identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
039B87C7FF82525BC6A9FD6EFDF93DC4.text	039B87C7FF82525BC6A9FD6EFDF93DC4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Incolacridinae Tinkham 1940	<div><p>Subfamily Incolacridinae Tinkham, 1940</p> <p>Incolacri Tinkham, 1940: 276, 302.</p> <p>Incolacridini: Storozhenko, 2018: 63.</p> <p>Type genus: Incolacris C. Willemse, 1932.</p> <p>Description. Body robust, medium to small size (Figs. 1, 25–28). Head rugose, considerably shorter than pronotum. Face in profile reclinated; frontal ridge below the antennal bases straight (Figs. 4, 8) or excised (Fig. 1). Fastigium of vertex triangular, deeply sulcate and projecting forward at the distance less than the half of eye length; lateral margins bordered by low carinulae reaching apical margin of the compound eyes; foveolae shallow, short triangular. Vertex between eyes considerably narrower than an eye (Fig. 9). Eyes large, oval. Frontal ridge distinct, sulcate, with margins almost parallel (Figs. 3, 5) or narrowed between antennal bases (Fig. 27). Antennae filiform. Pronotum rugose, crossed by three transverse furrows; prozona distinctly longer than metazona; median carina vestigial; lateral carinae absent; anterior margin of prozona almost straight; posterior margin of metazona triangularly rounded (Figs. 9, 28). Prosternal spine short, straight, conical, with broad base and pointed apex. Mesosternal lobes widely separated, subsquare or trapezoidal; mesosternal interspace transverse; metasternal lobes widely separated (Fig. 31). Tegmina and hind wings always present; apterous forms unknown. Hind femora stout; both dorsal and ventral median carinae finely serrated; dorso-median carina terminating in a small tooth; ventral genicular lobes of hind knee with distinct apical spine (Fig. 12) or narrowly rounded at apex (Fig. 14). Hind tibiae not apically expanded but provided by weakly keeled margins; outer apical dorsal spine small. Hind tarsi short, not longer than the half of hind tibiae; third segment (without claws) as long as two others together or slightly shorter; arolium large, reaching the middle or apex of claws. Tympanum large, oval. Male 10th abdominal tergite with distinct furculae or without it; supra-anal plate triangular, trapezoidal or long-oval, with a median basal impression; cerci compressed, short, conical, with pointed or blunt apex, unarmed or provided by tooth on the inner side; subgenital plate short (Figs. 10, 11. 13). Female supra-anal plate elongated, triangular, with a median basal impression; cerci conical with pointed apex; subgenital plate with long triangle pointed apex; basivalvular plates wide (Fig. 7). Ovipositor short or elongated; dorsal valves shorter than lower ones, widened apically, dentate only at apex; ventral valves slightly curved and also dentate or crenulate at apex only (Fig. 6). Male genitalia: epiphallus bridge-shaped, asymmetrical; bridge completely divided; left part of epiphallus larger what right one; oval sclerite present (Figs. 15–19, 22); valves of cingulum varied from long and narrow (Fig. 23) to short and stout (Fig. 21); apical valves of penis conical or lobe-like; basal valves of penis connected with apical ones by unbroken flexure (Figs. 21, 33).</p> <p>Differential diagnosis. Incolacridinae differs from all other subfamilies of Acrididae in the completely divided asymmetrical epiphallus and in specific shape of ovipositor, which widened apically and dentate only at apex (both characters are the brightest synapomorphies of this subfamily). The divided epiphallus is also typical for the subfamily Oxyinae and the tribe Mesambriini of the subfamily Catantopinae (Hollis, 1975; Storozhenko, 2018) but in the above mentioned taxa the epiphallus is always symmetrical and ovipositor valves are hook-like or spined/serrate along the full length. Ventral genicular lobes of hind knee in the subfamily Catantopinae are broadly rounded, in Oxyinae with distinct apical spine, while in Incolacridinae spined or narrowly rounded at apex. Mesosternal interspace is narrowly longitudinal in Oxyinae, but in Mesambriini and Incolacridinae it is transverse. Body in Incolacridinae is rugose and similar in this aspect to the tribe Mesambriini, while in Oxyinae body with smooth integument. In modern classification of grasshoppers the tribe Incolacridini occupies intermediate position between Catantopinae and Oxyinae; therefore it considered here as a separate subfamily.