3, 2a 4–5, 3a 6–9, 3b 5–6. Alveoli of 1b, 2b present.</p> <p>Legs. Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Leg I. Femur I, d minute, l′ 2, v″ 3–4; genu I, l′ 2, l″ 2; tibia I, d 60–65, l’ 11–12, l’’ 10–12, v’ 4–5, v’’ 14–17, k 3–4, φ 10; tarsus I, tc’ 16–18, tc’’ 18, pl’ 14–17, pl’’ 19–25, pv′ minute, pv’’ m–2, ω 4–5, s 5, u′′ minute. Leg II. Genu II, l′ 2–3; tibia II, d 40–45, l’ 8–9, v’ 17–18, v’’ 22; tarsus II, tc′ 3–4, tc ″ m–2, pl ″ 32–38, pv’’ m–2, u’ 4–5, u′′ m–2. Leg III. Genu III, l′ 2; tibia III, d 35–39, l’ 6–7, v’ 20–22, v’’ 23–29; tarsus III, tc’ 3–5, tc ″ m–2, pl ″ 35–43, pv’’ m–2, u’ 4–5, u′′ minute.</p> <p>Differential diagnosis. Eutarsopolipus despoticus sp. nov. is most similar to E. hebronae sp. nov., but differs by having a smaller gnathosoma, cheliceral stylets and setae tc″ on tarsi II-III. See below the Differential diagnosis of E. hebronae sp. nov. for further details.</p> <p>Etymology. The host’s specific name porphyriaca refers to its purple wings, and as the colour purple was associated with rulers, the name despoticus (a despot, a tyrant) alludes to this species living beneath the purple wings of its host while engaged in a parasitic life style.</p> </div>	http://treatment.plazi.org/id/4438245CFFD0096987AEF9D1FD11A04D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Seeman, Owen D.	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFD2096787AEFB69FC50A2D8.text	4438245CFFD2096787AEFB69FC50A2D8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eutarsopolipus hebronae Seeman 2021	<div><p>Eutarsopolipus hebronae sp. nov.</p> <p>(Figs 25–26)</p> <p>Diagnosis. Adult female: Gnathosomal length 61–65, width 60–67, cheliceral stylets 75–84; tarsi II-III setae tc′′ 3–4. Larval female: Gnathosomal length 50–55, width 50–53, cheliceral stylets 64–66; tarsi II-III setae tc′′ 2–3.</p> <p>Material examined. Holotype female, ex Castelnaudia cordata. <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=152.68028&amp;materialsCitation.latitude=-27.048056" title="Search Plazi for locations around (long 152.68028/lat -27.048056)">Mt Mee</a> SF, 6 km NNW barracks, 27° 2ʹ 53ʺ S 152° 40ʹ 49ʺ E, 8 Jan-3 Mar 1992, D. Cook, flight intercept trap, IN5148, HR# T 68607 (QMS 113463). Paratypes. All ex C. cordata. 6 females, 3 larvae, as follows: all ex. C. cordata. 3 females, 1 larva, same data as holotype (QMS113464–66, ANIC 52-003964); 1 larva, McIntosh Rd, Unumbar SF, 28° 25ʹ 19ʺ S 152° 42ʹ 14ʺ E, 25 Nov 1972, G. Monteith, IN6467, HR# T 64329 (ANIC 52-003965); 1 female, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=152.84805&amp;materialsCitation.latitude=-28.315832" title="Search Plazi for locations around (long 152.84805/lat -28.315832)">Lower Coomera</a>, 28° 12ʹ 20ʺ S 153° 10ʹ 46ʺ E, 9 Jan-6 Apr 1995, G. Monteith, pitfall trap, IN5998, HR# T 68610 (QMS 113467); 2 females, 1 larva, Lever’s Plateau, 28° 18ʹ 57ʺ S 152° 50ʹ 53ʺ E, 22 Feb-8 May 1976, G. &amp; S. R. Monteith, pitfall trap, rainforest, IN5485, HR# T 68619 (QMS 113468–69, ZMH-A0015194). 1 female, 1 larva in ANIC; 1 female in ZMH.</p> <p>Other material examined. 4 females from type locality, mounted with specimens of E. umbonatus. Not included as paratypes.</p> <p>Notes: 1 female from each of Mt Mee and Lever’s Plateau are E. hebronae but too poor to be considered paratypes. There is also a pharate female from McIntosh Rd.</p> <p>Description. FEMALE (Figs 25–26; n = 6).</p> <p>Gnathosoma (Fig. 25B). Length 61 (61–65), width 60 (60–67). Cheliceral stylets 78 (75–84), setae ch 35 (29–38), su 17 (13–15).</p> <p>Idiosoma. Length 420 (345–430), width 290 (255–340). Prodorsal plate eroded mid-anteriorly; setae slender; setae v1 6 (5–7), v2 alveolar remnant, sc1 10 (6–10), sc2 95 (85–105). Distance between setae v1–v1 56 (54–66), sc1–sc1 105 (93–105), v1–sc1 33 (21–30), v2–v2 64 (67–69), sc2–sc2 140 (135–150), sc1–sc2 40 (33–43). Opisthosomal setae slender. Plate C setae c1 9 (7–9), c2 12 (11–13), distance between setae c1–c1 82 (69–85). Plate D setae d 10 (7–10), d–d 145 (130–150), cupuli ia well apart from d, anterolaterad d. Plate EF setae f 10 (9–13), f–f 86 (79–95), cupuli im well apart from f, anterolaterad f. Plate H divided or fused weakly, seta h minute. Venter: coxisterna 1 well defined, coxisterna 2 with some folds, coxisterna 3 weakly defined; coxal setae slender, 1a 3 (2–3), 2a 5 (4–5), 3a 7 (6–7), 3b 5 (5). Alveoli of 1b, 2b present.</p> <p>Legs (Figs 26 D-E). Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Leg I. Femur I, d minute, l’ 3 (3), vʺ 8 (6–7); genu I, l′ 3 (3), l″ 3 (3–4); tibia I, d 50 (50–65), l’ 11 (11–13), lʺ 3 (3–4), v’ 8 (5–7), vʺ 12 (10–15), k 5 (5–7), φ 10 (10–12); tarsus I, tc’ 22 (19–20), tcʺ 18 (16–20), pl’ 15 (12–16), pl’’ 23 (20–23), pv′ 2 (2), pvʺ 2 (2), ω 5 (4–5), s 5 (5–6), u′′ 2 (m–2). Leg II. Genu II, l′ 4 (4–6); tibia II, d 45 (45–50), l’ 11 (9–12), v’ 19 (17–19), vʺ 21 (22–29); tarsus II, tc′ 4 (3–4), tc″ 4 (3–4), pl″ 52 (38–50), pvʺ 2 (2), u’ 6 (6–7), u′′ 2 (m–2). Leg III. Genu III, l′ 5 (4–5); tibia III, d 40 (35–50), l’ 8 (8–10), v’ 15 (14–18), vʺ 30 (26–30); tarsus III, tc’ 3 (2–3), tc″ 4 (3–4), pl″ 40 (40–50), pvʺ 2 (2), u’ 6 (6), u′′ m (m).</p> <p>LARVIFORM MALE. Unknown.</p> <p>LARVAL FEMALE (n = 3).</p> <p>Gnathosoma. Length 50–55, width 50–53. Cheliceral stylets 64–66, setae ch 35–36, su 8–10.</p> <p>Idiosoma. Length 210–270, width 165–210. Prodorsal plate with setae v1 6–7, v2 alveolar remnant, sc1 7–9, sc2 90–120. Distance between setae v1–v1 33–40, sc1–sc1 74–86, v1–sc1 29–34, v2–v2 51–60, sc2–sc2 80–90, sc1–sc2 25–30. Plate C setae c1 7–9, c2 10–13, distance between setae c1–c1 50–59. Plate D setae d 8–10, d–d 61–74, cupuli ia anterolaterad d. Plate EF setae f 9–11, f–f 43–52, cupuli im anteriad to anterolaterad f. Plate H long and narrow, 16–20 wide, ca. 30–40 long bearing setae h1 ca. 150, h2 8–10. Venter: coxisterna 1-2 moderately well defined, with few folds, coxisterna 3 poorly defined, membranous; coxal setae slender, 1a 3, 2a 5, 3a 5, 3b 5–6. Alveoli of 1b, 2b present.</p> <p>Legs. Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Leg I. Femur I, d minute, l′ minute, v″ 5–7; genu I, l′ 3, l″ 2–3; tibia I, d 75–80, l’ 3, l’’ 2–3, v’ 6–8, v’’ 9–12, k 3–5, φ 11–12; tarsus I, tc’ 15–16, tc’’ 16, pl’ 9–13, pl’’ 18–20, pv′ m–2, pv’’ 2, ω 3–4, s 5–6, u′′ minute. Leg II. Genu II, l′ 3–4; tibia II, d 45–50, l’ 9–10, v’ 13–18, v’’ 23–26; tarsus II, tc′ 2–3, tc ″ 2–3, pl ″ 45–50, pv’’ 2, u’ 5–6, u′′ minute. Leg III. Genu III, l′ 3–4; tibia III, d 40–42, l’ 9–13, v’ 12–15, v’’ 20–26; tarsus III, tc’ 2, tc ″ 2–3, pl ″ 40–50, pv’’ 2, u’ 5–6, u′′ m–2.</p> <p>Differential diagnosis. Eutarsopolipus hebronae sp. nov. differs from E. despoticus sp. nov. by having a much larger gnathosoma (61–65 × 60–67 versus 45–51 × 47–51) and cheliceral stylets (75–84 versus 50–55). The species also differ in the size of setae tc″ on tarsi II-III, being much larger in E. hebronae (3–4) than E. despoticus (2–3). Even though these setae are only about 1 μm longer, they are obviously larger, being around twice as massive in E. hebronae. These differences in gnathosomal size and tarsal setae are present in the adult and larval female.</p> <p>Etymology. It is with great pleasure that I name this species for Wendy Hebron, with whom I have shared the role of collection manager for 14 years. Not only has her welcome and proficient assistance in the Queensland Museum arachnology collection been greatly appreciated, but her fine efforts have allowed me to dedicate more time to acarology.