identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03DF1874FF8BFFCCFF5081ADFA75FBF0.text	03DF1874FF8BFFCCFF5081ADFA75FBF0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Calciphilopteris Yesilyurt & H. Schneider 2010	<div><p>Calciphilopteris Yesilyurt &amp; H.Schneider, gen. nov. — Figs. 1A–C</p> <p>Calciphilopteris gen. nov. Doryopteridis affinis sed rhizomate breviter vel longe repenti, paleis rhizomatum lanceolatis vel anguste lanceolatis clathratis vel semiclathratis, petiolo tereti fasciculum vascularem unum tantum continenti, petiolo laminaque fibrillas (pilos squamiformes) atque squamas non clathratas vel semiclathratas ferentibus, sporangio plerumque longipedicellato 32 sporas continenti, plantis plerumque rupicola locus calcareos incolentibus differt.</p> <p>Type species:— Calciphilopteris ludens (Wall. ex Hook.) Yesilyurt &amp; H.Schneid. = Pteris ludens Wallich ex. Hooker (1858: 210). Holotype: INDIA: Wallich 88 (K).</p> <p>Plants terrestrial or rupicolous; essentially growing on limestone. Rhizome slender, short- to long-creeping, scales up to 3 mm, linear to ovate-lanceolate, acuminate to caudate, predominantly bicoloured, with clathrate to semi-clathrate centre, and paler, narrow to broad border, margin entire to dentate towards the apex. Petiole terete to slightly sulcate (towards) at the base of the lamina; one vascular bundle; dark brown, atropurpureous to black, base with scales as those of the rhizome, higher up the petiole with indumentum (generally dense towards the base of the lamina) usually scaly, scales semi- to non-clathrate, with fibrils, rarely with white hairs or glabrous. Sterile and fertile fronds dimorphic (architecture of fertile lamina more complex), sub-coriaceous to coriaceous; glabrescent to glabrous, indumentum on the abaxial side with glandular hairs and/or fibrils and non-clathrate scales at the base of the lamina; veins anastomosing, without included free veinlets. Sterile fronds up to 45 cm; lamina up to 17 cm, ranging from simple, entire or 3-lobed to pedate; oblong-oval, cordate, sagittate, hastate, lobed to pentagonal in outline; pinnae/segments1–2(–4) pairs; the margins usually with a whitish or brown cartilagineous border, entire; hydathodes generally present on the upper surface; veins ending free near the margin (occasionally confluent in C. allenae); basal pinnae lobed basiscopically or not. Fertile fronds up to 65 cm; lamina up to 20(–25) cm; sagittate, hastate, pentagonal to broad-cordate or suborbicular in outline, 3-lobed to pedate, pedate-pinnatifid (pedate-2-pinnatifid); pinnae/segments 2–5(–6) pairs; ascending to patent, deltoid to long-lanceolate, acute or acuminate, predominantly with entire margins; basal pinna/segment with acroscopic and/or with basiscopic segments/lobes; first segment/lobe divided or not, falcate to lanceolate. Sori marginal, usually on a continuous vascular commissure (connecting the vein-tips), rarely interrupted at sinuses, covered by a continuous modified marginal indusium, without paraphyses; receptacle and indusium continuous, including in the sinuses; indusium with margin entire to slightly eroded. Sporangium with pedicel 1–2(–2.5) times the capsule length; capsule without basal cell; annulus with 17–34 indurate cells; with 32 spores per sporangium. Spores trilete, tetrahedral to globose, echinate or cristate, without equatorial flange. Chromosome number n = 29, 30 or multiples thereof.