</p> <p>Composition. This small subfamily consists of four genera from a nominotypical tribe Incolacridini Tinkham, 1940 distributed in the Oriental region.</p> </div>	https://treatment.plazi.org/id/039B87C7FF82525BC6A9FD6EFDF93DC4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Storozhenko, Sergey Yu.	Storozhenko, Sergey Yu. (2021): On the exact position of the tribe Incolacridini in the modern classification of grasshoppers (Orthoptera: Acrididae). Zootaxa 4970 (1): 106-118, DOI: https://doi.org/10.11646/zootaxa.4970.1.3
039B87C7FF81525BC6A9FB9EFD1D3ACF.text	039B87C7FF81525BC6A9FB9EFD1D3ACF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Incolacris C. Willemse 1932	<div><p>Genus Incolacris C. Willemse, 1932, stat. resurr.</p> <p>Incolacris C. Willemse, 1932: 150 (type species: Incolacris rubritarsi C. Willemse, 1932, by original designation); C. Willemse, 1939: 89; Tinkham, 1940: 302; C. Willemse, 1956: 185.</p> <p>Stolzia (part.): Hollis, 1975: 211; Otte, 1995: 338; Yin et al., 1996: 679; Li et al., 2006: 55.</p> <p>Description. Head in frontal view elongated; frontal ridge broad, complete, almost reaching clypeus (Fig. 5). Face in lateral view reclinated; frontal ridge almost straight (Figs. 4, 8). Ventral genicular lobes of the hind femora triangular with distinct apical spine (Fig. 12). Male 10th abdominal tergite without furculae (Fig. 10). Male supra-anal plate triangular, with a median basal impression, lateral margins with two teeth in the basal half on each side or without it. Male cerci surpassing the supra-anal plate, compressed, conical, with pointed apex, without teeth on the inner side (Fig. 11). Valves of ovipositor short, with apex rounded and strongly dentate (Fig. 6). Male genitalia: epiphallus with broad bridge, ancorae present (Fig. 15); the shape of other parts of genitalia unknown.</p> <p>Species included. The genus consists of five species distributed in the Philippines, South China and Malaysia.</p> <p>Remarks. Hollis (1975) synonymized Incolacris under Stolzia based the only on the asymmetrical shape of epiphallus but I disagree with such synonymy and consider it as a distinct genus differs from other genera of Incolacridinae in the specific shape of frontal ridge.</p> </div>	https://treatment.plazi.org/id/039B87C7FF81525BC6A9FB9EFD1D3ACF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Storozhenko, Sergey Yu.	Storozhenko, Sergey Yu. (2021): On the exact position of the tribe Incolacridini in the modern classification of grasshoppers (Orthoptera: Acrididae). Zootaxa 4970 (1): 106-118, DOI: https://doi.org/10.11646/zootaxa.4970.1.3
039B87C7FF81525BC6A9F90FFBEC3B9D.text	039B87C7FF81525BC6A9F90FFBEC3B9D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Incolacris flavomaculata C. Willemse 1939	<div><p>Incolacris flavomaculata C. Willemse, 1939</p> <p>Incolacris flavomaculata Willemse, 1939: 89, 90, fig. 3 (holotype —female, Peninsular Malaysia, Malaca: Dinding Island; in NHME Maastricht); C. Willemse, 1956: 188, fig. 104.</p> <p>Stolzia flavomaculata: Hollis, 1975: 216; Otte, 1995: 339; Yin et al., 1996: 679.</p> <p>Distribution. Malaysia (Dinding Island near the Malay Peninsula).</p> <p>Remarks. Male unknown. Herein this species is returned to the genus Incolacris.</p> </div>	https://treatment.plazi.org/id/039B87C7FF81525BC6A9F90FFBEC3B9D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Storozhenko, Sergey Yu.	Storozhenko, Sergey Yu. (2021): On the exact position of the tribe Incolacridini in the modern classification of grasshoppers (Orthoptera: Acrididae). Zootaxa 4970 (1): 106-118, DOI: https://doi.org/10.11646/zootaxa.4970.1.3
