taxonID	type	description	language	source
B510B805C1405B1081A1B04ACA258316.taxon	diagnosis	Diagnosis. Otoba scottmorii is similar to O. cyclobasis from Ecuador. However, it differs in having smaller leaves (5.5 - 8.5 [- 11.2] vs. 9 - 14 [- 18] cm long) with an attenuate to acuminate apex (vs. cuspidate), inconspicuous marginal and secondary veins (vs. conspicuous), fewer lateral veins (4 - 7 vs. 14 - 17), straight axes of staminate inflorescences (vs. zig zag), a longer filament column ([1.3 -] 1.5 - 1.6 vs. 1 mm long), larger fruits (1.8 - 2.1 vs. 1.5 - 2.5 mm long) with thinner pericarp (0.68 - 0.89 vs. 1 - 2 mm thick), and seeds that are gibbose at the apex (vs. near-basal and lateral gibba).	en	Santamaria-Aguilar, Daniel, Lagomarsino, Laura P. (2021): Two new species of Otoba (Myristicaceae) from Colombia. PhytoKeys 178: 147-170, DOI: http://dx.doi.org/10.3897/phytokeys.178.64564, URL: http://dx.doi.org/10.3897/phytokeys.178.64564
B510B805C1405B1081A1B04ACA258316.taxon	description	Description. Tree (2 -) 15 - 20 m tall x 45 cm diam., external and internal bark not described. Exudate once described as red in the bark, transparent and without specifying from where, or without exudate. Twigs 0.88 - 0.1 cm thick, terete to slightly flattened laterally, the external bark brown to grayish, with malpighiaceous trichomes that are 0.2 - 0.3 mm long, brown to ferruginous or sometimes glabrescent. Young foliar bud 0.6 - 1.2 cm long, densely pubescent. Leaves: petiole 0.5 - 1.1 x 0.055 - 0.1 cm, canaliculate, not winged; lamina 5.5 - 8.5 (- 11.2) x 1.7 - 2.5 (- 3.7) cm, elliptic; adaxial side glabrous, drying blackish, sometimes brown in young leaves, the surface muricate; abaxial side drying brown to whitish grayish, the surface wrinkled to muricate, sparsely pubescent to glabrescent, with malpighiaceous trichomes that are 0.2 - 0.4 mm long, sessile, and ferruginous, and squamate trichomes that are ca. 0.08 mm diam. with the central portion dark and sides lighter (sometimes these sides appearing absent), with whitish crystals; vernation line absent; midvein flat to very slightly ribbed on adaxial side, the same color as the surface, abaxially 0.2 - 0.4 mm wide, slightly raised, darker than the surface; secondary veins brochidodromous, the loops 0.1 - 0.3 cm from the margin (sometimes the loops not visible), lateral veins 4 - 7 per side, 3 - 5 veins per 3 cm, on adaxial side flat and visible, on abaxial side flat and not very conspicuous, arcuate distally, the marginal vein not visible on adaxial side, slightly visible on abaxial side; tertiary veins indistinct; base attenuate, not revolute; margin entire, not revolute; apex attenuate to acuminate, the acumen 0.6 - 1 (- 2) cm long. Staminate inflorescence: axillary (only very young twigs) or supraaxillary, with 1 - 2 (- 3) main axes, spiciform, these axes 1.5 - 4.3 cm x 0.41 - 0.1 mm, pubescent, the trichomes ferruginous to coppery, each axis compound with 2 - 3 (- 6) fascicles of flowers, each fascicle with 1 - 5 (- 8) flowers, alternate; bracts not seen; pedicel 3 - 5 mm long, pubescent; bracteoles absent. Staminate flowers: flower bud 1 - 2.5 x 0.6 - 0.1 mm, elliptic to lanceolate; perianth 2 - 2.5 mm long, yellow or orange (in fresh material), sub-membranaceous, connate by 0.5 - 0.7 (- 1) mm; lobes 3 (4), 1.5 - 1.8 x 0.8 - 1.1 mm, without resinous punctuations or lines, pubescent outside, the trichomes ferruginous to coppery, inside glabrous, the apex acute to obtuse, without inflexed-apiculo, the edges flat or slightly turned inwards distally, without a swollen lobed ring; filament column (1.3 -) 1.5 - 1.6 mm long, bowling pin-shaped, fleshy, connate, glabrous; anthers 3, 0.2 - 0.4 mm long, free, lanceolate to oblong, apex slightly incurved. Pistillate inflorescence and flowers: not seen. Infructescence 2 - 4 cm long, with a solitary fruit; pedicel 1.2 - 1.6 cm long. Fruits 1.8 - 2.1 x 1.5 - 1.7 cm, green when fresh, globose, surface glabrous, colliculate-rugose, sometimes with whitish lenticels (J. Brand 682), the line of dehiscence smooth, the base obtuse, apex apiculate, the acumen 0.1 - 0.3 cm long; pericarp 0.68 - 0.89 mm thick; seed ca. 1.6 x 1.5 cm, similar in shape to the fruit, brownish (in dry material), gibbose at the apex (ca. 0.3 mm wide), the testa ca. 0.4 mm thick; aril described once as red (E. Renteria 4680), brownish to yellowish translucent in dry material, dry and membranous in texture.	en	Santamaria-Aguilar, Daniel, Lagomarsino, Laura P. (2021): Two new species of Otoba (Myristicaceae) from Colombia. PhytoKeys 178: 147-170, DOI: http://dx.doi.org/10.3897/phytokeys.178.64564, URL: http://dx.doi.org/10.3897/phytokeys.178.64564
B510B805C1405B1081A1B04ACA258316.taxon	description	Distinctive characters. Otoba scottmorii is recognized by a combination of leaf, inflorescence, and fruit traits. Its small leaf blades have long apices and thin petioles, and lack vernation lines on the abaxial surface. The staminate inflorescences are delicate with flowers on relatively thin, long pedicels; these flowers have sub-membranaceous perianth and lack a swollen-lobed ring, a bowling pin-shaped filament column, and anthers that are lanceolate to oblong. Finally, fruits are small with thin pericarp and membranous aril.	en	Santamaria-Aguilar, Daniel, Lagomarsino, Laura P. (2021): Two new species of Otoba (Myristicaceae) from Colombia. PhytoKeys 178: 147-170, DOI: http://dx.doi.org/10.3897/phytokeys.178.64564, URL: http://dx.doi.org/10.3897/phytokeys.178.64564
B510B805C1405B1081A1B04ACA258316.taxon	etymology	Etymology. Is a pleasure to name this species after Dr. Scott A. Mori (1941 - 2020), a wonderful person and skilled botanist; a dedicated explorer of Central and South America humid forests (where this species occurs), especially in the Guianas and the Amazon basin; and an authority on Neotropical Lecythidaceae. His taxonomic and ecological publications gave great inspiration to the first author, as did Dr. Mori's personal support. For an account of Dr. Mori's legacy, see Boom (2020), Forget (2020), and Prance et al. (2021).	en	Santamaria-Aguilar, Daniel, Lagomarsino, Laura P. (2021): Two new species of Otoba (Myristicaceae) from Colombia. PhytoKeys 178: 147-170, DOI: http://dx.doi.org/10.3897/phytokeys.178.64564, URL: http://dx.doi.org/10.3897/phytokeys.178.64564
