taxonID	type	description	language	source
052487CFFFC0E52846C6B1DB8FECFA28.taxon	diagnosis	Diagnosis. Arthrosolaenomeridini males differ from others Chelodesminae males by the combination of the following characteristics: presence of a row of setae 2 + 2 at the apex of the head (Fig. 29 A); presence of two pairs of ventral projections in the fifth sternite; presence of lateral macrosetae, in mesal aspect, on gonocoxae (Figs 32 E – F); laminated telopodite with a cingulum on the lateral margin (Figs 34 A – C); prefemoral process of the gonopod long and thin (Figs 33 B – E). Descriptive notes. Body length between 39 − 76 mm. Gonopod: composed of a basal portion (gonocoxae), containing lateral macrosetae; and a distal portion (telopod) divided into two well-developed narrow branches: the external branch (solenomere) presented as a single branch containing a variable-position cingulum and an evident spermatic groove, and the internal branch (prefemoral process) varying in size and format according to genera.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFC0E52846C6B1DB8FECFA28.taxon	distribution	Distribution. The tribe is distributed in Brazil in the northern region (states of Pará and Tocantins), central-west region (states of Goiás, Mato Grosso and Mato Grosso do Sul) and southeast (state of São Paulo). Composition. Arthrosolaenomeris Schubart, 1943; Gangugia Schubart, 1947; Angelodesmus Schubart, 1962; Abiliodesmus n. gen.; Folcloredesmus n. gen.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFC1E52946C6B2C68F8DFA64.taxon	type_taxon	Type species: A. chapadensis Schubart, 1943, by original designation; Schubart, 1958: 205; Schubart, 1960: 453; Jeekel, 1971: 249; Hoffman, 1976: 177; Hoffman, 1980: 151.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFC1E52946C6B2C68F8DFA64.taxon	diagnosis	Diagnosis. Males of Arthrosolaenomeris differ from those of other Arthrosolaenomeridini genera by the combination of the following characters: rectangular ventral projections in the post-gonopodal sternites (Fig. 30 E) (acuminated in A. curupira n. sp. and A. caipora n. sp.); posterior edge of the gonopod aperture with a small dentiform process; presence of a secondary process in medial portion in the prefemoral process of the gonopod (Fig. 4 D – F) (absent in A. iara n. sp.); presence of a rounded extension of the gonocoxae, in oral view (Fig. 32 D); and apex of the solenomere abruptly descending (Fig. 4 D).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFC1E52946C6B2C68F8DFA64.taxon	description	Description. General characters: Body length between 47.78 mm (A. caipora n. sp.) and 69.42 mm (A. iara n. sp.). Coloration (specimens long preserved in 70 % ethanol) variable between species, ranging from ocher to reddish brown, paranota tip and posterior border of the metaterga slightly whitish (some specimens of A. pantanalensis show a purple coloration for the whole body). Gnathochilarium: lingual plate covered by several setae; mentum centrally covered by setae, with smooth edges and stipes covered by thick setae (sparsely covered in A. curupira n. sp. and A. caipora n. sp.). Body rings: cuticle slightly rough; alignment of paranota in posterior view curved ventrally (straight in A. caipora n. sp.). Sternite of the fourth segment with a pair of projections, sternite of the fifth segment with two pairs of projections covered with setae. Sternite of sixth body ring with two pairs of ventral projections; seventh body ring with a pair of ventral projections (absent in A. chapadensis) and post-gonopodal sternites with two pairs of rectangular ventral projections (acuminate in A. curupira n. sp. and A. caipora n. sp.). Paranota form: round to slightly rectangular (Figs 4 B, 8 B). Legs: podomeres with several thick ventral setae (Fig. 31 A) and granules in the tibia and tarsus (Fig. 31 F) (except in A. curupira n. sp. and A. caipora n. sp.), dorsal lobe of prefemur conspicuous of the anterior legs (Fig. 31 A) and a ventroapical projection in the prefemur of the post-gonopod legs (Fig. 31 E). Male characters: Leg pair of the third body ring with the coxae presenting the rectangular genital papilla. Gonopod aperture on seventh body ring with the posterior margin excavated and with a small process. Gonopods: gonocoxae equivalent to about half the length of the telopodite, cylindrical, without a spiniform process. Prefemoral process (Figs 4 D – F): elongate with a dorso-basal process and a secondary process in its medial portion (long in A. chapadensis, A. pantanalensis, A. saci n. sp., A. iara n. sp. and short in A. curupira n. sp., A. caipora n. sp.). Cingulum in an apical position (Fig. 4 F) (middle position in A. caipora n. sp.). Solenomere long and thin with its apex abruptly descending (Fig. 4 D – F). Female characters: Vulvae: oval-shaped, having equal proportions in lateral view, with an operculum advanced (Figs 10 A – B).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFC1E52946C6B2C68F8DFA64.taxon	distribution	Distribution. States of Mato Grosso, Mato Grosso do Sul and São Paulo (Fig. 11). Composition. Arthrosolaenomeris chapadensis Schubart, 1943; A. pantanalensis Schubart, 1943; A. saci n. sp.; A. curupira n. sp.; A. caipora n. sp.; A. iara n. sp.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFC6E52E46C6B2C6896FFB76.taxon	description	Figures 4, 10 A – B, 11	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFC6E52E46C6B2C6896FFB76.taxon	materials_examined	Additional material. BRAZIL: São Paulo: Rosana (22 ° 34 ’ 47 ” S, 53 ° 03 ’ 32 ” W), Usina Hidrelétrica Engenheiro Sérgio Motta (Porto Primavera), 1 ♂, 1 ♀ and one juvenile, 1999 – 2000, Equipe Butantan coll. (IBSP 1049).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFC6E52E46C6B2C6896FFB76.taxon	diagnosis	Diagnosis. Adult males differ from other species of the genus by the combination of the following characters: presence of indentations in the prefemoral process (Fig. 4 D) and absence of a conspicuous groove in the apex of the prefemoral process (Fig. 4 D).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFC6E52E46C6B2C6896FFB76.taxon	description	Description. Male (Holotype, MZSP 1092). Coloration (long-preserved in 70 % ethanol): Head and antennae ocher. Body ocher and paranota tip with the same color as the body (Figs 4 A – C). Legs ocher. Telson ocher. Total length: 65.98. Total width: 10.02. Collum, length 3.10, width 9.33. Antennomere lengths (1> 7): 0.71; 1.91; 1.82; 1.62; 1.56; 1.51; 0.39. Podomeres lengths (1> 7): 1.03; 1.60; 2.78; 1.52; 1.96; 1.97; 0.68. Gonopod aperture, length 1.98, width 3.35. Telson, length 0.77. Gonopod: length 2.86, width 2.07. Gonocoxae: length 1.11, width 1.02. Telopodite: length 2.77, width 1.00. Prefemoral process long (greater than or equal to the solenomere) and narrow (Figs 4 D – F). Presence of a large secondary process positioned in its medial portion (Fig. 4 D). Posterior portion to the secondary process with several indentations. Solenomere long and narrow (Fig. 4 E). Cingulum positioned centroposteriorly (Fig. 4 F). Apex of the solenomere abruptly descending (Fig. 4 D). Female (IBSP 1049). Body as in male. Total length: 57.60. Total width: 9.35. Antennomere lengths (1> 7): 0.87; 1.72; 1.54; 1.36; 1.37; 1.38; 0.33. Podomeres lengths (1> 7): 0.82; 1.26; 2.62; 1.04; 1.12; 1.63; 0.55. Telson, length 1.38. Epigyne rectangular shaped. Vulvae (Figs 10 A – B): length 1.34, width 0.68. External valve: length 0.98, width 0.34. Internal valve: length 0.94, width 0.28. Operculum: length 0.39, width. 0.56.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFC6E52E46C6B2C6896FFB76.taxon	distribution	Distribution. States of Mato Grosso do Sul and São Paulo, Brazil (Fig. 11).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFC6E52D46C6B6828E61FE26.taxon	description	Figures 5, 10 C – D, 11	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFC6E52D46C6B6828E61FE26.taxon	materials_examined	Additional material. BRAZIL: Mato Grosso: Cáceres (16 ° 04 ’ 15 ” S, 57 ° 40 ’ 44 ” W), 18 ind., XII. 1997, M. Callefo coll. (IBSP 530); 7 ♂, X. 1917, E. Garbe coll. (MZSP 410); Porto Estrela (15 ° 19 ’ 28 ” S, 57 ° 13 ’ 39 ” W), Estação Ecológica Serra das Araras, 2 ♂ and 2 ♀, 12. XI. 2017, T. F. Conceição coll. (CZUFMT 847); 2 ♂, 19. XI. 2017, N. Barrozo coll. (CZUFMT 848); 5 ♂, 19. XI. 2017, T. F. Conceição coll. (CZUFMT 842); 1 ♂, 16. XI. 2015, A. Chagas-Jr and A. B. Kury coll. (CZUFMT 845); Chapada dos Guimarães (15 ° 27 ’ 39 ” S, 55 ° 44 ’ 58 ” W), Chapada Aventura, 2 ♂ and 1 ♀, 07. XI. 2015, A. Chagas-Jr et al. col. (CZUFMT 840); Fazenda Morro da Lage, 1 ♂, 28. X. 2017, M. Moreno coll. (CZUFMT 846); Cuiabá (15 ° 35 ’ 47 ” S, 56 ° 05 ’ 48 ” W), Bairro Nova Esperança I, 1 ♂, 17. XII. 2015, E. Santos coll. (CZUFMT 844); campus UFMT, 1 ♂, Marques and Hallis coll. (CZUFMT 832); Sede EESA — Arquidiocesana de Cuiabá, 4 ♀, 16. XI. 2015 (CZUFMT 843); Nova Xavantina (14 ° 40 ’ 24 ” S, 52 ° 21 ’ 11 ” W), 2 ♂ and 2 ♀, XII. 1946, H. Sick coll. (MZSP 1087); 1 ♂, XII. 1946, H. Sick coll. (MZSP 1088).