039B87C7FF80525AC6A9FF4BFBC53DF1.text	039B87C7FF80525AC6A9FF4BFBC53DF1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Incolacris hainanensis Tinkham 1940	<div><p>Incolacris hainanensis Tinkham, 1940</p> <p>Incolacris hainanensis Tinkham, 1940: 302, pl. 11, fig. 2 (holotype —male, China, China Southeast, Guangdong, Hainan, Ta Hian; in ICRI Zhongshan).</p> <p>Stolzia hainanensis: Hollis, 1975: 216; Zheng &amp; Ma, 1989: 329; Otte, 1995: 339; Yin et al., 1996: 679; Li et al., 2006: 55, fig. 27.</p> <p>Distribution. China (Hainan Island).</p> <p>Remarks. Female unknown. Herein this species is returned to the genus Incolacris.</p> </div>	https://treatment.plazi.org/id/039B87C7FF80525AC6A9FF4BFBC53DF1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Storozhenko, Sergey Yu.	Storozhenko, Sergey Yu. (2021): On the exact position of the tribe Incolacridini in the modern classification of grasshoppers (Orthoptera: Acrididae). Zootaxa 4970 (1): 106-118, DOI: https://doi.org/10.11646/zootaxa.4970.1.3
039B87C7FF80525AC6A9FDD1FB183F57.text	039B87C7FF80525AC6A9FDD1FB183F57.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Incolacris jianfengensis (Zheng et Ma 1989) Storozhenko 2021	<div><p>Incolacris jianfengensis (Zheng et Ma, 1989), comb. nov.</p> <p>Figs. 8–12</p> <p>Stolzia jianfengensis Zheng &amp; Ma, 1989: 325, 329, figs. 1–6 (holotype —male, China, Hainan, Ledong, Jianfengling Mountain; in SNNU Shaanxi); Otte, 1995: 339; Yin et al., 1996: 679; Li et al., 2006: 57, fig. 28.</p> <p>Distribution. China (Hainan Island).</p> <p>Remarks. Female unknown. Herein this species is transferred from Stolzia to Incolacris.</p> </div>	https://treatment.plazi.org/id/039B87C7FF80525AC6A9FDD1FB183F57	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Storozhenko, Sergey Yu.	Storozhenko, Sergey Yu. (2021): On the exact position of the tribe Incolacridini in the modern classification of grasshoppers (Orthoptera: Acrididae). Zootaxa 4970 (1): 106-118, DOI: https://doi.org/10.11646/zootaxa.4970.1.3
039B87C7FF80525AC6A9FCB7FE233895.text	039B87C7FF80525AC6A9FCB7FE233895.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Incolacris rubritarsi C. Willemse 1932	<div><p>Incolacris rubritarsi C. Willemse, 1932</p> <p>Figs. 4–7</p> <p>Incolacris rubritarsi C. Willemse, 1932: 151, pl. XI, fig. 1 (holotype —female, Philippines, Siargao Island; in NHME Maastricht); Sjöstedt, 1933: 33; C. Willemse, 1939: 90; Tinkham, 1940: 302; C. Willemse, 1956: 188; Petersen &amp; Gaedike, 1970: 169.</p> <p>Stolzia rubritarsi: Hollis, 1975: 216, figs 36, 37; Otte, 1995: 339; Yin et al., 1996: 679.</p> <p>Distribution. Philippines (Siargao Island).</p> <p>Remarks. Male unknown. The photos of paratype female are available in the Orthoptera Species File database (Cigliano et al., 2021).</p> </div>	https://treatment.plazi.org/id/039B87C7FF80525AC6A9FCB7FE233895	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Storozhenko, Sergey Yu.	Storozhenko, Sergey Yu. (2021): On the exact position of the tribe Incolacridini in the modern classification of grasshoppers (Orthoptera: Acrididae). Zootaxa 4970 (1): 106-118, DOI: https://doi.org/10.11646/zootaxa.4970.1.3
039B87C7FF80525AC6A9FAF5FB923A35.text	039B87C7FF80525AC6A9FAF5FB923A35.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Incolacris trifasciata C. Willemse 1932	<div><p>Incolacris trifasciata C. Willemse, 1932</p> <p>Fig. 15</p> <p>Incolacris trifasciata C. Willemse, 1932: 152 (holotype —male, Philippines, Siargao Island; in NHME Maastricht); Sjöstedt, 1933: 33; C. Willemse, 1939: 89; Tinkham, 1940: 302; C. Willemse, 1956: 187; Petersen &amp; Gaedike, 1970: 17.</p> <p>Stolzia trifasciata: Hollis, 1975: 216, fig. 34; Otte, 1995: 339; Yin et al., 1996: 679.</p> <p>Distribution. Philippines (Siargao Island).</p> <p>Remarks. Female unknown. Herein this species is returned to the genus Incolacris. The photos of paratype male are available in the Orthoptera Species File database (Cigliano et al., 2021).</p> </div>	https://treatment.plazi.org/id/039B87C7FF80525AC6A9FAF5FB923A35	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Storozhenko, Sergey Yu.	Storozhenko, Sergey Yu. (2021): On the exact position of the tribe Incolacridini in the modern classification of grasshoppers (Orthoptera: Acrididae). Zootaxa 4970 (1): 106-118, DOI: https://doi.org/10.11646/zootaxa.4970.1.3