B510B805C1405B1081A1B04ACA258316.taxon	distribution	Distribution. Otoba scottmorii is known only from the humid forests in the Department of Antioquia in northwestern Colombia in the Municipios of Mutata, Segovia, Taraza, Turbo, and Valdivia (Fig. 4). Three collections were collected under 100 m elevation (20 - 80 m), and three others above 400 m elevation (410 - 730 m).	en	Santamaria-Aguilar, Daniel, Lagomarsino, Laura P. (2021): Two new species of Otoba (Myristicaceae) from Colombia. PhytoKeys 178: 147-170, DOI: http://dx.doi.org/10.3897/phytokeys.178.64564, URL: http://dx.doi.org/10.3897/phytokeys.178.64564
B510B805C1405B1081A1B04ACA258316.taxon	discussion	Discussion. Otoba scottmorii is similar to O. acuminata and O. vespertilio D. Santam. & J. E. Jimenez, from Costa Rica and Panama, and O. cyclobasis T. S. Jaram. & Balslev from Ecuador. They all have relatively small leaf blades of similar shape and thin petioles, and usually lack vernation lines (Figs 5, 6); staminate flowers with short perianth and small anthers; and similarly sized fruit with thin pericarp (Fig. 5). The four species can be distinguished by the characteristics in Table 2. As mentioned in the introduction, Otoba scottmorii was confused with the Mesoamerican species O. acuminata, which we now consider to be endemic to the Caribbean slope of Costa Rica and Panama. The specimens referred to as O. acuminata from Colombia in recent floras or checklists (e. g., Cogollo et al. 2007; Cogollo 2011; Ulloa Ulloa et al. 2017; Jaramillo-Vivanco and Balslev 2020) correspond to O. scottmorii. The confusion between these two species is likely driven by the similar morphology of the leaf blades (i. e., size and shape, usually without vernation lines, and thin petioles). However, O. scottmorii differs from O. acuminata in having narrower leaf blades (Fig. 5 B, A respectively), staminate flowers with short perianth and narrow perianth lobes, a longer column of filaments, and fruits with a smooth surface and thin pericarp (Table 2). Although we have not observed the pistillate flowers of O. scottmorii, it is likely that the ovary is glabrous as the fruits do not have traces of trichomes, while in O. acuminata the ovary is pubescent; see for example: A. Rodriguez et al. 1559 (CR- 2 sheets, MO); A. Estrada et al. 4829 (CR); B. Hammel & M. Grayum 14288 (CR, INPA [digital image], MEXU [digital image], MO). For a list of specimens that correspond to O. acuminata, see Santamaria-Aguilar et al. (2019).	en	Santamaria-Aguilar, Daniel, Lagomarsino, Laura P. (2021): Two new species of Otoba (Myristicaceae) from Colombia. PhytoKeys 178: 147-170, DOI: http://dx.doi.org/10.3897/phytokeys.178.64564, URL: http://dx.doi.org/10.3897/phytokeys.178.64564
B510B805C1405B1081A1B04ACA258316.taxon	materials_examined	Specimens examined. Colombia. Antioquia: Mutata, Sitio Rio Surambay, 12 km N de Mutata, 07 ° 20 ' N, 076 ° 30 ' W, 30 - 80 m, 21 Nov. 1987 (♂ fl), R. Callejas et al. 5752 (INPA digital image, MO, NY); ibid, 21 Nov. 1987 (fr), R. Callejas et al. 5789 (INPA digital image, MO, NY); Turbo, Carretera Tapon del Darien, Sector Rio Leon-Lomas Aisladas, km 37, [07 ° 39 ' 11 " N, 076 ° 58 ' 02 " W], 20 m, 29 Nov. 1993 (fr), J. Brand & M. Narvaez 682 (COL [n. v.], JAUM digital image, MO); rio Cianura, Paso de la Reina, 730 m, 13 Mar 1986 (fr), E. Renteria et al. 4680 (JAUM digital image); Taraza, Corregimiento El 12, 210 kms, NE de Medellin, via El 12 - Barroblanco, km 3, 07 ° 30 ' N, 075 ° 20 ' W, [450 m], 09 Nov. 1987 (♂ fl), R. Callejas et al. 5509 (INPA digital image, NY); Valdivia, corregimiento Puerto Valdivia, km 5 de Puerto Valdivia hacia " El 12 ", colecciones a lo largo del Rio Pescado, 07 ° 20 ' N, 075 ° 20 ' W, 410 m, 14 May. 1987 (♂ fl), R. Callejas et al. 3440 (MO- 2 sheets, NY).	en	Santamaria-Aguilar, Daniel, Lagomarsino, Laura P. (2021): Two new species of Otoba (Myristicaceae) from Colombia. PhytoKeys 178: 147-170, DOI: http://dx.doi.org/10.3897/phytokeys.178.64564, URL: http://dx.doi.org/10.3897/phytokeys.178.64564
