taxonID	type	description	language	source
652887C7B354BF71587E886FFCA01D95.taxon	description	(Fig. 1)	en	Ahrens, Dirk (2000): New species of Calloserica Brenske, 1894 from Sikkim and Nepal (Coleoptera, Melolonthidae, Sericini). Zoosystema 22 (4): 815-824, DOI: http://doi.org/10.5281/zenodo.4689031
652887C7B354BF71587E886FFCA01D95.taxon	materials_examined	TYPE MATERIAL. — Holotype:, Sikkim de Padamtsin [Padamchen] à Lingtou Juillet 1901 / Muséum Paris ex. Coll. R. Oberthur (MNHN). Paratype: 1, Sikkim de Padamtsin [Padamchen] à Lingtou Juillet 1901 / Muséum Paris ex. Coll. R. Oberthur (CA).	en	Ahrens, Dirk (2000): New species of Calloserica Brenske, 1894 from Sikkim and Nepal (Coleoptera, Melolonthidae, Sericini). Zoosystema 22 (4): 815-824, DOI: http://doi.org/10.5281/zenodo.4689031
652887C7B354BF71587E886FFCA01D95.taxon	etymology	ETYMOLOGY. — The species is named in honour to Nicole Berti (MNHN), who was of great assistance with the search for types during my visits at MNHN.	en	Ahrens, Dirk (2000): New species of Calloserica Brenske, 1894 from Sikkim and Nepal (Coleoptera, Melolonthidae, Sericini). Zoosystema 22 (4): 815-824, DOI: http://doi.org/10.5281/zenodo.4689031
652887C7B354BF71587E886FFCA01D95.taxon	description	DESCRIPTION Length: 11.1 - 12.1 mm, width: 5.3 - 5.4 mm, length of elytra: 8.7 - 9.4 mm. Body oblong, chestnut with opaque tomentum, elytral striae lighter, labroclypeus shiny, dorsal surface erectly hairy. Labroclypeus shiny, transverse, sides convexly narrowed anteriorly, all borders strongly reflexed and anterior margin deeply sinuate at middle, front angles abruptly rounded; labrum broadly trapezoidal and anteriorly produced, longest extension medially; clypeal surface densely punctate, punctures slightly elevated, with thin setae. Frontoclypeal suture feebly elevated and weakly curved; smooth area in front of eye 1.5 time as wide as long, ocular canthus long and slender, with few short setae. Punctures on frons vanishing under thick opaque tomentum, only recognizable by their long erect setae. Antenna yellow, 10 - segmented; club in male 4 - segmented, a trifle longer than combined lengths of remaining segments, seventh segment one-third shorter than club, in female 3 - segmented, a trifle shorter than combined lengths of remaining segments. Eyes small, ratio diameter / interocular width: 0.5 (,). Pronotum transverse, widest at base, lateral margin straightly convergent from base to middle, at middle strongly curved, anteriorly behind bluntly rounded, not protudent anterior angles almost straight, anterior margin straight; general shape similar to that of C. tigrina, but sides only posteriorly feebly sinuate before obtusely rounded hind angles; disc with shallow punctation, double erect pilosity directed forward, mixed short and two times longer setae. Proepisternum weakly grooved, posterior border produces with pronotal base shortly rounded angle. Scutellum oblong, triangular, densely punctate, basally with a deep impression. Elytra oblong, intervals flat, mixed coarse punctures bearing long, erect, forward directed setae with groups of fine punctures bearing short white scales; apex membranous. Ventral surface including metacoxal plates, femora, and abdominal sternites clothed with fine, recumbent, yellow setae. Abdominal sternites without any elevation. Mesosternum between mesocoxae equal in width to slender mesofemur. Ratio of length metepisternum / metacoxa: 1 / 1.23. Pygidium in male evenly convex, densely and shallowly punctate, with double pilosity, long setae