039B87C7FF80525DC6A9F915FE293DE0.text	039B87C7FF80525DC6A9F915FE293DE0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stolzia C. Willemse 1930	<div><p>Genus Stolzia C. Willemse, 1930</p> <p>Stolzia C. Willemse, 1930: 104, 117 (type species: Stolzia rubromaculata C. Willemse, 1930, by original designation); C. Willemse, 1956: 116; Hollis, 1975: 211; Otte, 1995: 338; Yin et al., 1996: 679; Li et al., 2006: 55; Storozhenko, 2020: 16.</p> <p>Butonacris C. Willemse, 1933:118 (type species: Butonacris fasciata C. Willemse, 1933, by original designation); C. Willemse, 1938: 84; C. Willemse, 1956: 190; synonymized with Stolzia by Hollis (1975).</p> <p>Description. Head in frontal view oval; frontal ridge broad, disappear below the lower margins of eyes, not reaching clypeus (Fig. 5). Face in lateral view slightly reclinated; frontal ridge weakly excised below lower margin of eyes (Fig. 2). Ventral genicular lobes of the hind femora triangular with distinct apical spine. Male 10th abdominal tergite with or without furculae. Male supra-anal plate triangular, with a median basal impression, lateral margins sometimes with a blunt tooth on each side near the apex. Male cerci surpassing the supra-anal plate, compressed, conical, with pointed apex, with or without teeth on the inner side. Valves of ovipositor short, with apex rounded and strongly dentate. Male genitalia: epiphallus with broad bridge and large ancorae (Figs. 16–18); the shape of other parts of genitalia unknown.</p> <p>Species included. The genus consists of six species distributed over the Greater Sunda Islands (Java, Sumatra, Sulawesi, and Borneo).</p> </div>	https://treatment.plazi.org/id/039B87C7FF80525DC6A9F915FE293DE0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Storozhenko, Sergey Yu.	Storozhenko, Sergey Yu. (2021): On the exact position of the tribe Incolacridini in the modern classification of grasshoppers (Orthoptera: Acrididae). Zootaxa 4970 (1): 106-118, DOI: https://doi.org/10.11646/zootaxa.4970.1.3
039B87C7FF87525DC6A9FE22FDBF3F64.text	039B87C7FF87525DC6A9FE22FDBF3F64.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stolzia aberrans (C. Willemse 1938)	<div><p>Stolzia aberrans (C. Willemse, 1938)</p> <p>Fig. 17</p> <p>Butonacris aberrans C. Willemse, 1938: 84, fig. 2 (holotype —male, Malaysia, Central East Borneo; in BMNH London); C. Willemse, 1956: 194, fig. 108.</p> <p>Stolzia aberrans: Hollis, 1975: 216, fig. 32; Otte, 1995: 339; Yin et al., 1996: 679.</p> <p>Distribution. Malaysia (Borneo Island).</p> <p>Remarks. Female unknown.</p></div> 	https://treatment.plazi.org/id/039B87C7FF87525DC6A9FE22FDBF3F64	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Storozhenko, Sergey Yu.	Storozhenko, Sergey Yu. (2021): On the exact position of the tribe Incolacridini in the modern classification of grasshoppers (Orthoptera: Acrididae). Zootaxa 4970 (1): 106-118, DOI: https://doi.org/10.11646/zootaxa.4970.1.3
039B87C7FF87525DC6A9FCA6FDBF38CC.text	039B87C7FF87525DC6A9FCA6FDBF38CC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stolzia borneensis (C. Willemse 1938)	<div><p>Stolzia borneensis (C. Willemse, 1938)</p> <p>Butonacris borneensis C. Willemse, 1938: 83, fig. 1 (holotype —male, Malaysia, Borneo; in NBC Leiden); C. Willemse, 1956: 192, fig. 106.</p> <p>Stolzia borneensis: Hollis, 1975: 216, fig. 31; Otte, 1995: 339; Yin et al., 1996: 679.</p> <p>Distribution. Malaysia (Borneo Island).</p> <p>Remarks. Female unknown.</p></div> 	https://treatment.plazi.org/id/039B87C7FF87525DC6A9FCA6FDBF38CC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Storozhenko, Sergey Yu.	Storozhenko, Sergey Yu. (2021): On the exact position of the tribe Incolacridini in the modern classification of grasshoppers (Orthoptera: Acrididae). Zootaxa 4970 (1): 106-118, DOI: https://doi.org/10.11646/zootaxa.4970.1.3