</p> <p>Remarks. This species was found co-infesting its host species with E. umbonatus sp. nov. and an undescribed species of Eutarsopolipus close to E. echinatus Seeman, 2019a (Tables 1, 2).</p> <p>The ochoai species group - basiatus subgroup</p> <p>Diagnosis. Female with dorsal idiosomal setae fine, slender; setae h minute or alveolar vestiges. All life stages with gnathosomal apex with small paired lobes.</p></div> 	http://treatment.plazi.org/id/4438245CFFD2096787AEFB69FC50A2D8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Seeman, Owen D.	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFDC096787AEF8DDFDD1A3B0.text	4438245CFFDC096787AEF8DDFDD1A3B0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eutarsopolipus basiatus Seeman 2021	<div><p>The ochoai species group - basiatus subgroup</p> <p>Diagnosis. Female with dorsal idiosomal setae fine, slender; setae h minute or alveolar vestiges. All life stages with gnathosomal apex with small paired lobes.</p></div> 	http://treatment.plazi.org/id/4438245CFFDC096787AEF8DDFDD1A3B0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Seeman, Owen D.	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFDD096087AEFF5AFE69A749.text	4438245CFFDD096087AEFF5AFE69A749.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eutarsopolipus basiatus Seeman 2021	<div><p>Eutarsopolipus basiatus sp. nov.</p> <p>(Figs 28–31)</p> <p>Diagnosis. Adult female: sc1 5–7, c1 6–9, c2 9–14, d 7–9, f 6–7, v2-v2 48–62, sc1-sc1 36-46, d-d 91–115, f-f 76–92; genua II-III setae l′ 3–4, tibia I setae l′ 4–6, v′ 5–6, tarsi II-III setae tc′′ 2–3. Adult male: leg I with two claws; c1 3–4, c2 4–5, v2-v2 26–33; tarsi II-III setae tc″ m–2. Larval female: c1 6–8, c2 8–10, d 6 – 10, f 7–9, sc2–sc2 64–71, v1-sc1 27–32; tarsi II-III setae tc′′ minute.</p> <p>Material examined. Holotype female, ex Castelnaudia porphyriaca. Conondale Range, 27 Nov. 1974, G. Monteith, unregistered specimen. Paratypes. All ex C. porphyriaca. 4 females, 4 males, 5 larvae, as follows: 1 female, 2 males, 2 larvae, same data as holotype (QMS 113518–20, ANIC 52-003966 – 67); 3 females, 2 males, 2 larvae, 1.5 km from Mt Cabinet, via <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=152.56361&amp;materialsCitation.latitude=-26.710835" title="Search Plazi for locations around (long 152.56361/lat -26.710835)">Jimna</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=152.56361&amp;materialsCitation.latitude=-26.710835" title="Search Plazi for locations around (long 152.56361/lat -26.710835)">Sunday Ck</a>, 26° 42ʹ 39ʺ S 152° 33ʹ 49ʺE, 9 Sep-30 Nov 1974, G. &amp; S. Monteith, pitfall trap, IN5370, HR# T 57070 (QMS 113521–25, ANIC 52-003968, ZMH-A0015195); 1 female, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=152.6189&amp;materialsCitation.latitude=-26.712778" title="Search Plazi for locations around (long 152.6189/lat -26.712778)">Conondale Range</a>, site 1, 26° 42ʹ 46ʺ S 152° 37ʹ 08ʺE, 20 Apr–2 May 1995, B. Nomann &amp; G. Thompson, pitfall trap, IN7291, HR# T 68573 (QMS 113526). All in QM except 1 female, 1 male, 1 larva in ANIC and 1 female in ZMH.</p> <p>Note: 1 female from Conondale has a fully developed larva in its body; measurements from this larva are included where features could be measured.</p> <p>Description. FEMALE (Figs 28–29; n = 5).</p> <p>Gnathosoma (Fig. 29D). Length 45 (45–46), width 44 (42–45), apex indented, terminating in small but distinct lobes. Cheliceral stylets 40 (38–44), setae ch 20 (16–24), su 9 (8–12).</p> <p>Idiosoma (Fig. 28). Length 315 (275–325) width 405 (295–470). Prodorsal plate not eroded anteriorly; setae slender, v1 6 (5–7), v2 alveolar remnant, sc1 6 (5–7), sc2 76 (69–82). Distance between setae v1–v1 41 (34–38), sc1–sc1 100 (82–94), v1–sc1 42 (36–37; asymmetry in one specimen: 36/46), v2–v2 62 (48–58), sc2–sc2 110 (90–105), sc1–sc2 26 (27–32). Opisthosomal setae slender. Plate C setae c1 9 (6–8), c2 13 (9–14), distance between setae c1–c1 83 (69–77). Plate D setae d 9 (7–9), d–d 115 (91–110), cupuli ia anterolaterad d. Plate EF setae f 7 (6–7), f–f 85 (76–92), cupuli im anteriad to anterolaterad f. Plate H divided, seta h minute. Venter: coxisterna 1-2 moderately well defined, with few folds, coxisterna 3 membranous; coxal setae slender, 1a 3 (2–3), 2a 4 (4–5), 3a 5 (4–6), 3b 6 (5–6). Alveoli of 1b, 2b present.</p> <p>Legs (Figs 29 A-C). Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Leg I. Femur I, d minute, l’ m (m–2), v″ 4 (4–5); genu I, l′ 3 (2–3), l″ 2 (2–3); tibia I, d 60 (45–55), l’ 4 (5–6), l’’ 7 (7–8), v’ 6 (5–6), v’’ 18 (14–17), k 5 (5–6), φ 10 (9–10); tarsus I, tc’ 17 (14–17), tc’’ 16 (15–16), pl’ 10 (8–9), pl’’ 20 (15–21), pv′ m (m), pv’’ 2 (2), ω 5 (4–5), s 5 (5), u′′ m (m). Leg II. Genu II, l′ 4 (3–4); tibia II, d 35 (30–42), l’ 5 (6–8), v’ 21 (19–21), v’’ 28 (21–28); tarsus II, tc′ 2 (2–4), tc ″ 2 (2), pl ″ 45 (37–46), pv’’ 2 (2), u’ 5 (5–6), u′′ m (2). Leg III. Genu III, l′ 4 (3–4); tibia III, d 35 (32–43), l’ 7 (5–8), v’ 19 (15–21), v’’ 24 (19–25); tarsus III, tc’ 3 (2–4), tc ″ 2 (2–3), pl ″ 35 (30–37), pv’’ 2 (2), u’ 5 (4–5), u′′ m (m).</p> <p>LARVIFORM ADULT MALE (Fig. 30; n = 4).</p> <p>Gnathosoma. Length 28–30, width 29–31, apex indented, terminating in small but distinct lobes. Cheliceral stylets 17–22, setae ch 5–7, su 4–7.</p> <p>Idiosoma (Fig. 30). Length 140–175, width 105–120. Prodorsal plate with setae v1 4–5, v2 alveolar remnant, sc1 3–5, sc2 55–65. Distance between setae v1–v1 19–24, sc1–sc1 42–46, v1–sc1 23–26, v2–v2 26–32, sc2–sc2 53–58, sc1–sc2 16–17 (one asymmetrical specimen 14/19). Plate CD setae c1 3–4, c2 4–5, d 4–5, distance between setae c1–c1 43–51, c1–c2 23–26, d–d 37–42; cupuli ia laterad setae d. Plate EF slightly eroded medially; with setae f 4–5, distance f–f 27–28; cupuli im anteriad setae f. Genital capsule length 24–27, width 37–41, setae h m–2. Venter: coxal setae 1a 3–4, 2a 3–4, 3a 3–5, 3b 4–6.</p> <p>Legs. Leg I with two claws. Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Leg I. Femur I, d minute, v″ 3–4; genu I l′ 2–3, l″ 5–6; tibia I, d 29–45, l’ 5, l’’ 2, v ’ 3–4, v ’’ 8–13, k 3–4, φ 8–9; tarsus I, tc’ 14–15, tc’’ 13–15, pl’ 7–10, pl’’ 11–15, pv ′ minute, pv’’ m–2, ω 4–5, s 4, u ′′ minute. Leg II. Genu II, l′ 2–3; tibia II, d 29–38, l’ 4–6, v’ 10–14, v’’ 15–19; tarsus II, tc′ 2–3, tc ″ m–2, pl ″ 26–32, pv’’ minute, u’ 4–5, u″ minute. Leg III. Genu III, l′ 2; tibia III, d 18–26, l’ 3–5, v’ 10–14, v’’ 15–22; tarsus III, tc′ 2–3, tc ″ m–2, pl ″ 24–30, pv’’ minute, u’ 4, u″ minute.</p> <p>LARVAL FEMALE (Fig. 31; n = 6).</p> <p>Gnathosoma. Length 36–41, width 37–39, apex indented, terminating in small but distinct lobes. Cheliceral stylets 34–37, setae ch 34–37, su 7–10.</p> <p>Idiosoma (Fig. 31). Length 220–245, width 150–175. Prodorsal plate with setae v1 4–8, v2 alveolar remnant, sc1 3–6, sc2 70–85. Distance between setae v1–v1 29–32, sc1–sc1 63–73, v1–sc1 31–32 (asymmetrical in one specimen, 33/27), v2–v2 42–45, sc2–sc2 64–71, sc1–sc2 25–26 (asymmetrical in one specimen, 27/23). Plate C setae c1 6–8, c2 8–10, distance between setae c1–c1 51–57. Plate D setae d 6–10, d–d 48–54, cupuli ia anterolaterad setae d. Plate EF setae f 7–9 (one specimen expressing two f setae on left, possibly in place of cupule im), f–f 40–51, cupuli im anteriad f. Plate H narrow, large, 23–29 wide, 35–38 long, bearing setae h1 ca. 170, h2 6–8. Venter: coxisterna 1-2 moderately well defined, with some folds, coxisterna 3 membranous; coxal setae slender, 1a 3, 2a 3–5, 3a 5–7, 3b 4–5. Alveoli of 1b, 2b present.