four times longer than the short ones, in female less convex, with a longitudinal median elevation, which is apically more distinctive. Metatibia slender, ratio width / length: 1 / 4.75; dorsal edge apically only moderately carinate, with two groups of spines, the proximal one situated at middle of metatibial length, basally few small setae on a longitudinal elevation; outside feebly longitudinally grooved and finely scarcely scattered punctate, ventral edge with four tubercles bearing a strong seta. Metatarsal segments laterally carinate, first metatarsomere almost as long as following two segments combined, onefourth longer than superior metatibial spur; mesotarsal segments laterally not carinate, dorsally without longitudinal impressions. Anterior tibia short, bidentate, external border with a blunt median extension.	en	Ahrens, Dirk (2000): New species of Calloserica Brenske, 1894 from Sikkim and Nepal (Coleoptera, Melolonthidae, Sericini). Zoosystema 22 (4): 815-824, DOI: http://doi.org/10.5281/zenodo.4689031
652887C7B354BF71587E886FFCA01D95.taxon	discussion	REMARKS The species is externally very similar to C. tigrina Brenske, but is readily distinguished by the genital features of the male (Fig. 1) (Ahrens 1995: 138, figs 1 - 3).	en	Ahrens, Dirk (2000): New species of Calloserica Brenske, 1894 from Sikkim and Nepal (Coleoptera, Melolonthidae, Sericini). Zoosystema 22 (4): 815-824, DOI: http://doi.org/10.5281/zenodo.4689031
652887C7B357BF7058588999FCF71E95.taxon	description	(Fig. 2)	en	Ahrens, Dirk (2000): New species of Calloserica Brenske, 1894 from Sikkim and Nepal (Coleoptera, Melolonthidae, Sericini). Zoosystema 22 (4): 815-824, DOI: http://doi.org/10.5281/zenodo.4689031
652887C7B357BF7058588999FCF71E95.taxon	materials_examined	TYPE MATERIAL. — Holotype:, British India Sikkim Lachen-Lachung, VIII. 1933 / Muséum Paris ex. Coll. R. Oberthur (MNHN).	en	Ahrens, Dirk (2000): New species of Calloserica Brenske, 1894 from Sikkim and Nepal (Coleoptera, Melolonthidae, Sericini). Zoosystema 22 (4): 815-824, DOI: http://doi.org/10.5281/zenodo.4689031
652887C7B357BF7058588999FCF71E95.taxon	etymology	ETYMOLOGY. — The species is named after locality Lachung. A C	en	Ahrens, Dirk (2000): New species of Calloserica Brenske, 1894 from Sikkim and Nepal (Coleoptera, Melolonthidae, Sericini). Zoosystema 22 (4): 815-824, DOI: http://doi.org/10.5281/zenodo.4689031
652887C7B357BF7058588999FCF71E95.taxon	description	DESCRIPTION Length: 9.3 mm, width: 5.3 mm, length of elytra: 7.4 mm. Body oblong, chestnut, with opaque tomentum, elytral striae lighter, labroclypeus shiny, dorsal surface erectly hairy. Labroclypeus shiny, transverse, sides convexly narrowed anteriorly, all borders strongly reflexed and anterior margin deeply sinuate at middle, front angles abruptly rounded; labrum broadly trapezoidal and anteriorly produced, longest extension medially; clypeal surface densely punctate, punctures slightly elevated, with fine setae. Frontoclypeal suture weakly curved; smooth area in front of eye 1.5 time as wide as long, ocular canthus long and slender, with few setae. Punctures on frons vanishing under thick opaque tomentum, only recognizable by their long erect setae. Antenna yellow, 10 - segmented; club 4 - segmented, a trifle longer than combined lengths of remaining segments, seventh segment one-third shorter than club. Eyes small, ratio diameter / interocular width: 0.56. Pronotum transverse, widest at base, lateral margin subparallel from base to middle, anteriorly strongly curved, anterior angles