039B87C7FF87525DC6A9FB0EFD093A14.text	039B87C7FF87525DC6A9FB0EFD093A14.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stolzia fasciata (C. Willemse 1933)	<div><p>Stolzia fasciata (C. Willemse, 1933)</p> <p>Fig. 18</p> <p>Butonacris fasciata C. Willemse, 1933: 118, fig. 4 (holotype —male, Malaysia, Sulawesi, Buton Island; in BMNH London); C. Willemse, 1938: 84, fig. 3; C. Willemse, 1956: 193, fig. 107.</p> <p>Stolzia fasciata: Hollis, 1975: 216, fig. 35; Otte, 1995: 339; Yin et al., 1996: 679.</p> <p>Incolacris atrifrons C. Willemse, 1956: 186 (holotype —female, Asia-Tropical, Boetonboligo; in NBC Leiden); synonymized with Stolzia fasciata by Hollis (1975).</p> <p>Stolzia atrifrons: Otte, 1995: 339.</p> <p>Distribution. Indonesia (Buton Island located off the southeast peninsula of Sulawesi Island).</p> <p>Remarks. Both male and female are known.</p></div> 	https://treatment.plazi.org/id/039B87C7FF87525DC6A9FB0EFD093A14	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Storozhenko, Sergey Yu.	Storozhenko, Sergey Yu. (2021): On the exact position of the tribe Incolacridini in the modern classification of grasshoppers (Orthoptera: Acrididae). Zootaxa 4970 (1): 106-118, DOI: https://doi.org/10.11646/zootaxa.4970.1.3
039B87C7FF87525DC6A9F976FE4A3B97.text	039B87C7FF87525DC6A9F976FE4A3B97.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stolzia javana Ramme 1941	<div><p>Stolzia javana Ramme, 1941</p> <p>Figs. 3, 16</p> <p>Stolzia javana Ramme, 1941: 101, pl. XIII, fig. 5 (holotype —female, Indonesia, Java, Malang; in MfN Berlin); C. Willemse, 1956: 117; Hollis, 1975: 216, fig. 33; Otte, 1995: 339; Yin et al., 1996: 679.</p> <p>Distribution. Indonesia (Java Island).</p> <p>Remarks. Both sexes are known. The photos of holotype female are available in the Orthoptera Species File database (Cigliano et al., 2021).</p> </div>	https://treatment.plazi.org/id/039B87C7FF87525DC6A9F976FE4A3B97	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Storozhenko, Sergey Yu.	Storozhenko, Sergey Yu. (2021): On the exact position of the tribe Incolacridini in the modern classification of grasshoppers (Orthoptera: Acrididae). Zootaxa 4970 (1): 106-118, DOI: https://doi.org/10.11646/zootaxa.4970.1.3
039B87C7FF86525CC6A9FCB7FF583998.text	039B87C7FF86525CC6A9FCB7FF583998.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bettotania C. Willemse 1933	<div><p>Genus Bettotania C. Willemse, 1933</p> <p>Bettotania C. Willemse, 1933: 75 (type species: Bettotania maculata C. Willemse, 1933, by original designation); C. Willemse, 1956: 112; Ingrisch, 1989: 227; Otte, 1995: 277; Yin et al., 1996: 96.</p> <p>Paracelebesia Miller, 1935a: 697 (type species: Paracelebesia festiva Miller, 1935, by original designation); synonymized with Bettotania by C. Willemse (1956).</p> <p>Description. Head in frontal view oval; frontal ridge narrow, almost reaching the clypeus. Face in lateral view reclinated; frontal ridge considerably projecting between the antennae and forming a widened rostrum (Fig. 1). Ventral genicular lobes of the hind femora narrowly triangular with apex obtuse, without spine. Male 10th abdominal tergite with furculae. Male supra-anal plate long-oval, with a median basal impression, lateral margins with a pointed tooth somewhat behind the middle, apex rounded. Male cerci compressed, conical, apex pointed, inner side without tooth. Valves of ovipositor relatively long, with apex rounded and crenulate (Fig. 1). Male genitalia: epiphallus divided and asymmetrical; bridge relatively broad; left lophus long, finger-shaped; right lophus small, rounded; ancorae absent; oval sclerites attached to epiphallus (Fig. 19); apodemes broad; valves of cingulum short; basal valves of penis connected with apical ones by a sinuate, long, unbroken flexure (Fig. 21).</p> <p>Species included. The genus consists of four species distributed in Malaysia (Peninsular Malaysia and Borneo Island).</p></div> 	https://treatment.plazi.org/id/039B87C7FF86525CC6A9FCB7FF583998	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Storozhenko, Sergey Yu.	Storozhenko, Sergey Yu. (2021): On the exact position of the tribe Incolacridini in the modern classification of grasshoppers (Orthoptera: Acrididae). Zootaxa 4970 (1): 106-118, DOI: https://doi.org/10.11646/zootaxa.4970.1.3
