identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
8545EB45EC0AA01AFF3F886EFEA88330.text	8545EB45EC0AA01AFF3F886EFEA88330.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gahrliepia mirabilis Schluger, Grochovskaja, Ngu, Hoe and Tung 1960	<div><p>Gahrliepia mirabilis Schluger, Grochovskaja, Ngu, Hoe and Tung, 1960</p> <p>(Figs. 1A, B, 2–4)</p> <p>Gahrliepia mirabilis Schluger, Grochovskaja, Ngu, Hoe and Tung, 1960c: 472, figs. 54–60.</p> <p>Gahrliepia (Gateria) mirabilis: Vercammen-Grandjean 1968: 116.</p> <p>Gahrliepia (Gahrliepia) mirabilis: Lakshana 1973: 23; Kudryashova 2004: 46; Chau et al. 2007: 157, fig. 75.</p> <p>Diagnosis. SIF = 4B-N-3-2110.0000; fsp = 7.6.6; fCx = 1.1.1; fSt = 2.2; fPp = B/N/NNN; fSc: PL&gt; AL; Ip = 653–766; fD = 2H-4-7-8-6-4-4-3; DS = 28–36; V = 44–50; NDV = 74–83. Scutal puncta of two types: sparse large and dense small. Two pairs of usurped setae on scutum. Standard measurements of type series given in Table 2.</p> <p>Description (larva) [based on lectotype and 5 paralectotypes]. IDIOSOMA (Figs. 1A, B, 2). Eyes 2 + 2, posterior pair very small, almost reduced; 28–36 barbed dorsal idiosomal setae including one pair of humeral setae; 4 sternal setae; 44–50 ventral setae; NDV = 74–83.</p> <p>GNATHOSOMA (Fig. 3A, B). Cheliceral blade with tricuspid cap; cheliceral base with dense puncta; gnathobase with dense puncta; palpal femur with sparse puncta; gnathobase with 1 pair of branched setae; galeala nude; palpal claw with 3 prongs; seta on palpal femur branched; setae on palpal genu and tibia nude; palpal tarsus with 4 branched setae and tarsala.</p> <p>SCUTUM (Fig. 1A). Hexagonal, longer than wide, with 1 pair of ALs, 1 pair of PLs, and 2 pairs of usurped setae (dorsal idiosomal setae situated on the scutum as a result of its expansion in the posterior direction), widest between PLs and anterior pair of usurped setae, posterior scutal margin almost straight; anterior usurped setae situated far from lateral scutal margins, posterior usurped setae situated in posterior scutal angles, distance between anterior usurped setae about same as distance between posterior usurped setae; PLs slightly longer than ALs; ALs and PLs thicker than usurped setae; sensilla fusiform, covered with spikes; sensillum bases situated closer to level of ALs than to level of PLs; scutal puncta of two types: sparse large and dense small.</p> <p>LEGS (Fig. 3C–E). All with 1 pair of claws and claw-like empodium. Leg I: 7-segmented, coxa with 1 branched seta (1B); trochanter 1B; basifemur 1B; telofemur 5B; genu 4B, 2 genualae, microgenuala; tibia 8B, 2 tibialae, microtibiala; tarsus 21B, tarsala, microtarsala proximal to tarsala, subterminala, parasubterminala, pretarsala. Leg II: 6-segmented, coxa 1B; trochanter 1B; femur 6B; genu 3B, genuala; tibia 6B, 2 tibialae; tarsus 16B, tarsala, microtarsala near and proximal to tarsala, pretarsala. Leg III: 6-segmented, coxa 1B; trochanter 1B; femur 4B, nude ventral femorala; genu 3B, genuala; tibia 6B; tarsus 15B.</p> <p>Hosts. Rhizomys pruinosus Blyth, 1851 (Rodentia: Spalacidae), Leopoldamys edwardsi (Thomas, 1882) (= Rattus edwardsi camphaensis D. Tien (nomen nudum)), Rattus tanezumi Temminck, 1844 (= R. rattus khyensis) (Rodentia: Muridae), Tupaia sp. (Scandentia: Tupaiidae) (Schluger et al. 1960c), Tupaia glis (Diard and Duvaucel, 1820) (Scandentia: Tupaiidae), Hylomys suillus S. Müller,1840 (Erinaceomorpha: Erinaceidae), Crocidura horsfieldii (Tomes, 1856) (Soricomorpha, Soricidae), Bandicota indica (Bechstein, 1800), Berylmys berdmorei (Blyth, 1851), Maxomys surifer (Miller, 1900), Rattus rattus (L., 1758) (Rodentia: Muridae), Menetes berdmorei (Blyth, 1849) (Rodentia: Sciuridae), Cannomys badius (Hodgson, 1841) (Rodentia: Spalacidae) (Lakshana 1973), Suncus murinus (L., 1766), (Soricomorpha, Soricidae), Paradoxurus hermaphroditus (Pallas, 1777) (Carnivora: Viverridae), Herpestes urva (Hodgson, 1836) (Carnivora: Herpestidae), Bandicota bengalensis (Gray, 1835), Berylmys bowersi (Anderson, 1879), Niviventer fulvescens (Gray, 1847) (= Rattus fulvescens), N. niviventer (Hodgson, 1836) (= Rattus niviventer), Rattus andamanensis (Blyth, 1860) (= R. koratensis Kloss, 1919), R. norvegicus (Berkenhout, 1769), R. tanezumi Temminck, 1844 (= R. flavipectus (Milne-Edwards, 1872)) (Rodentia: Muridae) (Chau et al. 2007).</p> <p>Distribution. Thailand (Lakshana 1973), Vietnam.</p> <p>Type material examined (Fig. 4). Lectotype (here designated): larva ZMMU Tdt-3223 (specimen 1) ex R. tanezumi No 42018, Vietnam, Quang Tri Province, Vinh Linh, 18 August 1956, coll. I.M. Grochovskaja. Paralectotypes: two larvae, ZMMU Tdt-3224 (specimen 1) and ZMMU Tdt-3225 (specimen 4), same collection data as for lectotype; one larva ZMMU Tdt-3226 (specimen 4) ex unknown host, Vietnam, Nghe An Province, Phu Quy, 9 September 1956, coll. I.M. Grochovskaja; one larva ZMMU Tdt-3227 (specimen 1) ex L. edwardsi (labeled as Rattus edwardsi camphaensis), Vietnam, Quang Ninh Province, Ha Lam, 1 February 1956, coll. I.M. Grochovskaja; one larva ZMMU Tdt-3228 (specimen 3) ex “tupaia”, Vietnam, Quang Ninh Province, Cam Pha, 2 March 1956, coll. I.M. Grochovskaja.</p> <p>Remarks. According to the original description, in this species “all scutal surface with small puncta” (Schluger et al. 1960c); thus, Schluger did not recognize two types of scutal puncta in G. mirabilis, obviously due to insufficiently precise optics. However, she described such punctation in G. tenella Traub and Morrow, 1955, where this trait is more distinct (Schluger et al. 1960c, Fig. 63).