</p> <p>Legs. Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Leg I. Femur I, d minute, l′ minute, v″ 3–4; genu I, l′ 2, l″ 2; tibia I, d 50–55, l’ 5–8, l’’ 8–10, v’ 5–6, v’’ 11–14, k 4–5, φ 10–11; tarsus I, tc’ 15–16, tc’’ 14–15, pl’ 10–13, pl’’ 20–23, pv′ minute, pv’’ minute, ω 5, s 4–5, u′′ minute. Leg II. Genu II, l′ 2–3; tibia II, d 32–40, l’ 6–7, v’ 15–18, v’’ 17–27; tarsus II, tc′ m–2, tc ″ minute, pl ″ 26–38, pv’’ m–2, u’ 4–5, u′′ minute. Leg III. Genu III, l′ 2; tibia III, d 31–40, l’ 5–7, v’ 16–17, v’’ 21–23; tarsus III, tc’ 2, tc ″ minute, pl ″ 31–38, pv’’ m–2, u’ 4–5, u′′ minute.</p> <p>Differential diagnosis. The most obvious difference between E. basiatus sp. nov. and all other species on Castelnaudia is the retention of two claws on leg I of the male. This species is also distinctive within the basiatus subgroup by having: tiny setae tc″ on tarsi II-III (minute-2 versus distinctly larger (3–5) in all other species, being at least twice as massive); and shorter idiosomal setae (e.g., females with short d (7–9) versus 12–23 in the other species).</p> <p>Etymology. The specific name basiatus (a kisser) refers to the distinctive lip-like protrusion of the gnathosoma found in this species group.</p> </div>	http://treatment.plazi.org/id/4438245CFFDD096087AEFF5AFE69A749	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Seeman, Owen D.	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFDB097B87AEFC6DFEB7A3EE.text	4438245CFFDB097B87AEFC6DFEB7A3EE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eutarsopolipus labiatus Seeman 2021	<div><p>Eutarsopolipus labiatus sp. nov.</p> <p>(Figs 29, 32–36)</p> <p>Diagnosis. Adult female: sc1 15–19, c1 20–22, c2 21–24, d 20–24, f 10–16, v2-v2 57–64, sc1-sc1 84-92, d-d 135– 150, f-f 100–120; genua II-III setae l′ 2–3, tibia I setae l′ 7–8, v′ 3–4, tarsi II-III setae tc′′ 4–5. Adult male: leg I with one claw; c1 7–12, c2 8–10, v2-v2 34–40; tarsi II-III setae tc″ 3–4. Larval female: c1 15–18, c2 19–24, d 17 – 19, f 11–15, sc2–sc2 83–100, v1-sc1 22–28; tarsi II-III setae tc′′ 3–4.</p> <p>Material examined. Holotype female, ex. Castelnaudia marginifera. Mt Mistake Plateau, via Goomburra, 27° 58ʹ 37ʺ S 152° 22ʹ 58ʺ E, 17-22 Nov 1987, G. Monteith, rainforest, IN6919, HR# T 88508 (QMS 113432). Paratypes. All ex. C. marginifera. 27 females, 10 males, 11 larvae, as follows: 6 females, 2 males, 2 larvae, same data as holotype (QMS 113433–39); 8 females, 4 males, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=152.84805&amp;materialsCitation.latitude=-28.315832" title="Search Plazi for locations around (long 152.84805/lat -28.315832)">Bald Mtn area</a>, via Emu Vale, 28° 13ʹ 57ʺ S 152° 24ʹ 54ʺ E, 26 Jan-30 Jan 1973, G. Monteith, IN9131, HR# T 88513 (QMS 113440–49, ANIC 52-003969); 13 females, 4 males, 9 larvae, Lever’s Plateau, 28° 18ʹ 57ʺ S 152° 50ʹ 53ʺ E, 29 Feb-11 Apr 1992, D. Cook, pitfall trap, rainforest, IN5139, HR# T 64339 (QMS 113450–62, ANIC 52-003970 – 71, ZMH-A0015196–98). All in QM except 1 female, 1 male, 1 larva in ANIC and ZMH.</p> <p>Other material examined. 1 pharate female, same data as holotype.</p> <p>Description. FEMALE (Figs 29, 32–33; n = 27, 7 measured).</p> <p>Gnathosoma (Fig. 29E). Length 49 (47–51), width 42 (41–43), apex indented, terminating in small but distinct lobes. Cheliceral stylets 41 (40–45), setae ch 17 (17–21), su 10 (7–12).</p> <p>Idiosoma (Figs 32–33). Length 305 (300–370) width 235 (265–300). Prodorsal plate not eroded anteriorly; setae slender, v1 7 (7–10), v2 alveolar remnant, sc1 asymmetrical lengths, 17 and 12 (15–19, not asymmetrical), sc2 80 (75–84). Distance between setae v1–v1 45 (43–48), sc1–sc1 88 (85–92), v1–sc1 24 (27–33), v2–v2 57 (58–64), sc2– sc2 150 (140–155), sc1–sc2 48 (38–48). Opisthosomal setae slender. Plate C setae c1 21 (20–22), c2 22 (21–24), distance between setae c1–c1 83 (72–97). Plate D setae d 20 (21–24), d–d 140 (135–150), cupuli ia anterolaterad d. Plate EF setae f 11 (10–16), f–f 110 (100–120), cupuli im anteriad to anterolaterad f. Plate H divided, seta h minute. Venter: coxisterna 1-2 moderately well defined, with few folds, coxisterna 3 defined but membranous; coxal setae slender, 1a 3 (3), 2a 5 (4–5), 3a 6 (6–7), 3b 6 (6–7). Alveoli of 1b, 2b present.</p> <p>Legs (Figs 33 B-D). Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Ranges given only when holotype measurement different from E. basiatus. Leg I. Femur I, d minute, l’ 2, v″ 6 (5–8); genu I, l′ 2, l″ 2; tibia I, d 45 (40–50), l’ 8 (7–8), l’’ 14 (10–12), v’ 4 (3–4), v’’ 16, k 4 (4–5), φ 11 (10–11); tarsus I, tc’ 14, tc’’ 15, pl’ 10, pl’’ 19, pv′ 2 (m–2), pv’’ 2, ω 4, s 5, u′′ m (m). Leg II. Genu II, l′ 3; tibia II, d 40, l’ 7, v’ 13 (13–15), v’’ 40 (35–40); tarsus II, tc′ 2, tc ″ 5 (4–5), pl ″ 40, pv’’ 2, u’ 7 (6–7), u′′ minute. Leg III. Genu III, l′ 2 (2); tibia III, d 35, l’ 5, v’ 18, v’’ 21; tarsus III, tc’ 2 (2), tc ″ 5 (4–5), pl ″ 30, pv’’ 2, u’ 7 (6–7), u′′ minute.</p> <p>LARVIFORM ADULT MALE (Fig. 34; n = 10, 5 measured).</p> <p>Gnathosoma. Length 31–33, width 29–30, apex indented, terminating in small but distinct lobes; pharynx distorted in drawn specimen. Cheliceral stylets 20–23, ch 5–6, su 4–5.</p> <p>Idiosoma. Length 180–200, width 145–175. Prodorsal plate with setae v1 3–5, v2 alveolar remnant, sc1 5–7, sc2 50–65. Distance between setae v1–v1 28–32, sc1–sc1 47–53, v1–sc1 17–22 (asymmetry in one specimen, 11/19), v2–v2 34–40, sc2–sc2 69–78, sc1–sc2 16–25 (one asymmetrical specimen 32/25). Plate CD setae c1 7–12, c2 8–10, d 6–10 (asymmetrical in one specimen, 7/13), distance between setae c1–c1 48–57, c1–c2 34–42 (asymmetrical in one specimen, 25/39), d–d 44–53; cupuli ia anterolaterad setae d. Plate EF slightly eroded medially; with setae f 4–5, distance f–f 26–32; cupuli im anterolaterad setae f. Genital capsule length 28–29, width 38–42, setae h minute. Venter: coxal setae 1a 3, 2a 4–5, 3a 2–3, 3b 4–5.</p> <p>Legs. Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Ranges given only when first measurement different from E. basiatus. Leg I. Femur I, d minute, l′ minute, v″ 4–6; genu I l′ m–2, l″ m–2; tibia I, d 45, l’ m–2, l’’ m–2, v ’ 4–5, v ’’ 9–14, k 3, φ 8; tarsus I, tc’ 13–14, tc’’ 13–14, pl’ 10–11, pl’’ 12, pv ′ m–2, pv’’ m–2, ω 2–3, s 4–5, u ′′ minute. Leg II. Genu II, l′ 2; tibia II, d 21–30, l’ 6–7, v’ 14, v’’ 16; tarsus II, tc′ 2, tc ″ 3–4, pl ″ 26, pv’’ 2, u’ 5, u″ minute. Leg III. Genu III, l′ minute; tibia III, d 19, l’ 5–6, v’ 14, v’’ 14–18; tarsus III, tc′ 2, tc ″ 3–4, pl ″ 26, pv’’ 2, u’ 5, u′′ minute.</p> <p>LARVAL FEMALE (Figs 35–36; n = 11, 5 measured).</p> <p>Gnathosoma. Length 42–44, width 37–40, apex indented, terminating in small but distinct lobes. Cheliceral stylets 36–43, setae ch 20–24, su 5–9.</p> <p>Idiosoma (Fig. 35). Length 220–265, width 155–205. Prodorsal plate with setae v1 5–7, v2 alveolar remnant, sc1 6–8, sc2 75–100. Distance between setae v1–v1 36–46, sc1–sc1 66–81, v1–sc1 22–28 (asymmetrical in one specimen: v2 and sc1 reversed in position), v2–v2 48–64, sc2–sc2 83–100, sc1–sc2 28–32. Plate C setae c1 15–18, c2 19–24, distance between setae c1–c1 49–60. Plate D setae d 17–19, d–d 60–80, cupuli ia closely anteriad to anterolaterad setae d. Plate EF setae f 11–15, f–f 46–64, cupuli im anteriad to anterolaterad f. Plate H narrow, large, 20–24 wide, 34–42 long, bearing setae h1 ca. 170, h2 10–17. Venter: coxisterna 1-2 moderately well defined, with some folds, coxisterna 3 membranous; coxal setae slender, 1a 2–3, 2a 4–5, 3a 4–5, 3b 5–6. Alveoli of 1b, 2b present.