identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03AE87A4700AFFCCFCECFAD5FCAAF90D.text	03AE87A4700AFFCCFCECFAD5FCAAF90D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cephalogale Jourdan 1862	<div><p>Cephalogale Jourdan, 1862</p> <p>DIAGNOSIS. — Medium sized to large Cephalogalini n. tr., relatively short and robust P2-P3 and p2-p4, without pac or pacd or with a reduced pacd in p4.</p> <p>TYPE SPECIES. — C. geoffroyi Jourdan, 1862 by original designation.</p> <p>OTHER SPECIES. — C. ginesticus Kuss, 1962; C. gergoviensis Viret, 1929; C. shareri (Wang, Hunt, Tedford &amp; Lander, 2009).</p> </div>	http://treatment.plazi.org/id/03AE87A4700AFFCCFCECFAD5FCAAF90D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bonis, Louis De	Bonis, Louis De (2013): Ursidae (Mammalia, Carnivora) from the Late Oligocene of the “ Phosphorites du Quercy ” (France) and a reappraisal of the genus Cephalogale Geoffroy, 1862. Geodiversitas 35 (4): 787-814, DOI: 10.5252/g2013n4a4, URL: http://dx.doi.org/10.5252/g2013n4a4
03AE87A4700AFFCCFF15FAF5FC0DFAA2.text	03AE87A4700AFFCCFF15FAF5FC0DFAA2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cephalogalini de Bonis 2013	<div><p>Tribe CEPHALOGALINI n. tr.</p> <p>TYPE GENUS. — Cephalogale Jourdan, 1962 by present designation.</p> <p>DIAGNOSIS. — Partially coming from the descriptions of Beaumont (1965), Bonis (1973), Hunt (1998), and Teilhard de Chardin (1915).</p> <p>Digitigrade Hemicyoninae containing several lineages of small (Early Oligocene) to medium sized (Late Oligocene-Early Miocene) primitive ursids (skull length of about 10-20 cm). Dental formula = I3/2-3, C1/1, P4/4, M2/3. Ŋe m2 has a metaconid larger or equal to the protoconid like in the Amphicynodontidae Simpson, 1945 but the Cephalogalini n. tr. are distinguished from them by the m1 which is larger relatively to the premolars and whose metaconid is reduced (open trigonid), trigonid is more bladelike and talonid has a robust elongated hypoconid continuing by a low crest which encloses the talonid basin and continues in the lingual side before gently joining the metaconid without any notch; sometimes a very small entoconid is present in the crest; m2 has a reduced or absent paraconid and a talonid similar to that of m1. Ŋere is an overall evolutionary trend toward an increasing of size, reduction of metaconid of m1 and correlative increasing distance between m1 paraconid and metaconid, increasing relative length of the m2 talonid. Some lineages may be distinguished by the size and shape of the lower premolars which can be laterally compressed and bladelike to somewhat robust and transversely widened depending on the lineage; p1 elongated and knoblike, posterior accessory cuspid often present in p2-3, generally present, although sometimes very small, in p4. No posterior accessory cusps on P1-3. P4 is short relative to the molars with a well developed, more or less posteriorly situated protocone and a short metastyle and sometimes with a very small parastyle. Low cuspidated M1 with postprotocrista joining directly the distal border, except in Adelpharctos Bonis, 1971, without any contact with the metacone thus the trigone basin is posteriorly open; there is a trend to a sub-quadrate occlusal outline with inflation of the metaconule; large lingual and variable buccal cingulum; M2 smaller and lower cuspidated than M1 with a variable occlusal outline from triangular to oval, buccal cusps being separated or quite fused; ursid basicranium with Type A bulla (Hunt 1974), well ossified, completely enclosing middle ear, with inflation of medial part of bulla in some species.</p> <p>GENERA INCLUDED. — Cephalogale, Adelpharctos Bonis, 1971, Filholictis n. gen., Phoberogale Ginsburg &amp; Morales, 1995, Cyonarctos n. gen.</p></div> 	http://treatment.plazi.org/id/03AE87A4700AFFCCFF15FAF5FC0DFAA2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bonis, Louis De	Bonis, Louis De (2013): Ursidae (Mammalia, Carnivora) from the Late Oligocene of the “ Phosphorites du Quercy ” (France) and a reappraisal of the genus Cephalogale Geoffroy, 1862. Geodiversitas 35 (4): 787-814, DOI: 10.5252/g2013n4a4, URL: http://dx.doi.org/10.5252/g2013n4a4