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFC6E52D46C6B6828E61FE26.taxon	diagnosis	Diagnosis. Adult males differ from all other species of the genus by the combination of the following characters: absence of indentations in the prefemoral process (Fig. 5 D) and a long secondary process in the prefemoral process (Figs 5 D – F).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFC6E52D46C6B6828E61FE26.taxon	description	Description. Male (Holotype, MZSP 1085). Coloration (long-preserved in 70 % ethanol): head reddish yellow and antennae ocher. Body reddish brown and paranota tip with the same color as the body (Figs 5 A – C). Legs reddish brown. Telson ocher. Total length: 60.16. Total width: 9.05. Collum, length 2.73, width 8.74. Antennomere lengths (1> 7): 0.77; 1.80; 1.82; 1.45; 1.43; 1.40; 0.42. Podomere lengths (1> 7): 1.02; 1.26; 2.25; 1.32; 1.60; 1.57; 0.52. Gonopod aperture, length 1.76, width 2.86. Telson, length 1.12. Gonopod: length 2.90, width 2.17. Gonocoxae: length 1.01, width 1.14. Telopodite: length 2.76, width 1.06. Prefemoral process long (greater than the solenomere) and narrow (Fig. 5 E). Presence of a large secondary process positioned in its medial portion (Fig. 5 D). Posterior portion to the secondary process without indentations. Solenomere long and narrow (Fig. 5 F). Cingulum positioned centroposteriorly (Fig. 5 F). Apex of the solenomere abruptly descending (Fig. 5 D). Female (CZUFMT 847). Body as in male. Total length: 50.48. Total width: 6.92. Antennomere lengths (1> 7): 0.54; 1.39; 1.12; 1.06; 1.13; 1.04; 0.25. Podomeres lengths (1> 7): 0.62; 0.82; 1.87; 0.71; 0.76; 1.22; 0.39. Telson, length 0.97. Epigyne triangular shaped. Vulvae (Fig. 10 C – D): length 1.06, width 0.55. External valve: length 0.76, width 0.24. Internal valve: length 0.69, width 0.26. Operculum: length 0.30, width 0.50.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFC6E52D46C6B6828E61FE26.taxon	distribution	Distribution. State of Mato Grosso, Brazil (Fig. 11).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFC5E52D46C6B0448C46F933.taxon	description	Figures 6, 10 E – F, 11 urn: lsid: zoobank. org: act: 6 DDB 84 AA- 29 B 2 - 4237 - 8 F 86 - E 3 C 9647 FB 57 E	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFC5E52D46C6B0448C46F933.taxon	materials_examined	Type material. Male holotype from Bonito (21 ° 07 ’ 16 ” S, 56 ° 28 ’ 55 ” W), Mato Grosso do Sul, Brazil, 14 – 23. X. 2002, Equipe Biota coll., deposited in IBSP 2571. Paratypes: four males and one female with the same data of the holotype, deposited in IBSP 2577 (female), IBSP 2568 (two males) and IBSP 2581 (two males). Additional material. BRAZIL: Mato Grosso do Sul: Bonito (21 ° 07 ’ 16 ” S, 56 ° 28 ’ 55 ” W), 1 ♂, 1 ♀ and one juvenile, 14. X. 1993, A. Eterovic coll. (IBSP 1148); Miranda (20 ° 14 ’ 01 ” S, 56 ° 21 ’ 59 ” W), Fazenda Cayman, 1 ♂, X. 1992, A. Eterovic coll. (IBSP 7538).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFC5E52D46C6B0448C46F933.taxon	diagnosis	Diagnosis. Adult males differ from all other species of the genus by the combination of the following characters: presence of indentations in the prefemoral process (Fig. 6 D) and presence of a conspicuous groove in the apex of the prefemoral process (Fig. 6 D).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFC5E52D46C6B0448C46F933.taxon	description	Description. Male (Holotype, IBSP 2571). Coloration (long-preserved in 70 % ethanol): Head ocher, with brownish spots. Antennae ocher becoming more whitish along each antennomere towards the apex, the seventh antennomere totally brownish. Body reddish brown (Figs 6 A – C). In the anterior segments the posterior portion of the metazonite is ochre and the edge of the paranota with the same coloration as the body, except for the slightly clearer peritremata. Legs ochre. Telson reddish brown. Total length: 64.22. Total width: 10.09. Collum, length 2.52, width, 9.54. Antennomere lengths (1> 7): 0.69; 1.87; 1.92; 1.64; 1.68; 1.40; 0.32. Podomeres lengths (1> 7): 1.00; 1.41; 2.69; 1.49; 1.81; 1.85; 0.60. Gonopod aperture, length 1.80, width 3.27. Telson, length 1.31. Gonopod: length 2.97, width 2.19. Gonocoxae: length 1.02, width 1.34. Telopodite: length 2.84, width 0.84. Prefemoral region 1 / 3 the size of telopodite. Prefemoral process (Figs 6 D – E) long, presenting a dorso-basal lobe, starting from an angle of 135 °, ascending parallel to the solenomere and partially covering it in mesal view, in the middle part of the process it presents a long secondary process with dagger-shape, with a series of indentations above the secondary process (Fig. 6 D). The terminal part presents a conspicuous concavity, the process ending in an acute apex (Fig. 6 D). Solenomere long and narrow (Fig. 6 F). Cingulum positioned centroposteriorly (Fig. 6 F). Apex of the solenomere acute and abruptly descending (Figs 6 D, F). Female (Paratype, IBSP 2577). Body as in male. Total length: 56.58. Total width: 9.15. Epigyne triangular shaped. Vulva (Fig. 10 E – F): length 1.35, width 0.68. External valve: length 0.91, width 0.32. Internal valve: length 0.90, width 0.34. Operculum: length 0.50, width 0.55.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFC5E52D46C6B0448C46F933.taxon	distribution	Distribution. Mato Grosso do Sul, Brazil (Fig. 11).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFC5E52D46C6B0448C46F933.taxon	etymology	Etymology. The species epithet refers to the Brazilian folkloric character “ Saci ”. According to the indigenous legend, Saci is a boy with only one leg, characterized by his antics, who smokes a pipe and carries a red cap. Noun in apposition.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFC5E53046C6B5598901FEEA.taxon	description	Figures 7, 11 urn: lsid: zoobank. org: act: 5 C 4 BD 0 F 8 - 892 D- 4 C 88 - 9 D 0 B- 24460 FE 62524	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFC5E53046C6B5598901FEEA.taxon	materials_examined	Type material. Male holotype from Nova Xavantina (14 ° 40 ’ 24 ” S, 52 ° 21 ’ 11 ” W), Mato Grosso, Brazil, XI. 1949, Instituto Butantan Expedition coll., deposited in IBSP 59. Paratype: one male with the same data of the holotype, deposited in IBSP 7786.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFC5E53046C6B5598901FEEA.taxon	diagnosis	Diagnosis. Adult males differ from all other species of the genus by the combination of the following characters: short (about half the length of secondary processes categorized as “ elongated ”) secondary process of the prefemoral process (Fig. 7 D) and presence of a lateral row of macrosetae in the gonocoxae, in mesal view (Fig. 7 D).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFC5E53046C6B5598901FEEA.taxon	description	Description. Male (Holotype, IBSP 59). Coloration (long-preserved in 70 % ethanol): Head and antennae ochre. Body ocher with the edge of the paranota yellow whitish (Figs 7 A – C). Legs ocher whitish. Telson ocher. Total length: 59.40. Total width: 9.76. Collum, length 2.98, width 9.56 Antennomere lengths (1> 7): 0.69; 1.85; 1.50; 1.37; 1.38; 1.55; 0.37. Podomeres lengths (1> 7): 0.89; 1.33; 2.84; 1.18; 1.59; 1.99; 0.45. Gonopod aperture, length 1.55, width 3.03. Telson, length 1.68. Gonopod: length 3.17, width 1.88. Gonocoxae: length 0.90, width 1.03. Telopodite: length 3.04, width 0.90. Prefemoral region 1 / 3 the size of telopodite. Prefemoral process (Fig. 7 D) length, broad and without indentations, presenting a dorso-basal lobe, starting from an angle of 135 °, ascending parallel to the solenomere and partially covering it in mesal view, in the middle part of the process presents a short trianguliform secondary process, acute apex (Fig. 7 D). Solenomere long and narrow (Fig. 7 F). Cingulum positioned centroposteriorly (Fig. 7 F). Apex of the solenomere acute and abruptly descending (Fig. 7 F). Female. Unknown.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFC5E53046C6B5598901FEEA.taxon	distribution	Distribution. Known only from the type locality (Fig. 11).