</p> <p>Distribution:—Southeast Asia, from Indo-China to New Guinea (and Australia).</p> <p>Habitat &amp; Ecology:—exclusively on limestone outcrops.</p> <p>Etymology:—the genus name is a composite from the Greek calx (limestone), philus (loving) and pteris (fern) and emphasizes the exclusivity of limestone outcrops as the natural habitat for this fern.</p> <p>Diagnostic key to the species of Calciphilopteris</p> <p>1. Basal pinna/segment with 2–4 basiscopic segments, all ascendant; first basiscopic segment with 1–3 lobes (Fig. 1B), conspicuous or not, ascendant; sinus (between pinnae/segments) usually narrow, acute or round (seldom slightly rectangular); fertile lamina pedate-pinnatifid to pedate-2(–3)-pinnatifid, broadly ovate to cordate-ovate, pinnae/segments 3–7 (Fig 1A), all equally ascendant, long lanceolate to linear lanceolate (except basal pinna)........ C. papuana</p> <p>- Basal pinnae/segment with 1–3 basiscopic segments, ascendant or not; first segment very rarely lobed; sinus (between pinnae/segments) rounded to slightly rectangular; fertile lamina pedate to pedate-pinnatifid, hastate to trilobed, ovate-lanceolate, slightly pentagonal or obovate, pinnae/segments 1–4, all equally ascendant or not, usually lanceolate (except basal one)........................................................................................................................................ 2</p> <p>2. Scales of rhizome and base of petiole long lanceolate to linear lanceolate, caudate, margin (half way towards apex) dentate; fertile lamina generally sagittate to trilobed (rarely pedate), hydathodes absent; marginal veins regularly anastomosing along the leaf margin............................................................................................................. C. alleniae</p> <p>- Scales of rhizome and base of petiole lanceolate to long lanceolate, not caudate, margin (half way towards apex) remotely dentate to entire; fertile lamina hastate, pedate to pedate-bipinnatifid, hydathodes present (sometimes obscure); marginal veins ending free along the leaf margin......................................................................................... 3</p> <p>3. Basal pinnae with acroscopic and basiscopic segments; scales margin (half way towards apex) entire; fertile lamina commonly pedate-pinnatifid to pedate-2-pinnatifid, pentagonal, pinnae/segments 2–4, all equally ascendant; basal pinna/segment with 1(–2) acroscopic segment and 2–3 basiscopic segments/lobes (Fig. 1A), all ascendant; first basiscopic segment/lobe generally unlobed.......................................................................................................... C. ludens</p> <p>- Basal pinnae with acroscopic pinnae only; scales margin (half way towards apex) dentate to occasionally dentate; fertile lamina commonly hastate to pedate (pedate-pinnatifid), ovate-lanceolate, slightly pentagonal, pinnae/segments 1–2(–3), basal pinna/segment ascendant, other segments/lobes patent to slightly ascendant; basal pinna/segment with 1(–2) basiscopic segment/lobe, which is usually unlobed and generally parallel to the petiole (Fig. 1C)..................................................................................................................................................................... C. wallichii</p></div> 	http://treatment.plazi.org/id/03DF1874FF8BFFCCFF5081ADFA75FBF0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Yesilyurt, Jovita C.;Schneider, Harald	Yesilyurt, Jovita C., Schneider, Harald (2010): The new fern genus Calciphilopteris (Pteridaceae). Phytotaxa 7: 52-59, DOI: 10.11646/phytotaxa.7.1.7