03AE87A4700BFFCDFF06FF61FDB5FDF2.text	03AE87A4700BFFCDFF06FF61FDB5FDF2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Adelpharctos Bonis 1971	<div><p>Adelpharctos Bonis, 1971</p> <p>EMENDED DIAGNOSIS. — Cephalogalini n. tr. with a more cutting dentition than the other genera of the subfamily, m1 shorter relative to the premolar row with a less reduced metaconid and a talonid tapering backward. P2-3 with three roots, P4 short and robust, triangular shaped M1 with a metaconule more buccal relatively to the protocone than in other Cephalogalini n. tr. M1, hypometacrista joining the base of the metacone and with a modest linguo-distal cingulum.</p> <p>TYPE SPECIES. — A. mirus Bonis, 1971.</p> <p>OTHER SPECIES. — A. ginsburgi Bonis, 2011.</p> </div>	http://treatment.plazi.org/id/03AE87A4700BFFCDFF06FF61FDB5FDF2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bonis, Louis De	Bonis, Louis De (2013): Ursidae (Mammalia, Carnivora) from the Late Oligocene of the “ Phosphorites du Quercy ” (France) and a reappraisal of the genus Cephalogale Geoffroy, 1862. Geodiversitas 35 (4): 787-814, DOI: 10.5252/g2013n4a4, URL: http://dx.doi.org/10.5252/g2013n4a4
03AE87A4700BFFCDFCBCFDCDFB26FBD9.text	03AE87A4700BFFCDFCBCFDCDFB26FBD9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyonarctos Bonis 2013	<div><p>Cyonarctos n. gen.</p> <p>ETYMOLOGY. — From the Greek cyon (dog) and arctos (bear) for an ursid looking like a canid.</p> <p>TYPE SPECIES. — Cyonarctos dessei n. sp. by present designation.</p> <p>DIAGNOSIS. — Ŋat of the type species.</p> <p>DIFFERENTIAL DIAGNOSIS. — Adelpharctos differs by the larger size, the relatively higher trigonid and the short talonid of m1, the three rooted P2-P3 and the more V shaped M1. Cephalogale lacks a pacd in p2-p3, p4 is lower than the paraconid of m1, the premolar crowns more mas - sive, diastemas between p2-p3 and p3-p4. Phoberogale has lower and more elongated P2-P3 and p2-p4, and robust cingula and cingulids Filholictis n. gen. is more primitive with m1 shorter relative to the premolars and with its metaconid in contact with the paraconid, and a short m2.</p> </div>	http://treatment.plazi.org/id/03AE87A4700BFFCDFCBCFDCDFB26FBD9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bonis, Louis De	Bonis, Louis De (2013): Ursidae (Mammalia, Carnivora) from the Late Oligocene of the “ Phosphorites du Quercy ” (France) and a reappraisal of the genus Cephalogale Geoffroy, 1862. Geodiversitas 35 (4): 787-814, DOI: 10.5252/g2013n4a4, URL: http://dx.doi.org/10.5252/g2013n4a4
03AE87A4700BFFCDFED2FD0AFDDFFC36.text	03AE87A4700BFFCDFED2FD0AFDDFFC36.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Filholictis Bonis 2013	<div><p>Filholictis n. gen.</p> <p>TYPE AND UNIQUE SPECIES. — Canis filholi Munier- Chalmas in Filhol, 1877 by original designation.</p> <p>DERIVATIO NOMINIS. — Dedicated to H. Filhol who published many Quercy fossils.</p> <p>DIAGNOSIS. — Same as that of the type species.</p> <p>Filholictis filholi</p> <p>(Munier-Chalmas in Filhol, 1877)</p> <p>Canis filholi Munier-Chalmas in Filhol, 1877: 319, figs 123, 124. — Beaumont 1965: 23.