</p> <p>Gahrliepia mirabilis is similar to G. yangchenensis Chen and Hsu, 1957 and differs from the latter by the presence of nude ventral palpal tibial seta vs. having slender branches; the presence of two types of scutal puncta vs. one type (however, the original description of G. yangchenensis could be inexact in this point); and by a slightly larger scutum (AW = 49–56 vs. 42–51, PW = 70–86 vs. 67–79, ASB = 21–30 vs. 19, and PSB = 112–126 vs. 109) (Chen &amp; Hsu 1957; Li et al. 1997). Gahrliepia mirabilis is also similar to G. eurypunctata Jeu, Yu and Wan, 1983 and differs from it by the presence of nude palpal genual and ventral palpal tibial setae (fPp = B/N/NNN vs. B/B/ NNB), slightly smaller scutum (PW = 70–86 vs. 86–90, PSB = 112–126 vs. 136–141, SD = 134–155 vs. 161–170, AP = 40–45 vs. 46–51), and much shorter scutal setae (AL = 28–36 vs. 45–53 and PL = 32–38 vs. 52–54) (Jeu et al. 1983).</p> </div>	http://treatment.plazi.org/id/8545EB45EC0AA01AFF3F886EFEA88330	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Antonovskaia, Anastasia A.;Stekolnikov, Alexandr A.	Antonovskaia, Anastasia A., Stekolnikov, Alexandr A. (2021): Redescriptions of ten chigger mite species (Acariformes: Trombiculidae) from Vietnam. Zootaxa 4969 (1): 1-53, DOI: https://doi.org/10.11646/zootaxa.4969.1.1
8545EB45EC04A01EFF3F8FBFFAF083D0.text	8545EB45EC04A01EFF3F8FBFFAF083D0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Walchia delicatula (Schluger, Grochovskaja, Ngu, Hoe and Tung 1960)	<div><p>Walchia delicatula (Schluger, Grochovskaja, Ngu, Hoe and Tung, 1960)</p> <p>(Figs. 1C, D, 5–7)</p> <p>Gahrliepia (Walchia) delicatula Schluger, Grochovskaja, Ngu, Hoe and Tung, 1960c: 470, figs. 46–53.</p> <p>Gahrliepia (Walchia) delicatula: Chau et al. 2007: 167, fig. 82.</p> <p>Walchia (Walchia) delicatula: Vercammen-Grandjean 1968: 111.</p> <p>Walchia delicatula: Kudryashova 2004: 45.</p> <p>Walchia tianguangshanensis Zhao, 1981: 324, figs. 1–5, syn. nov.</p> <p>Diagnosis. SIF = 4B-N-3-2110.0000; fsp = 7.6.6; fCx = 1.1.1; fSt = 2.2; fPp = N/N/NNN; fSc: AL&gt; PL; Ip = 552–639; fD = 2H-6-6-6-6-4-3-2; DS = 31–38; V = 38–49; NDV = 69–87. Eyes 2 + 2. Standard measurements of type series given in Table 3.</p> <p>Description (larva) [based on lectotype and 11 paralectotypes]. IDIOSOMA (Figs. 1C, D, 5). Eyes 2 + 2, posterior pair smaller; 31–38 barbed dorsal idiosomal setae including one pair of humeral setae; 4 sternal setae; 38–49 ventral setae; NDV = 69–87.</p> <p>GNATHOSOMA (Fig. 6A, B). Cheliceral blade with tricuspid cap; cheliceral base moderately covered with puncta; gnathobase with dense puncta and 1 pair of branched setae; galeala nude; palpal claw with 3 prongs; setae on palpal femur, genu, and tibia nude; palpal tarsus with 4 branched setae and tarsala.</p> <p>SCUTUM (Fig. 1C). Pentagonal, longer than width, with pointed posterior angle, densely covered with small puncta, with 1 pair of ALs and 1 pair of PLs; sensilla fusiform, covered with spikes; sensillum bases situated at equal distances from levels of ALs and PLs.</p> <p>LEGS (Fig. 6C–E). All with 1 pair of claws and claw-like empodium. Leg I: 7-segmented, coxa with 1 branched seta (1B); trochanter 1B; basifemur 1B; telofemur 5B; genu 4B, 2 genualae, microgenuala; tibia 8B, 2 tibialae, microtibiala; tarsus 22B, tarsala, microtarsala proximal to tarsala, subterminala, parasubterminala, pretarsala. Leg II: 6-segmented, coxa 1B; trochanter 1B; femur 6B; genu 3B, genuala; tibia 6B, 2 tibialae; tarsus 16B, tarsala, microtarsala near and proximal to tarsala, pretarsala. Leg III: 6-segmented, coxa 1B; trochanter 1B; femur 4B, nude ventral femorala; genu 3B, genuala; tibia 6B; tarsus 15B.</p> <p>Hosts. Tupaia hongaensis (nomen nudum) (Scandentia: Tupaiidae), Rattus norvegicus, Rhizomys pruinosus (Schluger et al. 1960c), Rattus tanezumi (= R. flavipectus) (Chau et al. 2007), Berylmys bowersi (this study).</p> <p>Distribution. China (this study), Vietnam.</p> <p>Type material examined (Fig. 7). Lectotype (here designated): larva ZMMU Tdt-3240 (specimen 3) ex R. pruinosus No 208, Vietnam, Ha Giang Province, Ha Giang, 19 May 1956, coll. I.M. Grochovskaja. Paralectotypes: seven larvae, ZMMU Tdt-3237 (specimen 3), ZMMU Tdt-3238 (specimens 2, 3), ZMMU Tdt-3239 (specimen 2), ZMMU Tdt-3240 (specimen 5), ZMMU Tdt-3241 (specimen 3), ZMMU Tdt-3242 (specimen 3), same collection data as for lectotype; five larvae, ZMMU Tdt-3234 (specimens 1, 2, 4), ZMMU Tdt-3236 (specimens 2, 5), ex R. pruinosus No 258, Vietnam, Ha Giang Province, Ha Giang, 3 June 1956, coll. I.M. Grochovskaja.</p> <p>Remarks. According to the original description, W. delicatula is similar to W. neosinensis (Hsu and Wen, 1956), but differs in the “shape of trichobothria (sensilla), position of seta on leg coxa III, nude outer lateral seta on palpal tibia, wider scutum, and other characters” (Schluger et al. 1960c). The “outer lateral seta on palpal tibia” is actually ventral palpal tibial seta. The difference in its shape (fPp = N/N/NNN in W. delicatula and fPp = N/N/NNB in W. neosinensis) is the most clear difference between these two species. We did not find the difference between these species in the position of coxal seta III. The difference in the shape of sensilla (narrower in W. delicatula than in W. neosinensis) seems reliable, although rather slight. According to our data, the metric differences between these species are as follows: AW = 41–49, PW = 52–69, SB = 34–45, ASB = 22–31, SD = 83–99, and AP = 42–52 in the type series of W. delicatula vs. 33, 41, 29, 20, 82, and 38 in W. neosinensis, respectively (Hsu &amp; Wen 1956; Li et al. 1997).