taxonID	type	description	language	source
19FAC2E7F4AA5F7FAC79CF7EFE9A2E05.taxon	diagnosis	Diagnosis. Tortopsis andaki sp. nov., known from imagos of both sexes and nymph, can be distinguished from all other species of the genus by: In adults, 1) fore wing length 9.2 - 9.6 mm (male), 9.5 - 9.7 mm (female); 2) wings hyaline slightly tinted with purplish gray, veins purplish gray; 3) parastylus curved dorsally, more markedly on apical third, with a longitudinal ventral furrow (Figs 33 - 34); 4) penis slender, apical spine rounded and flattened (Figs 36 - 38); 5) parastyli receptors on sternum VIII with V-shaped sockets (Figs 39 - 40); 6) head shaded gray among ocelli, occiput with small gray marks (Fig. 27). The nymphs are characterized by: 1) mandibles with 7 stout small spines on inner margin of tusk see arrows in Fig. 44; 2) occiput whitish without gray markings (Fig. 31); 3) wingbuds completely whitish (Fig. 32). Male imago. Length (mm): body, 10.8 - 11.0; fore wing, 9.2 - 9.6; hind wing, 4.0; foreleg, 4.9 - 5.0; cerci, 25.0 - 26.0. General coloration whitish shaded dorsally with light purplish gray (Figs 27). Head. Whitish shaded with light brownish gray among ocelli and on scape and pedicel, flagellum hyaline. Thorax. Pronotum hyaline shaded with purplish gray dorsally as in Fig. 27. Mesonotum whitish yellow shaded with purplish gray along medioparapsidal sutures and contiguous area, medial zone paler. Metanotum whitish yellow shaded gray dorsally. Thoracic pleura and sterna whitish. Legs whitish shaded with light purplish gray on fore tibia and fore tarsi. Wings: Membrane hyaline very lightly tinted with purplish gray, longitudinal and cross veins purplish gray. Abdomen. Translucent white, shaded slightly but extensively with purplish gray on terga, including medial area; some markings are darker (Fig. 27) and a thin medial dark line is present along most terga; terga VIII-X shaded with darker gray. Abdominal sterna whitish; sternum IX shaded very slightly with gray on median area. Genitalia (Figs 28 - 29, 33 - 38): whitish except apex of parastylus yellowish; parastylus curved dorsally more markedly on apical third (in lateral view, Figs 28 and 34 - 35) and with a longitudinal ventral furrow along its entire length (Fig. 33); forceps and penis translucent white, shaded very slightly gray on forceps; penis slender, spine at apex of penis rounded and flattened (Figs 36 - 38). Caudal filament whitish translucent. Female imago. Length (mm): body, 10.0 - 10.1; fore wing, 9.5 - 9.7; hind wing, 3.9 - 4.0; cerci, 3.0. General coloration as in male but shading more strongly marked. Head black between ocelli, with small light gray marks on occiput. Wings membrane hyaline slightly tinged with whitish yellow, veins brownish. Abdomen shaded with gray on terga. Parastyli receptors on sternum VIII with sockets (Figs 39 - 40) with sinuous lateral margin and acute distal corner (V-shaped). Nymph (mature female). Length: body, 17.0 mm. General coloration yellowish white shaded with gray dorsally (Fig. 30). Head. Whitish shaded with gray among ocelli, occiput completely white without marks (Fig. 31). Antennae and mouthparts whitish except apex of tusks brownish, and spines yellowish. Finger-like gill present near base of maxillae (Figs 47 - 48). Mandibular tusks with 7 small stout spines on inner margin, basal to subdistal tubercle (see arrows in Fig. 44); outer margin of tusk with row of 16 - 17 stout spines (Figs 42 - 44). Thorax. Pronotum, anterior ring shaded gray widely including the anterolateral pointed projections, posterior ring as in Fig. 30, both with a median pale line. Meso- and metanotum with gray and brown marks, lighter on median area; wing buds whitish, longitudinal veins hyaline, except at the basal region, with the anal and costal margins brownish. Thoracic pleura and sterna whitish. Legs whitish, with yellowish setae and apex of tarsal claws yellowish (Figs 49 - 52). Abdomen. Abdominal color pattern as described for the adults (Fig. 30). Gills: vestigial gills I translucent; gills II-VII well developed, formed by a pair of large whitish lamellae, the outer (dorsal) lamellae of each pair