identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
9D481962FF93B21B4B29F9E4FC953A19.text	9D481962FF93B21B4B29F9E4FC953A19.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Goniurosaurus huuliensis Orlov, Ryabov, Nguyen, Nguyen & Ho 2008	<div><p>Goniurosaurus huuliensis Orlov, Ryabov, Nguyen, Nguyen &amp; Ho, 2008</p> <p>Fig. 3</p> <p>Diagnosis</p> <p>Body splayed and gracile, SVL 97.2–134.6 mm; external nares bordered by 6–8 nasal scales; supraorbital region with a row of enlarged tubercles; outer surface of upper eyelid composed of granular scales, about one half the size of those on top of head and without enlarged tubercles; internasal 1 or 2 (rarely absent); supralabials 9–12; infralabials 9–12; preorbital scales 14–20; eyelid fringe scales 51–59; postmentals 2–4; gular region below lower jaws with enlarged tubercles; paravertebral tubercles 31–37; scale rows around midbody 118–130, granular scales surrounding tubercles 11–13; axillary pockets deep; subdigital lamellae under fourth toe 21–25; precloacal pores in males 25–30; iris reddish brown; dorsal ground color of head, body and limbs dark brown, without small dark brown blotches (dark blotches present only on lower zone of flanks); nuchal loop thin, posteriorly protracted (in V-shape); dorsal body bands between limb insertions 3, thin, immaculate yellow; gular region with brown spots (modified after Orlov et al. 2008; Nguyen 2011).</p> <p>Description (Supp. file 1: Table S2)</p> <p>Body splayed and gracile, adult males SVL 97.2–132.2 mm (mean ± SE: 118.9± 1.4 mm, n =32), TaL 36.7–108.6 mm (72.7 ± 3.7 mm), adult females SVL 97.4–134.6 mm (121.1 ± 1.2 mm, n =46), TaL 48.4–106.7 mm (78.2 ± 2.7 mm), juveniles SVL 73.5–74.4 mm (n =2), TaL 62.5–62.7 mm (Supp. file 1: Table S2); head triangular, wider than neck, covered by uniform granular scales interspersed with tubercles in temporal and occipital regions; scales on rostrum slightly larger and flatter; enlarged supraorbital tubercles in a conspicuous row; middorsal portion of rostral partially sutured dorsomedially, bordered laterally by first supralabial on each side, dorsolaterally by prenasal on each side, and dorsally by internasal and two supranasals; internasal 1–3 (rarely absent); external nares bordered by 5–7 nasals (prenasal, anterior and posterior supranasals, 2 slightly enlarged postnasal, and 1 or 2 granular scales); preorbital scales 14–20; supralabials 9–12, grading into granular scales posteriorly; infralabials 9–12; eyes large, pupils vertical; eyelid fringe scales 51–59, those of upper eyelid slightly enlarged; outer surface of upper eyelid composed of granular scales of about one half the size of those on top of head, without enlarged tubercles; fold of skin originating from suborbital region extends posteroventrally across angle of jaw; external auditory meatus elliptical with long axis directed dorsoventrally; tympanum deeply recessed; mental triangular, bordered laterally by first infralabial on each side and posteriorly by 2–4 postmentals; postmentals bordered by 7–10 gular scales; gular region below lower jaws with enlarged tubercles; gular scales juxtaposed and granular, grading posteriorly into flat hexagonal pectoral scales and even larger ventral scales.</p> <p>Neck narrower than body, covered with uniform granular scales interspersed with several sharply pointed conical tubercles on nape; tubercles on flanks conical, those of vertebral region somewhat flatter; dorsal body tubercles surrounded by 11–13 granular scales; dorsal tubercle rows at midbody 19–24; paravertebral tubercles between limb insertions 31–37, distinct vertebral row of tubercles absent; scales around midbody 118–130; ventral scales large; males with 25–30 precloacal pores in continuous series, females without precloacal pores (but pitted scales present); region posterior to vent covered by flat juxtaposed scales and greatly swollen, containing 1 or 2 enlarged tubercles on each side at level of vent; tail thick at base; light band on tail base in width of 9–12 granular scales and with 8–10 tubercles in transversal series.</p> <p>Limbs relatively long and thin, covered dorsally with granular scales interspersed with several tubercles and ventrally with flat juxtaposed to subimbricate scales; dorsal granular scales grade into slightly flattened subimbricate scales on top of pes and manus; hind limbs larger than forelimbs; axillary pockets deep; subdigital lamellae wide, 10 or 11 under first finger, 18–21 under fourth finger, 11 or 12 under first toe, 21–25 under fourth toe; digits laterally compressed, increasing in length from first to fourth, fifth shorter than fourth (modified after Orlov et al. 2008; Nguyen 2011).