</p> <p>Cephalogale filholi – Schlosser 1888: 103; 1904-1905: 293. — Bonis 1973: 82.</p> <p>HOLOTYPE. — Mandible figured by Filhol (1877: figs 123, 124); (MNHN.F.nn).</p> <p>DIAGNOSIS. — Primitive middle sized Cephalogalini n.tr. differing from the Amphicynodontidae by the large lower carnassial relative to the premolars and from the other Cephalogalini n. tr. by a less reduced metaconid of m 1 in contact with the paraconid and closing the central valley of the trigonid. High lower premolars without pacd, p3 as tall as p4 both being higher than the paraconid of m1; m2 relatively short for a Hemicyoninae, paraconid present but metaconid more developed than the protoconid.</p> <p>TYPE LOCALITY. — Unknown, “Phosphorites du Quercy ”.</p> <p>GEOLOGICAL AGE. — Unknown but probably Oligocene.</p> <p>Phoberogale</p> <p>Ginsburg &amp; Morales, 1995</p> <p>DIAGNOSIS. — Medium sized to large Cephalogalini n. tr., low and elongated P2-P3 and p2-p4, p2-p4 with developed pacd, robust cingula and cingulids in all the cheek teeth.</p> <p>TYPE SPECIES. — Cephalogale depereti Viret, 1929 by original designation.</p> <p>OTHER SPECIES. — P. gracile (Viret, 1929); P. bonali (Helbing, 1928); P. minor (Filhol, 1877).</p> </div>	http://treatment.plazi.org/id/03AE87A4700BFFCDFED2FD0AFDDFFC36	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bonis, Louis De	Bonis, Louis De (2013): Ursidae (Mammalia, Carnivora) from the Late Oligocene of the “ Phosphorites du Quercy ” (France) and a reappraisal of the genus Cephalogale Geoffroy, 1862. Geodiversitas 35 (4): 787-814, DOI: 10.5252/g2013n4a4, URL: http://dx.doi.org/10.5252/g2013n4a4
03AE87A4700BFFCDFED0FC56FD1AF90D.text	03AE87A4700BFFCDFED0FC56FD1AF90D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Filholictis filholi (Filhol 1877)	<div><p>Filholictis filholi</p> <p>(Munier-Chalmas in Filhol, 1877)</p> <p>Canis filholi Munier-Chalmas in Filhol, 1877: 319, figs 123, 124. — Beaumont 1965: 23.</p> <p>Cephalogale filholi – Schlosser 1888: 103; 1904-1905: 293. — Bonis 1973: 82.</p> <p>HOLOTYPE. — Mandible figured by Filhol (1877: figs 123, 124); (MNHN.F.nn).</p> <p>DIAGNOSIS. — Primitive middle sized Cephalogalini n.tr. differing from the Amphicynodontidae by the large lower carnassial relative to the premolars and from the other Cephalogalini n. tr. by a less reduced metaconid of m 1 in contact with the paraconid and closing the central valley of the trigonid. High lower premolars without pacd, p3 as tall as p4 both being higher than the paraconid of m1; m2 relatively short for a Hemicyoninae, paraconid present but metaconid more developed than the protoconid.</p> <p>TYPE LOCALITY. — Unknown, “Phosphorites du Quercy ”.</p> <p>GEOLOGICAL AGE. — Unknown but probably Oligocene.</p> <p>Phoberogale</p> <p>Ginsburg &amp; Morales, 1995</p> <p>DIAGNOSIS. — Medium sized to large Cephalogalini n. tr., low and elongated P2-P3 and p2-p4, p2-p4 with developed pacd, robust cingula and cingulids in all the cheek teeth.</p> <p>TYPE SPECIES. — Cephalogale depereti Viret, 1929 by original designation.</p> <p>OTHER SPECIES. — P. gracile (Viret, 1929); P. bonali (Helbing, 1928); P. minor (Filhol, 1877).</p> </div>	http://treatment.plazi.org/id/03AE87A4700BFFCDFED0FC56FD1AF90D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bonis, Louis De	Bonis, Louis De (2013): Ursidae (Mammalia, Carnivora) from the Late Oligocene of the “ Phosphorites du Quercy ” (France) and a reappraisal of the genus Cephalogale Geoffroy, 1862. Geodiversitas 35 (4): 787-814, DOI: 10.5252/g2013n4a4, URL: http://dx.doi.org/10.5252/g2013n4a4