bluntly rounded, not protudent, anterior margin straight; general shape similar to that of C. tigrina, but sides posteriorly not sinuate before obtusely rounded hind angles; disc with shallow punctation, double erect pilosity directed forward, mixed short and two times longer setae. Proepisternum weakly grooved, posterior border produces with pronotal base a sharp, distinctive angle. Scutellum oblong, triangular, densely punctate, basally with a deep transverse impression. Elytra oblong, intervals flat, mixed coarse punctures bearing long, erect, forward directed setae with groups of fine punctures bearing short white scales; apex membranous. Ventral surface including metacoxal plates, femora, and abdominal sternites clothed with fine, recumbent, yellow setae. Abdominal sternites without any elevation. Mesosternum between mesocoxae equal in width to slender mesofemur. Ratio of length metepisternum / metacoxa: 1 / 1.2. Pygidium convex, densely and shallowly punctate, with double pilosity, long setae four times longer than the short. Metatibia slender, ratio width / length: 1 / 4.2; dorsally longitudinally carinate, with two groups of spines, the proximal one situated at middle of metatibial length, basally few small setae on a longitudinal elevation; outside longitudinally grooved, dorsally coarsly wrinkled, ventral edge with four tubercles bearing a strong seta. Metatarsal segments laterally carinate, first metatarsomere almost as long as following two segments combined, one-third longer than superior metatibial spur; mesotarsal segments laterally not carinate, dorsally without longitudinal impressions. Anterior tibia short, bidentate, external border with a blunt median extension.	en	Ahrens, Dirk (2000): New species of Calloserica Brenske, 1894 from Sikkim and Nepal (Coleoptera, Melolonthidae, Sericini). Zoosystema 22 (4): 815-824, DOI: http://doi.org/10.5281/zenodo.4689031
652887C7B357BF7058588999FCF71E95.taxon	discussion	REMARKS The species is externally similar to C. brendelli Ahrens, 1999. It differs by ratio of length of first metatarsomere with the combined lengths of the second and third metatarsomeres. C. bertiae n. sp. (Sikkim) and C. lachungensis n. sp. seem closely related because of their similar features of male genitalia. They differ from each other by size of parameres and aedeagus, both can be distinguished from the genitalitary very similar C. brendelli Ahrens, 1999 by the shorter parameres (Fig. 2) (Ahrens 1999: 187, figs 6 - 8). Female unknown.	en	Ahrens, Dirk (2000): New species of Calloserica Brenske, 1894 from Sikkim and Nepal (Coleoptera, Melolonthidae, Sericini). Zoosystema 22 (4): 815-824, DOI: http://doi.org/10.5281/zenodo.4689031
652887C7B351BF765BB58C9CFED81CB5.taxon	description	(Fig. 3)	en	Ahrens, Dirk (2000): New species of Calloserica Brenske, 1894 from Sikkim and Nepal (Coleoptera, Melolonthidae, Sericini). Zoosystema 22 (4): 815-824, DOI: http://doi.org/10.5281/zenodo.4689031
652887C7B351BF765BB58C9CFED81CB5.taxon	materials_examined	TYPE MATERIAL. — Holotype:, Nepal: 8800 ’ [feet], Kathmandu Dist., Phulcoki / 6. VII. 84 / Lt. Col. M. G. Allen, Brit. Mus. 1983 - 254 (BMNH). Paratypes: 3, 2, same data as holotype (BMNH, CA).	en	Ahrens, Dirk (2000): New species of Calloserica Brenske, 1894 from Sikkim and Nepal (Coleoptera, Melolonthidae, Sericini). Zoosystema 22 (4): 815-824, DOI: http://doi.org/10.5281/zenodo.4689031