95FD5676998159B1A17DBF9200F2466E.taxon	diagnosis	Diagnosis. Otoba squamosa is similar to O. gordoniifolia from Colombia and Ecuador, and both species grow in montane forest in the Andes. However, it differs in its leaves with shorter petioles (1.7 - 2.7 [- 3.8] vs. [3 -] 5 - 7 cm long) and smaller lamina (6.7 - 14.5 vs. [13 -] 24 - 34 cm long), staminate flowers with a perianth with a swollen ring (vs. without) and smaller anthers (0.5 - 0.7 vs. 0.7 - 1.5 mm long), pistillate flowers with a glabrous ovary (vs. pubescent), and fruits with thin pericarp (2.6 - 3 vs. 3 - 5 mm thick).	en	Santamaria-Aguilar, Daniel, Lagomarsino, Laura P. (2021): Two new species of Otoba (Myristicaceae) from Colombia. PhytoKeys 178: 147-170, DOI: http://dx.doi.org/10.3897/phytokeys.178.64564, URL: http://dx.doi.org/10.3897/phytokeys.178.64564
95FD5676998159B1A17DBF9200F2466E.taxon	description	Description. Tree 8 - 18 (- 24) m tall x 11.9 - 34.7 cm diam., external and internal bark not described. Exudate hyaline, oxidizing reddish, only reported from in flowers and fruits. Twigs 0.13 - 0.27 cm thick, terete to slightly flattened laterally, the external bark brown to blackish, with malpighiaceous trichomes 0.2 - 0.6 mm long, brown to ferruginous, the indument denser in young parts. Young foliar bud 1.6 - 4.3 (- 5.8) cm long, densely pubescent. Leaves: petiole 1.7 - 2.7 (- 3.8) x 0.1 - 0.2 (- 0.25) cm, canaliculate, very short-winged; lamina 6.7 - 14.5 x 3.4 - 6 (- 8) cm, elliptic, rarely widely elliptic; adaxial side glabrous, usually drying dark brown to blackish, the surface muricate-reticulate; abaxial side usually drying pale to dark brown, the surface muricate, sparsely pubescent, with malpighiaceous trichomes 0.3 - 0.6 mm long, sessile, ferruginous, and scale-like trichomes ca. 0.1 diam. with the central part dark, contrasting with the lighter sides, crystals generally absent and if present very few; vernation line imprints 2 parallel lines, 0.7 - 1.5 (- 1.8) cm from the margin, the panel area 1.8 - 2.5 (- 3.4) cm wide (in the central portion), the same color as the surface; midvein flat on adaxial side, the same color as the surface or blackish, abaxially 0.5 - 0.9 mm wide, raised, a little darker than the surface; secondary veins brochidodromous, the loops 0.2 - 0.3 cm from the margin, lateral veins 13 - 17 per side, (3 -) 4 - 6 veins per 3 cm, on adaxial side slightly caniculate, on abaxial side flat to slightly raised, not very conspicuous, arcuate distally, the marginal vein not visible on adaxial side, slightly visible on abaxial side; tertiary veins indistinct; base acute to cuneate, not revolute; margin entire, not revolute; apex acute, the acumen 0.3 - 0.9 cm long. Staminate inflorescence: axillary and / or ramiflorus, with 1 - 2 main axes, spiciform, these axes 2 - 7 cm x 0.6 - 1.2 mm, pubescent, the trichomes