</p> <p>Coloration in life (Fig. 3)</p> <p>Dorsal ground color of head, body and limbs signal brown (juveniles), grey-brown (young adults, SVL: 97.2–116 mm) and signal brown (adults), without small dark brown blotches (dark blotches present only on lower zone of flanks); dull white tubercles on dorsal body, limbs in juveniles, few orange tubercles on limbs in adults; iris orange or red brown; five bands on the dorsal ground, thin, immaculate without dark spots, slight bisque in juveniles, orange brown or yellow in adults, all edged anteriorly and posteriorly by thin dark brown bands, including one thin nuchal loop extending from posterior corners of eyes and posteriorly protracted (in V-shape), three body bands between limb insertions, and another one on tail base; ground color of tail dark brown, and signal brown at mottled tail base; 3–6 immaculate white caudal bands; ventral surfaces of head, body and limbs dull white and immaculate except for few dark brown spots on margin regions of gular and limbs.</p></div> 	http://treatment.plazi.org/id/9D481962FF93B21B4B29F9E4FC953A19	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ngo, Hai Ngoc;Nguyen, Huy Quoc;Tran, Hieu Minh;Ngo, Hanh Thi;Gewiss, Laurenz Rafael;Schingen-Khan, Mona van;Nguyen, Truong Quang;Ziegler, Thomas	Ngo, Hai Ngoc, Nguyen, Huy Quoc, Tran, Hieu Minh, Ngo, Hanh Thi, Gewiss, Laurenz Rafael, Schingen-Khan, Mona van, Nguyen, Truong Quang, Ziegler, Thomas (2021): A morphological and molecular review of the genus Goniurosaurus, including an identification key. European Journal of Taxonomy 751 (1): 38-67, DOI: https://doi.org/10.5852/ejt.2021.751.1379, URL: http://dx.doi.org/10.5852/ejt.2021.751.1379
9D481962FF9CB2174B91FD66FD503F87.text	9D481962FF9CB2174B91FD66FD503F87.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Goniurosaurus araneus Grismer, Viets & Boyle 1999	<div><p>Goniurosaurus araneus Grismer, Viets &amp; Boyle, 1999</p> <p>Diagnosis</p> <p>Body splayed and gracile, SVL 108.9–124.0 mm; external nares bordered by 6–8 nasal scales; supraorbital region with a row of enlarged tubercles; outer surface of upper eyelid composed of small granular</p> <p>scales, one-half the size of those on top of head; internasal single; supralabials 8–10; infralabials 8 or 9; preorbital scales 13–18; eyelid fringe scales 52–67; postmentals 4–6; dorsal body scales elongate; paravertebral tubercles 29–38; scales around midbody 129–147; scales surrounding dorsal tubercles 10– 14; axillary pockets deep; subdigital lamellae under fourth toe 23–25; precloacal pores in males 18–23; iris dark brown; dorsal ground color of head, body and limbs immaculate dull yellow-gray; nuchal loop posteriorly protracted (in V-shape); dorsal body bands between limb insertions 3, wide, edged anteriorly and posteriorly by wide dark brown bands; light band on tail base in width of 15 or 16 granular scales; ground color of tail black, caudal bands 5, white, completed ventrally; ventral surface of head, body and limbs dull white and immaculate (Grismer et al. 1999; Nguyen 2011; Chen et al. 2014).</p> <p>Remarks</p> <p>During our field work in northern Vietnam, including the type locality in Cao Bang Province, no specimen of Goniurosaurus araneus was seen. Therefore, the diagnosis was solely based on the descriptions of Grismer et al. (1999) and Chen et al. (2014).</p> </div>	http://treatment.plazi.org/id/9D481962FF9CB2174B91FD66FD503F87	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ngo, Hai Ngoc;Nguyen, Huy Quoc;Tran, Hieu Minh;Ngo, Hanh Thi;Gewiss, Laurenz Rafael;Schingen-Khan, Mona van;Nguyen, Truong Quang;Ziegler, Thomas	Ngo, Hai Ngoc, Nguyen, Huy Quoc, Tran, Hieu Minh, Ngo, Hanh Thi, Gewiss, Laurenz Rafael, Schingen-Khan, Mona van, Nguyen, Truong Quang, Ziegler, Thomas (2021): A morphological and molecular review of the genus Goniurosaurus, including an identification key. European Journal of Taxonomy 751 (1): 38-67, DOI: https://doi.org/10.5852/ejt.2021.751.1379, URL: http://dx.doi.org/10.5852/ejt.2021.751.1379
9D481962FF9DB2194B13FCC4FB6E3B67.text	9D481962FF9DB2194B13FCC4FB6E3B67.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Goniurosaurus catbaensis Ziegler, Nguyen, Schmitz, Stenke & Rosler 2008	<div><p>Goniurosaurus catbaensis Ziegler, Nguyen, Schmitz, Stenke &amp; Rösler, 2008</p> <p>Fig. 2</p> <p>Diagnosis</p> <p>Body splayed and gracile, SVL 89.3–125.3 mm; external nares bordered by 6–8 nasal scales; supraorbital region with a row of enlarged tubercles; outer surface of upper eyelid composed of granular scales, about the same size of those on top of head and with a row of 6–10 enlarged tubercles; internasals absent; supralabials 8–11; infralabials 7–10; eyelid fringe scales 45–56; postmentals 2–5; gular region below lower jaws without enlarged tubercles; paravertebral tubercles 31–38; scale rows around midbody 112– 127, granular scales surrounding tubercles 8–11; axillary pockets deep; subdigital lamellae under fourth toe 22–25; precloacal pores in males 16–23; iris orange-brown; dorsal ground color of head, body and limbs gray-brown to pale brown and mottled with dark brown blotches; nuchal loop thin, posteriorly protracted (in V-shape); dorsal body bands between limb insertions 3–4, thin, yellow, without dark spots; light band on tail base in width of 8–9 scales; ground color of tail black, caudal bands 5, white, completed ventrally; ventral surface of head, body and limbs dull white and immaculate, gular region with brown spots (modified after Ziegler et al. 2008; Nguyen 2011).