039B87C7FF86525CC6A9F9FAFE473B1D.text	039B87C7FF86525CC6A9F9FAFE473B1D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bettotania cinctifemur (Miller 1935)	<div><p>Bettotania cinctifemur (Miller, 1935)</p> <p>Fig. 1</p> <p>Paracelebesia cinctifemur Miller, 1935b: 710, fig. 1 (holotype —female, Malaysia, Borneo, Mt. Kinabalu, Kabayau; in BMNH London).</p> <p>Bettotania cinctifemur: C. Willemse, 1956: 114; Ingrisch, 1989: 228; Otte, 1995: 277.</p> <p>Distribution. Malaysia (Borneo Island).</p> <p>Remarks. Male unknown.</p></div> 	https://treatment.plazi.org/id/039B87C7FF86525CC6A9F9FAFE473B1D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Storozhenko, Sergey Yu.	Storozhenko, Sergey Yu. (2021): On the exact position of the tribe Incolacridini in the modern classification of grasshoppers (Orthoptera: Acrididae). Zootaxa 4970 (1): 106-118, DOI: https://doi.org/10.11646/zootaxa.4970.1.3
039B87C7FF86525CC6A9FF4BFDBF3DD0.text	039B87C7FF86525CC6A9FF4BFDBF3DD0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stolzia nigromaculata (C. Willemse 1938)	<div><p>Stolzia nigromaculata (C. Willemse, 1938)</p> <p>Butonacris nigromaculata C. Willemse, 1938: 120, figs. 4, 5 (holotype —male, Malaysia, Borneo, Sarawak, Kuching; in UMO Oxford); C. Willemse, 1956: 191, fig. 105.</p> <p>Stolzia nigromaculata: Hollis, 1975: 216; Otte, 1995: 339; Yin et al., 1996: 679.</p> <p>Distribution. Malaysia (Borneo Island).</p> <p>Remarks. Female unknown.</p></div> 	https://treatment.plazi.org/id/039B87C7FF86525CC6A9FF4BFDBF3DD0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Storozhenko, Sergey Yu.	Storozhenko, Sergey Yu. (2021): On the exact position of the tribe Incolacridini in the modern classification of grasshoppers (Orthoptera: Acrididae). Zootaxa 4970 (1): 106-118, DOI: https://doi.org/10.11646/zootaxa.4970.1.3
039B87C7FF86525CC6A9FE33FD093F54.text	039B87C7FF86525CC6A9FE33FD093F54.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stolzia rubromaculata C. Willemse 1930	<div><p>Stolzia rubromaculata C. Willemse, 1930</p> <p>Fig. 2</p> <p>Stolzia rubromaculata C. Willemse, 1930: 118, 209, fig. 58 (holotype —male, Indonesia, Sumatra, Solok; in NHME Maastricht); Ramme, 1941: 100 (description of female); C. Willemse, 1956: 117; Hollis, 1975: 216; Otte, 1995: 339; Yin et al., 1996: 679.</p> <p>Distribution. Indonesia (Sumatra Island).</p> <p>Remarks. Both male and female are known.</p></div> 	https://treatment.plazi.org/id/039B87C7FF86525CC6A9FE33FD093F54	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Storozhenko, Sergey Yu.	Storozhenko, Sergey Yu. (2021): On the exact position of the tribe Incolacridini in the modern classification of grasshoppers (Orthoptera: Acrididae). Zootaxa 4970 (1): 106-118, DOI: https://doi.org/10.11646/zootaxa.4970.1.3
039B87C7FF84525EC6A9FB59FB273AFB.text	039B87C7FF84525EC6A9FB59FB273AFB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Asymmetritania Storozhenko 2021	<div><p>Genus Asymmetritania Storozhenko, gen. nov.</p> <p>Type species: Stolzia vietnamensis Storozhenko, 2020, here designated.</p> <p>Description. Head in frontal view oval; frontal ridge narrow, almost reaching the clypeus (Fig. 27). Face in lateral view reclinate; frontal ridge projecting between the antennae but forming a relatively short rostrum (Figs. 25, 26). Ventral genicular lobes of the hind femora narrowly triangular with apex obtuse, without spine (Fig. 14). Male 10th abdominal tergite with furculae. Male supra-anal plate long, with a median basal impression, lateral margins with a pointed tooth in apical third, apex rounded (Figs. 13, 30). Male cerci compressed, short, conical, with or without a small preapical tooth. Valves of ovipositor short, with apex rounded and strongly dentate (Fig. 29). Male genitalia: epiphallus strongly asymmetrical; bridge narrow; left lophus large, extending; right lophus small; ancorae absent; oval sclerites separated from epiphallus (Figs. 22, 32); apodemes long and narrow; valves of cingulum and apical valves of penis long, completely covered by ectophallic sheath of penis; basal valves of penis connected with apical ones by a short sclerotized or membranous flexure (Figs. 21, 33).