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFC5E53046C6B5598901FEEA.taxon	etymology	Etymology. The species epithet is a reference of the Brazilian folkloric character “ Curupira ”. According to the indigenous legend, Curupira is the protector of the Brazilian forests. Noun in apposition.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFD8E53146C6B33088EDFC52.taxon	description	Figures 8, 11 urn: lsid: zoobank. org: act: 7 CE 267 B 4 - BDD 7 - 47 C 1 - 8 A 11 - C 17 E 03 AABF 02	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFD8E53146C6B33088EDFC52.taxon	materials_examined	Type material. Male holotype from Usina Hidrelétrica Guaporé (15 ° 07 ’ 00 ” S, 58 ° 58 ’ 00 ” W), Vale São Domingos and Pontes de Lacerda, Mato Grosso, Brazil, 08. X. 2002, I. Knysak coll., deposited in IBSP 1569.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFD8E53146C6B33088EDFC52.taxon	diagnosis	Diagnosis. Adult males differ from all other species of the genus by the combination of the following characters: short (about half the length of secondary processes categorized as “ elongated ”) secondary process of the prefemoral process (Fig. 8 D), absence of a lateral row of macrosetae in the gonocoxae, in mesal view (Fig. 8 D) and prefemoral process with a more flat aspect in relation to the other species of the genus (Fig. 8 E).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFD8E53146C6B33088EDFC52.taxon	description	Description. Male (Holotype, IBSP 1569). Coloration (long-preserved in 70 % ethanol): Head and antennae ocher whitish. Body ocher and paranota tip with the same color as the body (Figs 8 A – C). Legs ocher whitish. Telson ocher. Total length: 47.78. Total width: 7.06. Collum, length 2.07, width 6.69. Antennomere lengths (1> 7): 0.60; 1.34; 1.39; 1.26; 1.44; 1.37; 0.20. Podomeres lengths (1> 7): 0.67; 1.00; 1.96; 0.848; 1.09; 1.48; 0.47. Gonopod aperture, length 1.23, width 2.08. Telson, length 1.07. Gonopod: length 2.00, width 1.65. Gonocoxae: length 0.74, width 1.03. Telopodite: length 1.98, width 0.63. Prefemoral region 1 / 3 the size of telopodite. Prefemoral process (Figs 8 D – F) long and ventrally wide, presenting a dorso-basal lobe, starting from an angle of 135 °, ascending parallel to the solenomere and partially covering it in ventral view, in the middle part of the process presents a short secondary process in oval form (Fig. 8 D). Prefemoral process ending in an acute apex (Figs 8 D – E). Solenomere long, but not reaching the apex of the prefemoral process (Fig. 8 F). Cingulum in medial position (Fig. 8 F). Apex of the solenomere abruptly descending (Fig. 8 D). Female. Unknown.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFD8E53146C6B33088EDFC52.taxon	distribution	Distribution. Known only from the type locality (Fig. 11).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFD8E53146C6B33088EDFC52.taxon	etymology	Etymology. The species epithet is a reference of the Brazilian folkloric character “ Caipora ”. According to the indigenous legend, Caipora is considered the protector of the animals and guardian of the forests. Its origin is in the Tupi-Guarani indigenous mythology and means “ inhabitant of the bush ” in the Tupi language. Noun in apposition.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFD9E53646C6B1A78C62F84B.taxon	description	Figures 9, 10 G – H, 11 urn: lsid: zoobank. org: act: AEBF 860 A- 7309 - 4 E 68 - BE 80 - F 2 A 27710 CDD 0	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFD9E53646C6B1A78C62F84B.taxon	materials_examined	Type material. Male holotype from Usina Hidrelétrica Guaporé, Vale São Domingos and Pontes de Lacerda (15 ° 11 ’ 60 ” S, 59 ° 22 ’ 00 ” W), Mato Grosso, Brazil, 03. X. 2002, G. Marçal coll., deposited in IBSP 1577. Paratypes: one male and one female from Chapada dos Parecis, Diamantino (14 ° 09 ’ 43 ” S, 57 ° 08 ’ 01 ” W), Mato Grosso, Brazil, XI. 1993, D. Novais and R. Pardini coll., deposited in IBSP 7537. Additional material. BRAZIL: Mato Grosso: Vila Bela da Santíssima Trindade (15 ° 00 ’ 00 ” S, 59 ° 57 ’ 00 ” W), Fazenda Barranco Alto, 4 ♂, 2 ♀ and one juvenile, 14. XI. 2015, A. Chagas-Jr and A. B. Kury coll. (CZUFMT 841); Porto Esperidião (15 ° 51 ’ 11 ” S, 58 ° 27 ’ 36 ” W), 1 ♂, 18 – 22. XII. 1976, P. E. Vanzolini coll. (MZSP 1081).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFD9E53646C6B1A78C62F84B.taxon	diagnosis	Diagnosis. Adult males differ from all other species of the genus by the absence of the secondary process in the prefemoral process (Fig. 9 D).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFD9E53646C6B1A78C62F84B.taxon	description	Description. Male (Holotype, IBSP 1577). Coloration (long-preserved in 70 % ethanol): Head and antennae ochre with the seventh antennomere darker. Body ocher and paranota tip with the same color as the body (Figs 9 A – C). Legs ocher whitish. Telson ocher. Total length: 68.40. Total width: 10.13 mm. Collum 2.93 length, 9.89 width. Antennomere lengths (1> 7): 0.77; 2.12; 1.93; 1.62; 1.70; 1.49; 0.44. Podomeres lengths (1> 7): 1.24; 1.70; 2.64; 1.51; 1.96; 2.00; 0.56. Gonopod aperture 2.13 length, 3.83 width. Telson 1.43 length. Gonopod: 2.92 length, 2.90 width. Gonocoxae: 1.18 length, 1.50 width. Telopodite: 2.92 length, 1.48 width. Prefemoral region 1 / 3 the size of telopodite. Prefemoral process long and acute, starting from an angle of 180 °, ascending parallel to the solenomere and partially covering it in mesal view (Fig. 9 D). Solenomere long, wide in the middle zone due to the presence of the cingulum and narrowing at the apex (Fig. 9 F). Cingulum in medial position (Fig. 9 F). Apex of the solenomere abruptly descending (Fig. 9 D). Female (paratype, IBSP 7537). Body as in male, except for sternites with triangular projections, legs showing dorsal projections not conspicuous and without granules. Total length: 69.42. Total width: 9.49. Collum 3.15 length, 8.82 width. Antennomere lengths (1> 7): 0.73; 1.95; 1,45; 1.30; 1.35; 1.22; 0.35. Podomeres lengths (1> 7): 0.83; 1.06; 2.06; 1.04; 1.44; 1.50; 0.46. Telson 1.46 length. Epigyne rounded with irregular edges. Vulvae (Figs 10 G – H): 1.42 length, 0.67 width. External valve: 1.01 length, 0.30 width. Internal valve: 0.99 length, 0.28 width. Operculum: 0.44 length, 0.54 width.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFD9E53646C6B1A78C62F84B.taxon	distribution	Distribution. Mato Grosso, Brazil (Fig. 11).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFD9E53646C6B1A78C62F84B.taxon	etymology	Etymology. The species epithet is a reference of the Brazilian folkloric character “ Iara ”. According to the indigenous legend, Iara is a beautiful mermaid that inhabits the rivers of the north of the country. Noun in apposition.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFDCE53546C6B7E08F1CFCE2.taxon	type_taxon	Type species: G. tapirapensis Schubart, 1947, by original designation; Schubart, 1958: 205; Jeekel, 1971: 264; Hoffman, 1976: 177; Hoffman, 1980: 151.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFDCE53546C6B7E08F1CFCE2.taxon	diagnosis	Diagnosis. Males of Gangugia differ from those of other Arthrosolaenomeridini genera by the combination of the following characters: gonocoxae globose (Fig. 32 B); prefemoral process 225 ° descending in its initial portion (Fig. 12 D) (except G. cuca n. sp.); prefemoral process narrow and sharply curved, in ventral view (Fig. 12 E).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFDCE53546C6B7E08F1CFCE2.taxon	description	Description. General characters: Body length between 39.07 mm (G. boitata n. sp.) and 69.42 mm (G. mula n. sp.). Coloration (specimens long preserved in 70 % ethanol) variable between species, ranging from ocher to brown, in general the paranota tip and posterior border of the metaterga slightly whitish (G. cuca n. sp. show the coloration of the prozonite reddish brown). Gnathochilarium: lingual plate covered by several setae; mentum only centrally covered by setae, with smooth edges and stipes sparsely covered. Body rings: cuticle slightly rough; alignment of paranota in posterior view curved ventrally (straight in G. mula n. sp.). Sternite of the fourth segment with a pair of projections, sternite of the fifth segment with two pairs of projections covered with setae and sixth segment with two pairs of intumescences also covered by setae, the posterior pair being less conspicuous; absence of a pair on the seventh sternite (present in G. boto n. sp.) and post-gonopodal sternites with two pairs of small rounded ventral projections (Fig. 30 D) (acuminate in G. boitata n. sp.; G. cuca n. sp.). Paranota form: round to slightly rectangular (Figs 12 B, 14 B). Legs: podomeres without thick ventral setae, dorsal lobe of prefemur conspicuous, ventral projection on the coxae (Fig. 31 B) present only in G. boto n. sp. and G. mula n. sp., presence of granules in the tibia and tarsus (Fig. 31 F) (absent in G. cuca n. sp.) and a ventroapical projection in the prefemur of the postgonopod legs (Fig. 31 E). Male characters: Leg pair of the third body ring with the coxae presenting the rectangular genital papilla. Posterior edge of the gonopod aperture excavated and smooth. Gonopods: gonocoxae equivalent to about half the length of the telopodite, globose, without a spiniform process. Prefemoral process (Figs 12 D – F): elongate and thin, completely covering the solenomere in ectal view (except in G. simplex and G. cuca n. sp.) presence of a dorsobasal process (rounded in Gangugia tapirapensis, G. boto n. sp., G. mula n. sp. and sub-triangular in Gangugia simplex, G. boitata n. sp., G. cuca n. sp.) and a basal secondary process is present in G. boto n. sp., G. mula n. sp.). Cingulum in basal position (middle position in G. simplex and G. cuca n. sp.). Solenomere long and thin with a hook-shaped apex (Figs 13 D – F). Female characters: Vulvae: oval-shaped. In lateral view: rounded in G. mula n. sp. and rectangular in G. tapirapensis.