is shaded with gray on a medio longitudinal band (Fig. 30), the inner (ventral) lamellae of each pair present only a thin brownish line along trachea. Caudal filaments yellowish white. Egg. Subcircular, white. Length, 330 - 350 μm; width, 280 - 300 μm.	en	Molineri, Carlos, Dias, Lucimar G., Zuniga, Maria del Carmen (2021): New insights into the phylogeny of Tortopus Needham and Murphy and Tortopsis Molineri (Ephemeroptera, Polymitarcyidae) with description of three new species. Arthropod Systematics & amp; Phylogeny 79: 151-170, DOI: http://dx.doi.org/10.3897/asp.79.e62735, URL: http://dx.doi.org/10.3897/asp.79.e62735
19FAC2E7F4AA5F7FAC79CF7EFE9A2E05.taxon	etymology	Etymology. This species is dedicated to the Andaki indigenous people, an American ethnic group that inhabited the upper Caqueta River basin.	en	Molineri, Carlos, Dias, Lucimar G., Zuniga, Maria del Carmen (2021): New insights into the phylogeny of Tortopus Needham and Murphy and Tortopsis Molineri (Ephemeroptera, Polymitarcyidae) with description of three new species. Arthropod Systematics & amp; Phylogeny 79: 151-170, DOI: http://dx.doi.org/10.3897/asp.79.e62735, URL: http://dx.doi.org/10.3897/asp.79.e62735
19FAC2E7F4AA5F7FAC79CF7EFE9A2E05.taxon	distribution	Distribution. Known only from the type locality.	en	Molineri, Carlos, Dias, Lucimar G., Zuniga, Maria del Carmen (2021): New insights into the phylogeny of Tortopus Needham and Murphy and Tortopsis Molineri (Ephemeroptera, Polymitarcyidae) with description of three new species. Arthropod Systematics & amp; Phylogeny 79: 151-170, DOI: http://dx.doi.org/10.3897/asp.79.e62735, URL: http://dx.doi.org/10.3897/asp.79.e62735
DE6AE29A6439559E8DAC27DE20E1470D.taxon	materials_examined	Material. 1 male imago (CEBUC) from Colombia, Caqueta, Curillo, rio Caqueta, 220 m, 01 ° 01 ' 43.5 " N, 75 ° 55 ' 8.6 " W, 20. ix. 2018, LG Dias, MC Zuniga, B Toro, JP Chaux and C Molineri cols. - Notes: The single male collected in Caqueta River (Colombia) presents a straight parastyli only curved at the apex (Figs 54 - 55), a feature that characterizes T. limoncocha. Body and wing size, and coloration also fit the diagnosis of this species (Molineri 2010).	en	Molineri, Carlos, Dias, Lucimar G., Zuniga, Maria del Carmen (2021): New insights into the phylogeny of Tortopus Needham and Murphy and Tortopsis Molineri (Ephemeroptera, Polymitarcyidae) with description of three new species. Arthropod Systematics & amp; Phylogeny 79: 151-170, DOI: http://dx.doi.org/10.3897/asp.79.e62735, URL: http://dx.doi.org/10.3897/asp.79.e62735
F1E5B4CD8E9955169A748D682313B230.taxon	diagnosis	Diagnosis. Tortopsis toro sp. nov., known from imagos of both sexes and nymphs, can be distinguished from all other species of the genus by: In adults, 1) forewing length ranges between 12.2 - 13.2 mm (male) and 15.5 - 17.0 mm (female); 2) hyaline wings, with brownish veins and costal margin shaded slightly gray; 3) parastyli almost straight in lateral view, slightly curved medially, with a longitudinal ventral furrow (Figs 2 - 3, 19 - 21); 4) penis apically widened and flattened, with rounded semicircular apical spine (Fig. 20); 5) female parastyli receptors with rounded and elongated sockets (Fig. 22); 6) head shaded black among ocelli, occiput with very light gray anastomosed lines (Figs 1, 6); 7) coloration as in Fig. 1. The nymphs can be separated from the others of the genus by: 1) mandibles at most with 2 stout small spines on inner margin of tusk (basally to large subdistal tubercle, see arrow in Fig. 24); 2) outer margin of tusk, just before distal spur, with a marked indentation (arrow in Fig. 25); 3) head shaded with gray on occiput as in Fig. 6; 4) developing fore wing buds pigmented with gray slightly along costal margin and base of longitudinal veins (similar to T. puella in Molineri 2008: figure 16). Male imago. Length (mm): body, 14.0 - 14.5; forewing, 12.2 - 13.2; hindwing, 5.9 - 6.1; foreleg, 5.3 - 5.9; cerci, 32.0 - 36.0. General coloration yellowish white shaded with gray dorsally (Fig. 1). Head. Coloration yellowish white except black area among ocelli (Fig. 1). Antennae: scape and pedicel yellowish white completely