03AE87A4700BFFCDFC82FF61FBABFDBC.text	03AE87A4700BFFCDFC82FF61FBABFDBC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phoberogale Ginsburg & Morales 1995	<div><p>Phoberogale</p> <p>Ginsburg &amp; Morales, 1995</p> <p>DIAGNOSIS. — Medium sized to large Cephalogalini n. tr., low and elongated P2-P3 and p2-p4, p2-p4 with developed pacd, robust cingula and cingulids in all the cheek teeth.</p> <p>TYPE SPECIES. — Cephalogale depereti Viret, 1929 by original designation.</p> <p>OTHER SPECIES. — P. gracile (Viret, 1929); P. bonali (Helbing, 1928); P. minor (Filhol, 1877).</p> </div>	http://treatment.plazi.org/id/03AE87A4700BFFCDFC82FF61FBABFDBC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bonis, Louis De	Bonis, Louis De (2013): Ursidae (Mammalia, Carnivora) from the Late Oligocene of the “ Phosphorites du Quercy ” (France) and a reappraisal of the genus Cephalogale Geoffroy, 1862. Geodiversitas 35 (4): 787-814, DOI: 10.5252/g2013n4a4, URL: http://dx.doi.org/10.5252/g2013n4a4
03AE87A4700BFFC7FCC1FB74FE35F90D.text	03AE87A4700BFFC7FCC1FB74FE35F90D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyonarctos DESSEI N.	<div><p>Cyonarctos dessei n. sp.</p> <p>Cephalogale cf. minor – Bonis 1974: 28. Non Cephalogale minor Filhol, 1882.</p> <p>Cephalogale minor – Remy et al. 1987: 187. — Biochrom’ 97: 793.</p> <p>Cephalogale cadurcensis – Remy et al. 1987: 187. Non Canis cadurcensis Filhol, 1877.</p> <p>Cephalogale sp. – Remy et al. 1987: 187.</p> <p>HOLOTYPE. — Left hemi-mandible with p2-m2 (PD 477) from Pech Desse.</p> <p>PARATYPES. — Left hemi-mandible with broken p2, p3-m3 (PD 475), left hemi-mandible with c, p2-m2 (PD 273), fragment of right maxilla with P2-M1, (PD 130), frag-</p> <p>Dimension 1; Eigenvalue:,00031 (52,34% of Inertia)</p> <p>ment of right maxilla with P4-M2 (PD 507), left M2 (PD 514), right M1 (PD 137) and other labelled material from Pech Desse which are identified as the components of the original type series (ICZN: art. 73 D).</p> <p>LOCALITY. — Pech Desse near Mouillac, Quercy, France.</p> <p>REFERRED SPECIMENS. — See material from Pech du Fraysse.</p> <p>DATING. — MP 28 (about 24.9-24.5 Ma).</p> <p>DISTRIBUTION. — Quercy (South western France).</p> <p>ETYMOLOGY. — Dedicated to my colleague Jean Desse who gave his name to the locality.</p> <p>DIAGNOSIS. — Small,(minimum length of m1 = 14.5 mm), to medium-sized (maximum length of m1 = 18.2 mm) Cephalogalini n. tr. P2-P3 without posterior accessory cusp (pac); P2 slightly asymmetrical; P3 symmetrical; P2-3 with tall and acute crowns; P4 with moderately distally displaced large protocone, moderate cingulum, very small parastyle, short metastyle blade; triangular M1with paracone higher than metacone, V shaped protocone of which the anterior arm joins the mesial cingulum while the posterior one turns distally and thus opening the trigone basin; oval M2 with metacone smaller than or similar to the paracone and close to or separated from the paracone; small one-rooted p1; p1-p3 asymmetrical without posterior accessory cuspid (pacd); p4 more symmetrical with a developed pacd; p2 as tall as p3 and p4 taller than the m1 paraconid; m1 with moderately reduced metaconid and a “ Cephalogale like” talonid; m2 with a small but clear paraconid and a relatively short talonid; small m3 with a clear metaconid and a distinct talonid.</p> <p>BIOMETRY OF CYONARCTOS DESSEI N. GEN., N. SP.</p> <p>Ŋe morphological characters of the material from Pech du Fraysse and Pech Desse correspond to those of the tribe Cephalogalini n. tr. It can be divided in each locality in two populations clearly separated by the size but attributed to one species.