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFDCE53546C6B7E08F1CFCE2.taxon	distribution	Distribution. States of Pará, Tocantins, Goiás and Mato Grosso, Brazil (Fig. 18). Composition. Gangugia tapirapensis Schubart, 1947; Gangugia simplex Schubart, 1958; G. boitata n. sp.; G. cuca n. sp.; G. boto n. sp.; G. mula n. sp.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFDDE53B46C6B74D8E2DFE7E.taxon	description	Figures 12, 18, 24 A – B	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFDDE53B46C6B74D8E2DFE7E.taxon	materials_examined	Additional material. BRAZIL: Mato Grosso: Barra do Tapirapé (Now known as Santa Terezinha); (- 10.471803 °; - 50.515290 °); 3 ♂, 2 ♀ and one juvenile, XI – XII. 1939, A. L. de Carvalho coll. (MNRJ 12008); 1 ♂, I – II. 1940, A. L. de Carvalho coll. (MNRJ 11821); four juveniles, xii. 1839 – ii. 1940, A. L. de Carvalho coll. (MNRJ 11820, destroyed after the Museu Nacional fire in 2018); 84 juveniles (MNRJ 11677, destroyed after the Museu Nacional fire in 2018); 94 individuals, xi – xii. 1839, A. L. de Carvalho coll. (MNRJ 11679, destroyed after the Museu Nacional fire in 2018); 42 individuals, (MNRJ 11680, destroyed after the Museu Nacional fire in 2018); 1 ♂ and 1 ♀, I. 1940, A. L. de Carvalho coll. (MZSP 1072); 1 ♀, I – II. 1940, A. L. de Carvalho coll. (MZSP 1073).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFDDE53B46C6B74D8E2DFE7E.taxon	diagnosis	Diagnosis. Adult males differ from all other species of the genus by the combination of the following characters: presence of indentations in the prefemoral process (Fig. 12 D) and presence of a sigmoid curvature in the apex of the solenomere (Fig. 12 F).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFDDE53B46C6B74D8E2DFE7E.taxon	description	Description. Male (Lectotype, MZSP 1076). Coloration (long-preserved in 70 % ethanol): Head and antennae ochre. Body ocher and with the lateral border of the paranota and the posterior edge of the metaterga more whitish (Figs 12 A – C). Legs ochre. Telson ocher. Total length: 62.08. Total width: 9.78. Collum, length 2.82, width 9.11. Antennomere lengths (1> 7): 0.81; 2.19; 1.57; 1.45; 1.37; 1.43; 0.37. Podomeres lengths (1> 7): 1.03; 1.30; 2.83; 1.25; 1.58; 1.69; 0.47. Gonopod aperture, length 2.00, width 4.32. Telson, length 1.22. Gonopod: length 2.97, width 2.79. Gonocoxae: length 1.37, width 0.98. Telopodite: length 2.25, width 1.85. Prefemoral process long (larger than the solenomere), narrow and curved, terminal portion presenting numerous indentations (Figs 12 D – E). Solenomere long, narrow and curved (Fig. 12 D). Cingulum in basal position (Fig. 12 F). Apex of the solenomere with descending sigmoid curvature (Fig. 12 F). Female (Paralectotype, MZSP 1070). Body as in male, except for the legs; without modifications, with only the presence of a slight dorsal lobe in the prefemur of the anterior legs. Total length: 56.19. Total width: 9.00. Antennomere lengths (1> 7): 0.65; 1.67; 1.29; 1.22; 1.23; 1.16; 0.36. Podomeres lengths (1> 7): 0.69; 1.08; 2.61; 0.92; 1.03; 1.47; 0.43. Telson, length 1.06. Epigyne triangular with irregular edges. Vulvae (Figs 24 A – B): length 1.57, width 0.83. External valve: length 1.12, width 0.37. Internal valve: length 1.11, width 0.42. Operculum: length 0.45, width 0.63.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFDDE53B46C6B74D8E2DFE7E.taxon	distribution	Distribution. Known only from the type locality (Fig. 18).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFD3E53B46C6B39C8F1DFAE8.taxon	description	Figures 13, 18	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFD3E53B46C6B39C8F1DFAE8.taxon	materials_examined	Additional material. BRAZIL: Goiás: Jataí (- 10.471803 °; - 50.515290 °), Fazenda Nova Orlândia, 2 ♂ and 1 ♀, I. 1964, Exped. Dep. Zool. coll. (VMNH 110614).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFD3E53B46C6B39C8F1DFAE8.taxon	diagnosis	Diagnosis. Adult males differ from all other species of the genus by the combination of the following characters: presence of indentations in the prefemoral process (Fig. 13 D) and the prefemoral process not completely involving the solenomere, in ectal view (Fig. 13 E – F).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFD3E53B46C6B39C8F1DFAE8.taxon	description	Description. Male (Holotype, MZSP 1077). Coloration (long-preserved in 70 % ethanol): Head and antennae ocher yellowish. Body ocher and with the border of the paranota and posterior margin of the metaterga slightly whitened (Figs 13 A – C). Legs ochre yellowish. Gonopod aperture on seventh body ring broken. Telson ocher. Total length: 65.67. Total width: 9.43. Collum, length 2.55, width 8.75. Antennomere lengths (1> 7): 0.74; 2.00; 1.78; 1.47; 1.36; 1.47; 0.30. Podomeres lengths (1> 7): 1.08; 1.44; 2.61; 1.24; 1.50; 1.48; 0.55. Gonopod aperture, length 1.80, width 3.73. Telson, length 0.93. Gonopod: length 2.98, width 2.39. Gonocoxae: length 1.17, width 1.09. Telopodite: length 2.83, width 1.34. Prefemoral process long (smaller than the solenomere), narrow and curved, terminal portion presenting several indentations (Fig. 13 D). Long, narrow and curved solenomere (Figs 13 E – F). Cingulum in medial position (Fig. 13 F). Apex of the solenomere sickle-shaped (Fig. 13 F). Female (VMNH 110614). See Hoffman (1976): 179, for more details.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFD3E53B46C6B39C8F1DFAE8.taxon	distribution	Distribution. Goiás, Brazil (Fig. 18).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFD3E53846C6B7358F2AFE7E.taxon	description	Figures 14, 18 urn: lsid: zoobank. org: act: 47505136 - DC 93 - 40 BB-A 9 A 8 - E 0821 B 2 F 7 B 03	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFD3E53846C6B7358F2AFE7E.taxon	materials_examined	Type material. Male holotype from Parque Nacional das Emas (18 ° 22 ’ 01 ” S, 52 ° 47 ’ 59 ” W), Mineiros, Goiás, Brazil, X. 1992, M. Barroso coll., deposited in IBSP 1061.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFD3E53846C6B7358F2AFE7E.taxon	diagnosis	Diagnosis. Adult males differ from all other species of the genus by the combination of the following characters: gonocoxae conspicuously greater than half the size of the telopodite (Fig. 14 D); absence of indentations in the prefemoral process (Fig. 14 D); absence of a secondary process of the prefemoral process and presence of a lobe in the solenomere, visible in ventral view (Fig. 14 E).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFD3E53846C6B7358F2AFE7E.taxon	description	Description. Male (Holotype, IBSP 1061). Coloration (long-preserved in 70 % ethanol): Head and antennae ochre. Body ocher and paranota tip with the same color as the body (Figs 14 A – C). Legs ocher whitish. Telson ocher. Total length: 39.07. Total width: 5.57. Collum, length 2.30, width 5.19. Antennomere lengths (1> 7): 0.48; 1.35; 1.10; 1.06; 0.99; 1.00; 0.24. Podomeres lengths (1> 7): 0.66; 0.88; 1.50; 0.70; 0.89; 1.10; 0.33. Gonopod aperture, length 1.33, width 2.19. Telson, length 1.05. Gonopod: length 1.58, width 1.68. Gonocoxae: length 0.96, width 1.06. Gonocoxae globose, about 1 / 2 the size of the telopodite. Telopodite: length 1.44, width 0.74. Prefemoral region short, 1 / 3 the size of telopodite. Prefemoral process long and falciform, presenting a dorso-basal lobe, starting from an angle of 225 °, involving the solenomere in its initial portion, subsequently ascending parallel to the solenomere and partially covering it in ectal view (Figs 14 D – F). Solenomere long and falciform, not reaching the apex of the prefemoral process (Fig. 14 E). Cingulum in basal position (Fig. 14 F). Median portion of the solenomere with a rounded lobe, visible in ventral view (Fig. 14 E). Apex of the solenomere sickle-shaped (Fig. 14 E). Female. Unknown.