</p> <p>The species W. tianguangshanensis Zhao, 1981, which was described from a single specimen ex B. bowersi from Guangdong Province of China, does not differ from W. delicatula, neither by morphology, nor by measurements (Table 2), and therefore is synonymized here with the latter.</p> <p>Specimens Nos 1 and 2 on the slide ZMMU Tdt-3237 belong to Walchia pacifica (Chen and Hsu, 1955).</p> </div>	http://treatment.plazi.org/id/8545EB45EC04A01EFF3F8FBFFAF083D0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Antonovskaia, Anastasia A.;Stekolnikov, Alexandr A.	Antonovskaia, Anastasia A., Stekolnikov, Alexandr A. (2021): Redescriptions of ten chigger mite species (Acariformes: Trombiculidae) from Vietnam. Zootaxa 4969 (1): 1-53, DOI: https://doi.org/10.11646/zootaxa.4969.1.1
8545EB45EC00A001FF3F8FF7FAA58082.text	8545EB45EC00A001FF3F8FF7FAA58082.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Walchia dismina (Schluger, Grochovskaja, Ngu, Hoe and Tung 1960)	<div><p>Walchia dismina (Schluger, Grochovskaja, Ngu, Hoe and Tung, 1960)</p> <p>(Figs. 8A, B, 9–11)</p> <p>Gahrliepia (Walchia) dismina Schluger, Grochovskaja, Ngu, Hoe and Tung, 1960c: 468, figs. 31–38.</p> <p>Gahrliepia (Walchia) dismina: Lakshana 1973: 24; Chau et al. 2007: 168, fig. 83.</p> <p>Walchia (Walchia) dismina: Vercammen-Grandjean 1968: 112.</p> <p>Walchia dismina: Kudryashova 2004: 45; Chaisiri et al. 2016: 335.</p> <p>Diagnosis. SIF = 4B-N-2-2110.0000; fsp = 7.6.6; fCx = 1.1.2; fSt = 2.2; fPp = N/N/NNN; fSc: PL&gt; AL; Ip = 688– 739; fD = 2H-6-[6-2]-6-[4-2]-4-5; DS = 32–39; V = 44–51; NDV = 77–90. Eyes absent. Standard measurements of type series given in Table 4.</p> <p>Description (larva) [based on lectotype and 6 paralectotypes]. IDIOSOMA (Figs. 8A, B, 9). Eyes absent; 32–39 dorsal idiosomal setae covered with long barbs, including one pair of humeral setae; 4 sternal setae; 44–51 ventral setae; NDV = 77–90.</p> <p>GNATHOSOMA (Fig. 10D, E). Cheliceral blade with tricuspid cap; cheliceral base with sparse puncta; gnathobase with sparse puncta and 1 pair of branched setae; galeala nude; palpal claw with 2 prongs; setae on palpal femur, genu, and tibia nude; palpal tarsus with 4 branched setae and tarsala.</p> <p>SCUTUM (Fig. 8A). Pentagonal, longer than width, with angulate posterior margin, moderately covered with small puncta, with 1 pair of ALs and 1 pair of PLs; PLs longer ALs; sensilla claviform, covered with spikes; sensillum bases situated at equal distances from levels of ALs and PLs.</p> <p>LEGS (Fig. 10A–C). All with 1 pair of claws and claw-like empodium. Leg I: 7-segmented, coxa with 1 branched seta (1B); trochanter 1B; basifemur 1B; telofemur 5B; genu 4B, 2 genualae, microgenuala; tibia 8B, 2 tibialae, microtibiala; tarsus 22B, tarsala, microtarsala proximal to tarsala, subterminala, parasubterminala, pretarsala. Leg II: 6-segmented, coxa 1B; trochanter 1B; femur 6B; genu 3B, genuala; tibia 6B, 2 tibialae; tarsus 16B, tarsala, microtarsala far proximal to tarsala, pretarsala. Leg III: 6-segmented, coxa 2B; trochanter 1B; femur 4B, nude ventral femorala; genu 3B, genuala; tibia 6B; tarsus 14B.</p> <p>Hosts. Rattus sp. (Schluger et al. 1960c), Leopoldamys sabanus (Thomas, 1887) (= Rattus sabanus), Mus cervicolor Hodgson, 1845 (Rodentia: Muridae), Rattus tanezumi (= R. sladeni (Anderson, 1879)), Bandicota indica, Maxomys surifer, Menetes berdmorei, Niviventer niviventer, Rattus rattus, Tupaia glis (Lakshana 1973), Mus cookii Ryley, 1914, Rattus losea (Swinhoe, 1871) (Rodentia: Muridae) (Chaisiri et al. 2016).</p> <p>Distribution. Thailand (Lakshana 1973; Chaisiri et al. 2016), Vietnam.</p> <p>Type material examined (Fig. 11). Lectotype (here designated): larva ZMMU Tdt-3252 (specimen 1) ex Rattus sp. No 542, Vietnam, Nghe An Province, Phu Quy, 8 September 1956, coll. I.M. Grochovskaja. Paralectotypes: two larvae, ZMMU Tdt-3252 (specimens 2, 3), same collection data as for lectotype; two larvae, ZMMU Tdt-3254 (specimen 1), ZMMU Tdt-3255 (specimen 2), ex Rattus sp. No 542, Vietnam, Nghe An Province, Phu Quy, 9 September 1956, coll. I.M. Grochovskaja; two larvae, ZMMU Tdt-3258 (specimens 3, 4), ex Rattus sp. No 595, Vietnam, Nghe An Province, Phu Quy, 13 September 1956, coll. I.M. Grochovskaja.</p> <p>Remarks. According to the original description, W. dismina is similar to W. kritochaeta Traub and Evans, 1957 and differs from it by longer legs and by the presence of two setae on leg coxa III (fCx = 1.1.2 vs. 1.1.1) (Schluger et al. 1960c). The latter difference is reliable, but the former one is unclear, as the lengths of legs were absent in the original description of W. kritochaeta (Traub &amp; Evans 1957). However, we found the following additional differences between these species. Walchia dismina has 2-pronged palpal claw, equal leg claws, and has no eyes, while W. kritochaeta has 3-pronged palpal claw, unequal leg claws (one claw is very thin, like the empodium), and one pair of eyes.</p> <p>Walchia dismina is also similar in the size and shape of scutum to W. isonychia Nadchatram and Traub, 1964, but differs in the presence of two setae on leg coxa III (vs. one seta), a 2-pronged palpal claw (vs. 3-pronged), and by the absence of eyes (vs. eyes 1 + 1).