</p> <p>Description (Supp. file 1: Table S2)</p> <p>Body splayed and gracile, adult males: SVL 92.9–125.3 mm (mean ± SE: 112.3± 0.8 mm, n =80), TaL 7.2– 97.9 mm (75.2 ± 1.9 mm); adult females: SVL 89.3–122.1 mm (111.8± 0.8 mm, n =93), TaL 17.5– 98.3 mm (70.3 ± 1.7 mm); juveniles: SVL 53.4–78.8 mm (68.8 ± 1.8 mm, n =21), TaL 11.8–61.2 mm (48.1 ± 2.6 mm) (Supp. file 1: Table S2); head triangular, wider than neck, covered by uniform granular scales interspersed with tubercles in temporal and occipital regions; scales on rostrum slightly larger and flatter; enlarged supraorbital tubercles in a conspicuous row; middorsal portion of rostral partially sutured dorsomedially, bordered laterally by first supralabial on each side, dorsolaterally by prenasal on each side, and dorsally by two supranasals; internasal (postrostral) scales absent; external nares bordered by 6–8 nasals: anteriorly by prenasal and supranasal, dorsally by supranasal, posteriorly by two slightly enlarged postnasals and 1 or 2 smaller granular scales, and ventrally by prenasal; prenasals with long recurved ventral portion; supranasals triangular, meeting in midline behind rostral suture; preorbital scales 10–13; supralabials 8–11; infralabials 7–10; eyes relatively large, pupils vertical; eyelid fringe scales 45–56, those of upper eyelid slightly enlarged; outer surface of upper eyelid composed of granular scales of about the same size of those on top of head, including a row of 6–10 enlarged tubercles; fold of skin originating from suborbital region extends posteroventrally across angle of jaw; external auditory meatus elliptical with long axis directed dorsoventrally; tympanum deeply recessed; mental triangular, bordered laterally by first infralabial on each side and posteriorly by 2–5 postmentals; postmentals bordered by 6–10 gular scales; gular region below lower jaws without enlarged tubercles; gular scales juxtaposed and granular, abruptly grading posteriorly into flat hexagonal pectoral scales and even larger ventral scales.</p> <p>Neck narrower than body, covered with uniform granular scales interspersed with several sharply pointed conical tubercles on nape; tubercles on flanks conical, those of vertebral region somewhat lower in profile; dorsal body tubercles surrounded by 9–11 granular scales; dorsal tubercle rows at midbody 19–24; paravertebral tubercles between limb insertions 31–38, distinct vertebral row of tubercles absent; scales around midbody 112–127; larger ventral scales grade abruptly into smaller granular scales immediately anterior to vent at level of preanal pores; males with 18–23 precloacal pores in a transverse continuous series; region posterior to vent covered by flat juxtaposed scales and great hemipenial bulges, containing 2 or 3 enlarged postcloacal tubercles laterally on each side at level of vent; tail long and thin, thick at base, anteriorly with whorls; dorsal tail scales flat, smooth, up to 1.5 times the size of dorsal scales, arranged in more or less regular transverse rows; light band on tail base in width of 8–9 granular scales and with 7–8 tubercles in a transversal series; subcaudals larger than those on dorsal surface of tail.</p> <p>Limbs relatively long and thin, covered dorsally with granular scales interspersed with several tubercles and ventrally with flat juxtaposed to subimbricate scales; dorsal granular scales grade into slightly flattened subimbricate scales on top of pes and manus; hind limbs slightly larger than forelimbs; larger granular scales on ventral surface of pes and manus; axillary pockets deep; subdigital lamellae wide, 9–11 under first finger, 18–21 under fourth finger, 9–12 under first toe, 22–25 under fourth toe; digits laterally compressed, increasing in length from first to fourth, fifth shorter than fourth (modified after Ziegler et al. 2008; Nguyen 2011).</p> <p>Coloration in life (Fig. 2)</p> <p>Dorsal ground color of head, body and limbs grey brown (in animals with SVL 53.4–105 mm) and blotches chestnut brown (in animals with SVL 110–125.3 mm), juveniles without small blotches, adults mottled with few circular blotches on body and limbs and long dark brown blotches on head; few yellow conical tubercles on franks of neck and body, and limbs in adults; iris light orange or red brown; five bands on the dorsal ground, thin, immaculate without dark spots, cream in juveniles and light orange or yellow in adults, all edged anteriorly and posteriorly by thin dark brown bands, including one thin nuchal loop extending from posterior corners of eyes and posteriorly protracted (in V-shape), three body bands between limb insertions, and another one on tail base; ground color of tail dark brown, and grey brown at mottled tail base; 3–5 immaculate white caudal bands, edged anteriorly and posteriorly in black; ventral surface of head, body and limbs dull white, juveniles absolutely immaculate, but adults with few dark spots on limbs, weak brown lateral spotting in gular region, venter and limbs.