ferruginous, each axis compound with 2 - 5 fascicles of flowers, each fascicle with 3 - 6 flowers, alternate; bracts ca. 1.5 - 1.6 x 1 mm (observed in very young inflorescences), densely pubescent outside, the trichomes ferruginous; pedicel 1.3 - 4 mm long, pubescent; bracteoles absent. Staminate flowers: flower bud 2 - 3 x 1 - 1.5 mm; perianth 3.5 - 4.7 mm long, yellowish to yellowish-green (in fresh material), fleshy (hardening near the base by the ring), connate by 1 - 1.7 mm; lobes 3 (4), (2 -) 2.5 - 3.4 x (0.9 -) 1.4 - 2 mm, without resinous punctuations or lines, pubescent outside, the trichomes ferruginous, inside glabrous, smooth to lightly spongy, the apex in some flowers with a minutely inflexed-apiculate, the margin edges slightly turned inwards, slightly wavy; ring present, 0.1 - 0.4 mm wide, lobed, smooth, or sometimes spongy; filament column 1.5 - 2.1 mm long, usually cylindrical, slightly narrow towards the apex, fleshy, glabrous; anthers 3 (4), 0.5 - 0.7 mm long, free, lanceolate to oblong, apex slightly incurved. Pistillate inflorescence: axillary, 1.4 - 3.2 cm long, pubescent, the trichomes ferruginous, each axis compound with 1 - 2 fascicles of flowers, each fascicle with 2 - 3 flowers, alternate; bracts not seen; pedicel 3 - 4 mm long, pubescent; bracteoles absent. Pistillate flowers: flower bud 3 - 4 x ca. 2 mm; perianth 3 - 5 mm long, the color and texture as in the staminate flower, connate by 1 - 1.5 mm; lobes 3 (4), 2 - 3.5 x 2 - 2.5 mm, without resinous punctuations or lines, pubescent outside, the trichomes ferruginous, inside glabrous; ring present; gynoecium 2 - 3 x 1.5 - 2.3 mm, glabrous, ovary sessile to short-stalked, ca. 0.6 mm long; stigma 2 - lipped, subsessile; stigmatic lips ca. 0.6 mm long. Infructescence probably with one fruit (fruits separated from the axis in all specimens observed); pedicel ca. 1 - 1.1 cm long. Fruits (2.5 -) 3.4 - 3.6 x (1.9 -) 2.8 - 3 cm, green, globose, the surface glabrous, rugose, sometimes with whitish to brownish lenticels, the line of dehiscence smooth, the base obtuse, and sometimes getting narrower towards the pedicel, apex obtuse or acute, the acumen 0.8 - 1 cm long; pericarp 2.6 - 3 mm thick; seed (2.1 -) 2.5 - 2.9 x (1.7 -) 2.3 - 2.4 cm, similar in shape to the fruit, whitish or brown (in dry material), gibbose at the apex or nearly so, the testa 0.4 - 0.6 mm thick; aril described once as white (D. Sanchez et al. 1529), brownish to white-yellowish in dry material, waxy to dry in texture.	en	Santamaria-Aguilar, Daniel, Lagomarsino, Laura P. (2021): Two new species of Otoba (Myristicaceae) from Colombia. PhytoKeys 178: 147-170, DOI: http://dx.doi.org/10.3897/phytokeys.178.64564, URL: http://dx.doi.org/10.3897/phytokeys.178.64564
95FD5676998159B1A17DBF9200F2466E.taxon	description	Distinctive characters. Otoba squamosa is recognized by a variety of leaf traits, including: squamate or scale-like indument mixed with malpighiaceous trichomes on the abaxial surface, lateral veins that are more or less conspicuous, forming a marginal vein, and vernation lines that parallel the midvein. Additionally, the staminate flowers have a perianth with swollen-lobed ring in the inner surface, a typically cylindrical filament column with lanceolate to oblong anthers and pistillate flowers that have a glabrous gynoecium. Finally, the fruits are relatively large, with thick pericarp.	en	Santamaria-Aguilar, Daniel, Lagomarsino, Laura P. (2021): Two new species of Otoba (Myristicaceae) from Colombia. PhytoKeys 178: 147-170, DOI: http://dx.doi.org/10.3897/phytokeys.178.64564, URL: http://dx.doi.org/10.3897/phytokeys.178.64564