</p> <p>Specimen No. 4 on the slide ZMMU Tdt-3254, specimens Nos 1, 3, 4, and 5 on the slide ZMMU Tdt-3255, specimens Nos 2 and 5 on the slide ZMMU Tdt-3258 belong to Walchia chinensis (Chen and Hsu, 1955). Specimen No. 2 on the slide ZMMU Tdt-3254 presumably belongs to Walchia disparunguis (Oudemans, 1929).</p> </div>	http://treatment.plazi.org/id/8545EB45EC00A001FF3F8FF7FAA58082	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Antonovskaia, Anastasia A.;Stekolnikov, Alexandr A.	Antonovskaia, Anastasia A., Stekolnikov, Alexandr A. (2021): Redescriptions of ten chigger mite species (Acariformes: Trombiculidae) from Vietnam. Zootaxa 4969 (1): 1-53, DOI: https://doi.org/10.11646/zootaxa.4969.1.1
8545EB45EC1CA005FF3F8A94FD41801E.text	8545EB45EC1CA005FF3F8A94FD41801E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cheladonta neda Schluger, Grochovskaja, Ngu, Hoe and Tung 1960	<div><p>Cheladonta neda Schluger, Grochovskaja, Ngu, Hoe and Tung, 1960</p> <p>(Figs. 8C, D, 12–14)</p> <p>Cheladonta neda Schluger, Grochovskaja, Ngu, Hoe and Tung, 1960a: 190, fig. 11.</p> <p>Schoutedenichia (Schoutedenichia) neda: Vercammen-Grandjean 1965b: 101; 1968: 96.</p> <p>Cheladonta (Cheladonta) neda: Lakshana 1973: 21; Chau et al. 2007: 151, fig. 73.</p> <p>Diagnosis. SIF = 4B-N-3-2110.0000; fsp = 7.7.7; fCx = 1.1.1; fSt = 2.2; fPp = B/b/BbB; fSc: PL ≥ AM&gt; AL; Ip = 473–531; fD = 4H-8-4-8-6-2-2; DS = 31–35; V = 27–31; NDV = 61–66. Eyes 2 + 2. Cheliceral blade with dorsal serration. Standard measurements of type series given in Table 5.</p> <p>Description (larva) [based on lectotype, 4 paralectotypes, and 1 additional specimen]. IDIOSOMA (Figs. 8C, D, 12). Eyes 2 + 2; 31–35 barbed dorsal idiosomal setae including two pairs of humeral setae; 4 sternal setae; 27–31 ventral setae; NDV = 61–66.</p> <p>GNATHOSOMA (Fig. 13A, B). Cheliceral blade with tricuspid cap and dorsal serration; cheliceral base with few puncta; gnathobase with sparse puncta and 1 pair of branched setae; galeala nude; palpal claw with 3 prongs; seta on palpal femur branched; seta on palpal genu with 1 branch; dorsal and ventral palpal tibial setae branched, lateral palpal tibial seta with one branch; palpal tarsus with 4 branched setae and tarsala.</p> <p>SCUTUM (Fig. 8C). Nearly rectangular, wider than long, with almost straight posterior margin, moderately covered with small puncta, with 1 AM, 1 pair of ALs and 1 pair of PLs; PLs longer than ALs; sensilla claviform, covered with spikes; sensillum bases situated anterior to level of PLs (PSB – P-PL = 5–9 µm).</p> <p>LEGS (Fig. 13C–E).All 7-segmented, with 1 pair of claws and claw-like empodium. Leg I: coxa with 1 branched seta (1B); trochanter 1B; basifemur 1B; telofemur 5B; genu 4B, 2 genualae, microgenuala; tibia 8B, 2 tibialae, microtibiala; tarsus 22B, tarsala, microtarsala proximal to tarsala, subterminala, parasubterminala, pretarsala. Leg II: coxa 1B; trochanter 1B; basifemur 2B; telofemur 4B; genu 3B, genuala; tibia 6B, 2 tibialae; tarsus 16B, tarsala, microtarsala at level of tarsala, pretarsala. Leg III: coxa 1B; trochanter 1B; basifemur 1B; telofemur 3B; genu 3B, genuala; tibia 6B; tarsus 14B.</p> <p>Hosts. Leopoldamys edwardsi (= Rattus edwardsi camphaensis D. Tien (nomen nudum)) (Schluger et al. 1960a), Menetes berdmorei, Tupaia glis (Lakshana 1973), Maxomys surifer (this study).</p> <p>Distribution. Thailand (Lakshana 1973), Vietnam.</p> <p>Type material examined (Fig. 14). Lectotype (here designated): larva ZMMU Tdt-3283 (specimen 5) ex Leopoldamys edwardsi No 595, Vietnam, Nghe An Province, Phu Quy, 13 September 1956, coll. I.M. Grochovskaja. Paralectotypes: two larvae, ZMMU Tdt-3283 (specimen 2), ZMMU Tdt-3282 (specimen 4), same collection data as for lectotype; one larva ZMMU Tdt-3277 (specimen 3) ex Leopoldamys edwardsi No 27, Vietnam, Quang Ninh Province, Ha Long (= Hon Gai), 1 February 1956, coll. I.M. Grochovskaja; one larva ZMMU Tdt-3284 (specimen 3) ex Leopoldamys edwardsi No 555, Vietnam, Nghe An Province, Phu Quy, 9 September 1956, coll. I.M. Grochovskaja.</p> <p>Additional material examined. One larva ZMMU XM14.63k1 (specimen 11) ex Maxomys surifer No CT-63, Vietnam, Dong Nai Province, Dong Nai <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.38571&amp;materialsCitation.latitude=11.411847" title="Search Plazi for locations around (long 107.38571/lat 11.411847)">National Reserve</a> (Cat Tien), 11°24′42.65″N, 107°23′8.58″E, 11 November 2014, coll. Yu. V. Lopatina.</p> <p>Remarks. The serrated cheliceral blade and shape of scutum (significantly wider than long, with the distance between sensillum bases almost equal to the distance from sensillum base to the lateral scutal margin) confirm that this species belongs to the genus Cheladonta Lipovsky, Crossley and Loomis, 1955, contrary to the opinion of Vercammen-Grandjean (1968).