</p> <p>Legs (Fig. 36). Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Ranges given only when first measurement different from E. basiatus. Leg I. Femur I, d minute, l′ minute, v″ 4–6; genu I, l′ m–2, l″ m–2; tibia I, d 50–60, l’ 8–12, l’’ 7–12, v’ 6–7, v’’ 11–15, k 4, φ 10; tarsus I, tc’ 12–14, tc’’ 13–14, pl’ 10, pl’’ 15–19, pv′ minute, pv’’ m–2, ω 3–4, s 5, u′′ minute. Leg II. Genu II, l′ m–2; tibia II, d 35, l’ 7, v’ 12–17, v’’ 19; tarsus II, tc′ 2–3, tc ″ 3–4, pl ″ 30, pv’’ 2, u’ 5, u′′ minute. Leg III. Genu III, l′ minute; tibia III, d 28–34, l’ 7, v’ 16, v’’ 18–25; tarsus III, tc’ 2, tc ″ 3–4, pl ″ 27–36, pv’’ 2, u’ 5–6, u′′ minute.</p> <p>Differential diagnosis. Eutarsopolipus labiatus sp. nov. is most similar to E. savatus sp. nov. by sharing long setae sc1 (exceeding 10) and greater distance d-d (135–150) and f-f (100–120). Females are very similar and differ by the shorter cheliceral seta (17–21 versus 24–39) and slightly longer idiosomal setae. The idiosomal setal lengths are however more distinctive in the larvae: c1 15–18, c2 19–24, d 17–19 in E. labiatus versus c1 6–9, c2 9–12, d 7–10 in E. savatus.</p> <p>Etymology. The specific name labiatus (lipped) refers to the protruding lip-like gnathosoma found in this species group.</p> </div>	http://treatment.plazi.org/id/4438245CFFDB097B87AEFC6DFEB7A3EE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Seeman, Owen D.	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFC1097887AEFF5AFD30A067.text	4438245CFFC1097887AEFF5AFD30A067.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eutarsopolipus osculum Seeman 2021	<div><p>Eutarsopolipus osculum sp. nov.</p> <p>(Figs 37–38)</p> <p>Diagnosis. Adult female: sc1 7–8, c1 15–17, c2 16–18, d 16–22, f 6–11, v2-v2 35–40, sc1-sc1 60–65, d-d 95–98, f-f 67–80; genua II-III setae l′ m–2, tibia I setae l′ 7–8, v′ 2–3, tarsi II-III setae tc′′ 3–4. Adult male: leg I with one claw; c1 5, c2 5, v2-v2 27; tarsi II-III setae tc″ 2–3. Larval female: c1 11, c2 9, d 10, f 7, sc2–sc2 71, v1-sc1 22; tarsi II-III setae tc′′ 3.</p> <p>Material examined. Holotype female, ex. Castelnaudia cordata. “Glen Witheren”, shelf scrub, 28° 2ʹ 58ʺ S 153° 6ʹ 58ʺ E, 30 Dec 1991 - 24 May 1992, G. Monteith, pitfall trap, IN5682, rainforest, HR# T 68606 (QMS 113471). <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=153.11612&amp;materialsCitation.latitude=-28.049444" title="Search Plazi for locations around (long 153.11612/lat -28.049444)">In</a> QM. Paratypes. All ex. C. cordata. 2 females, 1 male, 1 larva, as follows: 1 female, 1 male, 1 larva, same data as holotype (QMS 113472-74); 1 female, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=152.52583&amp;materialsCitation.latitude=-28.30861" title="Search Plazi for locations around (long 152.52583/lat -28.30861)">Mt Clunie</a> summit, 28° 18ʹ 31ʺ S 152° 31ʹ 33ʺ E, 16 Dec 1972, G. Monteith, rainforest, IN6361, HR# T 64324 (ANIC 52-003972). All in QM except 1 female in ANIC.</p> <p>Note: The holotype and paratype females from “Glen Witherin” are on one slide.</p> <p>Description. FEMALE (Figs 37–38; n = 3).</p> <p>Gnathosoma. Length 42 (44), width 40 (36–38), apex indented, terminating in small but distinct lobes. Cheliceral stylets 21 (23–25), setae ch 36 (35–41), su 8 (12–13).</p> <p>Idiosoma (Fig. 37). Length 275 (290–310) width 185 (180–200). Prodorsal plate not eroded anteriorly; setae slender, v1 6 (6–7), v2 alveolar remnant, sc1 7 (7–8), sc2 90 (80–85). Distance between setae v1–v1 33 (30–36), sc1–sc1 65 (60–61), v1–sc1 20 (16–22), v2–v2 38 (35–40), sc2–sc2 105 (93–105), sc1–sc2 35 (31–37). Opisthosomal setae slender. Plate C setae c1 17 (15–17), c2 18 (16), distance between setae c1–c1 75 (59–72). Plate D setae d 22 (16–20), d–d 98 (95–98), cupuli ia anterolaterad d. Plate EF setae f 10 (6–11), f–f 75 (67–80), cupuli im anterolaterad f. Plate H divided, seta h minute. Venter: coxisterna 1-2 moderately well defined, with few folds, coxisterna 3 membranous; coxal setae slender, 1a 3 (2–3), 2a 4 (4–5), 3a 5 (4–6), 3b 5 (5–6). Alveoli of 1b, 2b present.</p> <p>Legs (Fig. 38). Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Ranges given only when holotype measurement different from E. basiatus. Leg I. Femur I, d minute, l’ minute, v″ 5 (5–8); genu I, l′ 2 (2), l″ 2; tibia I, d 50, l’ 7 (7–8), l’’ 8, v’ 2 (2–3), v’’ 8 (8–10), k 5, φ 10; tarsus I, tc’ 15, tc’’ 13 (13–14), pl’ 8, pl’’ 21, pv′ 2 (2), pv’’ 2, ω 3 (3), s 5, u′′ m. Leg II. Genu II, l′ m (m); tibia II, d 36, l’ 6, v’ 15 (18), v’’ 22; tarsus II, tc′ 2 (m–2), tc ″ 4 (3–4), pl ″ 45, pv’’ 2, u’ 6, u′′ m. Leg III. Genu III, l′ m (m–2); tibia III, d 33, l’ 7, v’ 10 (10–17), v’’ 25; tarsus III, tc’ 2 (2), tc ″ 4 (3–4), pl ″ 45 (42–45), pv’’ 2, u’ 7 (6), u′′ m.</p> <p>LARVIFORM ADULT MALE (n = 1).</p> <p>Gnathosoma. Length 28, width 26, apex indented, terminating in small but distinct lobes. Cheliceral stylets 20, setae ch 6, su 5.</p> <p>Idiosoma. Length 140, width 110. Prodorsal plate with setae v1 4 (absent one side), v2 alveolar remnant, sc1 3, sc2 55. Distance between setae v1–v1 not measureable, sc1–sc1 41, v1–sc1 17, v2–v2 27, sc2–sc2 54, sc1–sc2 16. Plate CD setae c1 5, c2 5, d 6, distance between setae c1–c1 42, c1–c2 27, d–d 39; cupuli ia anteriad setae d. Plate EF slightly eroded medially; with setae f 4, distance f–f 25; cupuli im anteriad setae f. Genital capsule length 25, width 31, setae h minute. Venter: coxal setae 1a 2, 2a 3, 3a 3, 3b 3.</p> <p>Legs. Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Leg I. Femur I, d minute, l′ minute, v″ 3; genu I, l′ minute, l″ minute; tibia I, d 38, l’ 4, l’’ 4, v ’ 3, v ’’ not measurable, k 3, φ 8; tarsus I, tc’ 13, tc’’ 10, pl’ 5, pl’’ 15, pv ′ minute, pv’’ minute, ω 3, s 4, u ′′ minute. Leg II. Genu II, l′ 2; tibia II, d 19, l’ 4, v’ 11, v’’ 16; tarsus II, tc′ minute, tc ″ 2, pl ″ 31, pv’’ minute, u’ 4, u″ minute. Leg III. Genu III, l′ minute; tibia III, d 19, l’ 5, v’ 9, v’’ 13; tarsus III, tc′ minute, tc ″ 3, pl ″ 25, pv’’ minute, u’ 5, u″ minute.</p> <p>LARVAL FEMALE (n = 1).</p> <p>Gnathosoma. Length 40, width 35, apex indented, terminating in small but distinct lobes. Cheliceral stylets 35, setae ch 28, su 7.</p> <p>Idiosoma. Length 195, width 140. Prodorsal plate with setae v1 5, v2 alveolar remnant, sc1 6, sc2 95. Distance between setae v1–v1 28, sc1–sc1 58, v1–sc1 22, v2–v2 33, sc2–sc2 71, sc1–sc2 22. Plate C setae c1 11, c2 9, distance between setae c1–c1 46. Plate D setae d 10, d–d 51, cupuli ia anterolaterad setae d. Plate EF setae f 7, f–f 38, cupuli im anterolaterad f. Plate H narrow, large, 20 wide, 33 long, bearing setae h1, broken, h2 7. Venter: coxisterna 1-2 moderately well defined, with some folds, coxisterna 3 membranous; coxal setae slender, 1a 4, 2a 4, 3a 4, 3b 5. Alveoli of 1b, 2b present.</p> <p>Legs. Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Leg I. Femur I, d minute, l′ minute, v″ 5; genu I, l′ minute, l″ minute; tibia I, d 55, l’ 8, l’’ 8, v’ 3, v’’ 13, k 5, φ 10; tarsus I, tc’ 13, tc’’ 14, pl’ 6, pl’’ 20, pv′ minute, pv’’ 2, ω 3, s 5, u′′ minute. Leg II. Genu II, l′ minute; tibia II, d 27, l’ 6, v’ 12, v’’ 17; tarsus II, tc′ 2, tc ″ 3, pl ″ 35, pv’’ 2, u’ 5, u′′ minute. Leg III. Genu III, l′ 2; tibia III, d 34, l’ 7, v’ 13, v’’ 15; tarsus III, tc’ minute, tc ″ 3, pl ″ 34, pv’’ minute, u’ 6, u′′ minute.</p> <p>Differential diagnosis. Eutarsopolipus osculum sp. nov. is most similar to E. basiatus sp. nov. in having short setae sc1 (&lt;10) and a shorter distance between setae d-d (95–115) and f-f (67–92). In addition to the features noted in the Differential diagnosis of E. basiatus, E. osculum has a shorter seta c1 (15–17 versus 6–9).