652887C7B351BF765BB58C9CFED81CB5.taxon	etymology	ETYMOLOGY. — From the Latin delectabilis, “ tasty ”.	en	Ahrens, Dirk (2000): New species of Calloserica Brenske, 1894 from Sikkim and Nepal (Coleoptera, Melolonthidae, Sericini). Zoosystema 22 (4): 815-824, DOI: http://doi.org/10.5281/zenodo.4689031
652887C7B351BF765BB58C9CFED81CB5.taxon	description	DESCRIPTION Length: 8.1 - 9.4 mm, width: 4.6 - 5.3 mm, length of elytra: 6.4 - 6.6 mm. Body oblong, dark green with opaque tomentum, elytra with brownish striae, dorsal surface densely erectly hairy. Labroclypeus shiny, transverse, sides curved anteriorly, front angles rounded, anterior margin with large median sinuation; labrum broadly trapezoidal, with only few very shallow punctures posteriorly; clypeal surface with densely scattered punctures and erect setae. Frontoclypeal suture weakly curved, smooth area in front of eye 1.5 time as wide as long, ocular canthus long, slender and very finely punctate, with few long setae. Punctures on frons vanishing under thick, opaque tomentum, only recognizable by their long setae. Antenna yellow, 10 - segmented; club in male 4 - segmented, distinctly longer than combined length of remaining segments, seventh segment one-fifth shorter than club; in female club 3 - segmented, a trifle shorter than combined length of remaining segments, seventh segment transversely produced. Eyes rather small, ratio diameter / interocular width: 0.62 (), 0.48 (). Mentum elevated, anteriorly indistinctly flattened. Shape of pronotum similiar to C. poggii Ahrens, 1995, transverse, widest at base, sides from base to middle straightly convergent, at middle strongly curved inward, anteriorly convexly curved, front angles broadly rounded, not protuding, anterior margin straight; disc with shallow punctation, double, erect, forward directed pilosity mixed, short, straight and two times longer, at apex backwards crooked setae. Proepisternum grooved basally, posterior border produces with pronotal base a sharp, distinctive A C angle. Scutellum oblong, triangular, flat, densely punctate. Elytra slender, intervals flat, mixed coarse punctures bearing long, erect, forward directed setae with groups of fine punctures bearing short white scales; apex membranous. Ventral surface including metacoxal plates, femora and abdominal sternites clothed with fine, recumbent setae, on metasternum setae more erect; penultimate abdominal sternite posteriorly with two weak elevations, abdominal sternites in female simple. Mesosternum between mesocoxae equal in width to slender mesofemur. Ratio of length metepisternum / metacoxa: 1 / 1.23. Pygidium in male evenly convex, densely shallowly punctate, with short and long setae, punctures of long setae twice as large as those of short ones, pygidium in female moderately convex, but apically with a distinct convexity. Metatibia slender, long, ratio width / length: 1 / 4.1, outside longitudinally grooved at middle and smooth, only with few, minute punctures; dorsally carinate, with two external setigerous groups of spines, basal one at middle of metatibial length, basally few small setae on a longitudinal elevation, ventral edge with four tubercles bearing a strong seta. Metatarsal segments laterally carinate, first metatarsomere one-third longer than superior metatibial spur and as long as the following two segments combined. Mesotarsi laterally not carinate, dorsally without A C B longitudinal impressions. Anterior tibia moderately long, bidentate, all tarsal claws simple and symmetrical.	en	Ahrens, Dirk (2000): New species of Calloserica Brenske, 1894 from Sikkim and Nepal (Coleoptera, Melolonthidae, Sericini). Zoosystema 22 (4): 815-824, DOI: http://doi.org/10.5281/zenodo.4689031