03DF1874FF89FFCCFF50864EFEF9F865.text	03DF1874FF89FFCCFF50864EFEF9F865.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Calciphilopteris alleniae (R. M. Tryon) Yesilyurt & H. Schneider	<div><p>1. Calciphilopteris alleniae (R.M.Tryon) Yesilyurt &amp; H.Schneider, comb. nov.</p> <p>Basionym:— Doryopteris alleniae Tryon (1962: 91, fig 1–4), as ‘ allenae ’. Holotype: MALAYSIA: Perak: Gunong Idong, near Perak, 24 May 1962, B.M. Allen 4763 (GH). Paratypes: 12 January 1959, B.M. Allen 4103 (GH, US), 13 August 1959, B.M. Allen 4343 (GH).</p> <p>Rhizome predominantly short-creeping, scales 3 mm, ferrugineous, lanceolate to linear-lanceolate, caudate, margin dentate. Petiole terete, black, indument of fibrils, particularly dense towards the base of the lamina. Sterile and fertile fronds coriaceous; glabrescent, indument on the abaxial side with glandular hairs and/or fibrils, veins usually joined at the vein-ends (at the margin of the lamina). Sterile fronds up to 20 cm long; lamina up to 10 cm, simple, entire to 3-lobed, ovate to oblong-ovate, hydathodes absent. Fertile fronds up to 35 cm; lamina up to 15 cm; suborbicular, obovate to 3-lobed (rarely pedate); pinnae/segments conspicuously ascendant, long linear-lanceolate. Indusium with margin entire to more or less irregular. Sporangium with pedicel 1–1½ × the capsule length; annulus with up to 34 indurated cells. Spores light castaneous to dark castaneous, echinate.</p> <p>Distribution:—known only from a few localities on the Malay Peninsula (Perak: Gunong Idong, Gunong Temurong and Selangor: Gua Batu).</p> <p>Note:—there is an orthographic error in Doryopteris allenae which is regulated in the ICBN by Article 60.11, recommendation 60C.1 and it is here therefore, corrected (it should be Calciphilopteris alleniae instead of C. allenae).</p> <p>Selected specimens:— MALAYSIA: Ding Hou 709 (L); Molesworth-Allen 4103 (GH), 4393 (GH), 4763 (GH, US).</p></div> 	http://treatment.plazi.org/id/03DF1874FF89FFCCFF50864EFEF9F865	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Yesilyurt, Jovita C.;Schneider, Harald	Yesilyurt, Jovita C., Schneider, Harald (2010): The new fern genus Calciphilopteris (Pteridaceae). Phytotaxa 7: 52-59, DOI: 10.11646/phytotaxa.7.1.7
03DF1874FF8EFFCBFF5082A6FAFEFA98.text	03DF1874FF8EFFCBFF5082A6FAFEFA98.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Calciphilopteris ludens (Wall. ex Hook.) Yesilyurt & H. Schneider	<div><p>2. Calciphilopteris ludens (Wall. ex Hook.) Yesilyurt &amp; H.Schneider, comb. nov. — Fig. 1A.</p> <p>Basionym:— Pteris ludens Wallich (1828: 88), nomen nudum, Pteris ludens Wall. ex. Hooker (1858: 210), Lithobrochia ludens (Wall. ex Hook.) Beddome (1865: t. 27), Pellaea ludens (Wall. ex Hook.) Prantl (1882: 419), Doryopteris ludens (Wall. ex Hook.) Smith (1875: 289). Holotype: INDIA: Wallich 88 (K).