</p> <p>Metric analysis of the dental material included within the genus Cyonarctos n. gen. in the two studied localities, and especially m1 measurements, is used to decide if the fossils belong to a single species or not. Although the m1 length would be “the poorest predictor of body weight” (Van Valkenburgh 1990), it generally gives a good indication on the sexual dimorphism in extant species of carnivorans. A principal components analysis (PCA) based on m1 dimensions (Fig. 1A) shows a group of small specimens clearly separated from a group of larger specimens. Ŋe morphological characters of these groups are quite similar and the problem is to know if there are two size separated species or only one dimorphic species, generally carnivoran males being larger than females with some exceptions. A correspondence analysis (CA) shows that the two groups are mixed (Fig. 1B). Ŋe CA being based on the differences in proportions (Bonis &amp; Lebeau 1974), it indicates that both groups, small and large individuals, have the same proportions. Ŋe difference in size between the two groups is tested by comparisons with extant and fossil species.</p> <p>Metric comparisons with extant or fossil taxa were made from published data (Del Campana 1913; Spahni 1955; Bonifay 1971, 1972; Poplin 1972; Bonifay 1975; Davis 1977; Klein 1986; Berta 1988; Legendre &amp; Roth 1988; Van Valkenburgh 1990; Dayan et al. 1992; Kieser &amp; Kroeneveld 1992a, b; Koufos 1992; Fistani &amp; Crégut-Bonhoure 1993; Okarma &amp; Buchalczyk 1993; Argand 1995; Crégut-Bonnoure 1996; Rook &amp; Torre 1996; Baryshnikov 1996, 2012; Gittlleman &amp; Van Valkenburg 1997; Szuma 2000; Dayan et al. 2002; Palmqvist et al. 2002; Abramov &amp; Puzachenko 2005; Baryshnikov &amp; Tsukala 2010; Castel et al. 2010; Pérez-Rippol et al. 2010; Woldrich 1879) and personal data.</p> <p>A sexual dimorphism index (SDI 1) can be computed for m1 by the difference between male and female m1 length means × 100 divided by the sum of both means:</p> <p>(Lm 1a – Lm 1b) × 100</p> <p>Lm 1a + Lm 1b</p> <p>Ŋis ratio gives the difference between male and female specimens relatively to the size of m1. It has been computed for 152 extant populations of carnivorans analysed from data in the literature. Ŋe resulting histogram is J shaped (Fig. 2), the lowest values, (similar of slightly different sex sizes), are close to the vertical axis and the highest values (large difference between both sexes) are far from it. Most of the species indices are situated between the values zero to four (116 species or populations). Ŋe SDI 1 for the two Quercy localities (7.2) is between four and eight values with 26 extant species, eight species having higher indices. Ŋe largest differences may occur in large species as Panthera leo (Linnaeus, 1758), Panthera tigris (Linnaeus., 1758) or Ursus arctos Linnaeus, 1758 but also in medium sized as Gulo gulo Linnaeus, 1758 or Felis temmincki Vigors &amp; Horsfield, 1827 and small species as Mustela erminea Linnaeus, 1758. Indices can vary within a same genus and even within the same species depending on the origin of the sample (0.16 to 4.8 for Canis latrans Say, 1823, 2 to 10 for Ursus arctos, 3.25 to 10.4 for Panthera leo, 1.5 to 5.8 for Nasua nasua Linnaeus, 1766 and 0.3 to 5.1 for Vulpes vulpes (Linnaeus, 1758)). Ŋe SDI 1 of the Pech Desse-Pech du Fraysse sample is compatible with a single species hypothesis.