95FD5676998159B1A17DBF9200F2466E.taxon	etymology	Etymology. The specific epithet refers to the squamate or scale-like indument that is present with more typical malpighiaceous trichomes on the abaxial surface of the leaf blades. The squamate or scale-like indument is not unique to this species; it is also present in most specimens of O. acuminata, O. glycycarpa, O. scottmorii, and O. vespertilio; also Smith and Wodehouse (1938), they described for O. lehmannii (as Dialyanthera Lehmannii).	en	Santamaria-Aguilar, Daniel, Lagomarsino, Laura P. (2021): Two new species of Otoba (Myristicaceae) from Colombia. PhytoKeys 178: 147-170, DOI: http://dx.doi.org/10.3897/phytokeys.178.64564, URL: http://dx.doi.org/10.3897/phytokeys.178.64564
95FD5676998159B1A17DBF9200F2466E.taxon	distribution	Distribution. Otoba squamosa is known from the Cordillera Occidental of Colombia, specifically in the municipalities of Frontino and Urrao in the Department of Antioquia (Fig. 4). It grows in premontane forest between 1330 - 1450 m.	en	Santamaria-Aguilar, Daniel, Lagomarsino, Laura P. (2021): Two new species of Otoba (Myristicaceae) from Colombia. PhytoKeys 178: 147-170, DOI: http://dx.doi.org/10.3897/phytokeys.178.64564, URL: http://dx.doi.org/10.3897/phytokeys.178.64564
95FD5676998159B1A17DBF9200F2466E.taxon	discussion	Discussion. The collections now identified as Otoba squamosa were previously included under the concept of O. gordoniifolia (Fig. 8), and identified as such in herbarium specimens and in the literature (e. g., Cogollo 2011). These species share similarities: both grow in montane forests and have the leaf blades with conspicuous vernation lines (Figs 7 A, F, 8 B), long petioles (shorter in the new species), lanceolate to oblong anthers, and fruit with thick pericarp. However, Otoba squamosa differs in its smaller leaf blades with thinner petioles, staminate flowers with a shorter perianth with a swollen-lobed ring in the inner surface and smaller anthers, and pistillate flowers with glabrous ovaries. See Table 3 for a comparison of these distinguishing characteristics. In addition, O. squamosa ' s leaf blades are less pubescent abaxially, the terminal young foliar bud is shorter, and petioles are shorterwinged. Within Otoba, the swollen-lobed ring in the staminate perianth is shared between O. cyclobasis (and even gives this species its specific epithet; Jaramillo and Balslev 2001) and O. squamosa; in addition to this feature, both species have glabrous ovaries. However, the new species has longer petioles (1.7 - 2.7 [- 3.8] vs. 1 - 1.5 cm) and leaves (6.7 - 14.5 vs. 3.4 - 6 [- 8] cm), lanceolate to oblong anthers (vs. globose), and larger fruits ([2.5 -] 3.4 - 3.6 vs. 1.5 - 2.5 cm long); further, O. squamosa grows at higher elevations (1330 - 1450 vs. 150 - 300 m). In the " Key to the species (pistillate or fruiting plants) " from Jaramillo-Vivanco and Balslev (2020), Otoba squamosa matches the recently described O. vespertilio from Costa Rica and Panama, though it is clearly not conspecific. Otoba squamosa differs from O. vespertilio in its longer petioles (1.7 - 2.7 [- 3.8] vs. 0.8 - 2 cm long), abaxially conspicuous vernation lines (vs. absent), staminate perianth with a swollen-lobed ring (vs. without), and larger fruits ([2.5 -] 3.4 - 3.6 vs. 2.2 - 2.7 x 1.6 - 1.7 cm) with thicker pericarp (2.6 - 3 vs. 1.3 - 1.8 mm).	