</p> <p>Rhizome predominantly long-creeping, scales up to 3 mm, dark brown, lanceolate, margin entire. Petiole terete to sulcate adaxially towards the base of the lamina, dark brown to atropurpureous, indumentum usually with hairs (whitish), non-clathrate scales and fibrils. Sterile and fertile fronds glabrescent to glabrous, indumentum on the abaxial side with fibrils and non-clathrate microscales. Sterile fronds up to 35 cm; lamina up to 20 cm, sagittate to pedate; ovate to pentagonal; pinnae/segments1–2(–3) pairs; hydathodes generally present (sometimes obscure). Fertile fronds up to 45 cm; lamina up to 25 cm, pentagonal to broad-cordate, pedate, pedate-pinnatifid (to pedate-2-pinnatifid); pinnae/segments in 2–4(–5) pairs; basal pinna/segment falcate, with 1(–2) acroscopic lobes and 2–3 basiscopic segments/lobes, which are ascendant, falcate to lanceolate, rarely the first basiscopic segment/lobe is divided; other pinnae/segments al equally ascendant, lanceolate to long-lanceolate. Indusium with margin entire to slightly eroded. Sporangium with pedicel 1–2(– 2½) × the capsule length; capsule with annulus with 17–34 indurated cells. Spores dark castaneous, cristate.</p> <p>Distribution:— India, Indochina, southern China, western parts of Malesia (Flores, Java, Malay Peninsula, Simbawa, Sumatra), Thailand, and Australia (where it was once recorded from north-eastern Queensland by D.L. Jones s.n. (BRI), see: Bostock 1998).</p> <p>Note:—some specimens (Congdon 617, Molesworth-Allen 2785, E. Smith 250A, Kerr 250, 11365) show unique leaf and spore characters that are similar but not identical with characters observed in other species of Calciphilopteris. It was decided to assign these specimens here to C. ludens, because available collections are insufficient to make unequivocal taxonomic assignments. These specimens may represent either a unique form of C. ludens or an undiscovered species.</p> <p>Selected specimens:— MALAYSIA: Perlis: Molesworth-Allen 2816 (US, S). Kedah: Alphonso &amp; Samsuri 123 (BM); Henderson 29066 (BM); Littke 260 (L). INDONESIA: Java: Horsfield s.n. 110 (BM); Mosset 188 (C, GH, L, PE). THAILAND: Geesink et al. 5949 (BM, L); Hennipman 3029 (BM, L, NY, U, US); Kerr 5476 (BM); Kostermanns 762 (L); Larsen 5790 (BM, GH); Rock 503 (US); Shimizu et al. 13325 (C, L); Tagawa et al. 1903 (L, US);. MYANMAR: Dickason 7762 (GH); Kress et al. 97-6017 (US); J. F. Smith 89 (GH). VIETNAM (Cochinchina): Larsen 8964 (GH); Pierre s.n. (GH). SINGAPORE: Corner &amp; Henderson 22815 (C). CHINA: Yunan: Henry 13325 (MO, NY). INDIA: Assam: Mann s.n. (UC).</p></div> 	http://treatment.plazi.org/id/03DF1874FF8EFFCBFF5082A6FAFEFA98	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Yesilyurt, Jovita C.;Schneider, Harald	Yesilyurt, Jovita C., Schneider, Harald (2010): The new fern genus Calciphilopteris (Pteridaceae). Phytotaxa 7: 52-59, DOI: 10.11646/phytotaxa.7.1.7
03DF1874FF8EFFCAFF508793FB29FE98.text	03DF1874FF8EFFCAFF508793FB29FE98.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Calciphilopteris papuana (Copel.) Yesilyurt & H. Schneider	<div><p>3. Calciphilopteris papuana (Copel.) Yesilyurt &amp; H.Schneider, comb. nov. — Fig. 1B.</p> <p>Basionym:— Doryopteris papuana Copeland (1911: 86). Holotype: PAPUA NEW GUINEA: Goodenough Bay, King 208, (MICH, isotype BO).</p> <p>Rhizome slender, short- to long-creeping, scales up to 2.5(–3) mm, dark brown, lanceolate, to ovatelanceolate, margin entire. Petiole terete, dark brown to atropurpureous, with fibrils and usually scaly, scales semi- to non-clathrate. Sterile and fertile fronds glabrescent to glabrous, indumentum on the adaxial side with fibrils on the main veins only, abaxial side with fibrils throughout (smaller on the lamina, bigger on the main veins) and non-clathrate to semi-clathrate microscales on the main veins. Sterile fronds up to 28 cm; lamina up to 15 cm, pedate; ovate to pentagonal; pinnae/segments1–3 pairs; hydathodes present (sometimes obscure). Fertile