</p> <p>Composition. The genus consists of two species from continental part of South-East Asia.</p> <p>Etymology. From Latin asymmetrica and the genus name Bettotania. Genus feminine.</p> </div>	https://treatment.plazi.org/id/039B87C7FF84525EC6A9FB59FB273AFB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Storozhenko, Sergey Yu.	Storozhenko, Sergey Yu. (2021): On the exact position of the tribe Incolacridini in the modern classification of grasshoppers (Orthoptera: Acrididae). Zootaxa 4970 (1): 106-118, DOI: https://doi.org/10.11646/zootaxa.4970.1.3
039B87C7FF84525EC6A9FF4BFE473DD0.text	039B87C7FF84525EC6A9FF4BFE473DD0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bettotania festiva (Miller 1935)	<div><p>Bettotania festiva (Miller, 1935)</p> <p>Paracelebesia festiva Miller, 1935a: 698, fig. 6, Pl. XIV, fig. 7 (holotype —female, Peninsular Malaysia, Sungai Lui, Ulu Langat sub district, Selangor; in BMNH London).</p> <p>Bettotania festiva: C. Willemse, 1956: 115; Ingrisch, 1989: 228; Otte, 1995: 277.</p> <p>Distribution. Peninsular Malaysia.</p> <p>Remarks. Male unknown.</p></div> 	https://treatment.plazi.org/id/039B87C7FF84525EC6A9FF4BFE473DD0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Storozhenko, Sergey Yu.	Storozhenko, Sergey Yu. (2021): On the exact position of the tribe Incolacridini in the modern classification of grasshoppers (Orthoptera: Acrididae). Zootaxa 4970 (1): 106-118, DOI: https://doi.org/10.11646/zootaxa.4970.1.3
039B87C7FF84525EC6A9FE33FE473E9B.text	039B87C7FF84525EC6A9FE33FE473E9B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bettotania flavostriata F. Willemse 1963	<div><p>Bettotania flavostriata F. Willemse, 1963</p> <p>Bettotania flavostriata F. Willemse, 1963: 171 (holotype —female, Malaysia, East Borneo, “Tabang, Bengen River”; in NHME Maastricht); Ingrisch, 1989: 228; Otte, 1995: 277; Yin et al., 1996: 96.</p> <p>Distribution. Malaysia (Borneo Island).</p> <p>Remarks. Male unknown.</p></div> 	https://treatment.plazi.org/id/039B87C7FF84525EC6A9FE33FE473E9B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Storozhenko, Sergey Yu.	Storozhenko, Sergey Yu. (2021): On the exact position of the tribe Incolacridini in the modern classification of grasshoppers (Orthoptera: Acrididae). Zootaxa 4970 (1): 106-118, DOI: https://doi.org/10.11646/zootaxa.4970.1.3
039B87C7FF84525EC6A9FCF4FDF43879.text	039B87C7FF84525EC6A9FCF4FDF43879.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bettotania maculata C. Willemse 1933	<div><p>Bettotania maculata C. Willemse, 1933</p> <p>Figs. 19–21</p> <p>Bettotania maculata C. Willemse, 1933: 75, fig. 4 (holotype —male, Malaysia, Borneo, Sabah, Sandakan; in BMNH London); C. Willemse, 1956: 114, fig. 81; Ingrisch, 1989: 228, figs 96–99; Otte, 1995: 277; Yin et al., 1996: 96.</p> <p>Distribution. Malaysia (Borneo Island).</p> <p>Remarks. Both sexes are known.</p></div> 	https://treatment.plazi.org/id/039B87C7FF84525EC6A9FCF4FDF43879	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Storozhenko, Sergey Yu.	Storozhenko, Sergey Yu. (2021): On the exact position of the tribe Incolacridini in the modern classification of grasshoppers (Orthoptera: Acrididae). Zootaxa 4970 (1): 106-118, DOI: https://doi.org/10.11646/zootaxa.4970.1.3
039B87C7FF845251C6A9F8D4FC0B3D51.text	039B87C7FF845251C6A9F8D4FC0B3D51.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Asymmetritania asymmetrica (Ingrisch 1989) Storozhenko 2021	<div><p>Asymmetritania asymmetrica (Ingrisch, 1989), comb. nov.</p> <p>Figs. 13, 22–24</p> <p>Bettotania asymmetrica Ingrisch, 1989: 229, figs. 100–107 (holotype —male, Thailand, Chanthaburi, Khao Soi Dao; in ZSM Munich); Otte, 1995: 277; Yin et al., 1996: 96.</p> <p>Distribution. Thailand.