</p> <p>According to the original description, C. neda is similar to C. micheneri Lipovsky, Crossley and Loomis, 1955 and differs in “significantly lesser body size, number of dorsal setae, feathered setae on femur and tibia of palps and other characters” (Schluger et al. 1960a). Actually, C. neda is most similar to C. brevipalpis (André, 1946) described from a gerbil in Tunisia (Vercammen-Grandjean &amp; André 1967) and differs from the latter by the presence of a 3- pronged palpal claw vs. 4-pronged, two pairs of eyes vs. one pair, shorter legs (Ip = 473–531 vs. 577–613), slightly smaller scutum (PW = 49–55 and SD = 29–31 vs. 60–64 and 34–35, respectively), and shorter scutal setae (AL = 10–12 and PL = 21–24 vs. 19 and 26–30, respectively). Cheladonta neda is also similar to C. gouldi Lakshana, 1969 and differs from it by the presence of branched dorsal and lateral palpal tibial setae (fPp = B/b/BbB vs. B/B/ NNB), presence of tibiala III, a lesser number of idiosomal setae (DS = 31–35 and V = 27–31 vs. 40–46 and 40, respectively), narrower scutum (AW = 42–48 and PW = 49–55 vs. 52–55 and 68–72, respectively), shorter setae (AM = 17–23, AL = 10–12, PL = 21–24, and H = 22–27 vs. 25–27, 20–23, 33–36, and 34, respectively), and by shorter legs (Ip = 473–531 vs. 632) (Lakshana 1969).</p> </div>	http://treatment.plazi.org/id/8545EB45EC1CA005FF3F8A94FD41801E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Antonovskaia, Anastasia A.;Stekolnikov, Alexandr A.	Antonovskaia, Anastasia A., Stekolnikov, Alexandr A. (2021): Redescriptions of ten chigger mite species (Acariformes: Trombiculidae) from Vietnam. Zootaxa 4969 (1): 1-53, DOI: https://doi.org/10.11646/zootaxa.4969.1.1
8545EB45EC18A005FF3F8BC7FF458165.text	8545EB45EC18A005FF3F8BC7FF458165.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Doloisia Oudemans 1910	<div><p>Genus Doloisia Oudemans, 1910</p> <p>Remarks. We treate Traubacarus Audy and Nadchatram, 1957 as a subgenus of Doloisia following Stekolnikov (2021).</p> </div>	http://treatment.plazi.org/id/8545EB45EC18A005FF3F8BC7FF458165	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Antonovskaia, Anastasia A.;Stekolnikov, Alexandr A.	Antonovskaia, Anastasia A., Stekolnikov, Alexandr A. (2021): Redescriptions of ten chigger mite species (Acariformes: Trombiculidae) from Vietnam. Zootaxa 4969 (1): 1-53, DOI: https://doi.org/10.11646/zootaxa.4969.1.1
8545EB45EC18A008FF3F8A1FFD5A8555.text	8545EB45EC18A008FF3F8A1FFD5A8555.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Doloisia (Doloisia)	<div><p>Doloisia (Doloisia) alata Schluger, Grochovskaja, Ngu, Hoe and Tung, 1961</p> <p>(Figs. 15A, B, 16–18)</p> <p>Doloisia alata Schluger, Grochovskaja, Ngu, Hoe and Tung, 1961: 448, figs. 1–7.</p> <p>Doloisia alata: Wen 1984: 302; Li et al. 1997: 333, fig. 2-23-2; Kudryashova 2004: 37.</p> <p>Doloisia (Doloisia) alata: Vercammen-Grandjean 1968: 97; Chau et al. 2007: 142, fig. 67.</p> <p>Diagnosis. SIF = 3B1b-N-3-2110.0000; fsp = 7.7.7; fCx = 3.5.10; fSt = 2.2; fPp = B/B/NNB; fSc: PL &gt;&gt; AM&gt; AL; Ip = 688–741; fD = 4H-10-8-6-6-2; DS = 36–40; V = 38–41; NDV = 75–80. Eyes absent. Standard measurements of type series given in Table 6.</p> <p>Description (larva) [based on lectotype and 4 paralectotypes]. IDIOSOMA (Figs. 15A, B, 16). Eyes absent; 36–40 dorsal idiosomal setae covered with long thin barbs, including two pairs of humeral setae; 4 sternal setae; 38–41 ventral setae; NDV = 75–80.</p> <p>GNATHOSOMA (Fig. 17A, B). Cheliceral blade with two large hooks; gnathobase with moderate puncta and 1 pair of branched setae; galeala nude; palpal claw with 3 prongs, internal prong long and strongly curved; setae on palpal femur and genu branched; dorsal and lateral palpal tibial setae nude, ventral palpal tibial seta branched; palpal tarsus with 1 small dorsal seta having 1 branch, 3 heavily branched ventral setae, and tarsala.</p> <p>SCUTUM (Fig. 15A, 17C). Trapezoidal, wider than long, with straight posterior margin, moderately covered with small puncta, with 1 AM covered with short barbs, 1 pair of nude ALs and 1 pair of PLs covered with long thin barbs; AM anterior to level of ALs; PLs much longer AM, ALs shorter AM; sensilla bulbous, covered with spikes; sensillum bases situated at equal distances from levels of ALs and PLs.</p> <p>LEGS (Fig. 17D–F).All 7-segmented, with 1 pair of claws and claw-like empodium. Leg I: coxa with 3 branched setae (3B); trochanter 1B; basifemur 1B; telofemur 5B; genu 4B, 2 genualae, microgenuala; tibia 8B, 2 tibialae, microtibiala; tarsus 21B, tarsala, microtarsala distal to tarsala, subterminala, parasubterminala, pretarsala. Leg II: coxa 5B; trochanter 1B; basifemur 2B; telofemur 4B; genu 3B, genuala; tibia 6B, 2 tibialae; tarsus 18B, tarsala, microtarsala near and proximal to tarsala, pretarsala. Leg III: coxa 10B; trochanter 1B; basifemur 2B; telofemur 3B; genu 3B, genuala; tibia 6B; tarsus 15B.</p> <p>Hosts. Leopoldamys edwardsi [= Rattus grochovskii D. Tien (nomen nudum)] (Schluger et al. 1961), Rattus losea (Wen 1984).</p> <p>Distribution. China (Wen 1984; Li et al. 1997), Vietnam.</p> <p>Type material examined (Fig. 18). Lectotype (here designated): larva ZMMU Tdt-3174 (specimen 1) ex L. edwardsi No 44 [labeled as “ Rattus sp. n. (Rattus grochovskii)”], Vietnam, Quang Ninh Province, Ha Lam District, Ha Long (= Hon Gai), 6 February 1956, coll. I.M. Grochovskaja. Paralectotypes: four larvae, ZMMU Tdt-3157 (specimen 2), ZMMU Tdt-3171 (specimen 1), ZMMU Tdt-3173 (specimen 2), ZMMU Tdt-3176 (specimen 1), same collection data as for lectotype.</p> <p>Remarks. According to the original description, D. alata is similar to D. synoti Oudemans, 1910 and differs from it “by the absence of eyes, greater number of dorsal setae, and by other characters” (Schluger et al. 1961). In our opinion, D. alata is more similar to D. hopuensis Hsu and Chen, 1964 than to D. synoti, and differs from it by a slightly lower number of idiosomal setae (fD = 4H-10-8-6-6-2 vs. 4H-12-10-8-6-4-2, DS = 36–40 vs. 46–48) and by longer legs (Ip = 688–741 vs. 546) (Li et al. 1997).</p> </div>	http://treatment.plazi.org/id/8545EB45EC18A008FF3F8A1FFD5A8555	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Antonovskaia, Anastasia A.;Stekolnikov, Alexandr A.	Antonovskaia, Anastasia A., Stekolnikov, Alexandr A. (2021): Redescriptions of ten chigger mite species (Acariformes: Trombiculidae) from Vietnam. Zootaxa 4969 (1): 1-53, DOI: https://doi.org/10.11646/zootaxa.4969.1.1