shaded with gray, flagellum hyaline. Thorax. Pronotum hyaline, shaded with gray dorsally (Fig. 1). Mesonotum whitish yellow shaded with gray along sutures but not medially. Metanotum whitish yellow shaded gray medially. Thoracic pleura and sterna whitish yellow. Legs yellowish white, shaded with grayish on fore tibia and tarsus, rest of leg (including femur and claws, not shaded). Wings. Membrane hyaline, shaded slightly gray along C and Sc areas; longitudinal and cross veins brownish. Abdomen. Whitish translucent with gray shading dorsally, except on medial band. Terga I-III shaded gray on submedial rounded areas; terga IV-VII similarly shaded but a longitudinal darker strip is visible submedially (Fig. 1); terga VIII-IX more widely shaded except on small marks and sigilla (Fig. 1); tergum X with whitish median area and blackish margins. Abdominal sterna pale, without dark shadings, except sternum IX with a small median gray mark on posterior margin. Genitalia (Figs 2 - 3, 19 - 21): whitish except parastylus yellowish. Parastylus almost straight in lateral view, slightly curved medially, with a longitudinal ventrolateral furrow along its entire length; penis distally widened, semicircular " spine " at apex of penis. Caudal filament whitish translucent. Female imago. Length (mm): body, 18.0 - 18.5; forewing, 15.5 - 17.0; hindwing, 6.5 - 7.5; cerci, 4.3 - 4.8. General coloration as in male except darker markings. Head. Coloration black between ocelli, with posterior and submedian grayish lines on occiput (similar to Fig. 4). Thorax. Furcasterna shaded with light gray medially. Wings: Membrane hyaline slightly tinged with yellow. Abdomen. shaded more extensively with gray on terga. Parastyli receptors on sternum VIII with elongated and rounded sockets (Fig. 22). Light gray marks medially on sterna VIII-IX. Egg. Suboval. Length, 410 - 420 μm; width, 320 - 340 μm. Nymph (mature). Length of body (from apex of tusks to apex of abdominal tergum X): female, 22.0 - 26.0 mm; male, 14.0 - 19.0 mm. General coloration yellowish white with gray markings dorsally (Figs 4 - 6). Head. Coloration whitish shaded gray among ocelli, occiput with very light gray anastomosed lines submedially as Fig. 6. Antennae and mouthparts whitish except apex of tusk orangeish, and spines yellowish. Finger-like gill present near base of maxillae (arrow in Fig. 13). Mandibular tusks with 0 - 2 stout spines on inner margin, basal to subdistal tubercle (see arrow in Fig. 24); outer margin of tusk with row of 18 - 26 stout spines (Figs 8 - 10, 24 - 25). Thorax. Pronotum, anterior ring shaded black almost completely including the anterolateral pointed projections, posterior ring patterned as in Fig. 6, both with a median pale line. Meso- and metanotum with gray and black marks, lighter on median area; wing buds whitish, pigmented with gray slightly along costal margin and base of longitudinal veins. Thoracic pleura and sterna whitish. Legs whitish, with yellowish setae and apex of tarsal claws yellowish (Figs 15 - 18). Abdomen. Color pattern as described for the adults. Gills: vestigial gills I translucent; gills II-VII well developed, formed by a pair of large whitish lamellae, the outer (dorsal) lamellae of each pair is shaded with black on a medio longitudinal band (thinner on gill II), the inner (ventral) lamellae of each pair show only a thin brownish line along trachea. Genital rudiments of males showing distally widened penis. Caudal filaments yellowish white.	en	Molineri, Carlos, Dias, Lucimar G., Zuniga, Maria del Carmen (2021): New insights into the phylogeny of Tortopus Needham and Murphy and Tortopsis Molineri (Ephemeroptera, Polymitarcyidae) with description of three new species. Arthropod Systematics & amp; Phylogeny 79: 151-170, DOI: http://dx.doi.org/10.3897/asp.79.e62735, URL: http://dx.doi.org/10.3897/asp.79.e62735
