taxonID	type	description	language	source
03AA878CFA233034FF3F64E94D6E5C47.taxon	description	Tulipa albanica differt ab T. scardica et T. schrenkii foliis inferioribus valde undulatis et canaliculatis, superioribus apice longe attenuato-rostrato, tepalorum macula basali nigra absente. Ab T. scardica praeterea differt filamentis aureis (nec eburneis vel nigrescentibus), capsulis minoribus, seminibus majoribus.	en	Shuka, Lulëzim, Tan, Kit, Siljak-Yakovlev, Sonja (2010): Tulipa albanica (Liliaceae), a new species from northeastern Albania. Phytotaxa 10: 17-25, DOI: 10.11646/phytotaxa.10.1.2
03AA878CFA233034FF3F64E94D6E5C47.taxon	materials_examined	Type: — ALBANIA (Northeast Albania: UTM 34 T DM 1 and 34 T DM 3). Kukësi district: 6 km from Kolshi village to Surroj, N- and W-facing open stony serpentine slopes of Ahmet Xhindit, in clearings of Quercus pubescens woodland and Buxus sempervirens scrub, 625 m, 42 ° 02 ’ N, 20 ° 20 ’ E, 7 May 2009, flowers yellow to golden-yellow, Shuka 250 (holotype TIR, isotypes C, LD, private herbaria Kit and Shuka). Paratype: — loc. ibid., flowers scarlet, 7 May 2009, Shuka 251 (TIR, C, private herbaria Kit and Shuka).	en	Shuka, Lulëzim, Tan, Kit, Siljak-Yakovlev, Sonja (2010): Tulipa albanica (Liliaceae), a new species from northeastern Albania. Phytotaxa 10: 17-25, DOI: 10.11646/phytotaxa.10.1.2
03AA878CFA233034FF3F64E94D6E5C47.taxon	description	Perennial herb. Bulb ovoid to ovoid-globose, 1.5 – 2.5 cm in diameter, not stoloniferous; tunics dark reddish-brown, chartaceous, prolonged into short neck; inner surface densely covered with straight, 2.5 – 3.5 mm long unicellular hairs. Stem erect, 13 – 40 cm (including subterranean portion), glabrous, glaucous to greyish-green. Leaves 3 – 5, alternate, glabrous, glaucous to greyish-green, subacute to mucronulate. Lowermost leaves (at ground level) linear-lanceolate to broadly-lanceolate, patent to erecto-patent, 10 – 25 × 1 – 3.5 cm, canaliculate, strongly undulate. Upper cauline leaves much smaller, weakly canaliculate, slightly undulate, ciliate at base, narrowing and long-tapering at apex. Flowers solitary (very rarely 2), large, erect, campanulate, existing in two colour forms, yellow to golden-yellow or carmine-scarlet turning deep reddishmaroon. Perianth segments slightly unequal; outer segments elliptical to obovate, 4 – 7.3 × 2.3 – 3.6 cm, obtuse- or rounded-apiculate; inner segments elliptic-oblanceolate to obovate-spathulate, 3.5 – 7 × 2 – 4.2 cm, obtuse to subacute. Perianth in red-flowered forms with conspicuous yellow zone or blotch extending one-sixth to onethird the length of a segment, visible outside. Filaments golden-yellow, 7 – 14 (– 17) × 1.5 – 2.5 mm, glabrous, dilated at base. Anthers burgundy-red to blackish-maroon, oblong, 7 – 13 × 2 – 3.5 mm, c. equalling or shorter than filaments; pollen blackish-maroon. Ovary subsessile, light to yellowish-green, 12 – 23 mm long; stigmas 3, recurved-decurrent, rose-pink, yellow or pale green. Capsule pale brown, ellipsoid, 3 – 3.5 × 1.5 – 2 cm, apiculate, shortly stipitate, with distinct transverse veins throughout. Seeds flat, triangular-deltoid, 5.5 – 6.5 × 4.5 – 6 mm, greenish-brown, turning dark reddish-brown at maturity. Flowering late April to mid-May; capsules dehiscing by early July.	en	Shuka, Lulëzim, Tan, Kit, Siljak-Yakovlev, Sonja (2010): Tulipa albanica (Liliaceae), a new species from northeastern Albania. Phytotaxa 10: 17-25, DOI: 10.11646/phytotaxa.10.1.2
03AA878CFA223033FF3F66FA4F8D5E83.taxon	description	The serpentine habitat of Tulipa albanica is similar to that of T. scardica in Radusha (locus classicus in Macedonia) and T. scardica in Mirusha region in Kosovo. According to Mayer & Micevski (1970), plants of T. scardica show great variation in several morphological characters such as leaf form, flower colour, length of filaments and anthers in different parts of its distribution range. Our field and herbarium studies confirm this variation in and between populations. Thus specimens of T. scardica from Taorska Klisura (collected by Micevski on 9 May 1964, SKO) and specimens from Mirusha region (collected by Krasniqi on 11 May 2002, herb. Pristina) have flat, slightly undulate leaves and perianth segments with a faint black basal blotch edged pale or bright yellow. The filament colour varies from blackish in Radusha (locus classicus) to white or pale purplish in tulips from Mirusha. During our fieldwork to Mirusha region on 15 May 2010, we have observed plants without any marking on their perianth segments and plants with white basal markings on their perianth. This supports the establishment of four colour forms within the T. scardica complex by Mayer & Micevski (1970). We consider the recently described T. serbica Tatić & Krivošej (1997: 733) as falling within the range of variation of T. scardica. Specimens from Crveni Bregovi-Orlov rid (collected by Micevski on 20 May 1965, SKO) have undulate leaves and flowers with a distinct black basal blotch within and superficially resemble red-flowered forms of T. albanica, but the latter can easily be distinguished by the complete absence of a black basal blotch on its perianth segments. It is noteworthy that golden-yellow flowers, which are predominant in T. albanica (comprising more than 80 % of the population), have never been observed in T. scardica. As in Tulipa scardica, variation in the colour of the perianth and stamens has also been observed in T. schrenkii native to the steppes and semi-desert areas of Turkey, Iran, the Caucasus, Daghestan, Kazakhstan, Azarbaijan, western Siberia and the Ukraine (Botschantzeva 1982). In T. albanica we have only observed yellow and red flower forms and the stamen colour is always constant regardless of perianth colour, i. e., the filaments are always golden-yellow and the anthers dark blackish-maroon (Fig. 2 C). Tulipa albanica differs from T. scardica and T. schrenkii by its canaliculate, strongly undulate lower leaves, the upper leaves with a long-tapering apex (Fig. 3) and the absence of a black basal blotch on the perianth segments. From T. scardica it additionally differs by its golden-yellow (not whitish or blackish) filaments, smaller capsules and larger seeds. Selected morphological characters of Tulipa albanica, T. scardica and T. schrenkii are summarized in Table 1.	en	Shuka, Lulëzim, Tan, Kit, Siljak-Yakovlev, Sonja (2010): Tulipa albanica (Liliaceae), a new species from northeastern Albania. Phytotaxa 10: 17-25, DOI: 10.11646/phytotaxa.10.1.2