8545EB45EC15A00DFF3F8E0EFC4887ED.text	8545EB45EC15A00DFF3F8E0EFC4887ED.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Doloisia (Doloisia) fulminans Schluger, Grochovskaja, Ngu, Hoe and Tung 1961	<div><p>Doloisia (Doloisia) fulminans Schluger, Grochovskaja, Ngu, Hoe and Tung, 1961</p> <p>(Figs. 15C, D, 19–22)</p> <p>Doloisia fulminans Schluger, Grochovskaja, Ngu, Hoe and Tung, 1961: 449, figs. 8–16.</p> <p>Doloisia fulminans: Kudryashova 2004: 39.</p> <p>Doloisia (Doloisia) fulminans: Vercammen-Grandjean 1968: 97; Chau et al. 2007: 147, fig. 71.</p> <p>Diagnosis. SIF = 3B1N-N-3-2110.0000; fsp = 7.7.7; fCx = 3.5.10; fSt = 2.2; fPp = B/B/NNB; fSc: PL &gt;&gt; AM&gt; AL; Ip = 688–763; fD = 4H-7-8-6-6-3; DS = 31–36; V = 31–38; NDV = 67–72. Eyes 1 + 1. Telofemur of leg I with long nude seta. Standard measurements of type series given in Table 7.</p> <p>Description (larva) [based on lectotype and 36 paralectotypes]. IDIOSOMA (Figs. 15C, D, 19). Eyes 1 + 1; 31–36 dorsal idiosomal setae sparsely covered with short barbs, including two pairs of humeral setae; 4 sternal setae; 31–38 ventral setae; NDV = 67–72.</p> <p>GNATHOSOMA (Fig. 20A, B). Cheliceral blade with two large hooks; gnathobase with sparse puncta and 1 pair of branched setae; galeala nude; palpal claw with 3 prongs, internal prong long and strongly curved; setae on palpal femur and genu branched; dorsal and lateral palpal tibial setae nude, ventral palpal tibial seta branched; palpal tarsus with 1 nude dorsal seta, 3 branched ventral setae, and tarsala.</p> <p>SCUTUM (Figs. 15C, 20C). Trapezoidal, wider than long, with almost straight posterior margin, moderately covered with small puncta, with 1 AM, 1 pair of ALs, and 1 pair of PLs; all scutal setae sparsely covered with short barbs; AM at level of ALs; PLs much longer AM, ALs shorter AM; sensilla claviform, bearing few barbs; sensillum bases situated at equal distances from levels of ALs and PLs.</p> <p>LEGS (Fig. 21). All 7-segmented, with 1 pair of claws and claw-like empodium. Leg I: coxa with 3 branched setae (3B); trochanter 1B; basifemur 1B; telofemur 4B and 1 long nude seta; genu 4B, 2 genualae, microgenuala; tibia 8B, 2 tibialae, microtibiala; tarsus 20B, tarsala, microtarsala distal to tarsala, subterminala, parasubterminala, pretarsala. Leg II: coxa 5B; trochanter 1B; basifemur 2B; telofemur 4B; genu 3B, genuala; tibia 6B, 2 tibialae; tarsus 15B, tarsala, microtarsala near and proximal to tarsala, pretarsala. Leg III: coxa 10B; trochanter 1B; basifemur 2B; telofemur 3B; genu 3B, genuala; tibia 6B; tarsus 16B.</p> <p>Hosts. Leopoldamys edwardsi [= Rattus grochovskii D. Tien (nomen nudum)] (Schluger et al. 1961).</p> <p>Distribution. Vietnam.</p> <p>Type material examined (Fig. 22). Lectotype (here designated): larva ZMMU Tdt-3160 (specimen 3) ex L. edwardsi No 44 [labeled as “ Rattus sp. n. (Rattus grochovskii)”], Vietnam, Quang Ninh Province, Ha Lam District, Ha Long (= Hon Gai), 6 February 1956, coll. I.M. Grochovskaja. Paralectotypes: 36 larvae, ZMMU Tdt-3160 (specimen 2), ZMMU Tdt-3145 (specimen 3), ZMMU Tdt-3154 (specimens 1, 2, 3, 4), ZMMU Tdt-3155 (specimen 1), ZMMU Tdt-3156 (specimen 1), ZMMU Tdt-3157 (specimens 1, 3), ZMMU Tdt-3158 (specimens 1, 2, 3), ZMMU Tdt-3159 (specimen 2), ZMMU Tdt-3161 (specimen 4), ZMMU Tdt-3162 (specimen 1), ZMMU Tdt- 3164 (specimen 4), ZMMU Tdt-3166 (specimens 1, 2), ZMMU Tdt-3167 (specimens 2, 3, 4), ZMMU Tdt-3173 (specimens 3, 4), ZMMU Tdt-3174 (specimens 2, 3), ZMMU Tdt-3183 (specimen 4), ZMMU Tdt-3184 (specimens 1, 2), ZMMU Tdt-3186 (specimens 2, 3, 4), ZMMU Tdt-3187 (specimen 1), ZMMU Tdt-3188 (specimen 3), ZMMU Tdt-3190 (specimen 3), ZMMU Tdt-3191 (specimen 2), same collection data as for lectotype.</p> <p>Remarks. According to the original description, D. fulminans is similar to D. alata and differs from it “by the presence of eyes and nude seta on telofemur of leg I and by longer dorsal setae, shape of trichobothria, greater size of body and scutum” (Schluger et al. 1961). We confirm the similarity of these species and their difference by the presence of eyes (1 + 1 in D. fulminans and 0 in D. alata) and by the presence of long nude seta on telofemur of leg I in D. fulminans (absent in D. alata). Sensilla (trichobothria) bear a few barbs in D. fulminans and are densely covered with spikes in D. alata. Morphometric differences between these two species are as follows. Size of scutum: AW = 35–45, PW = 59–71, SB = 28–33, and PSB = 19–24 in D. fulminans vs. 29–35, 51–58, 24–26, and 17–19 in D. alata, respectively; lengths of setae: H = 52–60, D max = 45–60, and V max = 38–50 in D. fulminans vs. 40–42, 39–43, and 31–36 in D. alata, respectively. Doloisia fulminans also has a lower number of idiosomal setae (fD = 4H-6-8-6-6-3 and NDV = 67–72 vs. 4H-10-8-6-6-2 and 75–80 in D. alata).</p> </div>	http://treatment.plazi.org/id/8545EB45EC15A00DFF3F8E0EFC4887ED	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Antonovskaia, Anastasia A.;Stekolnikov, Alexandr A.	Antonovskaia, Anastasia A., Stekolnikov, Alexandr A. (2021): Redescriptions of ten chigger mite species (Acariformes: Trombiculidae) from Vietnam. Zootaxa 4969 (1): 1-53, DOI: https://doi.org/10.11646/zootaxa.4969.1.1