652887C7B351BF765BB58C9CFED81CB5.taxon	discussion	REMARKS The species is externally similar to C. brendelli Ahrens, it differs by the shallower medial labroclypeal sinuation. In contrast to all other Calloserica species, C. delectabilis n. sp. is characterized by a erect lamina on ventral portion of aedeagal phallobasis (Fig. 3).	en	Ahrens, Dirk (2000): New species of Calloserica Brenske, 1894 from Sikkim and Nepal (Coleoptera, Melolonthidae, Sericini). Zoosystema 22 (4): 815-824, DOI: http://doi.org/10.5281/zenodo.4689031
652887C7B350BF755BB68888FEA81BB5.taxon	description	(Fig. 4)	en	Ahrens, Dirk (2000): New species of Calloserica Brenske, 1894 from Sikkim and Nepal (Coleoptera, Melolonthidae, Sericini). Zoosystema 22 (4): 815-824, DOI: http://doi.org/10.5281/zenodo.4689031
652887C7B350BF755BB68888FEA81BB5.taxon	materials_examined	TYPE MATERIAL. — Holotype:, Sikkim de Padamtsin [Padamchen] à Lingtou Juillet 1901 / Muséum Paris ex. Coll. R. Oberthur (MNHN).	en	Ahrens, Dirk (2000): New species of Calloserica Brenske, 1894 from Sikkim and Nepal (Coleoptera, Melolonthidae, Sericini). Zoosystema 22 (4): 815-824, DOI: http://doi.org/10.5281/zenodo.4689031
652887C7B350BF755BB68888FEA81BB5.taxon	etymology	ETYMOLOGY. — The species is dedicated to Prof. Y. Cambefort (MNHN), who greatly supported my work during my visits at MNHN.	en	Ahrens, Dirk (2000): New species of Calloserica Brenske, 1894 from Sikkim and Nepal (Coleoptera, Melolonthidae, Sericini). Zoosystema 22 (4): 815-824, DOI: http://doi.org/10.5281/zenodo.4689031
652887C7B350BF755BB68888FEA81BB5.taxon	description	DESCRIPTION Length: 9.3 mm, width: 4.6 mm, length of elytra: 7.3 mm. Body slender and oblong, chestnut with opaque toment, elytral striae lighter, labroclypeus shiny, dorsal surface erectly hairy. Labroclypeus shiny, transverse, sides convexly narrowed anteriorly, all borders strongly reflexed and anteriorly deeply sinuate at middle, front angles abruptly rounded; labrum broadly trapezoidal and anteriorly produced, longest extension medially; clypeal surface densely punctate, punctures slightly elevated, with fine setae. Frontoclypeal suture finely incised and feebly elevated, weakly curved; smooth area in front of eye 1.5 time as wide as long, ocular canthus long and slender, with few setae. Punctures on frons vanishing under thick opaque tomentum, only recognizable by their long erect setae. Antenna yellow, 10 - segmented; club 4 - segmented, apically feebly curved outward, almost twice as long as remaining segments together, seventh segment only a trifle shorter than club. Eyes large, ratio diameter / interocular width: 0.7. Pronotum narrow, widest at base, lateral margin straightly convergent from base to middle, anteriorly strongly curved, front angles bluntly round- ed, not protudent, posterior angles largely rounded; disc with very fine and shallow punctation, double erect pilosity directed forward, mixed short and two times longer setae. Proepisternum weakly grooved, posterior border produce a broadly rounded angle with pronotal base. Scutellum oblong, triangular, densely punctate, basally with a deep transverse impression. Elytra oblong, intervals flat, mixed coarse punctures bearing long, erect, forward directed setae with groups of fine punctures bearing short white scales; apex membranous. Ventral surface including metacoxal plates, femora, and abdominal sternites clothed with fine, recumbent, yellow setae. Abdominal sternites without any elevations. Mesosternum between