</p></div> 	http://treatment.plazi.org/id/9D481962FF9DB2194B13FCC4FB6E3B67	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ngo, Hai Ngoc;Nguyen, Huy Quoc;Tran, Hieu Minh;Ngo, Hanh Thi;Gewiss, Laurenz Rafael;Schingen-Khan, Mona van;Nguyen, Truong Quang;Ziegler, Thomas	Ngo, Hai Ngoc, Nguyen, Huy Quoc, Tran, Hieu Minh, Ngo, Hanh Thi, Gewiss, Laurenz Rafael, Schingen-Khan, Mona van, Nguyen, Truong Quang, Ziegler, Thomas (2021): A morphological and molecular review of the genus Goniurosaurus, including an identification key. European Journal of Taxonomy 751 (1): 38-67, DOI: https://doi.org/10.5852/ejt.2021.751.1379, URL: http://dx.doi.org/10.5852/ejt.2021.751.1379
9D481962FF96B21E4BB4FED7FD693FA4.text	9D481962FF96B21E4BB4FED7FD693FA4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Goniurosaurus luii Grismer, Viets & Boyle 1999	<div><p>Goniurosaurus luii Grismer, Viets &amp; Boyle, 1999</p> <p>Fig. 4</p> <p>Diagnosis</p> <p>Body splayed and gracile, SVL 86.5–126.5 mm; external nares bordered by 5–8 nasal scales; supraorbital region with a row of enlarged tubercles; outer surface of upper eyelid composed of granular scales, about one half the size of those on top of head and without enlarged tubercles; internasals 1 or 2; supralabials 8–12; infralabials 8–11; preorbital scales 13–16; eyelid fringe scales 46–61; postmentals 2–6; gular region below lower jaws with enlarged tubercles; paravertebral tubercles 29–38; scale rows around midbody 119– 144, granular scales surrounding tubercles 9–14; axillary pockets deep; subdigital lamellae under fourth toe 20–26; precloacal pores in males 23–32; iris brown or bright orange; dorsal ground color of head, body and limbs pale brown to grey brown, mottled with small dark brown blotched; nuchal loop thin, posteriorly protracted (in V-shape); dorsal body bands between limb insertions 3, thin, immaculate yellow; gular region, belly, and ventral surface of limbs with brown spots (Grismer et al. 1999; Vu et al. 2006; Nguyen 2011).</p> <p>Description (Supp. file 1: Table S2)</p> <p>Body splayed and gracile, males: SVL 88.8–123.0 mm (mean ±SE: 109.3 ± 1.4 mm, n =34), TaL 2.8–96.8 mm (72.5 ± 3.8 mm); adult females: SVL 86.5–126.5 mm (112.4 ± 1.3 mm, n=43), TaL 43.3–102.2 mm (72.2± 2.1 mm); juveniles: SVL 55.0– 84.8 mm (74.8 ± 3.1 mm, n=11), TaL 44.2– 68.9 (58.7 ± 2.7 mm) (Supp. file 1: Table S2); head triangular, wider than neck, covered by uniform granular scales interspersed with tubercles in temporal and occipital regions; scales on rostrum slightly larger and flatter; enlarged supraorbital tubercles in a conspicuous row; middorsal portion of rostral partially sutured dorsomedially, bordered laterally by first supralabial on each side, dorsolaterally by prenasal on each side, and dorsally by 1 or 2 internasal and two supranasals; internasals 1 or 2; external nares bordered by 5–8 nasal scales (prenasal, anterior and posterior supranasals, 2 slightly enlarged postnasal, and 1–3 granular scales); preorbital scales 13–16; supralabials 8–12, grading into granular scales posteriorly; infralabials 8–11; eyes large, pupils vertical; eyelid fringe scales 46–56, those of upper eyelid slightly enlarged; outer surface of upper eyelid composed of granular scales, about one half the size of those on top of head and without enlarged tubercles; fold of skin originating in suborbital region extends posteroventrally across angle of jaw; external auditory meatus elliptical with long axis directed dorsoventrally; tympanum deeply recessed; mental triangular, bordered laterally by first infralabial on each side and posteriorly by 2–6 postmentals; postmentals bordered by 6–11 gular scales; gular region below lower jaws with enlarged tubercles; gular scales juxtaposed and granular, abruptly grading posteriorly into flat hexagonal pectoral scales and even larger ventral scales.</p> <p>Neck narrower than body, covered with uniform granular scales interspersed with several sharply pointed conical tubercles on nape; tubercles on flanks conical, those of vertebral region somewhat more flat; dorsal body tubercles surrounded by 11–13 granular scales; dorsal tubercle rows at midbody 20–24; paravertebral tubercles between limb insertions 29–38, distinct vertebral row of tubercles absent; scale rows around midbody 119–144; ventral scales large; males with 24–32 precloacal pores in a transverse continuous series, females without precloacal pores (but pitted scales present); region posterior to vent covered by flat juxtaposed scales and greatly swollen, containing 1–3 enlarged tubercles on each side at level of vent; tail thick at base, light band on tail base in width of 9–13 granular scales with 9–10 tubercles in transversal series; ventral caudals of tail base larger than dorsal caudals.