</p> <p>Ŋe observed range of the m1 length within a group of specimens corresponds to another index of sexual dimorphism (SDI 2): “(Lm1 of the smallest specimen/Lm1 of the largest specimen) × 100”. Here we compare the group of small specimens (females?) and the group of large specimens (males?) of the Fig. 1. In a sample of m1, if the difference in length between the two extreme specimens is low, the ratio will be close to 100, but if the difference is significant, the ratio will be lower. When the ratio of the difference in size of the m1s is computed in extant carnivorans whose sexes are known, we observe that this index, computed in large populations, is higher for each sex than when the two morphs are mixed, indicating the sexual dimorphism (Table 3). For example, in Meles meles Linnaeus, 1758 from Transcaucasia, the index is 76 for the males, 88 for the females but only 75 when the two sexes are mixed. Ŋat index reaches 96 for the group of small specimens (14.49/15.05) and 90 for the group of large specimens (16.46/18.22) of Pech Desse and Pech du Fraysse. It reaches only 79.5 when the two groups are mixed (14.49/18.22). Ŋe last value is close to those which are generally found in extant and fossil carnivorans when both sexes are mixed. Even if this index is partially depending on the number of specimens, it gives a quite good idea on the sex difference in a sample of extant or fossil carnivorans. In the case of the studied fossils, we conclude that the numbers are compatible with a single species hypothesis.</p> <p>Ŋus each index within both groups, small and large, of the Quercy material seems to be too high to belong to two different species but they are convenient with a sexual dimorphism. Ŋus, on these criteria, the whole material from Pech Desse and Pech du Fraysse seems to be within the range of a single species.</p> <p>DESCRIPTION OF CYONARCTOS DESSEI N. GEN., N. SP. Mandible</p> <p>Ŋere are four quite complete mandibles from Pech Desse, one from Pech du Fraysse (Fig.3) and several fragments from Pech Desse and Pech du Fraysse. Ŋe corpus is quite robust (PD 273) or more slender (PD 475, 476, 477, PF 420) but it is always anteriorly tapering. Ŋe ventral border is gently convex in all specimens and the maximum height of the mandible is under m1. Ŋere is one mental foramen beneath the mesial root of p2 and another one beneath p3. Ŋe coronoid process is high with a large masseteric fossa. Ŋe condyle, moderately extended laterally, is situated at the level of the apex of m1. Ŋe symphyseal plate has a defined area of smooth bone above the rugose portion (Figs 3B, 4B). It is interpreted as corresponding to the presence of a fibrocartilage pad as in the wolf and allowing a quite flexible mandible symphysis and acting as a cushion during the displacements of the two branches of the mandible during chew- ing (Scapino 1965, 1981; Hunt 2009). Below the smooth plate, there are bony rugosities forming quite horizontal ridges converging forward. Two small foramina are ventrally situated beneath the canine for entry of nerves and blood vessels into this area (Scapino 1965: fig. 12). Ŋe smooth surface of the symphyseal plate is a horizontal rectangle in the large mandible PD 273 and a smaller elongated oblique oval in PD 477.</p> <p>Lower dentition (Table 1; Figs 3-6)</p> <p>Ŋe incisors are missing in all the specimens but two very small alveoli, corresponding to i2 and i3 are present in PD 315 and PD 477 mesio-buccally of the canine; the former is mesio-distally elongated and narrow, the latter is rounded and just a little enlarged. Ŋere is no place for a first incisor alveolus. Ŋe canine, whose crown is partially broken in PD 273, has a robust root but must have been quite low. Ŋe single rooted p1 is lacking in all the specimens. Ŋe other premolars increase in size from p2 to p4. Ŋe former is low, slightly elongated, without any accessory cusp and with a distal half longer than