</p> <p>Etymology. The name osculum (little mouth or little kiss) alludes to both the lip-like protrusion of the gnathosoma but also the smaller size of this species.</p> </div>	http://treatment.plazi.org/id/4438245CFFC1097887AEFF5AFD30A067	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Seeman, Owen D.	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFC3097587AEFB07FE1AA7F9.text	4438245CFFC3097587AEFB07FE1AA7F9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eutarsopolipus savatus Seeman 2021	<div><p>Eutarsopolipus savatus sp. nov.</p> <p>(Figs 39–40)</p> <p>Diagnosis. Adult female: sc1 11–19, c1 11–19, c2 14–20, d 12–19, f 8–12, v2-v2 48–58, sc1-sc1 76–88, d-d 120– 160, f-f 95–120; genua II-III setae l′ 2–4, tibia I setae l′ 8–10, v′ 4–5, tarsi II-III setae tc′′ 4–5. Adult male: leg I with one claw; c1 4–5, c2 5–6, v2-v2 34–41; tarsi II-III setae tc″ 3–4. Larval female: c1 6–9, c2 9–12, d 7–10, f 7, sc2-sc2 78–100, v1-sc1 22–24; tarsi II-III setae tc′′ 3.</p> <p>Material examined. Holotype female, ex. Castelnaudia eungella. Pease’s Lookout, Eungella, 21° 6ʹ 53ʺ S 148° 31ʹ 6ʺ E, 17 Nov 1992 - 15 Apr 1993, D. Cook, G. Monteith, flight intercept &amp; pitfall trap, IN5934, HR# T 82502 (QMS 113497). Paratypes. All ex. C. eungella. 9 females, 4 males, 11 larvae, as follows: 1 female, 4 larvae same data as holotype (QMS 113498, 113500–503); 2 larvae, same data except another beetle, HR# T 82503 (QMS 113504, 113578); 2 females, 1 male, 2 larvae, same data except another beetle, HR# T 82506 (QMS 113577, 11379–80, ANIC 52-003973, ZMH-A0015199); 2 females, 1 male, 1 larva, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=148.63388&amp;materialsCitation.latitude=-21.026945" title="Search Plazi for locations around (long 148.63388/lat -21.026945)">Mt Dalrymple</a>, 21° 1ʹ 37ʺ S 148° 38ʹ 2ʺ E, 21 Dec 1992 – 10 Jan 1993, ANZSES, HR# T 82501 (QMS 113581–83, ZMH-A0015200); 1 female, 2 larvae, Eungella National Park, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=148.59917&amp;materialsCitation.latitude=-21.018055" title="Search Plazi for locations around (long 148.59917/lat -21.018055)">Mt Dalrymple</a> track, 1200 m site, 21.026° S 148.627°E, 23 Nov 2013, C. Burwell, rainforest, sampcode 25933, beetle CB6 (QMS 113584–85, ANIC 52-003974); 1 female, 1 male, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=148.59917&amp;materialsCitation.latitude=-21.018055" title="Search Plazi for locations around (long 148.59917/lat -21.018055)">Mt William</a>, 21° 1ʹ 5ʺ S 148° 35ʹ 57ʺ E, 21 Dec 1992 – 10 Jan 1993, ANZSES, HR# T 82489 (QMS 113586–87); 2 females, 1 male, Upper Cattle Creek, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=148.60306&amp;materialsCitation.latitude=-21.028055" title="Search Plazi for locations around (long 148.60306/lat -21.028055)">Eungella</a>, 21° 1ʹ 41ʺ S 148° 36ʹ 11ʺ E, 17 Nov 1992, G. Monteith, G. Thompson, H. Janetzki, HR# T 82513 (QMS 113588–90). All in QM except 1 female and 1 larva in each of ANIC and ZMH.</p> <p>Other material examined. 1 pharate female ex HR# T 82503 (QMS 113505).</p> <p>Description. FEMALE (Figs 39–40; n = 10).</p> <p>Gnathosoma. Length 43 (43–48), width 41 (40–45), apex indented, terminating in small but distinct lobes. Cheliceral stylets 44 (38–40), setae ch 31 (24–39), su 14 (13–16).</p> <p>Idiosoma (Fig. 39). Length 360 (305–385), width 405 (250–350). Prodorsal plate not eroded anteriorly; setae slender, v1 9 (6–9), v2 alveolar remnant, sc1 16 (11–19; one specimen asymmetrical 10/15), sc2 80 (80–110). Distance between setae v1–v1 45 (37–45), sc1–sc1 82 (73–88), v1–sc1 26 (often asymmetrical, 23–32), v2–v2 58 (48–58), sc2–sc2 130 (125–135), sc1–sc2 38 (often asymmetrical, 31–46). Opisthosomal setae slender. Plate C setae c1 19 (9–16), c2 19 (14–20), distance between setae c1–c1 85 (80–115). Plate D setae d 19 (12–19), d–d 140 (120–160), cupuli ia anterolaterad d. Plate EF setae f 10 (8–12), f–f 115 (95–120), cupuli im anteriad to anterolaterad f. Plate H divided, seta h minute. Venter: coxisterna 1-2 moderately well defined, with few folds, coxisterna 3 membranous; coxal setae slender, 1a 4 (3–4), 2a 5 (4–5), 3a 6 (5–6), 3b 6 (6–7). Alveoli of 1b, 2b present.</p> <p>Legs (Fig. 40). Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Paratype measurement given only when holotype measurement different from E. basiatus. Leg I. Femur I, d minute, l’ 3, v″ 8 (7–10); genu I, l′ 3, l″ 3; tibia I, d 60, l’ 10 (8–10), l’’ 9 (8–9), v’ 5 (4–5), v’’ 11 (10–17), k 5, φ 9; tarsus I, tc’ 15, tc’’ 13 (13–15), pl’ 9, pl’’ 16 (15–18), pv′ 2 (2), pv’’ 2, ω 4, s 6 (5), u′′ m. Leg II. Genu II, l′ 4; tibia II, d 48 (48–55), l’ 7, v’ 17 (14–15), v’’ 17 (20–27); tarsus II, tc′ 2, tc ″ 4 (5), pl ″ 59 (50), pv’’ 2, u’ 6, u′′ minute. Leg III. Genu III, l′ 3; tibia III, d 55 (40–49), l’ 7, v’ 16, v’’ 17 (22–29); tarsus III, tc’ 2, tc ″ 5 (4–5), pl ″ 39 (37–40), pv’’ 2, u’ 7 (6–7), u′′ minute.</p> <p>LARVIFORM ADULT MALE (n = 4).</p> <p>Gnathosoma. Length 31–39, width 28–36, apex indented, terminating in small but distinct lobes. Cheliceral stylets 20–22, setae ch 6–7, su 6–7.</p> <p>Idiosoma. Length 150–180, width 125–150. Prodorsal plate with setae v1 2–3, v2 alveolar remnant, sc1 3–5, sc2 60–70. Distance between setae v1–v1 30–36, sc1–sc1 43–50, v1–sc1 15–16, v2–v2 34–41, sc2–sc2 63–77, sc1–sc2 19–25. Plate CD setae c1 4–5, c2 5–6, d 5–6, distance between setae c1–c1 46–54, c1–c2 32–38, d–d 40–46; cupuli ia anterolaterad setae d. Plate EF eroded medially; with setae f 4–5, distance f–f 25–29; cupuli im anterolaterad setae f. Genital capsule length 27–28, width 38–40, setae h minute. Venter: coxal setae 1a 2–3, 2a 3–4, 3a 2–3, 3b 4–5.</p> <p>Legs. Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Leg I. Femur I, d minute, l′ minute, v″ 4–6; genu I, l′ minute, l″ minute; tibia I, d 42–52, l’ 2–3, l’’ 3–4, v ’ 3, v ’’ 10, k 3, φ 8; tarsus I, tc’ 12–14, tc’’ 12–14, pl’ 9, pl’’ 10–14, pv ′ m–2, pv’’ m–2, ω 3–4, s 4–5, u ′′ minute. Leg II. Genu II, l′ minute; tibia II, d 26–30, l’ 5, v’ 11, v’’ 18–21; tarsus II, tc′ 2, tc ″ 3–4, pl ″ 25–28, pv’’ minute, u’ 4, u″ minute. Leg III. Genu III, l′ minute; tibia III, d 26–28, l’ 4–6, v’ 8–12, v’’ 16; tarsus III, tc′ 2, tc ″ 4, pl ″ 29–31, pv’’ 2, u’ 4, u″ minute.</p> <p>LARVAL FEMALE (n = 11).</p> <p>Gnathosoma. Length 38–41, width 33–36, apex indented, terminating in small but distinct lobes. Cheliceral stylets 30–35, setae ch 22–27, su 6–7.</p> <p>Idiosoma. Length 185–260, width 140–220. Prodorsal plate with setae v1 4–5, v2 alveolar remnant, sc1 5–6, sc2 105–110. Distance between setae v1–v1 32–35, sc1–sc1 59–65, v1–sc1 22–24, v2–v2 44–46, sc2–sc2 78–100, sc1–sc2 22–34. Plate C setae c1 6–9, c2 9–10, distance between setae c1–c1 50–71. Plate D setae d 7–10, d–d 54–72, cupuli ia anteriad to anteromesad d. Plate EF setae f 7–10, f–f 47–59, cupuli im anteriad to anterolaterad f. Plate H narrow, large, 18–21 wide, 31–36 long, bearing setae h1 ca. 160, h2 6–8. Venter: coxisterna 1-2 moderately well defined, with some folds, coxisterna 3 membranous; coxal setae slender, 1a 3, 2a 4, 3a 2–4, 3b 5 (one specimen lacking setae 3a and 3b expressed as alveoli). Alveoli of 1b, 2b present.</p> <p>Legs. Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Ranges given only when first measurement different from E. basiatus. Leg I. Femur I, d minute, l′ minute, v″ 4–6; genu I, l′ m–2, l″ 2; tibia I, d 55–65, l’ 7–11, l’’ 8–11, v’ 4–5, v’’ 8–10, k 4, φ 10; tarsus I, tc’ 15, tc’’ 15, pl’ 6–7, pl’’ 14–17, pv′ minute, pv’’ minute, ω 4, s 5, u′′ minute. Leg II. Genu II, l′ m–2; tibia II, d 40, l’ 6, v’ 10–14, v’’ 21; tarsus II, tc′ minute, tc ″ 3, pl ″ 28, pv’’ minute, u’ 5, u′′ minute. Leg III. Genu III, l′ 3; tibia III, d 36, l’ 7, v’ 10–15, v’’ 21; tarsus III, tc’ m–2, tc ″ 3, pl ″ 38–43, pv’’ minute, u’ 5, u′′ minute.</p> <p>Differential diagnosis. Eutarsopolipus savatus sp. nov. is most similar to E. labiatus and the distinguishing features are discussed under the Differential Diagnosis for the latter species.</p> <p>Etymology. The name savatus (a tender kiss) alludes to both the lip-like protrusion of the gnathosoma but also alludes to the larger size of this species.</p> <p>Remarks. One larva similar to E. echinatus Seeman, 2019a was found on HR# T82502; one male closest to the nahmani subgroup group was found on HR# T82502 (QMS 113499) (Tables 1, 2).</p> <p>The leytei species group</p></div> 	http://treatment.plazi.org/id/4438245CFFC3097587AEFB07FE1AA7F9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Seeman, Owen D.	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFCE097187AEFBD9FA39A3AD.text	4438245CFFCE097187AEFBD9FA39A3AD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eutarsopolipus biuncatus Seeman 2021	<div><p>Eutarsopolipus biuncatus sp. nov.</p> <p>(Figs 41–43)</p> <p>Diagnosis. Adult female: leg I with two claws; v1 8–10, sc2 15–20; genu I with two setae (l″ present); tarsus II setae tc″ 5–6, u′ 6–7. Adult male: setae sc2 2–3. Larval female: sc2 51–55.</p> <p>Material examined. Holotype female, ex. Notonomus transitus. Blue Mt, 0.6 km SE, 21° 36ʹ 19ʺ S 148° 58ʹ 21ʺ E, 3–4 Oct 1999, G. Monteith, D. Cook, C. Burwell, S. Evans, rainforest, IN7801, HR# T 89396 (QMS 113608). Paratypes. All ex. Notonomus transitus. 5 females, 5 males, 14 larvae, as follows: 2 females, 2 males, 10 larvae, same data as holotype (QMS 113609–18, ANIC 52-003975 – 77, ZMH-A0015201); 1 female, 1 male, 1 larva, 50 mi. W of Mackay, 21° 08ʹ 12ʺ S 148° 29ʹ 30ʺ E, 6 Jan 1973, J. Hammond, IN7864, HR# T 81753 (QMS 113621–23); 2 females, 2 males, 1 larva, Broken River, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=148.50833&amp;materialsCitation.latitude=-21.168055" title="Search Plazi for locations around (long 148.50833/lat -21.168055)">Eungella</a>, 21° 10ʹ 5ʺ S 148° 30ʹ 30ʺ E, 4 Dec 1992 – 30 Apr 1993, P. Lawless, M. Shaw, R. Raven, IN 9781, HR# T110084 (QMS 113624–26, ZMH-A0015202–03); 2 larvae, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=148.58527&amp;materialsCitation.latitude=-21.028889" title="Search Plazi for locations around (long 148.58527/lat -21.028889)">Dalrymple Rd</a>, Eungella NP, 21° 1ʹ 44ʺ S 148° 35ʹ 7ʺ E, 9 Nov 1991 – 29 Jul 1992, P. Lawless, R. Raven, M. Shaw, rainforest, IN7768, HR# T 81768 (QMS 113627–28). All in QM except 1 female, 1 male, 1 larva in ANIC and ZMH.</p> <p>Other material examined. 2 larvae, same data as holotype, in poor condition (QMS 113619–20).</p> <p>Description. FEMALE (Fig. 41; n = 6).</p> <p>Gnathosoma. Length 45 (44–46), width 39 (34–37). Palp with alveolar remnant of femoral seta. Cheliceral stylets 38 (35–36), setae ch 22 (14–16), su 11 (8–12).</p> <p>Idiosoma (Fig. 41A). Length ca. 500 (270–430), width ca. 300 (215–310), bigger specimens swollen, distorted. Stigmata apparent, large, in soft cuticle anteriad setae v1 to posterolaterad gnathosoma (varies according to mount), respiratory system weakly developed, tracheae not extending past small atrium, tracheae and atrium sometimes not visible. Prodorsal plate with setae v1 10 (8–9), v2 alveolar remnant, sc1 9 (7–8), sc2 20 (15–17). Distance between setae v1–v1 44 (39–51), sc1–sc1 69 (64–72), v1–sc1 22 (23–25), v2–v2 52 (53–57), sc2–sc2 84 (83–110), sc1–sc2 34 (24–32). Plate C appearing divided, each plate bearing its slender setae at their margins, c1 6 (5–7), c2 6 (4–5). Plate D setae d 5 (4–5), d–d 92 (85–95), cupuli ia anteriad d. Plate EF setae f 4 (4–5), f–f broken (49–60), cupuli im anterolaterad f. Plate H absent. Venter: coxisterna 1-3 well defined; coxal setae slender, 1a 5 (5–6), 2a 5 (4–5), 3a 6 (5–6), 3b 5 (4–5). Alveoli of 1b, 2b present.</p> <p>Legs (Figs 41 B-C). Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-9(+ ω), 0-1-4-7(+ ω), 0-1-4-7. Leg I. Femur I, d 1 (1–3), l’ 5 (4–5), v″ 8 (9–10); genu I, l’ 3 (2–3), l″ 2 (2–3); tibia I, d 16 (14–19), l’ 3 (2–3), l’’ 5 (5–6), v’ 3 (3–5), v’’ 9 (8–14), k 3 (3–4), φ 6 (6–7); tarsus I, tc’ 9 (9–10), tc’’ 9 (9–10), pl’ 8 (8–10), pl’’ 10 (8–12), pv′ 2 (2–3), pv’’ 2 (2–3), ω 3 (3–4), s 6 (5–6), u′′ minute (m–2), p′ m (m–2). Leg II. Genu II, l’ 3 (2–3); tibia II, d 9 (7–9), l’ 5 (3–5), v’ 8 (8–11), v’’ 14 (13–16); tarsus II, tc′ 8 (6–8), tc ″ 5 (5–6), pl ″ 10 (7–9), pv′ 3 (3–4), pv’’ 4 (4–5), ω 3 (2–3), u’ 7 (6–7), u″ 2 (2). Leg III. Genu III, l’ 3 (2–3); tibia III, d 4 (5–6), l’ 4 (3–4), v’ 8 (6–9), v’’ 11 (13–15); tarsus III, tc’ 6 (6–7), tc ″ 5 (5–6), pl ″ 8 (9–10), pv′ 3 (2–3), pv’’ 2 (2–3), u’ 6 (6–7), u″ 2 (2). Claws on legs I-III well developed and paired.</p> <p>LARVIFORM ADULT MALE (Fig. 42; n = 5).</p> <p>Gnathosoma. Length 28–31, width 28–29. Palp with alveolar remnant of femoral seta. Cheliceral stylets 17–18, setae ch 1–2, su 4–6.</p> <p>Idiosoma. Length 155–165, width 110–120. Prodorsal plate with setae v 1 m–2, v2 alveolar remnant, sc1 minute, sc2 2–3, thickened. Distance between setae v1–v1 19–22, sc1–sc1 45–48, v1–sc1 19–21, v2–v2 34–36, sc2–sc2 54–56, sc1–sc2 22–23. Plate CD incised posterolaterally, setae c1, c2, d minute, distance between setae c1–c1 44–46, c1–c2 26–35, d–d 31–32; cupuli ia laterad to anterolaterad setae d. Plate EF with setae f minute, distance f–f 21–23; cupuli im anterolaterad setae f. Genital capsule length 22–25, width 34–36, setae h minute. Venter: coxal setae slender, 1a 2–3, 2a 2–3, 3a 2, 3b 2–3.</p> <p>Legs. Setal counts legs I–III, femur-tarsus: 2-0-6(+ φ)-8(+ ω), 0-0-4-7(+ ω), 0-0-4-7. Compared to female: Femur I seta v″ absent, genu I l′ absent, genua I-III l″ absent; tarsus I p′ absent. Leg I. Femur I, d minute, l’ 2; genu I, l′ minute; tibia I, d 4–5, l’ minute, l’’ m–2, v ’ 2, v ’’ 6–7, k minute, φ 4–5; tarsus I, tc’ 7–8, tc’’ 8–9, pl’ 4–6, pl’’ 6–9, pv ′ 2, pv’’ 2–3, ω 3, s 4–5, u ′′ 1–2. Leg II. Tibia II, d 3–5, l’ 2–3, v’ 6–7, v’’ 7–8; tarsus II, tc′ 3, tc ″ 4, pl ″ 5–6, pv ′ 2, pv’’ 2–3, ω 2–3, u’ 5–6, u″ minute. Leg III. Tibia III, d 2, l’ 2, v’ 6–8, vʺ 7–9; tarsus III, tc′ 2–3, tc ″ 4, pl ″ 4–5, pv ′ 2, pv’’ 2, u ’ 5, u″ minute. Claws on leg II-III present. Setal form similar to that in female, but with tibia I setae l′, l″, v′ as tiny spines.</p> <p>LARVAL FEMALE (Fig. 43; n = 14).</p> <p>Gnathosoma. Length 30–34, width 27–30. Palp with alveolar remnant of femoral seta. Cheliceral stylets 25–27, setae ch 19–23, su 8–10.</p> <p>Idiosoma (Fig. 43). Length 165–225, width 105–185; larger specimens swollen laterally between prodorsal shield and plate CD. Prodorsal plate with setae v1 12–14, v2 alveolar remnant, sc1 5–6, sc2 51–55. Distance between setae v1–v1 20–23, sc1–sc1 48–51, v1–sc1 25–28, v2–v2 29–35, sc2–sc2 49–54, sc1–sc2 27–33. Plates C and D fused medially; line of fusion visible. Plate CD setae c1 5–6, c2 3–4, d 4–5, cupuli ia anteriad to anteromesad d, distance between setae c1–c1 26–27, d–d 24–28. Plate EF setae f 4–5, f–f 23–26, cupuli im anteriad f. Plate H poorly expressed, caudal cone 22–30 wide, 8–13 long, bearing setae h1 ca. 65–70, h2 3–4. Venter: coxisterna 1-3 well-defined; coxal setae slender, 1a 2–3, 2a 3–4, 3a 4–5, 3b 3–4. Alveoli of 1b, 2b present.