</p> <p>Limbs relatively long and thin, covered dorsally with granular scales interspersed with several tubercles and ventrally with flat juxtaposed to subimbricate scales; dorsal granular scales grade into slightly flattened subimbricate scales on top of pes and manus; hind limbs larger than forelimbs; axillary pockets deep; subdigital lamellae wide, 9–12 under first finger, 17–22 under fourth finger, 10–12 under first toe,</p> <p>20–26 under fourth toe; digits laterally compressed, increasing in length from first to fourth, fifth shorter than fourth (Grismer et al. 1999; Vu et al. 2006; Nguyen 2011).</p> <p>Coloration in life (Fig. 4)</p> <p>Dorsal ground color of head, body and limbs pale brown to grey brown, mottled with small dark brown circle blotches, juveniles without small dark brown blotches (except juveniles SVL ≥ 72 mm initially present on head); iris red brown or bright orange; five bands on dorsal ground, thin, immaculate without dark spots, slight bisque in juveniles, orange brown or yellow in adults, all edged anteriorly and posteriorly by thin dark brown bands, including one thin nuchal loop extending from posterior corners of eyes and posteriorly protracted (in V-shape), three body bands between limb insertions, and another one on tail base; ground color of tail nearly solid black, and grey brown at mottled tail base; 3–6 immaculate white caudal bands; ventral surfaces of head, body and limbs dull white with a few dark brown spots in gular region, on belly and limbs.</p></div> 	http://treatment.plazi.org/id/9D481962FF96B21E4BB4FED7FD693FA4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ngo, Hai Ngoc;Nguyen, Huy Quoc;Tran, Hieu Minh;Ngo, Hanh Thi;Gewiss, Laurenz Rafael;Schingen-Khan, Mona van;Nguyen, Truong Quang;Ziegler, Thomas	Ngo, Hai Ngoc, Nguyen, Huy Quoc, Tran, Hieu Minh, Ngo, Hanh Thi, Gewiss, Laurenz Rafael, Schingen-Khan, Mona van, Nguyen, Truong Quang, Ziegler, Thomas (2021): A morphological and molecular review of the genus Goniurosaurus, including an identification key. European Journal of Taxonomy 751 (1): 38-67, DOI: https://doi.org/10.5852/ejt.2021.751.1379, URL: http://dx.doi.org/10.5852/ejt.2021.751.1379
9D481962FF94B2044BBCFD29FEA93932.text	9D481962FF94B2044BBCFD29FEA93932.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Goniurosaurus lichtenfelderi (Mocquard 1897)	<div><p>Goniurosaurus lichtenfelderi (Mocquard, 1897)</p> <p>Fig. 5</p> <p>Diagnosis</p> <p>Body robust; SVL 80.6−113.5 mm; external nares bordered by 5–10 nasal scales; supraorbital region with a row of slightly enlarged tubercles; outer surface of upper eyelid composed of granular scales, about one-half the size of those on top of head and without enlarged tubercles; internasals 1–5 (rarely 1:2; 2:2 or 2:3); supralabials 7–10; infralabials 6–9; preorbital scales 12–19; eyelid fringe scales 43–58; postmentals 2–6; paravertebral tubercles 22−33; scale rows around midbody 117–131, granular scales surrounding tubercles 10−13; axillary pockets shallow; subdigital lamellae under fourth toe 17−21; precloacal pores in males 25–33, in females 17−21; dorsal ground color of head, body and limbs dark purple-brown, without small dark brown blotches; transverse body bands 4, nuchal loop thin, posteriorly rounded, in U-shape; dorsal body bands between limb insertions 2, thin, light yellow; gular region and without dark spots; ventral surfaces of head, body and limbs dull white with a few dark dots on margin regions (modified after Grismer 2000; Grismer et al. 2002; Nguyen 2011).</p> <p>Description (Supp. file 1: Table S2)</p> <p>Body robust, adult males: SVL 80.6–113.5 mm (mean ± SE: 97.8 ± 0.7 mm, n =92), TaL 6.2– 84.8 mm (61.5± 1.7 mm); adult females: SVL 81.0– 105.5 mm (96.0± 0.7 mm, n=72), TaL 22.7– 81.2 mm (58.4 ± 1.4 mm); juveniles: SVL 41.1–77.1 mm (65.4± 2.9 mm, n=14), TaL 20.9–64.7 mm (48.3 ± 3.4 mm) (Supp. file 1: Table S2); head triangular, wider than neck, covered by uniform granular scales interspersed with tubercles in temporal and occipital regions; scales on rostrum slightly larger and flatter; enlarged supraorbital tubercles in a conspicuous row; middorsal portion of rostral partially sutured dorsomedially, bordered laterally by first supralabial on each side, dorsolaterally by prenasal on each side, and dorsally by 1 or 2 internasals and two supranasals; internasals 1–5 (rarely 1:2; 2:2 or 2:3); external nares bordered by 5–10 nasals; preorbital scales 12–18; supralabials 7–10, grading into granular scales posteriorly; infralabials 6–9; eyes large, pupils vertical; eyelid fringe scales 47–58, those of upper eyelid slightly enlarged; outer surface of upper eyelid composed of granular scales of about one-half the size of those on top of head, without enlarged tubercles; fold of skin originating in the suborbital region extends posteroventrally across angle of jaw; external auditory meatus elliptical; tympanum deeply recessed; mental triangular, bordered laterally by first infralabial on each side and posteriorly by 2–5 postmentals; postmentals bordered by 7–10 gular scales; gular region below lower jaws without enlarged tubercles; gular scales juxtaposed and granular, abruptly grading posteriorly into flat hexagonal scales and even larger ventral scales.