the mesial one and a faint distal crest from apex to the base. Separated by a short diastema (1 to 4 mm), p3 is quite similar to p2 but the difference in length between mesial and distal halves of the crown is lesser and there is possibly a very minute posterior accessory cuspid (pacd) and a mesial basal cuspid (PD 273); a slight crest runs distally from the apex to the base. Ŋe crown of p4 is more symmetrical, with both mesial and distal basal cuspids and with a well developed posterior accessory cuspid in the distal crest. Ŋe premolars are quite tall, p3 being as tall as the paraconid of m1 and p4 being taller than it. In m1, the protoconid is taller than the paraconid; the metaconid is reduced and posteriorly displaced, clearly separated from the paraconid. Ŋe talonid has an elongated and dominant hypoconid prolonged by a crest which encloses the narrow talonid basin and joins the metaconid without any notch. Ŋe metaconid of m2 is the taller cuspid of this tooth; the paraconid is present and a small mesio-buccal crest encloses the small trigonid basin; the talonid is similar t in shape to that of m1. Ŋe m3 is oval, the metaconid higher than the protoconid and the paraconid is very small, a small hypoconid is present in the buccal crest. Ŋe cingulids are present but weak in premolars and molars.</p> <p>Upper dentition (Table 2; Fig. 7)</p> <p>A piece of maxilla with P3 and part of the mesial alveoli of P4 (PF 46), another one with P3-P4 (PF 37) and a third with P2-M1 (PD 130) are the witnesses of ante-carnassial premolars; their crowns are simple, acute, without accessory cusp, slightly elongated, lower than the paracone of P4 and the mesial portion is shorter than the distal one, especially in P3; there are two small crests, a mesial one from the apex to the mesio-lingual corner and a distal one from the apex to the distal border; a weak cingulum surrounds the base of the crown of P3. Ŋe carnassial is short relatively to the molars with a relatively short and quite robust metastyle; the protocone is a large semi-circular basined platform surrounded by a moderate cingulum and situated quite anteriorly for a hemicyonine; the moderate cingulum is surrounding the whole crown; a mesial crest runs from the apex of the paracone to the cingulum which is turning up as a very small parastyle and separated from the protocone by a shallow notch. In M1, the V shaped protocone is anteriorly prolonged by the preprotocrista to the base of the mesial surface of the paracone and posteriorly by a very tiny, quite indistinct metaconule, the large trigone basin being distally open; the paracone is larger and taller than the metacone and its mesial width is larger than its distal one; a distinct cingulum surrounds the buccal face ending mesially by a small bump corresponding to a tiny parastyle. Ŋe lingual cingulum is developed around the protocone but more distolingually. M2 has an oval occlusal outline; the paracone generally is larger and slightly taller than the metacone; the other characters are similar to those of M1 although the V is more open, its distal crest being almost parallel to the sagittal plane, and the cingulum better spread around the protocone although posteriorly thicker.</p></div> 	http://treatment.plazi.org/id/03AE87A4700BFFC7FCC1FB74FE35F90D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bonis, Louis De	Bonis, Louis De (2013): Ursidae (Mammalia, Carnivora) from the Late Oligocene of the “ Phosphorites du Quercy ” (France) and a reappraisal of the genus Cephalogale Geoffroy, 1862. Geodiversitas 35 (4): 787-814, DOI: 10.5252/g2013n4a4, URL: http://dx.doi.org/10.5252/g2013n4a4