</p> <p>Legs. Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6(+ ω), 0-1-4-6 (setae pv′ absent on tarsi II- III). Leg I. Femur I, d minute, l’ 2–3, v’’ 5–7; genu I, l’ 2–3, l″ m–2; tibia I, d 17–21, l’ 6–7, l’’ 5–7, v’ 3–4, v’’ 7–9, k 2–3, φ 6–7; tarsus I, tc’ 8–9, tc’’ 9–10, pl’ 5–6, pl’’ 9–10, pv′ 2, pv’’ 2, ω 3–4, s 4–5, u′′ minute. Leg II. Genu II, l’ 2–3; tibia II, d 10–12, l’ 4–5, v’ 7–9, v’’ 11–13; tarsus II, tc′ 3–4, tc ″ 4–5, pl ″ 7–9, pv’’ 3–4, ω 3–4, u’ 5–6, u′′ minute. Leg III. Genu III, l’ 2; tibia III, d 10–11, l’ 4–5, v’ 7–8, v’’ 11–12; tarsus III, tc’ 4–5, tc ″ 4–5, pl ″ 8–10, u’ 5–6, pv’’ 5–6, u′′ minute.</p> <p>Differential diagnosis. Eutarsopolipus biuncatus sp. nov. and E. janus sp. nov. are the only members of the leytei species group to retain paired claws on tarsus I in females. The only other species of Eutarsopolipus with females retaining two claws on tarsus I is Eutarsopolipus diunculosus Eidelberg, 1994, a member of the pterostichi species group (lacking both a respiratory system and setae on genua II-III).</p> <p>In addition to the diagnostic features mentioned for E. janus, female specimens appear to have divided plate C. None of these specimens are of sufficient quality to be sure—they may have split when mounted—but the division of dorsal plates occurs elsewhere in Eutarsopolipus (e.g., Regenfuss 1968; Husband 2000).</p> <p>Etymology. The specific name biuncatus (two-hooked) refers to the two claws on tarsus I of females.</p> <p>Remarks. All females had waste matter that obscured morphological features. This was partially removed by clearing some specimens in 5% KOH, but no female was of high quality. The holotype was chosen on the basis of having the best quality prodorsum and legs I-II, where most diagnostic features occur.</p> <p>Both E. biuncatus sp. nov. and E. janus sp. nov. are unusual in retaining setae pv′ on tarsi II-III in the male, a seta usually absent in Tarsonemoidea (Lindquist 1986; Seeman 2019a).</p> <p>Another specimen of N. transitus from the type locality (HR# T95636) had an unusual male of another species of Eutarsopolipus (lacking all genual setae, large, blunt-tipped Y-shaped claws). Males are difficult to place in species groups as they always lack respiratory systems and sometimes lack genual setae that are present in the female.</p> </div>	http://treatment.plazi.org/id/4438245CFFCE097187AEFBD9FA39A3AD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Seeman, Owen D.	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
4438245CFFCB090A87AEFF5AFE85A5C5.text	4438245CFFCB090A87AEFF5AFE85A5C5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eutarsopolipus janus Seeman 2021	<div><p>Eutarsopolipus janus sp. nov.</p> <p>(Figs 44–48)</p> <p>Diagnosis. Adult female: leg I with two claws; v1 19–24, sc2 36–40; genu I with two setae (l″ present); tarsus II setae tc″ 8–9, u′ 8–10. Adult male: setae sc2 27–34. Larval female: sc2 70–80.</p> <p>Material examined. Holotype female, ex. Notonomus flos. <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=145.46194&amp;materialsCitation.latitude=-16.230833" title="Search Plazi for locations around (long 145.46194/lat -16.230833)">Mt Lewis Rd</a>, 29 km from Highway, Cow Bay, 16° 13ʹ 51ʺ S 145° 27ʹ 43ʺ E, 18 Nov 1997 – 9 Feb 1998, G. Monteith &amp; D. Cook, pitfall trap, rainforest, IN1702, HR# T 65790 (QMS 113527). Paratypes. All ex Notonomus flos. 7 females, 7 males, 9 larvae, as follows: 7 females, 7 males, 7 larvae, same data as holotype (QMS 113528–42, ANIC 52-003978 – 80, ZMH-A0015204–05); 2 larvae, same data except different beetle, HR# T 65789 (QMS 113543, ZMH-A0015206). All in QM except 1 female, 1 male, 1 larva in each of ANIC and ZMH.</p> <p>Other material examined. Ex N. aurifer: 7 females, 5 males, 7 larvae, as follows: 6 females, 2 males, 4 larvae, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=145.6425&amp;materialsCitation.latitude=-17.023056" title="Search Plazi for locations around (long 145.6425/lat -17.023056)">Lamb’s Head</a>, 20 km SW Cairns, 17° 1ʹ 23ʺ S 145° 38ʹ 33ʺ E, 10–12 Dec 1989, G. Monteith, G. Thompson, H. Janetzki, HR# T 20079 (QMS 113544–54, 1 female ANIC); 2 males, 1 larva, same data except different beetle, HR# T 20078 (QMS 113555–57); 1 female, 1 male, 2 larvae, same data except different beetle, HR# T 20081 (QMS 113558–61).</p> <p><a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=145.28667&amp;materialsCitation.latitude=-16.460001" title="Search Plazi for locations around (long 145.28667/lat -16.460001)">Ex</a> N. spurgeoni: 5 females, 3 males, 4 larvae, as follows: 1 female, Mossman Bluff Track site 8, 16° 27ʹ 36ʺ S 145° 17ʹ 12ʺ E, 17–19 Dec 1988, G. Monteith, G. Thompson, ANZSES, HR# T 20799 (QMS 113591); 1 female, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=145.3&amp;materialsCitation.latitude=-16.466667" title="Search Plazi for locations around (long 145.3/lat -16.466667)">Mossman Bluff Track</a> general, 16° 28ʹ S 145° 18ʹ E, 20 Dec 1989 – 15 Jan 1990, G. Monteith, G. Thompson, ANZSES, HR# T 20863 (QMS 113592); 1 male, 4 larvae, same data except different beetle, HR# T 20849 (QMS 113598–602); 3 females, 2 males, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=145.21361&amp;materialsCitation.latitude=-16.375278" title="Search Plazi for locations around (long 145.21361/lat -16.375278)">Mt Spurgeon</a>, 7 km N, camp 2, 16° 22ʹ 31ʺ S 145° 12ʹ 49ʺ E, 17–19 Oct 1991, G. Monteith, H. Janetzki, D. Cook, L. Roberts, HR# T 20957 (QMS 113593-97).</p> <p>Ex N. dimorphicus: 1 female, 1 male, 3 larvae, as follows: 1 female, 1 larva, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=145.27444&amp;materialsCitation.latitude=-16.594168" title="Search Plazi for locations around (long 145.27444/lat -16.594168)">Mt Lewis</a>, barracks, 16° 35ʹ 39ʺ S 145° 16ʹ 28ʺ E, no date, G. Wood, ex HR# T 20722 (QMS 113603); 1 male, 1 larva, Mt Lewis Rd, 22 km from Highway, ANZSES site 3, 16° 32ʹ 48ʺ S 145° 16ʹ 53ʺ E, 29 Nov 1997 – 9 Feb 1998, G. Monteith, D. Cook, rainforest, IN1701, HR# T 65741 (QMS 113605–06); 1 larva, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=145.25&amp;materialsCitation.latitude=-16.55" title="Search Plazi for locations around (long 145.25/lat -16.55)">Mt Lewis</a>, 2.5 km N, via Julatten, 16° 33ʹ S 145° 15ʹ E, 3 Nov 1983, D. Yeates, G. Thompson, HR# T 20721 (QMS 113607).</p> <p>Note: One male is fully developed within a broken egg and is included as a paratype.</p> <p>Description (excluding specimens from N. aurifer).</p> <p>FEMALE (Figs 44–46; n = 21).</p> <p>Gnathosoma. Length 55 (52–57), width 51 (45–51). Palp with alveolar remnant of femoral seta. Cheliceral stylets 45 (45–49), setae ch 25 (25–35), su 18 (15–20).</p> <p>Idiosoma (Figs 44–45). Length ca. 340 (290–600), width ca. 240 (230–450), bigger specimens swollen, distorted. Stigmata apparent, large, in soft cuticle anteriad setae v1 to posterolaterad gnathosoma (varies according to mount), respiratory system not visible in most mounts but sometimes narrow trachea and small atrium apparent. Prodorsal plate with setae v1 20 (19–24), v2 alveolar remnant, sc1 10 (9–11), sc2 38 (36–40). Distance between setae v1–v1 56 (53–60), sc1–sc1 88 (87–96), v1–sc1 31 (34–37), v2–v2 69 (68–75), sc2–sc2 95 (91–95), sc1–sc2 37 (32–36). Plate C entire, setae c1 8 (8–9), c2 8 (8–10). Plate D setae d 9 (8–10), d–d 105 (110–125), cupuli ia anterolaterad d. Plate EF setae f 9 (7–8), f–f 71 (80–87), cupuli im anterolaterad to laterad f. Plate H present as small remnant. Venter: coxisterna 1-2 poorly defined, slightly broader than apodemes; coxisterna 3 well-defined; coxal setae slender, 1a 6 (6–7), 2a 6 (6–7), 3a 6 (7–9), 3b 6 (6–7). Alveoli of 1b, 2b present.