</p> <p>Neck narrower than body, covered with uniform granular scales interspersed with several sharply conical tubercles on nape; tubercles on flanks conical, those of vertebral region somewhat more flat; dorsal body tubercles surrounded by 10–13 granular scales; dorsal tubercle rows at midbody 15–22; paravertebral tubercles between limb insertions 22–33, distinct vertebral row of tubercles absent; scales around midbody 117–130; ventral scales large; males with 24–33 precloacal pores in a transverse continuous series, females without distinct precloacal pores (but 25–33 pitted scales present); region posterior to vent covered by flat juxtaposed scales and greatly swollen, containing 1 (rarely 2) enlarged tubercles on each side at the level of vent; tail thick at base; light band on tail base in width of 4–7 scales and with 6–10 tubercles in a transversal series.</p> <p>Limbs robust, covered dorsally with granular scales interspersed with several tubercles and ventrally with flat juxtaposed to subimbricate scales; dorsal granular scales grade into slightly flattened subimbricate scales on top of pes and manus; hind limbs larger than forelimbs; axillary pockets shallow; subdigital lamellae wide, 6–9 under first finger, 13–17 under fourth finger, 6–10 under first toe, 16–20 under fourth toe; digits laterally compressed, increasing in length from first to fourth, fifth shorter than fourth (modified after Grismer 2000; Grismer et al. 2002; Nguyen 2011).</p> <p>Coloration in life (Fig. 5)</p> <p>Dorsal ground color of head, body and limbs grey and grey brown in juveniles without blotches and chestnut brown in adults with dark brown blotches (in animals with SVL&gt; 80 mm); iris orange or redish brown; four bands on the dorsal ground, thin, yellow in both juveniles and adults, immaculate without dark spots (few dark brown spots present in some animals), all edged anteriorly and posteriorly by thin dark brown bands, including one thin nuchal loop extending from posterior corners of eyes and posteriorly rounded (in U-shape), two body bands between limb insertions, and another one on tail base; ground color of tail dark brown, and grey brown at mottled tail base; 3–5 immaculate white caudal bands except first band slightly yellow, and some animals with regenerated tail present immaculate grey brown without white caudal bands; ventral surfaces of head, body and limbs dull white with a few dark dots on margin regions of belly, limbs, gular region, but immaculate dull white in juveniles (SVL ≤ 65 mm) without dark spots.</p> <p>Morphological comparisons</p> <p>Our morphological comparisons of the tiger gecko species (except for G. araneus) from Vietnam revealed an overall significant difference in the snout-vent length (SVL). As the result, G. huuliensis had the largest SVL, while G. lichtenfelderi had the shortest SVL in both adult males and females, compared to the other tiger geckos (P &lt;0.05). However, the SVL of G. catbaensis was not significantly different from that of G. luii (P&gt; 0.05, Supp. file 2: Fig. S1).</p> <p>The PCA of 18 selected morphometric characters estimated the first (PC1) and second principal (PC2) components to explain 82.5% (74.5% and 8.0%, respectively) of the variance among the four investigated species of Goniurosaurus in Vietnam (Supp. file 2: Fig. S2). The extracted PC1 and PC2 scores of the PCA differ significantly among the four species (Kruskal-Wallis’s test, P &lt;0.05), of which the values and morphometric spaces of G. lichtenfelderi are not concordant with the three remainders (Fig. S2). Three morphological factors, namely head length (HL), head width (HW), and mouth width (MW) highly account for the overall difference in the PC1 score, while the PC2 score is mainly explained by the body height (BH) and body width (BW) (Supp. file 2: Fig. S2).</p> <p>Regarding the comparisons of the 17 species of Goniurosaurus from China and Vietnam, the multiple correspondence analysis (MCA) clustered them separately into three groups (G. lichtenfelderi group, G. luii group and G. yingdeensis group) represented with different meristic spaces on the spatial coordinate of Dim 1 and Dim 2 (Fig. 6A, Table 1). The characters of fourth toe (LT4), body limb shape and body bands highly account for the Dim1 score, whereas the characters of precloacal pores (PP) and granular scales surrounding dorsal tubercles (GST) are the most correlated with the Dim 2 score (Fig. 6B).</p> </div>	http://treatment.plazi.org/id/9D481962FF94B2044BBCFD29FEA93932	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ngo, Hai Ngoc;Nguyen, Huy Quoc;Tran, Hieu Minh;Ngo, Hanh Thi;Gewiss, Laurenz Rafael;Schingen-Khan, Mona van;Nguyen, Truong Quang;Ziegler, Thomas	Ngo, Hai Ngoc, Nguyen, Huy Quoc, Tran, Hieu Minh, Ngo, Hanh Thi, Gewiss, Laurenz Rafael, Schingen-Khan, Mona van, Nguyen, Truong Quang, Ziegler, Thomas (2021): A morphological and molecular review of the genus Goniurosaurus, including an identification key. European Journal of Taxonomy 751 (1): 38-67, DOI: https://doi.org/10.5852/ejt.2021.751.1379, URL: http://dx.doi.org/10.5852/ejt.2021.751.1379
