taxonID	type	description	language	source
03AD87A7FFC1773BB69E27B6FA67FAF9.taxon	description	(Figs 3, 6 – 8)	en	Winterton, Shaun L., Balakrishnan, Suryanarayanan Thangalazhi, Chenthamarakshan, Bijoy (2021): A new species of Joguina Navás, 1912 from India (Neuroptera: Chrysopidae). Zootaxa 4970 (3): 577-585, DOI: 10.11646/zootaxa.4970.3.9
03AD87A7FFC1773BB69E27B6FA67FAF9.taxon	diagnosis	Diagnosis. Antennal pedicel constricted basally; both wings with numerous cross-veins throughout; forewing subcostal veinlets with 1 – 3 semi-regular rows of cross-veins in basal half of wing, 0 – 1 row in distal half; forewing with single row of cross-veins between RA and RP; forewing RP mostly straight; cross-veins between RA and RP behind pterostigma in both wings; both wings with 1 – 2 conspicuous pustules (dark markings with associated deformation of wing membrane and veins) in discal area, pterostigma lacking dark spot; both wings with gradate series obscured by irregular cross-veins, 3 gradate series when definable; Psc extending greater than 3 / 4 of total wing length towards apex; forewing cell c 2 short and broad, poorly defined due to numerous cross-veins; 1 A and 2 A forked; posterior marginal area of forewing with 1 – 4 rows of cross-veins, hind wing with 0 – 3 rows, both wings with two rows in distal third; spermatheca with lateral striations, spermathecal duct long, tightly coiled.	en	Winterton, Shaun L., Balakrishnan, Suryanarayanan Thangalazhi, Chenthamarakshan, Bijoy (2021): A new species of Joguina Navás, 1912 from India (Neuroptera: Chrysopidae). Zootaxa 4970 (3): 577-585, DOI: 10.11646/zootaxa.4970.3.9
03AD87A7FFC1773BB69E27B6FA67FAF9.taxon	description	Included species. Joguina borneensis Kimmins, 1952 (Borneo) (Fig. 6); J. malayana Banks, 1931 (Malaysia) (Fig. 8); J. nicobarica (Brauer, 1964) (India) (Fig. 7); J. unimaculata Winterton, Balakrishnan & Chenthamaraksan sp. n. (India) (Fig. 3).	en	Winterton, Shaun L., Balakrishnan, Suryanarayanan Thangalazhi, Chenthamarakshan, Bijoy (2021): A new species of Joguina Navás, 1912 from India (Neuroptera: Chrysopidae). Zootaxa 4970 (3): 577-585, DOI: 10.11646/zootaxa.4970.3.9
03AD87A7FFC1773BB69E27B6FA67FAF9.taxon	discussion	Comments. Joguina contains four relatively similar species that differ only in the number and arrangement of anterior and posterior marginal cross-veins and in the position and number of dark wing pustules. The distinctive features of this genus are the single row of cross-veins between RA and RP basally, one to many irregular rows of cross-veins of anterior and posterior marginal cross-veins, and 1 – 2 round pustules in the wing while the pterostigmata are unmarked. Joguina is restricted to South-East Asia.	en	Winterton, Shaun L., Balakrishnan, Suryanarayanan Thangalazhi, Chenthamarakshan, Bijoy (2021): A new species of Joguina Navás, 1912 from India (Neuroptera: Chrysopidae). Zootaxa 4970 (3): 577-585, DOI: 10.11646/zootaxa.4970.3.9
03AD87A7FFC1773CB69E2286FE2FFD5D.taxon	description	(Figs 1 – 5) Zoobank: urn: lsid: zoobank. org: act: 19 A 80 A 59 - 4650 - 4 BC 5 - AC 29 - F 9 E 3 DFE 614 D 5 Common name. One-spotted delicate lacewing.	en	Winterton, Shaun L., Balakrishnan, Suryanarayanan Thangalazhi, Chenthamarakshan, Bijoy (2021): A new species of Joguina Navás, 1912 from India (Neuroptera: Chrysopidae). Zootaxa 4970 (3): 577-585, DOI: 10.11646/zootaxa.4970.3.9
03AD87A7FFC1773CB69E2286FE2FFD5D.taxon	materials_examined	Type material. Holotype male INDIA: Kerala State: Palakkad District, Pudunagaram, elev. 104 m, (coordinates: 10.6853, 76.6858), Bijoy. C., 20. XI. 2020 (specimen Reg. No. ZSI / WGRC / IRINV 15593).	en	Winterton, Shaun L., Balakrishnan, Suryanarayanan Thangalazhi, Chenthamarakshan, Bijoy (2021): A new species of Joguina Navás, 1912 from India (Neuroptera: Chrysopidae). Zootaxa 4970 (3): 577-585, DOI: 10.11646/zootaxa.4970.3.9
03AD87A7FFC1773CB69E2286FE2FFD5D.taxon	diagnosis	Diagnosis. Forewing and hind wing with single dark infuscate pustule; forewing with single row of cross-veins between subcostal veinlets in basal half of wing, absent in distal half wing; relatively few cross-veins in posterior margin of wing and never in multiple rows.	en	Winterton, Shaun L., Balakrishnan, Suryanarayanan Thangalazhi, Chenthamarakshan, Bijoy (2021): A new species of Joguina Navás, 1912 from India (Neuroptera: Chrysopidae). Zootaxa 4970 (3): 577-585, DOI: 10.11646/zootaxa.4970.3.9