</p> <p>Legs (Fig. 46). Setal counts legs I– III, femur-tarsus: 3-2-6(+ φ)-9(+ ω), 0-1-4-7(+ ω), 0-1-4-7. Leg I. Femur I, d 2 (2), l’ 13 (13–16), v″ 14 (12–15); genu I, l’ 5 (5–6), l″ 3 (3); tibia I, d 25 (27–30), l’ 10 (7–9), l’’ 6 (6–8), v’ 7 (7–8), v’’ 14 (11–12), k 4 (4–5), φ 6 (7–8); tarsus I, tc’ 13 (11–12), tc’’ 13 (13–15), pl’ 15 (14–17), pl’’ 18 (15–20), pv′ 3 (3–4), pv’’ 3 (2–3), ω 4 (4–5), s 6 (6–7), u′′ minute (m–2), p′ 2 (m–2). Leg II. Genu II, l’ 8 (5–6); tibia II, d 16 (15–20), l’ 9 (8–9), v’ 15 (14–16), v’’ 21 (20–23); tarsus II, tc′ 9 (9–11), tc ″ 8 (8–9), pl ″ 18 (16–18), pv′ 2 (2–3), pv’’ 4 (3–4), ω 4 (4–5), u’ 9 (9–10), u″ m (m), difficult to discern next to large u′. Leg III. Genu III, l’ 5 (4–5); tibia III, d 14 (15–20), l’ 9 (7–9), v’ 12 (12–15), v’’ 17 (20–21); tarsus III, tc’ 8 (8–10), tc ″ 8 (8–9), pl ″ 20 (18–20), pv′ 4 (3–5), pv’’ 3 (3–5), u’ 8 (8–9), u″ m (m), difficult to discern next to large u′. Claws on legs I-III well developed and paired. Tarsi I-III setae pvʹ, pvʺ slender. Large blunt spur-like setae: tarsus I s; tarsi II– III u’, tcʺ. Setae u′ pointed, not bifid.</p> <p>LARVIFORM ADULT MALE (Fig. 47; n = 16).</p> <p>Gnathosoma. Length 28–37, width 29–33. Cheliceral stylets 25–28, setae ch 3–4, su 8–9.</p> <p>Idiosoma (Fig. 47). Length 175–190, width 135–155. Prodorsal plate with setae v1 4–9, v2 alveolar remnant, sc 1 m–2, sc2 27–34. Distance between setae v1–v1 24–30, sc1–sc1 61–66, v1–sc1 24–26, v2–v2 42–49, sc2–sc2 64–72, sc1–sc2 24–28. Plate CD eroded posterolaterally, setae c1 2–3, c2 2, d m–2, distance between setae c1–c1 56–65, c1–c2 32–35, d–d 42–47; cupuli ia laterad to anterolaterad setae d. Plate EF with setae f m–2, distance f–f 22–25; cupuli im laterad to anterolaterad setae f. Genital capsule length 25–27, width 35–38, setae h minute. Venter: coxal setae slender, 1a 4–5, 2a 4–5, 3a 5–6, 3b 5–6.</p> <p>Legs. Setal counts legs I– III, femur-tarsus: 2-0-6(+ φ)-8(+ ω), 0-0-4-7(+ ω), 0-0-4-7. Compared to female: Femur I seta v″ absent, genua I-III l′ and l″ absent. Leg I. Femur I, d 2, l’ 2–4; tibia I, d 19–25, l’ 2, l’’ 2–3, v ’ 2–3, v ’’ 9–10, k 2–3, φ 6–7; tarsus I, tc’ 8–9, tc’’ 10–11, pl’ 8–11, pl’’ 10–12, pv ′ 2, pv’’ 2, ω 3–4, s 5, u ′′ minute. Leg II. Tibia II, d 6–11, l’ 3–4, v’ 9–12, v’’ 11–14; tarsus II, tc′ 6–8, tc ″ 5–6, pl ″ 11–12, pv ′ 2, pv’’ 3–4, ω 3–4, u’ 6, u″ minute, difficult to discern. Leg III. Tibia III, d 2, l’ 3–5, v’ 8–11, vʺ 10–14; tarsus III, tc′ 3–4, tc ″ 6–7, pl ″ 9–11, pv ′ 2–3, pv’’ 3–5, u ’ 6–7, u″ minute, difficult to discern. Claws on leg I single, claws on leg II-III paired. Setal form similar to that in female, but with tibia I setae l′, l″ tiny spines and v′ slightly thickened.</p> <p>LARVAL FEMALE (Fig. 48).</p> <p>Gnathosoma. Length 38–46, width 31–34. Palp without alveolar remnant of femoral seta. Cheliceral stylets 36–42, setae ch 26–33, su 12–15.</p> <p>Idiosoma (Fig. 48). Length 180–225, width 135–240; larger specimens swollen evenly around body. Prodorsal plate with setae v1 15–18, v2 alveolar remnant, sc1 8–10, sc2 70–80. Distance between setae v1–v1 29–31, sc1–sc1 69–72, v1–sc1 31–33, v2–v2 50–54, sc2–sc2 72–78, sc1–sc2 33–37. Plates C and D fused medially; line of fusion visible. Plate C setae c1 7–10, c2 7–10, distance between setae c1–c1 31–35. Plate D setae d 7–9, d–d 30–35, cupuli ia anterolaterad d. Plate EF setae f 6–8, f–f 29–33, cupuli im anteriad to anterolaterad f. Plate H small, 18–22 wide, 13–18 long, bearing setae h1 ca. 100, h2 4–6. Venter: coxisterna 1-3 well-defined; coxal setae slender, 1a 4–5, 2a 4–5, 3a 7–8, 3b 5–6. Alveoli of 1b, 2b present.</p> <p>Legs. Setal counts legs I– III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6(+ ω), 0-1-4-6 (setae pv′ absent on tarsi II- III). Leg I. Femur I, d m–2, l’ 2–3, v’’ 9–13; genu I, l’ 3, l″ 2–3; tibia I, d 25–32, l’ 7–8, l’’ 7–9, v’ 4–5, v’’ 8–12, k 3–4, φ 7–8; tarsus I, tc’ 8–9, tc’’ 10–11, pl’ 8–10, pl’’ 13–15, pv′ 2–3, pv’’ 2–3, ω 4–5, s 5–6, u′′ 2. Leg II. Genu II, l’ 4–5; tibia II, d 13–17, l’ 8–9, v’ 9–12, v’’ 13–16; tarsus II, tc′ 7–8, tc ″ 5–6, pl ″ 13–17, pv’’ 4–5, ω 4–5, u’ 6–7, u′′ minute. Leg III. Genu III, l’ 3–4; tibia III, d 15–17, l’ 5–6, v’ 10–11, v’’ 16–17; tarsus III, tc’ 7–9, tc ″ 5–7, pl ″ 16–20, u’ 6–7, pv’’ 5–6, u′′ minute. Setal fprm similar to that in female.</p> <p>Note: one larva (#2) has an unusual asymmetrical reversal of tcʺ and plʺ.</p> <p>Differential diagnosis. Eutarsopolipus janus sp. nov. and E. biuncatus sp. nov. both have the unusual feature of paired claws on leg I in females. Female Eutarsopolipus janus differ by having longer setae sc2 (36–40 versus 15–20) and tarsi II-III with longer setae tc′ (8–11 versus 3–7) and tc′′ (8–9 versus 5–6). Males are even more distinctive, with seta sc2 being well-developed in E. janus (27–34) but greatly reduced in E. biuncatus (2–3).</p> <p>Etymology. The specific name janus refers to the god Janus, who symbolised beginnings and transitions, alluding to the questions regarding the evolution and speciation of this possible complex of cryptic species on several host species.</p> <p>Remarks. This species was found on four host species: the type host, N. flos, as well as N. aurifer, N. dimorphicus and N. spurgeoni. All these species are found in the Wet Tropics region of north Queensland and at least N. aurifer, N. dimorphicus and N. flos are closely related (Will 2015). Thus, these species may not have speciated when their hosts speciated, or they have done so cryptically. In support of the latter hypothesis, some minor differences were observed in N. aurifer. Females of this species differed by having: slightly longer cheliceral stylets (50–56 versus 43–50), slightly longer tibia I solenidion (6–8 versus 8–10), slightly longer seta tc′ on tarsus II (10–13 versus 8–10) and tarsus III (11–13 versus 8–10), slightly longer setae tcʺ on tarsus II (9–10 versus 8–9) and tarsus III (10–11 versus 8–9) and u′ on tarsus III (9–11 versus 8–9). Several of these differences are also apparent in the larvae. Larvae from N. aurifer have longer cheliceral stylets (43–49 versus 36–42), longer tarsal seta tc′ on tarsus III (10–12 versus 7–9) and tarsal seta tc″ on tarsus II (6–7 versus 5–6).</p> <p>The species N. aurifer is also geographically separated from the other three species: the former is from a large fragment of the Wet Tropics south-west of Cairns, while the others are from a larger fragment southwest to west of Mossman comprising Mt Lewis (N. flos), Mt Spurgeon (N. spurgeoni) and Lamb’s Head (N. dimorphicus). These regions are separated by a biogeographical barrier, the Black Mountain Corridor, which separates radiations of other endemic rainforest animals (e.g., Krosch et al. 2009; Edwards &amp; Melville 2010) and plants (e.g., Burke et al. 2013). Furthermore, N. dimorphicus and N. flos are more closely related to each other than they are to N. aurifer (Will 2015).</p> </div>	http://treatment.plazi.org/id/4438245CFFCB090A87AEFF5AFE85A5C5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Seeman, Owen D.	Seeman, Owen D. (2021): Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae). Zootaxa 4971 (1): 1-74, DOI: https://doi.org/10.11646/zootaxa.4971.1.1