9D481962FF8EB2064AFAFBB0FAEC3E75.text	9D481962FF8EB2064AFAFBB0FAEC3E75.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Goniurosaurus Grismer, Viets & Boyle 1999	<div><p>Key to the species of Goniurosaurus (Fig. 7, Table 1)</p> <p>Modified from Grismer et al. (1999, 2002), Vu et al. (2006), Orlov et al. (2008), Ziegler et al. (2008), Wang et al. (2010, 2013, 2014), Nguyen (2011), Chen et al. (2014), Yang &amp; Chan (2015), Honda &amp; Ota (2017), Zhou et al. (2018, 2020a, 2020b), Qi et al. (2020a, 2020b) and the present study.</p> <p>1. Precloacal pores in males present, claws are sheathed by scales...................................................... 7</p> <p>– Precloacal pores absent, unsheathed claws (G. kuroiwae group)..................................................... 2</p> <p>2. Yellow brown to gold iris; a single scale at the base of each digit which is occasionally slightly enlarged............................................................................................... G. yamashinae (Okada, 1936)</p> <p>– Blood-red iris; one to three enlarged scales at the base of each digit on the limbs.......................... 3</p> <p>3. Adults without interspace mottling................................................................................................... 4</p> <p>– Adults with interspace mottling........................................................................................................ 5</p> <p>4. Robust body stature; dorsal body tubercles between the limb insertions in cross-section are triangular to elliptical and sharply keeled anteriorly; 34–42 paravertebral tubercles; ventral scales are juxtaposed and sharply raised..................................... G. toyamai Grismer, Ota &amp; Tanaka, 1994</p> <p>– Slender body stature; dorsal body tubercles between the limb insertions in cross-section are smooth or very weakly keeled; 22–29 paravertebral tubercles; ventrals are flat, wide, and subimbricate to imbricate............................................................................ G. splendens (Nakamura &amp; Uéno, 1959)</p> <p>5. Dorsal banding absent or incomplete................................................... G. kuroiwae (Namiye, 1912)</p> <p>– Prominent dorsal pattern consisting of transverse bands between the nape of the neck and the caudal constriction....................................................................................................................................... 6</p> <p>6. No such reddish or pinkish tint in dorsal pattern and iris......................... G. orientalis (Maki, 1931)</p> <p>– Reddish or pinkish tint to some extent in dorsal pattern and iris.............................................................................................................................................................. G. sengokui (Honda &amp; Ota, 2017)</p> <p>7. Precloacal pores in males less than 16 (G. yingdeensis group)........................................................ 8</p> <p>– Precloacal pores in males more than 16..........................................................................................11</p> <p>8. Scales around midbody 121–128; longitudinal dorsal tubercle rows at midbody 16–17...................................................................................... G. gollum Qi, Wang, Grismer, Chen, Lyu &amp; Wang, 2020</p> <p>– Scales around midbody 99–115; longitudinal dorsal tubercle rows at midbody 20–28................... 9</p> <p>9. Nuchal loop and body bands with small dark blotches; enlarged row of supraorbital tubercles absent; trunk of body usually with a longitudinal light vertebral stripe................................................................................................................................. G. varius Qi, Grismer, Lyu, Zhang, Li &amp; Wang, 2020</p> <p>– Nuchal loop and body bands without dark blotches; enlarged row of supraorbital tubercles present; trunk of body without a longitudinal light vertebral stripe............................................................. 10</p> <p>10. Tubercles between orbits present; gular scales bordering the postmentals 2–4; preorbital scales 5–7............................................................................................ G. yingdeensis Wang, Yang &amp; Cui, 2010</p> <p>– Tubercles between orbits absent; gular scales bordering the postmentals 4–6; preorbital scales 7–9........................................................................................ G. zhelongi Wang, Jin, Li &amp; Grismer, 2014</p> <p>11. Body and limbs robust (G. lichtenfelderi group)............................................................................ 12</p> <p>– Body and limbs splayed gracile (G. luii group).............................................................................. 16</p> <p>12. Precloacal pores in males 37–46....... G. bawanglingensis Grismer, Haitao, Orlov &amp; Anajeva, 2002</p> <p>– Precloacal pores in males fewer than 37......................................................................................... 13</p> <p>13. Nuchal loop protracted posteriorly (in V-shape); number of body bands 4; axillary pockets deep; precloacal pores in males more than 33......................... G. zhoui Zhou, Wang, Chen &amp; Liang, 2018</p> <p>– Nuchal loop rounded posteriorly (in U-shape); number of body bands 3; axillary pockets shallow; precloacal pores in males fewer than 33......................................................................................... 14</p> <p>14. Eyelid fringe scales 55–70; enlarged row of supraorbital tubercles absent; scales of upper eyelid to top of head equal of those on the top of the head in size.................... G. hainanensis Barbour, 1908</p> <p>– Eyelid fringe scales 47–58; enlarged row of supraorbital tubercles present; scales of upper eyelid to top of head ½ of those on the top of the head in size..................................................................... 15</p> <p>15. Lateral spotting on belly absent; preorbital scales 18–21; scales around midbody 109–118; subdigital lamellae under the fourth finger 18–19.............................................. G. kwanghua Zhu &amp; He, 2020</p> <p>– Lateral spotting on belly present; preorbital scales 12–18; scales around midbody 117–130; subdigital lamellae under the fourth finger 13–17...................................... G. lichtenfelderi (Mocquard, 1897)</p> <p>16. Internasal absent........................... G. catbaensis Ziegler, Nguyen, Schmitz, Stenke &amp; Rösler, 2008</p> <p>– Internasal present............................................................................................................................ 17</p> <p>17. Postcloacal tubercles 3–6................................................. G. araneus Grismer, Viets &amp; Boyle, 1999</p> <p>– Postcloacal tubercles 1–3................................................................................................................ 18</p> <p>18. Adult body length (SVL)&gt; 126 mm; dorsum without dark blotches............................................................................................................... G. huuliensis Orlov, Ryabov, Nguyen, Nguyen &amp; Ho, 2008</p> <p>– Maximum body length (SVL) ≤ 126 mm; dorsum with dark blotches.......................................... 19</p> <p>19. Dorsum with many small dark blotches.................................. G. luii Grismer, Viets &amp; Boyle, 1999</p> <p>– Dorsum with rarely scattered dark blotches................................................................................... 20</p> <p>20. Outer surface of upper eyelid composed of granular scales, about the same size of those on top of head and with enlarged tubercles.......................................................... G. gezhi Zhu, He &amp; Li, 2020</p> <p>– Enlarged row of supraorbital tubercles present; scales of upper eyelid to top of head ½ of those on the top of the head in size............................................................................................................... 21</p> <p>21. Precloacal pores in males 31–33.............................................. G. kwangsiensis Yang &amp; Chan, 2015</p> <p>– Precloacal pores in males fewer than 31......................................................................................... 22</p> <p>22. Lateral spotting on belly present; nasal scales 6–7; paravertebral tubercles between limb insertions 30–34; precloacal pores 26–28............................................... G. kadoorieorum Yang &amp; Chan, 2015</p> <p>– Lateral spotting on belly absent, nasal scales 8–9; paravertebral tubercles between limb insertions 27–28; precloacal pores 23.............................................. G. liboensis Wang, Yang &amp; Grismer, 2013</p></div> 	http://treatment.plazi.org/id/9D481962FF8EB2064AFAFBB0FAEC3E75	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ngo, Hai Ngoc;Nguyen, Huy Quoc;Tran, Hieu Minh;Ngo, Hanh Thi;Gewiss, Laurenz Rafael;Schingen-Khan, Mona van;Nguyen, Truong Quang;Ziegler, Thomas	Ngo, Hai Ngoc, Nguyen, Huy Quoc, Tran, Hieu Minh, Ngo, Hanh Thi, Gewiss, Laurenz Rafael, Schingen-Khan, Mona van, Nguyen, Truong Quang, Ziegler, Thomas (2021): A morphological and molecular review of the genus Goniurosaurus, including an identification key. European Journal of Taxonomy 751 (1): 38-67, DOI: https://doi.org/10.5852/ejt.2021.751.1379, URL: http://dx.doi.org/10.5852/ejt.2021.751.1379