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFD3E53846C6B7358F2AFE7E.taxon	distribution	Distribution. Known only from the type locality (Fig. 18).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFD3E53846C6B7358F2AFE7E.taxon	etymology	Etymology. The species epithet is a reference of the Brazilian folkloric character “ Boitatá ”. According to the indigenous legend, Boitatá is a large fire serpent that protects the animals and the forests from people who try to burn the vegetation. Noun in apposition.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFD1E53F46C6B50B8F11FCE2.taxon	description	Figures 15, 18 urn: lsid: zoobank. org: act: 8 D 16558 E- 85 A 5 - 48 E 6 - B 402 - 24 B 2 CA 2 C 841 A	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFD1E53F46C6B50B8F11FCE2.taxon	materials_examined	Type material. Male holotype from Fazenda Sandoval, Porto Nacional (10 ° 42 ’ 30 ” S, 48 ° 25 ’ 01 ” W), Tocantins, Brazil, 28. XI. 2013, A. Chagas-Jr and A. Giupponi coll., deposited in IBSP 7600. Paratype: one male with the same data of the holotype, deposited in CZUFMT 838.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFD1E53F46C6B50B8F11FCE2.taxon	diagnosis	Diagnosis. Adult males differ from all other species of the genus by the combination of the following characters: absence of indentations in the prefemoral process (Figs 15 D); prefemoral process not reaching the apex of the solenomere (Figs 15 D – F) and the prefemoral process not completely involving the solenomere, in ectal view (Figs 15 E – F).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFD1E53F46C6B50B8F11FCE2.taxon	description	Description. Male (Holotype, IBSP 7600). Coloration (long-preserved in 70 % ethanol): Head ocher, with area around the antennae and the dorsal groove of the head with brownish coloration. Antennae ocher with the dorsal region of each antennomere presenting brown coloration, the seventh antennomere totally brownish. Body presenting metazonite with its anterior portion reddish brown and the posterior portion yellowish whitish, prozonite presenting the anterior portion whitish yellow and the posterior portion brown reddish (Figs 15 A – C). Legs whitish yellow. Telson whitish yellow. Total length: 51.57. Total width: 8.31. Collum, length 2.93, width 8.00. Antennomere lengths (1> 7): 0.69; 1.63; 1.65; 1.50; 1.58; 1.40; 0.30. Podomeres lengths (1> 7): 0.83; 1.63; 2.35; 0.97; 1.18; 1.44; 0.50. Gonopod aperture, length 1.42, width 2.91. Telson, length 0.97. Gonopod: length 2.33, width 1.75. Gonocoxae: length 0.83, width 1.05. Telopodite: length 2.23, width 0.78. Prefemoral region 1 / 3 the size of telopodite. Prefemoral process as a short blade, elliptical and acute, presenting a dorso-basal lobe, starting from an angle of 180 °, ascending parallel to the solenomere and partially covering it in mesal view (Figs 15 D – E). Solenomere long, broad in the middle portion due to the presence of the cingulum and narrowing at apex zone (Figs 15 D – E). Cingulum in medial position (Fig. 15 F). Apex of the solenomere sickle-shaped (Figs 15 E – F). Female. Unknown.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFD1E53F46C6B50B8F11FCE2.taxon	distribution	Distribution. Known only from the type locality (Fig. 18).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFD1E53F46C6B50B8F11FCE2.taxon	etymology	Etymology. The species epithet is a reference of the Brazilian folkloric character “ Cuca ”. According to the legend from Portuguese culture origin, it is a frightening old witch composed by an alligator head, and kidnaps disobedient children. Noun in apposition.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFD7E53F46C6B1368812F862.taxon	description	Figures 16, 18 urn: lsid: zoobank. org: act: 114 FC 306 - 4126 - 449 F- 812 F- 80 A 07 A 6 E 83 E 0	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFD7E53F46C6B1368812F862.taxon	materials_examined	Type material. Male holotype from FLONA Carajás (06 ° 00 ’ 32 ” S, 49 ° 58 ’ 13 ” W), Pará, Brazil, 19 – 24. i. 2012, R. Andrade et al. coll., deposited in IBSP 7539.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFD7E53F46C6B1368812F862.taxon	diagnosis	Diagnosis. Adult males differ from all other species of the genus by the combination of the following characters: legs from the fifth to the eighth segment with ventral projections in the coxae (Fig. 31 B); presence of a sigmoid curvature at the apex of the solenomere (Fig. 16 D) and presence of an elongated secondary process in the prefemoral process (Figs 16 D – E).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFD7E53F46C6B1368812F862.taxon	description	Description. Male (Holotype, IBSP 7539). Coloration (long-preserved in 70 % ethanol): Head ocher. Antennae ocher yellowish. Body brown, with the lateral border of the paranota and the posterior edge of the metaterga yellowish ocher (Figs 16 A – B). Legs ocher. Legs from the fifth to the eighth segment with ventral projections in the coxae being the posterior pair of the sixth segment extremely elongated. Gonopod aperture on seventh body ring broad (Fig. 16 C); the posterior margin excavated presenting a central sub triangular support. Telson lost. Partial length: 48.13 (until segment 16). Total width: 10.76. Collum, length 3.27, width 10.56. Antennomere lengths (1> 7): 0.96; 2.13; 1.69; 1.65; 1.52; 1.69; 0.40. Podomeres lengths (1> 7): 1.17; 1.27; 3.03; 1.28; 1.92; 2.00; 0.50. Gonopod aperture, length 2.90, width 4.55. Gonopod: length 3.69, width 2.64. Gonocoxae: length 1.70, width 1.17. Gonocoxae globose, about 1 / 2 the size of the telopodite. Telopodite: length 3.15, width 1.45. Prefemoral region short (Fig. 16 D). Prefemoral process (Figs 16 D – F) long and curved, presenting a dorso-basal lobe, starting from an angle of 225 °, involving the solenomere in its initial portion and subsequently ascending parallel to the solenomere and partially covering it in ectal view (Fig. 16 F); in its anterior portion presents an elongated, thin and acuminate secondary process (Fig. 16 D). Solenomere long and having a lateral expansion (Figs 16 D – E), but not reaching the apex of the prefemoral process. Cingulum in basal position, covered by the prefemoral process. Apex of the solenomere with descending sigmoid curvature (Fig. 16 E). Female. Unknown.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFD7E53F46C6B1368812F862.taxon	distribution	Distribution. Known only from the type locality (Fig. 18).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFD7E53F46C6B1368812F862.taxon	etymology	Etymology. The species epithet is a reference of the Brazilian folkloric character “ Boto cor-de-rosa ”. According to indigenous legend, the “ pink dolphin ”, which lives in the Amazonian’s rivers, turns into a beautiful man during the nights of full moon to get women pregnant. In the next day, it turns into a Boto again. Noun in apposition.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFD4E50346C6B50B8C62FF0E.taxon	description	Figures 17 – 18, 24 C – D urn: lsid: zoobank. org: act: C 71 B 69 A 8 - 40 C 3 - 4 E 39 - 8050 - 6 ECA 65 AB 930 D	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFD4E50346C6B50B8C62FF0E.taxon	materials_examined	Type material. Male holotype from UHE Tucuruí, Tucuruí (03 ° 42 ’ 01 ” S, 49 ° 42 ’ 00 ” W), Pará, Brazil, 1984, Instituto Butantan Expedition coll., deposited in IBSP 277. Paratype: three females with the same data of the holotype, deposited in IBSP 274, IBSP 276, and IBSP 278.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFD4E50346C6B50B8C62FF0E.taxon	diagnosis	Diagnosis. Adult males differ from all other species of the genus by the combination of the following characters: legs from the fifth to the eighth segment with ventral projections in the coxae (Fig. 31 B); presence of triangular lateral expansion near the apex of the solenomere (Fig. 17 F) and presence of a short secondary process in the prefemoral process (Fig. 17 D).