03AD87A7FFC1773CB69E2286FE2FFD5D.taxon	description	Description. Male: Forewing length: 29.5 mm; hind wing length: 29.9 mm. Overall colouration when live dark to pale green with dark red markings on head, and dark bands on abdomen (Fig. 1); wing membrane hyaline except for distinct round pustules and darkening of gradate-like cross-veins (Fig. 3). Head. Dark green, clypeus, frons and vertex dark red, green posteriorly; antennal scape dark red; pedicel yellow with small dark mark; flagellum uniform yellow, longer than wing. Thorax. Prothorax uniform light green; mesothorax and metathorax entirely dark green, unmarked; legs very pale whitish-green, unmarked, tarsi light brown; forewing hyaline, venation pale green, except for several irregular series dark cross-veins corresponding to gradates and distally on Psm, single round dark-brown pustule located on basal-most gradate series close to RP, single row of cross-veins joining subcostal veinlets in basal half of forewing, cross-veins lacking in distal half; single row of cross-veins connecting veinlets along entire posterior marginal area; hind wing similarly hyaline with pale green venation, cross-veins arranged in series in distal part of wing, single row of cross-veins connecting veinlets in posterior marginal area; single small round dark-brown pustule on basal gradate series close to RP. Abdomen. Uniform light to dark-green, brown band present on tergites 2 – 6; end of abdomen slightly bulbous. Terminalia (Fig. 5). Typical of apochrysines; not dissected. Female: unknown.	en	Winterton, Shaun L., Balakrishnan, Suryanarayanan Thangalazhi, Chenthamarakshan, Bijoy (2021): A new species of Joguina Navás, 1912 from India (Neuroptera: Chrysopidae). Zootaxa 4970 (3): 577-585, DOI: 10.11646/zootaxa.4970.3.9
03AD87A7FFC1773CB69E2286FE2FFD5D.taxon	etymology	Etymology. The species name is derived from the Latin — unus, one; maculatus, spot, stain, referring to the single pustule in each wing.	en	Winterton, Shaun L., Balakrishnan, Suryanarayanan Thangalazhi, Chenthamarakshan, Bijoy (2021): A new species of Joguina Navás, 1912 from India (Neuroptera: Chrysopidae). Zootaxa 4970 (3): 577-585, DOI: 10.11646/zootaxa.4970.3.9
03AD87A7FFC1773CB69E2286FE2FFD5D.taxon	discussion	Comments. The male specimen was collected at lights in a village household. The surrounding habitat is semiclosed coconut plantation (Fig. 2) and is typical habitat for members of this subfamily. The single dark spot in each wing and the single row of cross-veins connecting subcostal veinlets in the basal half of the forewing is distinctive for this species. As noted by previous authors (Brooks & Barnard, 1990; Winterton & Brooks, 2002; Winterton & Gupta, 2020) the male genitalia are remarkably similar throughout the entire subfamily. Unfortunately, only the holotype of J. unimaculata sp. n. is known, and considering the expected lack of specific morphological characters therein, it was not dissected.	en	Winterton, Shaun L., Balakrishnan, Suryanarayanan Thangalazhi, Chenthamarakshan, Bijoy (2021): A new species of Joguina Navás, 1912 from India (Neuroptera: Chrysopidae). Zootaxa 4970 (3): 577-585, DOI: 10.11646/zootaxa.4970.3.9
03AD87A7FFC7773FB69E21D2FEF3FB29.taxon	description	(Fig. 9) Lainius Navás 1913: 300. Type species: Lainius constellatus Navás, 1913: 300.	en	Winterton, Shaun L., Balakrishnan, Suryanarayanan Thangalazhi, Chenthamarakshan, Bijoy (2021): A new species of Joguina Navás, 1912 from India (Neuroptera: Chrysopidae). Zootaxa 4970 (3): 577-585, DOI: 10.11646/zootaxa.4970.3.9
03AD87A7FFC7773FB69E21D2FEF3FB29.taxon	diagnosis	Diagnosis. Antennal pedicel constricted basally; both wings with numerous cross-veins throughout; forewing subcostal veinlets with four or more irregular rows of cross-veins in basal half of wing, 1 – 2 regular rows in distal half; numerous irregular cross-veins between RA and RP in forewing; forewing RP sigmoid in shape; no cross-veins between RA and RP behind pterostigma in both wings; forewing with numerous conspicuous pustules of varying size in discal area, two small pustules in hind wing; pterostigma with a small dark spot in both wings; both wings with gradate series obscured by irregular cross-veins; Psc extending greater than 3 / 4 of total wing length towards apex; forewing cell c 2 short and broad, poorly defined due to numerous cross-veins; 1 A and 2 A forked; posterior marginal area of both wings with multiple semi-regular rows of cross-veins, cross-veins absent in distal third; spermatheca with lateral striations, spermathecal duct long, tightly coiled. Included species. Lainius constellatus Navás, 1913 (Guatemala); L. decoratas Navás, 1930 (northern South America).	en	Winterton, Shaun L., Balakrishnan, Suryanarayanan Thangalazhi, Chenthamarakshan, Bijoy (2021): A new species of Joguina Navás, 1912 from India (Neuroptera: Chrysopidae). Zootaxa 4970 (3): 577-585, DOI: 10.11646/zootaxa.4970.3.9