F1E5B4CD8E9955169A748D682313B230.taxon	etymology	Etymology. This species is dedicated to Dr. Beatriz Toro Restrepo, Universidad de Caldas, in recognition of her work in environmental education, friendship and contributions in fieldwork with aquatic insects of Colombia.	en	Molineri, Carlos, Dias, Lucimar G., Zuniga, Maria del Carmen (2021): New insights into the phylogeny of Tortopus Needham and Murphy and Tortopsis Molineri (Ephemeroptera, Polymitarcyidae) with description of three new species. Arthropod Systematics & amp; Phylogeny 79: 151-170, DOI: http://dx.doi.org/10.3897/asp.79.e62735, URL: http://dx.doi.org/10.3897/asp.79.e62735
F1E5B4CD8E9955169A748D682313B230.taxon	distribution	Distribution. Known only from the type locality.	en	Molineri, Carlos, Dias, Lucimar G., Zuniga, Maria del Carmen (2021): New insights into the phylogeny of Tortopus Needham and Murphy and Tortopsis Molineri (Ephemeroptera, Polymitarcyidae) with description of three new species. Arthropod Systematics & amp; Phylogeny 79: 151-170, DOI: http://dx.doi.org/10.3897/asp.79.e62735, URL: http://dx.doi.org/10.3897/asp.79.e62735
7CDA9F3FDA375BC68836FE781C77B169.taxon	diagnosis	Diagnosis. Tortopus coreguaje sp. nov., known from male imago and nymph, can be distinguished from all other species of the genus by the following combination of characters: In male imagos, 1) fore wing length 10.9 - 11.0 mm (male); 2) pale wings, veins translucent gray; 3) pedestal short with relatively large, straight and very long parastyli (Figs 56 - 57, 67 and 69) (parastylus 2.6 times the length of pedestal base, in lateral view the parastylus reach 0.37 of the length of forceps from base); 3) ventral knob on forceps base relatively small; 4) penis relatively thin at base (fused portion), not strongly expanded distally (Figs 56 and 67); and 5) relatively dark species, head shaded stronger between lateral ocelli, occiput shaded gray except on pale medial line (Fig. 65). In the nymph: 1) inner protuberances on mandibular tusk almost contiguous (Figs 59 - 61); 2) frontal tuft of setae with irregular sinuous form (arrow in Fig. 58); 3) ventral and dorsal projection at the apex of fore tarsus subequal in length (Fig. 63). Male imago. Length (mm): body, 11.0; fore wing, 10.9 - 11.0; hind wing, 4.8; cerci, 28.0. General coloration yellowish white shaded with gray (Fig. 65). Head. Coloration yellowish white shaded widely with gray on dorsum, shaded stronger between lateral ocelli, except around median ocellus and hind margin (Fig. 65). Antennae: scape and pedicel yellowish white completely shaded with gray, flagellum hyaline. Thorax. Pronotum with anterior ring hyaline, shaded with black posterolaterally (Fig. 65); posterior ring shaded gray except on sublateral areas and medial line. Mesonotum whitish yellow shaded with gray along medioparapsidal sutures and shaded with black on a V-shaped mark between posterior scutal protuberances (Fig. 65). Metanotum whitish yellow shaded gray medially. Thoracic pleura and sterna whitish yellow, with grayish small marks medially on furcasterna. Legs yellowish white shaded with grayish (fore legs broken off and lost). Wings: Membrane hyaline except basally slightly grayish, longitudinal and cross veins translucent shaded slightly with gray. Abdomen. Abdominal segments translucent-hyaline. Tergum I shaded gray anterolaterally (Fig. 65), terga II-IX shaded more extensively towards rear segments, except on: medial line of terga II-VII, and two pale marks (one submedian and one sublateral) in II-IX; tergum X shaded more completely with gray. Abdominal sterna shaded widely with brownish gray except on intersegmental membranes and around gill sclerites (Fig. 66). Genitalia (Figs 56 - 57, 67 - 69): sternum IX shaded with black on posterior margin and at both sides of the median furrow; relatively large parastyli orangeish, straight and long (Fig. 56); parastylus 2.6 times the length of pedestal base, in lateral view the parastylus reach 0.37 of the length of forceps from base; forceps and penis yellowish white, shaded gray on forceps. Caudal filament whitish translucent. Male nymph (immature). Length: body, 9.5 mm; cerci, 2.5 mm; caudal filament, 2.0 mm. General coloration whitish shaded with gray (Fig. 70). Head. Head shaded with gray in net-shaped pattern posteriorly to epicranial suture (Fig. 70); anteriorly to this suture with two small lateral irregular-sinuous-shaped tufts of short setae (anterior to lateral