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFD4E50346C6B50B8C62FF0E.taxon	description	Description. Male (Holotype, IBSP 277). Coloration (long-preserved in 70 % ethanol): Head light brown. Antennae ocher yellowish. Body brown, with the edge of the paranota lighter (Fig. 17 A – C). Legs ocher yellowish. Legs from the fifth to the eighth segment with ventral projections in the coxae being the posterior pair of the sixth segment extremely elongated. Telson ocher. Total length: 65.51. Total width: 11.50. Collum, length 3.44, width 10.82. Antennomere lengths (1> 7): 0.95; 2.38; 2.14; 1.88; 1.79; 1.70; 0.35. Podomeres lengths (1> 7): 1.19; 1.69; 2.99; 1.42; 1.86; 1.90; 0.45. Gonopod aperture, length 2.46, width 4.87. Telson, length 1.41. Gonopod: length 3.42, width 3.07. Gonocoxae: length 1.51, width 1.36. Gonocoxae globose, about 1 / 2 the size of the telopodite. Telopodite: length 3.33, width 1.62. Prefemoral region short (Fig. 17 D). Prefemoral process (Fig. 17 D – F) long and curve, presenting a dorso-basal lobe, starting from an angle of 225 °, involving the solenomere in its initial portion and subsequently ascending parallel to the solenomere and partially covering it in ectal view (Fig. 17 F); in its anterior portion presents an elongated, thin and acuminate secondary process (Fig. 17 D). Solenomere long, curved and with a lateral expansion (Fig. 17 F), but not reaching the apex of the prefemoral process. Cingulum in basal position, covered by the prefemoral process. Apex of the solenomere abruptly descending (Figs 17 D – E). Female (IBSP 276). Body as in male, except for the legs; without modifications. Total length: 75.74. Total width: 11.46. Antennomere lengths (1> 7): 0.74; 2.07; 1.98; 1.70; 1.66; 1.75; 0.35. Podomeres lengths (1> 7): 1.10; 1.29; 3.048; 1.040; 1.28; 1.62; 0.32. Telson, length 1.35. Epigyne rounded with irregular edges. Vulvae (Figs 24 C – D): length 2.20, width 0.88. External valve: length 1.17, width 0.39. Internal valve: length 1.17, width 0.43. Operculum: length 0.32, width 0.62.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFD4E50346C6B50B8C62FF0E.taxon	distribution	Distribution. Known only from the type locality (Fig. 18).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFD4E50346C6B50B8C62FF0E.taxon	etymology	Etymology. The species epithet is a reference of the Brazilian folkloric character “ Mula-sem-cabeça ”. According to the legend, of uncertain origin, it’s a mule of brown color with a torch of fire in the place of the head. Noun in apposition.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFEBE50346C6B7028E2AF830.taxon	description	Figures 19, 28	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFEBE50346C6B7028E2AF830.taxon	diagnosis	Diagnosis. Same for the genus.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFEBE50346C6B7028E2AF830.taxon	description	Description. Male (Holotype, MZSP). Coloration (long-preserved in 70 % ethanol): Head and antennae ocher whitish. Body ocher with the edges of the paranota whitish (Figs 19 A – C). Legs whitish. Telson ocher whitish. Total length: 52.69. Total width: 10.5. Collum, length 1.97, width 6.67. Antennomere lengths (1> 7): 0.66; 1.74; 1.53; 1.33; 1.29; 1.29; 0.37. Podomeres lengths (1> 7): 0.91; 1.13; 2.16; 0.99; 1.28; 1.51; 0.42. Gonopod aperture, length 1.10, width 2.02. Telson, length 1.22. Gonopod: length 2.14, width 1.50. Gonocoxae: length 0.68, width 0.91. Telopodite: length 2.14, width 0.72. Prefemoral process long (size equal to the solenomere), sinuous and narrow, covering the apex of the solenomere (Figs 19 D, F). Solenomere long and narrow (Fig. 19 D). Cingulum in medial position (Fig. 19 F). Apex of the solenomere acuminate and ascending with the terminal zone marked by a stretch mark (Figs 19 D, F, arrows). Female. Unknown.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFEBE50346C6B7028E2AF830.taxon	distribution	Distribution. Known only from the type locality (Fig. 28).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFEBE50346C6B31B89D0FAF5.taxon	type_taxon	Type species: A. costalimai Schubart, 1962, by original designation; Hoffman, 1976: 173; Hoffman, 1980: 151; Shelley et al. 2000: 84	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFEBE50346C6B31B89D0FAF5.taxon	diagnosis	Diagnosis. Males of Angelodesmus differ from those of other Arthrosolaenomeridini genera by the combination of the following characters: stipes of the gnathochilarium with lateral folds; gonocoxae acuminate in ectal view (Fig. 19 F); dorso-basal lobe in the prefemoral process absent (Fig. 19 D); apex of the solenomere with the terminal zone marked by a stretch mark (Figs 19 D, F, arrows).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFEBE50346C6B31B89D0FAF5.taxon	description	Description. General characters: Coloration (specimen long preserved in 70 % ethanol) whitish yellow, paranota tip and posterior border of the metaterga slightly whitish. Gnathochilarium: lingual plate covered completely by sparsely setae; mentum only centrally covered by setae, with smooth edges and stipes completely covered by sparsely setae and positioned mainly in the median region, presenting lateral folds. Body rings: cuticle slightly rough; alignment of paranota in posterior view straight; sternite of the fourth segment with a pair of projections, sternites of the fifth and sixth segments with two pairs of intumescences also covered by setae, the posterior pair less conspicuous; leg pair of the third body ring missing. Sternite of sixth body ring with two pairs of ventral projections; seventh body ring without projections and post-gonopodal sternites with two pairs of rounded ventral projections (Fig. 30 D). Paranota form: round (Fig. 19 B). Legs: without modifications only a slightly dorsal lobe on prefemur (Fig. 31 D). Male characters: Leg pair of the third body ring missing, as well as the genital papilla. Posterior edge of the gonopod aperture without projections, excavated. Gonopods: gonocoxae equivalent to about half the length of the telopodite, cylindrical and acuminated in ectal view, without a spiniform process (Fig. 19 F). Prefemoral process: elongate and thin, without a dorso-basal process (Fig. 19 D). Cingulum in a middle position (Fig. 19 F). Solenomere also long and thin with apex ascending (Fig. 19 D). Female characters: Unknown.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFEBE50346C6B31B89D0FAF5.taxon	distribution	Distribution. State of Goiás, Brazil (Fig. 28). Composition. Now considered as monotypic: Angelodesmus costalimai Schubart, 1962.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFE8E50146C6B50B8854FB56.taxon	materials_examined	Type species. Arthrosolaenomeris planaltensis Schubart, 1960.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFE8E50146C6B50B8854FB56.taxon	diagnosis	Diagnosis. Males of Abiliodesmus n. gen. differ from those of other Arthrosolaenomeridini genera by the combination of the following characters: slight dorsal lobe on leg prefemur (Figs 31 C – D); lateral macrosetae dispersed on the gonocoxae, in mesal view (Fig. 32 F); cingulum with a basal position (Fig. 20 F); and presence of a membranous lateral flap at the apex of the solenomere (Fig. 22 D) (absent in Abiliodesmus mapinguari n. sp.).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFE8E50146C6B50B8854FB56.taxon	description	Description. General characters: Body length between 34.63 mm (A. defensor comb. nov.) and 69.15 mm (A. mapinguari n. sp.). Coloration (specimens long preserved in 70 % ethanol) variable between species, ranging from ocher to reddish brown, paranota tip and posterior border of the metaterga slightly whitish. Gnathochilarium: lingual plate covered completely by sparsely setae; mentum only centrally covered by setae, with smooth edges and stipes covered by sparsely setae. Body alignment of paranota in posterior view curved ventrally (straight in A. cataractae comb. nov and A. defensor comb. nov.). Sternite of the fourth segment with a pair of projections, sternite of the fifth with two pairs of projections covered by setae, the posterior pair less conspicuous. Sternite of sixth body ring without ventral projections (present in A. mapinguari n. sp.); seventh with a pair of ventral projections (absent in A. cataractae comb. nov.) and post-gonopodal sternites with two pairs of acuminate ventral projections (Fig. 30 F) (absent in A. cataractae comb. nov. and rounded in A. mapinguari n. sp.). Paranota form: round to slightly rectangular (Figs 20 B, 23 B). Legs: without modifications beyond a slight dorsal lobe on prefemur; thin setae on the ventral part of the podomeres in the anterior legs (Fig. 31 D). Male characters: Leg pair of the third body ring with the coxae presenting the rectangular genital papilla. Gonopod aperture on seventh body ring with the posterior margin excavated and with a small projection (absent in A. mapinguari n. sp.). Gonopods: gonocoxae equivalent to about half the length of the telopodite, cylindrical, without a spiniform process. Prefemoral process: elongate and thin with a dorso-basal process (Fig. 20 D). Cingulum in a basal position (Fig. 20 F). Solenomere long with apex ascending and lateral membranous expansion (Fig. 21 D) (absent in A. mapinguari n. sp.). Female characters: Vulvae (Figs 24 E – F): oval-shaped, having different proportions in lateral view.