fronds up to 42(–45) cm; lamina up to 27 cm; pentagonal to broadly-cordate or suborbicular, pedatepinnatifid to pedate-bipinnatifid; pinnae/segments in 2–6(–7) pairs; basal pinnae/segment falcate, with 1–2 acroscopic lobes and 2–4 basiscopic segments/lobes, which are ascendant, falcate to lanceolate, first segment/ lobe divided into 1–3 lobes, lobes ascendant, lanceolate; other pinnae/segments conspicuously ascendant, lanceolate to linear-lanceolate. Indusium with margin entire to slightly eroded. Sporangium with pedicel 1(–</p> <p>1½) × the capsule length; capsule with annulus with 17–25 indurated cells. Spores castaneous to light castaneous, cristate.</p> <p>Distribution:—endemic to New Guinea, mainly in the eastern part.</p> <p>Selected specimens:— PAPUA NEW GUINEA: Brass 21853 (BM, GH, L, US); 21623 (GH, L); Carr 12372 (BM, L, NY); Kairo 629 (BM, L); King s.n. (BM, UC, PE); Hartley 10656 (GH, L).</p></div> 	http://treatment.plazi.org/id/03DF1874FF8EFFCAFF508793FB29FE98	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Yesilyurt, Jovita C.;Schneider, Harald	Yesilyurt, Jovita C., Schneider, Harald (2010): The new fern genus Calciphilopteris (Pteridaceae). Phytotaxa 7: 52-59, DOI: 10.11646/phytotaxa.7.1.7
03DF1874FF8FFFCAFF508397FCAFFB46.text	03DF1874FF8FFFCAFF508397FCAFFB46.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Calciphilopteris wallichii (J. Sm.) Yesilyurt & H. Schneider	<div><p>4. Calciphilopteris wallichii (J.Sm.) Yesilyurt &amp; H.Schneider, comb. nov. — Fig. 1C.</p> <p>Basionym:— Doryopteris wallichii Smith (1841: 404), Lithobrochia smithii T. Moore (1862: 342), nomen nudum, based on D. wallichii J.Sm. Holotype: PHILIPPINES: Luzon, Cuming 238 (K, isotypes BM, C, G, L).</p> <p>Rhizome usually short-creeping, scales 2.5(–3) mm, light brown, lanceolate to long-lanceolate, margin entire to rarely dentate. Petiole terete to slightly sulcate towards the base of the lamina; dark brown, atropurpureous, with fibrils. Sterile and fertile fronds sub-coriaceous; glabrescent, indumentum on the abaxial side with glandular hairs and fibrils on the main veins. Sterile fronds up to 30 cm; lamina up to 17 cm, ranging from simple, entire to sagittate, hastate, or basally lobed; hydathodes generally present on the upper surface only (sometimes obscure). Fertile fronds up to 40 cm; lamina up to 19 cm; sagittate, hastate, sub-pedate to rarely pedate, with 2(–3) pairs of pinnae/segments; basal pinna/segment falcate to lanceolate, conspicuously ascendant, with 1–2 basiscopic segment/lobes, which are ascendant to usually parallel with the petiole; other pinnae/segment broadly lanceolate to lanceolate, usually patente. Indusium usually with entire margin. Sporangium with pedicel 1–1½ × the capsule length; capsule with annulus with 20 (24) indurated cells. Spores usually castaneous to light castaneous, cristate.</p> <p>Distribution:— Philippines (mainly Luzon).</p> <p>Selected specimens:— PHILIPPINES: Copeland 95 (BM, C, S); Curran 15608 (BM, C). Elmer 6349 (C. NY); Price 2007 (BM, L, US); Ramos 33245 (BM, L); Topping 857 (C, GH, MO, NY, S); 1014 (GH, NY); 1247 (GH, US); 1219 (GH, NY); Williams 1501 (PE, UC).</p></div> 	http://treatment.plazi.org/id/03DF1874FF8FFFCAFF508397FCAFFB46	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Yesilyurt, Jovita C.;Schneider, Harald	Yesilyurt, Jovita C., Schneider, Harald (2010): The new fern genus Calciphilopteris (Pteridaceae). Phytotaxa 7: 52-59, DOI: 10.11646/phytotaxa.7.1.7