mesocoxae equal in width to slender mesofemur. Ratio of length metepisternum / metacoxa: 1 / 1.23. Pygidium moderately convex, densely and shallowly punctate, with double pilosity, long setae four times longer than the short ones. Metatibia slender, ratio width / length: 1 / 4.4; dorsally longitudinally carinate, with two groups of spines, the proximal one situated at middle of metatibial length, basally few small setae on a longitudinal elevation; outside longitudinally grooved, only with few very small punctures, ventral edge with four tubercles bearing a strong seta. Metatarsal segments laterally carinate, first metatarsomere almost as long as following two segments combined, almost twice as long as superior metatibial spur; mesotarsal segments laterally not carinate, dorsally without longitudinal impressions. Anterior tibia moderately long, bidentate.	en	Ahrens, Dirk (2000): New species of Calloserica Brenske, 1894 from Sikkim and Nepal (Coleoptera, Melolonthidae, Sericini). Zoosystema 22 (4): 815-824, DOI: http://doi.org/10.5281/zenodo.4689031
652887C7B350BF755BB68888FEA81BB5.taxon	discussion	REMARKS This species is externally similar to C. begnasia Ahrens, 1999. It differs by ratio of length of the first metatarsomere with the combined lengths of the second and third metatarsomeres as well as by genitalic features of the male (Fig. 4). In C. cambeforti n. sp. the lateral apophysis of aedeagal phallobasis is ventrally largely produced and curved, apically not produced, the parameres are shorter than those of C. trigrina Brenske. Female unknown.	en	Ahrens, Dirk (2000): New species of Calloserica Brenske, 1894 from Sikkim and Nepal (Coleoptera, Melolonthidae, Sericini). Zoosystema 22 (4): 815-824, DOI: http://doi.org/10.5281/zenodo.4689031
652887C7B353BF755B95895EFC2F1B13.taxon	materials_examined	MATERIAL EXAMINED. — 2 ex., Nepal, 6 800 ’ [feet] Kathmandu Dist. Kakani / Lt. Col. M. G. Allen Brit. Mus. 1983 - 254 (BMNH); 1 ex, Nepal, Kathmandu V. Godawari, Mahabharat, 15. V- 25. VI. 1995 Jiri Kolibac leg. (CP). Calloserica chiplingensis Ahrens, 1999 Calloserica chiplingensis Ahrens, 1999: 192. A B MATERIAL EXAMINED. — 2 ex., Nepal 8800 ’ [feet] Kathmandu Dist. Phulcoki / Lt. Col. M. G. Allen Brit. Mus., 1983 - 254 (BMNH).	en	Ahrens, Dirk (2000): New species of Calloserica Brenske, 1894 from Sikkim and Nepal (Coleoptera, Melolonthidae, Sericini). Zoosystema 22 (4): 815-824, DOI: http://doi.org/10.5281/zenodo.4689031
652887C7B353BF7559AE8F1BFC911C22.taxon	materials_examined	MATERIAL EXAMINED. — 1 ex., Darjeeling A. Desgodins. / Muséum Paris ex. coll. R. Oberthur (MNHN).	en	Ahrens, Dirk (2000): New species of Calloserica Brenske, 1894 from Sikkim and Nepal (Coleoptera, Melolonthidae, Sericini). Zoosystema 22 (4): 815-824, DOI: http://doi.org/10.5281/zenodo.4689031
652887C7B353BF755BE38877FDD01CE6.taxon	materials_examined	MATERIAL EXAMINED. — 1 ex., Nepal Ganesh Himal 2 kmW Gholjong 85 ° 17 ’ E, 28 ° 11 ’ N / 2420 m, 12. X. 1995 L. Pergovits & L. Ronkay (HNHM).	en	Ahrens, Dirk (2000): New species of Calloserica Brenske, 1894 from Sikkim and Nepal (Coleoptera, Melolonthidae, Sericini). Zoosystema 22 (4): 815-824, DOI: http://doi.org/10.5281/zenodo.4689031
652887C7B353BF7459F1881CFBDC18D5.taxon	description	C. gosainkundensis Ahrens, 1999: 195. C. hingstoni Ahrens, 1999: 192. C. lachungensis n. sp.	en	Ahrens, Dirk (2000): New species of Calloserica Brenske, 1894 from Sikkim and Nepal (Coleoptera, Melolonthidae, Sericini). Zoosystema 22 (4): 815-824, DOI: http://doi.org/10.5281/zenodo.4689031