8545EB45EC2EA036FF3F8FF7FC2E84B1.text	8545EB45EC2EA036FF3F8FF7FC2E84B1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Doloisia (Traubacarus) gigantea (Schluger, Grochovskaja, Ngu, Hoe and Tung 1961)	<div><p>Doloisia (Traubacarus) gigantea (Schluger, Grochovskaja, Ngu, Hoe and Tung, 1961)</p> <p>(Figs. 23A, B, 24–27)</p> <p>Traubacarus giganteus Schluger, Grochovskaja, Ngu, Hoe and Tung, 1961: 452, figs. 25–32.</p> <p>Traubacarus giganteus: Kudryashova 2004: 39.</p> <p>Doloisia (Doloisia) gigantea: Vercammen-Grandjean 1968: 97.</p> <p>Doloisia gigantea: Wen 1984: 302.</p> <p>Doloisia (Doloisia) gigantens (sic): Chau et al. 2007: 146, fig. 70.</p> <p>Doloisia outoensis Chen and Hsu, 1965: 286, figs. 8–10.</p> <p>Diagnosis. SIF = 4B-N-3-2110.0000; fsp = 7.7.7; fCx = 1.3.7(6); fSt = 2.2; fPp = B/B/NNN(b); fSc: PL&gt; AM&gt; AL; Ip = 938–972; fD = 4H-[6-2]-8-6-4-3; DS = 32–33; V = 37–38; NDV = 70. Eyes absent. Standard measurements of type series given in Table 8.</p> <p>Description (larva) [based on lectotype and 2 paralectotypes]. IDIOSOMA (Figs. 23A, B, 24). Eyes absent; 32–33 dorsal idiosomal setae covered with few short barbs, including two pairs of humeral setae; 4 sternal setae; 37–38 ventral setae; NDV = 70.</p> <p>GNATHOSOMA (Fig. 25). Cheliceral blade with two large hooks; gnathobase with sparse puncta and 1 pair of branched setae; palpal femur, genu, and tibia with sparse puncta; galeala nude; palpal claw with 3 prongs, internal prong long and strongly curved; setae on palpal femur and genu branched; dorsal and lateral palpal tibial setae nude, ventral palpal tibial seta nude or branched; palpal tarsus with 4 branched setae and tarsala.</p> <p>SCUTUM (Fig. 23A). Bell-shaped, longer than wide, with slightly concave posterior margin, densely covered with small puncta, with 1 AM covered with few inconspicuous cilia and 1 pair of nude ALs; 1 pair of PLs covered with few inconspicuous cilia and situated off scutum; AM anterior to level of ALs; PLs much longer AM, ALs shorter AM; sensilla bulbous, with few short barbs on its stalk and 6 long spikes terminally; sensillum bases situated much closer to ALs than to PLs.</p> <p>LEGS (Fig. 26). All 7-segmented, with 1 pair of claws and claw-like empodium. Leg I: coxa with 1 branched seta (1B); trochanter 1B; basifemur 1B; telofemur 5B; genu 4B, 2 genualae, microgenuala; tibia 8B, 2 tibialae, microtibiala; tarsus 22B, tarsala, microtarsala distal to tarsala, subterminala, parasubterminala, pretarsala. Leg II: coxa 3B; trochanter 1B; basifemur 2B; telofemur 4B; genu 3B, genuala; tibia 6B, 2 tibialae; tarsus 18B, tarsala, microtarsala near and proximal to tarsala, pretarsala. Leg III: coxa 7B or 6B; trochanter 1B; basifemur 2B; telofemur 3B; genu 3B, genuala; tibia 6B; tarsus 18B.</p> <p>Hosts. Leopoldamys edwardsi [= Rattus grochovskii D. Tien (nomen nudum)] (Schluger et al. 1961), Niviventer confucianus (Milne-Edwards, 1871) (Rodentia: Muridae), Berylmys bowersi (= Rattus bowersi latouchei), Rattus rattus (Wen 1984).</p> <p>Distribution. China (Wen 1984), Vietnam.</p> <p>Type material examined (Fig. 27). Lectotype (here designated): larva ZMMU Tdt-3179 (specimen 1) ex L. edwardsi No 44 [labeled as “ Rattus sp. n. (Rattus grochovskii)”], Vietnam, Quang Ninh Province, Ha Lam District, Ha Long (= Hon Gai), 6 February 1956, coll. I.M. Grochovskaja. Paralectotypes: two larvae, ZMMU Tdt-3176 (specimen 4), ZMMU Tdt-3178 (specimen 1), same collection data as for lectotype.</p> <p>Remarks. Doloisia gigantea is similar to D. varmai (Audy and Nadchatram, 1957) (Schluger et al. 1961), from which it differs by the shape of sensillum (with an expanded top bearing 6 long spikes vs. rounded top covered with multiple, but shorter setules), by the presence of 3 setae on coxa II, vs. 4, branched palpal genual seta, vs. nude, wider scutum (AW = 30–35 vs. 25–26 and SB = 29–34 vs. 23–25), and longer AM (39 vs. 26–29). This species is also similar to D. vercammeni (Audy and Nadchatram, 1957) by the very large size and by the shape of sensilla, but differs in the presence of 3 setae on coxa II, vs. 4, branched palpal genual seta, vs. nude, shorter scutal and idiosomal setae (AL = 18–24, PL = 44–47, and H = 54, vs. 31–34, 66–72, and 83, respectively), and lower number of idiosomal setae (NDV = 69–70 vs. 82–107).</p> <p>Doloisia outoensis was synonymized with D. gigantea by Wen (1984). According the original description of D. outoensis, it has 3–4 setae on coxa II and 6–8 setae on coxa III (Chen &amp; Hsu 1965); thus, variation of these characters in D. gigantea is obviously wider than in its type series.</p> </div>	http://treatment.plazi.org/id/8545EB45EC2EA036FF3F8FF7FC2E84B1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Antonovskaia, Anastasia A.;Stekolnikov, Alexandr A.	Antonovskaia, Anastasia A., Stekolnikov, Alexandr A. (2021): Redescriptions of ten chigger mite species (Acariformes: Trombiculidae) from Vietnam. Zootaxa 4969 (1): 1-53, DOI: https://doi.org/10.11646/zootaxa.4969.1.1