en	Santamaria-Aguilar, Daniel, Lagomarsino, Laura P. (2021): Two new species of Otoba (Myristicaceae) from Colombia. PhytoKeys 178: 147-170, DOI: http://dx.doi.org/10.3897/phytokeys.178.64564, URL: http://dx.doi.org/10.3897/phytokeys.178.64564
95FD5676998159B1A17DBF9200F2466E.taxon	materials_examined	Specimens examined. Colombia. Antioquia: Frontino, Corregimiento Nutibara, cuenca alta del Rio Cuevas, bosque al lado carretera a La Blanquita, 1100 m, 21 Sep. 1987 (♀ fl & fr), D. Sanchez et al. 1529 (MO); Urrao, Parque Nacional Natural " Las Orquideas ", Sector Calles, margen derecha del Rio Calles, 06 ° 32 ' N, 076 ° 19 ' W, 1420 m, 25 Mar. 1988 (fr), A. Cogollo et al. 2573 (MO); Vereda Calles, Parque Nacional Natural " Las Orquideas ", margen derecha Quebrada La Honda, 06 ° 32 ' N, 076 ° 19 ' W, 1330 - 1400 m, 03 May 1993 (♂ fl), A. Cogollo et al. 6074 (MO); ibid, 08 May 1993 (♂ fl), A. Cogollo et al. 6190 (K- 2 sheets [n. v.], MO); ibid, 08 May 1993 (♀ fl), A. Cogollo et al. 6198 (MO); ibid, 11 May 1993 (♂ fl), A. Cogollo et al. 6279 (MO, NY); ibid, 11 May 1993 (♂ fl), A. Cogollo et al. 6300 (K n. v., MO); Vereda Calles, Parque Nacional Natural " Las Orquideas ", margen derecha del Rio Calles, en el filo NW de la Cabana de Calles, 06 ° 32 ' N, 076 ° 19 ' W, 1450 m, 15 Oct. 1993 (fr), A. Cogollo et al. 6918 (MO); ibid, 18 Oct. 1993 (fr), A. Cogollo et al. 7107 (MO); ibid, 08 Dec. 1993 (fr), A. Cogollo et al. 7959 (GH digital image, K [n. v.], MO, NY); Las Orquideas, Vereda Calles, Parque Nacional Natural Las Orquideas, Quebrada Honda, 06 ° 29 ' N, 076 ° 14 ' W, 1300 m, 08 Dec. 1992 (fr), J. Pipoly et al. 16745 (MO); ibid, 1330 m, 08 Dec. 1992 (♀ fl), J. Pipoly et al. 16798 (MO); ibid, 10 Dec. 1992 (♂ fl), J. Pipoly et al. 16902 (MO); ibid, 1300 m, 11 Dec. 1992 (♂ fl), J. Pipoly et al. 17019 (K [n. v.], MO); Parque Nacional Natural Las Orquideas, Vereda Calles, margen derecha del Rio Calles, 06 ° 32 ' N, 076 ° 19 ' W, 1350 - 1450 m, 05 Dec. 1993 (fl bud), J. Pipoly et al. 17721 (MO). The identification key below is modified from Jaramillo-Vivanco and Balslev 2020. Except where specified otherwise, the information comes from Smith 1950 |, measurements of the digital image of the holotype ¶, or measurements of herbarium specimens #. The key was difficult to build, in part because of current limited access to physical specimens during the COVID- 19 pandemic and in part because we were not able to identify obvious characters to separate species in some cases (e. g., between O. gordoniifolia and O. lehmannii). This may mean that the key is difficult to use, but we have decided to include it in case it is of use to some.	en	Santamaria-Aguilar, Daniel, Lagomarsino, Laura P. (2021): Two new species of Otoba (Myristicaceae) from Colombia. PhytoKeys 178: 147-170, DOI: http://dx.doi.org/10.3897/phytokeys.178.64564, URL: http://dx.doi.org/10.3897/phytokeys.178.64564