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFE8E50146C6B50B8854FB56.taxon	distribution	Distribution. States of Tocantins, Goiás and São Paulo, Brazil (Fig. 25).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFE8E50146C6B50B8854FB56.taxon	etymology	Etymology. The genus name is taken as a noun in apposition. Name in honor of a very important person in the life of the first author, considered a great storyteller, with the combining stem “ - desmus, ” meaning band, commonly used in generic names in Polydesmida. Composition. Abiliodesmus planaltensis comb. nov. (Schubart, 1960); Abiliodesmus cataractae comb. nov. (Hoffman, 1976); Abiliodesmus defensor comb. nov. (Hoffman, 1976); Abiliodesmus mapinguari n. sp.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFE9E50746C6B4468FEDFEB6.taxon	description	Figures 20, 24 E – F, 25	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFE9E50746C6B4468FEDFEB6.taxon	materials_examined	Additional material. BRAZIL: São Paulo: Monte Aprazível (20 ° 46 ’ 21 ” S, 49 ° 42 ’ 51 ” W), Fazenda São José do Varjão, 3 ♀ and one juvenile, 09. I. 1950, O. Schubart coll. (MZSP 1084); Ribeirão Preto (21 ° 10 ’ 39 ” S, 47 ° 48 ’ 37 ” W), Mata Santa Tereza, 3 ♂ and 1 ♀, 06 – 11. I. 2006, I. Cizauskas coll. (IBSP 3569); 4 ♂ and 1 ♀, 06 – 11. I. 2006, I. Cizauskas coll. (IBSP 3570).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFE9E50746C6B4468FEDFEB6.taxon	diagnosis	Diagnosis. Adult males differ from all other species of the genus by the presence of a narrow lateral expansion in the solenomere (Figs 20 D – F).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFE9E50746C6B4468FEDFEB6.taxon	description	Description. Male (Holotype, MZSP 1086). Coloration (long-preserved in 70 % ethanol): Head and antennae ochre. Body ocher with the edges of the paranota whitish (Figs 20 A – C). Legs ochre. Telson ocher. Total length: 50.84. Total width: 7.48. Collum, length 1.87, width 6.68. Antennomere lengths (1> 7): 0.61; 1.67; 1.30; 1.12; 1.37; 1.30; 0.38. Podomeres lengths (1> 7): 0.70; 0.94; 2.20; 0.97; 1.12; 1.55; 0.43. Gonopod aperture, length 1.29, width 2.34. Telson, length 0.81. Gonopod: length 2.71, width 2.11. Gonocoxae: length 0.84, width 0.98. Telopodite: length 2.56, width 1.12. Prefemoral process long (slightly larger than the solenomere) and narrow (Fig. 20 D). Solenomere long (Fig. 20 F), presence of a lateral membranous expansion (Figs 20 E – F). Cingulum situated basally (Fig. 20 F). Apex of the solenomere acuminate and ascending (Fig. 20 D). Female (IBSP 3569). Body as in male. Total length: 54.76. Total width: 8.18. Antennomere lengths (1> 7): 0.49; 1.69; 1.38; 1.46; 1.34; 1.11; 0.30. Podomeres lengths (1> 7): 0.82; 0.99; 2.18; 0.78; 1.02; 1.43; 0.59. Telson, length 1.72. Epigyne rounded with regular edges. Vulvae (Fig. 24 E – F): length 1.20, width 0.57. External valve: length 0.81, width 0.26. Internal valve: length 0.78, width 0.27. Operculum: length 0.37, width 0.45.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFE9E50746C6B4468FEDFEB6.taxon	distribution	Distribution. São Paulo, Brazil (Fig. 25).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFEFE50746C6B3C38E2AFB9A.taxon	description	Figures 21, 24 G – H, 25	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFEFE50746C6B3C38E2AFB9A.taxon	diagnosis	Diagnosis. Adult males differ from all other species of the genus by the combination of the following characters: sternite of seventh body ring without ventral projections, presence of a large lateral expansion in the solenomere (Figs 21 D – E) and the prefemoral process not involving the solenomere, in ectal view (Fig. 21 E).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFEFE50746C6B3C38E2AFB9A.taxon	description	Description. Male (Holotype, MZSP 1065). Coloration (long-preserved in 70 % ethanol): Head and antennae ochre with the seventh antennomere having three well-demarcated invaginations and one slight invagination between the sensory cones. Body ocher with the edges of the paranota whitish (Figs 21 A – C). Legs ocher yellowish. Telson ocher. Total length: 37.14. Total width: 5.88. Collum, length 1.83, width 5.52. Antennomere lengths (1> 7): 0.52; 1.40; 1.25; 1.09; 1.03; 1.17; 0.30. Podomeres lengths (1> 7): 0.69; 0.88; 1.98; 0.83; 1.03; 1.39; 0.45. Gonopod aperture, length 1.14, width 1.93. Telson, length 0.95. Gonopods: lost. Female (Paratype, MZSP 1066). Body as in male. Total length: 35.90. Total width: 5.64. Antennomere lengths (1> 7): 0.51; 0.91; 1.06; 1.06; 0.83; 1.16; 0.22. Podomeres lengths (1> 7): 0.50; 0.74; 1.64; 0.55; 0.70; 1.13; 0.38. Telson, length 0.81. Epigyne rounded with regular edges. Vulvae (Figs 24 G – H): length 0.95, width 0.50. External valve: length 0.63, width 0.22. Internal valve: length 0.64, width 0.22. Operculum: length 0.32, width 0.39.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFEFE50746C6B3C38E2AFB9A.taxon	distribution	Distribution. Known only from the type locality (Fig. 25).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFEFE50746C6B6EF8E2AF863.taxon	description	Figures 22, 25	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFEFE50746C6B6EF8E2AF863.taxon	diagnosis	Diagnosis. Adult males differ from all other species of the genus by the combination of the following characters: presence of a wide lateral expansion in the solenomere (Fig. 22 D) and the prefemoral process completely involving the solenomere, in ectal view (Fig. 22 F).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFEFE50746C6B6EF8E2AF863.taxon	description	Description. Male (Holotype, MZSP 1067). Coloration (long-preserved in 70 % ethanol): Head and antennae ochre (both antennae of the holotype are broken). Body ocher with the edges of the paranota whitish (Figs 22 A – C). Legs ocher yellowish. Telson ocher. Total length: 34.63. Total width: 5.26. Collum, length 1.39, width 4.62. Antennomere lengths (1> 7): 0.46; 1.28; 1.09; 1.17; 1.18; 1.10; 0.34. Podomeres lengths (1> 7): 0.55; 0.70; 1.74; 0.68; 0.79; 1.27; 0.39. Gonopod aperture, length 1.03, width 1.68. Telson, length 0.93. Gonopod: length 1.99, width 0.94. Gonocoxae: length 0.58, width 0.29. Telopodite: length 1.99, width 0.65. Prefemoral process long (slightly larger than the solenomere) and narrow (Fig. 22 E). Solenomere long with a lateral membranous expansion (Fig. 22 D). Cingulum in basal position (Fig. 22 F). Apex of the solenomere acuminate and ascending (Figs 22 D). Female (Paratype, VMNH 110613.2). See Hoffman (1976): 176 to taxonomic notes.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFEFE50746C6B6EF8E2AF863.taxon	distribution	Distribution. Known only from the type locality (Fig. 25).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFEDE50946C6B51D8E3DFF0E.taxon	description	Figures 23, 25, 27 A – B urn: lsid: zoobank. org: act: 12 BBB 981 - 7 D 78 - 40 C 3 - A 78 B- 3314 E 5268 F 0 D	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFEDE50946C6B51D8E3DFF0E.taxon	materials_examined	Type material. Male holotype from Palmas (10 ° 12 ’ 47 ” S, 48 ° 21 ’ 37 ” W), Tocantins, Brazil, X. 2001, I. Knysak and R. Martins coll., deposited in IBSP 1699. Paratypes: two females with same data of the holotype, IBSP 7787; one male and one female from Fazenda Giovam, Porto Nacional (10 ° 42 ’ 30 ” S, 48 ° 25 ’ 01 ” W), Tocantins, Brazil, 01. XII. 2013, Chagas-Jr, A. and Giupponi, A. coll., deposited in CZUFMT 834. Additional material. BRAZIL: Tocantins: Porto Nacional (10 ° 42 ’ 30 ” S, 48 ° 25 ’ 01 ” W), Fazenda Giovam, 1 ♂, 02. XII. 2013, A. Chagas-Jr and A. Giupponi coll. (CZUFMT 839); woods near the to the Tocantins River, 1 ♂, 29. XI. 2013, A. Chagas-Jr and A. Giupponi coll. (CZUFMT 837); Taquaruçu (10 ° 18 ’ 21 ” S, 48 ° 10 ’ 17 ” W), Vai Quem Quer, 1 ♂, 30. XI. 2013, A. Chagas-Jr and A. Giupponi coll. (CZUFMT 836).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFEDE50946C6B51D8E3DFF0E.taxon	diagnosis	Diagnosis. Adult males differ from all other species of the genus by the combination of the following characters: presence of a secondary process at the apex of the prefemoral process (Figs 23 D); absence of a lateral expansion in the solenomere (Figs 23 D – E) and the prefemoral process involving the solenomere, in ectal view (Fig. 23 F).