ocelli, arrow in Fig. 58); frontal ridge relatively straight in dorsal view (Fig. 58); fronto-clypeal region acutely projected medially (Fig. 58); a tuft of ca. 10 long setae basally to antennal condyle; and a group of 3 long setae anterior to eye and 6 long curved setae posterior to eye (Fig. 58). Mandibular tusks straight (Figs 59 - 61, 70), with 2 subdistal rigid inner setae and ca. 12 weaker but long setae forming a basal arc (Figs 59 - 61); inner margin with two large and contiguous tubercles, the subdistal directed ventrally and the submedian directed medially (Figs 59 - 61), also a small blade-like projection present basally to distal spur (arrow in Fig. 61). Maxillae with small triangular ventral gill. Hypopharynx and labium whitish. Thorax. Pronotum with narrow anterior ring (0.28 of total length of pronotum); antero-lateral corners acutely projected; shaded with black laterally on anterior ring and medially on posterior ring. Meso and metanotum shaded black medially except medial line; wingbuds whitish shaded gray only at base of costal area. Legs. Whitish with yellowish setae and sclerotized portions. Foreleg with tibia-tarsus flattened, remnants of the suture between tarsus and tibia visible, tarsal dorso-distal projection 0.5 of total length of claw (Figs 62 - 63); ventral surface of fore tibia with rows of long filtering setae as in Fig. 62; fore femur with anterobasal short U-shaped row of filtering setae and an immediately posterior small group of ca. 7 long rigid and simple setae; apex of fore tarsus with ventral and dorsal projections subequal in length (but the ventral is much thinner, Fig. 63). Middle leg with long setae on anterior and posterior (functionally ventral and dorsal, respectively) margins of femur, anterior margin of tibia and tarsus; apical third of tibia and tarsus completely covered with strong setae, apex of tibia with a brush of thick setae ventrally. Hind leg with short strong setae on posterior margin, and transverse subdistal row of short setae on dorsal surface; hind tibia and tarsus with long setae on posterior margin, anterior margin covered with short and strong setae. All tarsal claws slender and curved, without denticles. Abdomen (Fig. 70). Gill I single, small and elongated, remaining gills well developed and double. Terga II-IX with medio-longitudinal row of setae; abdominal sterna with lateral margins strongly covered with setae, increasing in number posteriorly, sterna V-VI also with row of shorter setae on posterior margin; sternum IX with a row of few long setae on posterior half, along medial line. Cerci with rows of setae at each article, mainly on basal fourth; terminal filament much thinner and with whorls of setae almost on its entire length.	en	Molineri, Carlos, Dias, Lucimar G., Zuniga, Maria del Carmen (2021): New insights into the phylogeny of Tortopus Needham and Murphy and Tortopsis Molineri (Ephemeroptera, Polymitarcyidae) with description of three new species. Arthropod Systematics & amp; Phylogeny 79: 151-170, DOI: http://dx.doi.org/10.3897/asp.79.e62735, URL: http://dx.doi.org/10.3897/asp.79.e62735
7CDA9F3FDA375BC68836FE781C77B169.taxon	etymology	Etymology. This species is dedicated to the Coreguaje community inhabiting along the piemont and lower parts of the Caqueta River and its affluents.	en	Molineri, Carlos, Dias, Lucimar G., Zuniga, Maria del Carmen (2021): New insights into the phylogeny of Tortopus Needham and Murphy and Tortopsis Molineri (Ephemeroptera, Polymitarcyidae) with description of three new species. Arthropod Systematics & amp; Phylogeny 79: 151-170, DOI: http://dx.doi.org/10.3897/asp.79.e62735, URL: http://dx.doi.org/10.3897/asp.79.e62735
7CDA9F3FDA375BC68836FE781C77B169.taxon	distribution	Distribution. Known only from the type locality.	en	Molineri, Carlos, Dias, Lucimar G., Zuniga, Maria del Carmen (2021): New insights into the phylogeny of Tortopus Needham and Murphy and Tortopsis Molineri (Ephemeroptera, Polymitarcyidae) with description of three new species. Arthropod Systematics & amp; Phylogeny 79: 151-170, DOI: http://dx.doi.org/10.3897/asp.79.e62735, URL: http://dx.doi.org/10.3897/asp.79.e62735