</p> <p>Remarks. Female unknown. Herein this species is transferred from Bettotania to the new genus. The record of this species from Vietnam (Storozhenko, 1998) is result of misidentification (see below), wherefore A. asymmetrica must be excluded from the list of the Vietnamese Orthoptera (Kim &amp; Pham, 2014). The photos of holotype male are available in the Orthoptera Species File database (Cigliano et al., 2021).</p> </div>	https://treatment.plazi.org/id/039B87C7FF845251C6A9F8D4FC0B3D51	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Storozhenko, Sergey Yu.	Storozhenko, Sergey Yu. (2021): On the exact position of the tribe Incolacridini in the modern classification of grasshoppers (Orthoptera: Acrididae). Zootaxa 4970 (1): 106-118, DOI: https://doi.org/10.11646/zootaxa.4970.1.3
039B87C7FF8A5250C6A9FF4BFBB438B3.text	039B87C7FF8A5250C6A9FF4BFBB438B3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Asymmetritania vietnamensis (Storozhenko 2020) Storozhenko 2021	<div><p>Asymmetritania vietnamensis (Storozhenko, 2020), comb. nov.</p> <p>Figs. 14, 25–33</p> <p>Stolzia vietnamensis Storozhenko, 2020a: 16, figs. 1–5 (holotype —female, Vietnam, Dong Nai Province, Cat Tien National Park; in ZIN St. Petersburg).</p> <p>Bettotania asymmetrica: Storozhenko, 1998: 37 (misidentification).</p> <p>Non-type material examined. Southern Vietnam, Dong Nai province, Nam Cat Tien sector of the Cat Tien National Park (approximately 150 km north of Ho Chi Minh City), 8-15 December 1990, 1 male, leg. V. V. Badikov.</p> <p>Description. Male (previously unknown). Similar to female (Fig. 25) but smaller and brightly colored (Figs. 26–28). Head and pronotum as in female but upper half of the lateral lobes of pronotum completely black. Antennae 19-segmented. Pronotum rugose; prozona 1.3 times as long as metazona. Tegmina touching each other in rest position, reaching knees of hind femora (Fig. 26); median area widening apically, with numerous transverse veinlets forming intercalar vein in basal part of the area (Fig. 27). Hind femora stout, 3.1 times as long as their maximal width, olive green with yellow pregenicular ring; hind tibia with 8 outer and 9 inner dorsal spines, blackish olive green with black base and greenish ring below. Tenth abdominal tergite continuous, with pointed furculae. Supraanal plate long, with a median basal impression, lateral margins with a pointed tooth in apical third, apex rounded. Cerci compressed, conical, with pointed apex, preapical tooth absent. Apical abdominal sternites with tufts of setae on both sides of the middle. Epiphallus strongly asymmetrical; left lophus large, in lateral view lingua-like; right lophus enlarged, broadly rounded; ancorae absent. Phallus rather long and narrow (Fig. 32). Cingulum consisting of apodemes, zygoma, and rami. Apodemes completely sclerotized; ventral side of zygoma connected with arch of cingulum. Valves of cingulum completely covered by ectophallic sheath of penis. Basal valves of penis connected with apical ones by a short sclerotized flexure (Fig. 33).</p> <p>Measurements. Male (length in mm): body—14.8; antenna—9.5; pronotum—4.2; tegmen—9.5; hind femur— 8.5; hind tibia—7.0.</p> <p>Distribution. Vietnam.</p> <p>Remarks. The above described male from Vietnam was wrongly identified as Bettotania asymmetrica by me (Storozhenko, 1998). This male was collected in the Cat Tien National Park—a type locality of previously known by a single female Stolzia vietnamensis. Therefore I believe reasonably that both Vietnamese specimens are conspecific. The male of Asymmetritania vietnamensis differs from A. asymmetrica in the shape of cerci and epiphallus, as well as in the flexure connected basal and apical valves of penis (in A. asymmetrica, the cerci with a small preapical tooth, the right lophus of epiphallus very small, and the flexure membranous).</p> </div>	https://treatment.plazi.org/id/039B87C7FF8A5250C6A9FF4BFBB438B3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Storozhenko, Sergey Yu.	Storozhenko, Sergey Yu. (2021): On the exact position of the tribe Incolacridini in the modern classification of grasshoppers (Orthoptera: Acrididae). Zootaxa 4970 (1): 106-118, DOI: https://doi.org/10.11646/zootaxa.4970.1.3