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFEDE50946C6B51D8E3DFF0E.taxon	description	Description. Male (Holotype, IBSP 1699). Coloration (long-preserved in 70 % ethanol): Head reddish brown with the labrum yellowish. Antennae reddish brown. Body reddish brown with the edges of the paranota whitish (Figs 23 A – C). Legs ocher whitish. Telson reddish brown. Total length: 59.44. Total width: 9.64. Collum 3.04 length, 8.85 width. Antennomere lengths (1> 7): 0.72; 1.68; 1.52; 1.78; 1.73; 1.44; 0.29. Podomeres lengths (1> 7): 0.92; 1.25; 2.30; 0.95; 1.21; 1.56; 0.55. Gonopod aperture 1.70 length, 3.03 width. Telson 1.82 length. Gonopod: 3.15 length, 2.25 width. gonocoxae: 1.00 length, 1.27 width. Telopodite: 3.09 length, 0.95 width. Prefemoral region short, 1 / 4 the size of telopodite. Prefemoral process (Figs 23 D – F) long and narrow, presenting a dorso-basal lobe, starting from an angle of 180 °, involving the solenomere in its initial portion, subsequently ascending parallel to the solenomere and partially covering it in ectal view, apex dividing into two acute projections (Fig. 23 D). Solenomere long, but not reaching the apex of the prefemoral process (Fig. 23 D). Cingulum in basal position (Fig. 23 F). Apex of the solenomere sickle-shaped with sigmoid curvature (Fig. 23 D). Female (Paratype, IBSP 1699). Body as in male. Total length: 69.15. Total width: 9.62. Collum 3.43 length, 9.60 width. Antennomere lengths (1> 7): 0.72; 2.00; 1.77; 1.51; 1.61; 1.44; 0.42. Podomeres lengths (1> 7): 0.92; 1.12; 2.22; 1.00; 1.13; 1.60; 0.49. Telson 1.47 length. Epigyne rounded with regular edges. Vulvae (Figs 27 A – B): 1.50 length, 0.70 width. External valve: 1.16 length, 0.29 width. Internal valve: 1.04 length, 0.34 width. Operculum: 0.37 length, 0.54 width.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFEDE50946C6B51D8E3DFF0E.taxon	distribution	Distribution. Tocantins, Brazil (Fig. 25).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFEDE50946C6B51D8E3DFF0E.taxon	etymology	Etymology. The species epithet is a reference of the Brazilian folkloric character “ Mapinguari ”. According to the indigenous legend, the “ Mapinguari ” is a hairy giant with one eye on the forehead and the mouth at the navel, living in the Amazon rainforest of Brazil. Noun in apposition.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFE1E50F46C6B36C8E2CFEB6.taxon	materials_examined	Type species. Folcloredesmus thomasi n. sp.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFE1E50F46C6B36C8E2CFEB6.taxon	diagnosis	Diagnosis. Males of Folcloredesmus n. gen. differ from those of other Arthrosolaenomeridini genera by the combination of the following characters: paranota sub-triangular (Fig. 26 B); slight dorsal lobe on leg prefemur; presence of a spiniform process in gonocoxae (Figs 26 D, 32 C).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFE1E50F46C6B36C8E2CFEB6.taxon	description	Description. General characters: Coloration (specimen long preserved in 70 % ethanol) ocher with the paranota tip slightly whitish. Gnathochilarium: lingual plate covered by several setae; mentum only centrally covered by setae, with smooth edges and stipes completely covered by sparse setae. Body rings: cuticle slightly rough; alignment of paranota in posterior view straight. Sternite of the fourth segment with a pair of projections, sternite of the fifth segment with two pairs of projections covered with setae and sternite of sixth body ring with two pairs of ventral projections; seventh body ring with a pair of projections and post-gonopodal sternites with two pairs of rounded ventral projections (Fig. 30 D). Paranota form: sub-triangular (Fig. 26 B). Legs: slight dorsal lobe, a ventroapical projection in the prefemur of the anterior legs (Fig. 31 E), and presence of granules in the tibia and tarsus; thin setae on the ventral part of the podomeres in the anterior legs (Fig. 31 F). Male characters: Gonopod aperture on seventh body ring with the posterior margin excavated, concave and transverse sulci; without projections. Gonopods: gonocoxae equivalent to about half the length of the telopodite, cylindrical, with a spiniform process (Fig. 26 D). Prefemoral process: elongate and thin (Fig. 26 D) and with indentations in the apical portion, in ectal view (Fig. 34 B). Cingulum in medial position (Fig. 26 F). Solenomere also long and thin with apex slightly curved (Figs 26 D – F). Female characters: Vulvae: oval-shaped, sub-triangular in lateral view.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFE1E50F46C6B36C8E2CFEB6.taxon	distribution	Distribution. State of Mato Grosso (Fig. 28).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFE1E50F46C6B36C8E2CFEB6.taxon	etymology	Etymology. The genus name comes from the Portuguese word for “ Folclore ” (culture shared by a particular group of people, as the tales characters used as species epithet in this paper) with the combining stem “ - desmus ”, commonly used in generic names in Polydesmida. Composition. One species: Folcloredesmus thomasi n. sp.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFE7E50C46C6B40F8912F827.taxon	description	Figures 26, 27 C – D, 28 urn: lsid: zoobank. org: act: 26862968 - 716 B- 47 D 3 - 9090 - 416 B 8 C 209 ABA	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFE7E50C46C6B40F8912F827.taxon	materials_examined	Type material. Male holotype from Fazenda São Nicolau (09 ° 50 ’ 25 ” S, 58 ° 14 ’ 53 ” W), Cotriguaçu, Mato Grosso, Brazil, 02. XI. 2014, M. Karam-Gemael coll., deposited in IBSP 7788. Paratypes: four males and two females from Fazenda São Nicolau, Cotriguaçu, Mato Grosso, Brazil, 11. XII. 2009, L. D. Batirola coll., deposited in IBSP 5452 and IBSP 5453. Additional material. BRAZIL: Mato Grosso: Cotriguaçu, Fazenda São Nicolau (09 ° 50 ’ 25 ” S, 58 ° 14 ’ 53 ” W), 2 ♂, X. 2017 (CZUFMT 849); Cotriguaçu, Fazenda São Nicolau (09 ° 50 ’ 25 ” S, 58 ° 14 ’ 53 ” W), 10 ♂, 02. XI. 2014, M. Karam-Gemael coll. (CZUFMT 833).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFE7E50C46C6B40F8912F827.taxon	diagnosis	Diagnosis. The same characteristics of the genus.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFE7E50C46C6B40F8912F827.taxon	description	Description. Male (Holotype, IBSP 7788). Coloration (long-preserved in 70 % ethanol): Head and antennae ochre with the seventh antennomere broken. Body ocher with the edges of the paranota whitish (Fig. 26 A – C), however in recently collected specimens the body’s coloration is reddish brown with the edge of the paranota yellowish (Figs 1 A – B). Legs ochre whitish. Telson ocher. Total length: 55.69. Total width: 7.46. Collum 2.66 length, 7.65 width. Antennomere lengths (1> 7): 0.62; 1.62; 1.49; 1.54; 1.55; 1.64; 0.26. Podomeres lengths (1> 7): 0.99; 1.42; 2.33; 1.29; 1.52; 1.85; 0.47. Gonopod aperture 1.54 length, 3.01 width. Telson 1.20 length. Gonopod: 2.37 length, 2.37 width. Gonocoxae: 1.13 length, 1.26 width. A small spiniform process present (Fig. 26 D). Telopodite: 2.30 length, 0.87 width. Prefemoral region 1 / 3 the size of telopodite. Prefemoral process (Figs 26 D – F) long, narrow and falciform, starting from an angle of 135 °, ascending parallel to the solenomere and partially covering it in ventral view; in ectal view, with indentations in the apical portion. Cingulum medially positioned (Fig. 26 F). Solenomere long and thin with apex acute and slightly curved (Fig. 26 F). Female (Paratype, IBSP 5453). Body as in male. Total length: 67.74. Total width: 8.90. Collum 3.40 length, 8.27 width. Epigyne short and triangular with irregular edges. Vulvae (Figs 27 C – D): 1.21 length, 0.75 width. External valve: 0.89 length, 0.40 width. Internal valve: 0.88 length, 0.37 width. Operculum: 0.30 length, 0.35 width.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFE7E50C46C6B40F8912F827.taxon	distribution	Distribution. Known only from the type locality (Fig. 28).	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
052487CFFFE7E50C46C6B40F8912F827.taxon	etymology	Etymology. The species epithet is taken as a noun in apposition honoring a very important person in the life of the first author, who has a passion for stories of Brazilian folklore. Noun in apposition.	en	Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M., Brescovit, Antonio Domingos (2021): Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae). Zootaxa 4970 (2): 201-256, DOI: https://doi.org/10.11646/zootaxa.4970.2.1