8545EB45EC2BA03AFF3F8EF2FE278555.text	8545EB45EC2BA03AFF3F8EF2FE278555.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Schoutedenichia centralkwangtunga (Mo, Chen, Ho and Li 1959)	<div><p>Schoutedenichia centralkwangtunga (Mo, Chen, Ho and Li, 1959)</p> <p>(Figs. 23C–E, 28–30)</p> <p>Eushoengastia centralkwangtunga Mo, Chen, Ho and Li, 1959: 255, figs. 17–24.</p> <p>Schoutedenichia centralkwangtunga: Domrow 1962: 359, figs. 1–10; Lakshana 1973: 20; Wen 1984: 304; Li et al. 1997: 432, fig. 2-36-2; Chau et al. 2007: 149, fig. 72; Chaisiri et al. 2016: 332.</p> <p>Schoutedenichia (Schoutedenichia) centralkwangtunga: Vercammen-Grandjean 1965b: 102; 1968: 96.</p> <p>Schoutedenichia alongensis Schluger, Grochovskaja, Ngu, Hoe and Tung, 1960a: 191, fig. 12; Kudryashova 2004: 37.</p> <p>Diagnosis. SIF = 4BS-N-3-2110.0000; fsp = 7.7.7; fCx = 1.1.1; fSt = 2.2; fPp = N/N/NNN; fSc: PL&gt; AM&gt; AL; Ip = 548–672; fD = 4H-[7-3]-10-8-8-7-6-2; DS = 48–55; V = 45–48; NDV = 96–103. Eyes absent. Standard measurements of the type series of Schoutedenichia alongensis given in Table 9.</p> <p>Description (larva) [based on lectotype and 2 paralectotypes of S. alongensis]. IDIOSOMA (Figs. 23C–E, 28). Eyes absent; 48–55 barbed dorsal idiosomal setae, including two pairs of humeral setae; 4 sternal setae; 45–48 ventral setae; NDV = 96–103.</p> <p>GNATHOSOMA (Fig. 29A, B). Cheliceral blade with tricuspid cap; gnathobase with sparse puncta and 1 pair of branched setae; cheliceral base and palpal femur with sparse puncta; galeala nude; palpal claw with 3 prongs; setae on palpal femur, genu, and tibia nude; palpal tarsus with 4 branched setae, nude subterminala, and tarsala.</p> <p>SCUTUM (Fig. 23C, D). Trapezoidal, wider than long, with concave posterior margin, moderately covered with small puncta, with 1 AM, 1 pair of ALs, and 1 pair of PLs; all scutal setae uniformly barbed; AM at level of ALs; PLs much longer AM, ALs shorter AM; sensilla claviform, covered with setules; sensillum bases situated at equal distances from levels of ALs and PLs.</p> <p>LEGS (Fig. 29C–E). All 7-segmented, with 1 pair of claws and claw-like empodium. Leg I: coxa with 1 branched seta (1B); trochanter 1B; basifemur 1B; telofemur 5B; genu 4B, 2 genualae, microgenuala; tibia 8B, 2 tibialae, microtibiala; tarsus 22B, tarsala, microtarsala slightly proximal to tarsala, subterminala, parasubterminala, pretarsala. Leg II: coxa 1B; trochanter 1B; basifemur 2B; telofemur 4B; genu 3B, genuala; tibia 6B, 2 tibialae; tarsus 16B, tarsala, microtarsala near and proximal to tarsala, pretarsala. Leg III: coxa 1B; trochanter 1B; basifemur 1B; telofemur 3B; genu 3B, genuala; tibia 6B; tarsus 15B.</p> <p>Hosts. Suncus murinus (Mo et al. 1959), Leopoldamys edwardsi [= Rattus edwardsi (Domrow 1962); = Rattus grochovskii D. Tien (nomen nudum) (Schluger et al. 1960a)], Herpestes javanicus (É. Geoffroy, 1818) (Carnivora: Herpestidae), Tamiops mcclellandi i (Horsfield, 1840) (Rodentia: Sciuridae), Bandicota indica, Crocidura horsfieldi, Menetes berdmorei, Niviventer niviventer, Rattus rattus, Tupaia glis (Lakshana 1973), Niviventer confucianus, Rattus tanezumi (= Rattus flavipectus) (Wen 1984), Rattus andamanensis (= R. koratensis) (Chau et al. 2007).</p> <p>Distribution. China (Mo et al. 1959; Wen 1984), Thailand (Lakshana 1973; Chaisiri et al. 2016), Vietnam (Domrow 1962).</p> <p>Type material examined (Fig. 30). Lectotype of Schoutedenichia alongensis (here designated): larva ZMMU Tdt-3278 (specimen 3) ex L. edwardsi No 27 [labeled as “ Rattus sp. n. (Rattus grochovskii)”], Vietnam, Quang Ninh Province, Ha Lam District, Ha Long (= Hon Gai), 1 February 1956, coll. I.M. Grochovskaja. Paralectotypes of Schoutedenichia alongensis: one larva ZMMU Tdt-3277 (specimen 1), same collection data as for lectotype; one larva ZMMU Tdt-3279 (specimen 4) ex Rattus rattus No 421, Vietnam, Quang Tri Province, Vinh Linh, 18 August 1956, coll. I.M. Grochovskaja.</p> <p>Additional material examined. One larva ZIN 9844 ex Bandicota indica No. R 7112, Thailand, Tak Province, Mae Sot District, rice field (harvested), Lat. 16.48168, Long. 98.44047, 30 November 2013, coll. K. Chaisiri.</p> <p>Remarks. Schoutedenichia alongensis was synonymized with S. centralkwangtunga by Domrow (1962) who examined two of its specimens collected from Suncus murinus in Saigon (currently Ho Chi Minh City) in South Vietnam. He provided measurements of scutum and scutal setae of these specimens. Comparison of our measurements of the S. alongensis type series, measurements of one additional specimen of S. centralkwangtunga from Thailand, and the reference data provided by Mo et al. (1959) and Domrow (1962) shows that these data constitute a continuous range of variability for the most part of characters (Table 9). One exception is the width of scutum (AW, PW, and SB), which is clearly smaller in the type series of S. centralkwangtunga than in other materials. That could be a result of geographic variation or a deformation of the scutum during preparation in the latter set of materials.</p> </div>	http://treatment.plazi.org/id/8545EB45EC2BA03AFF3F8EF2FE278555	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Antonovskaia, Anastasia A.;Stekolnikov, Alexandr A.	Antonovskaia, Anastasia A., Stekolnikov, Alexandr A. (2021): Redescriptions of ten chigger mite species (Acariformes: Trombiculidae) from Vietnam. Zootaxa 4969 (1): 1-53, DOI: https://doi.org/10.11646/zootaxa.4969.1.1
8545EB45EC27A03EFF3F8E9EFCB482A4.text	8545EB45EC27A03EFF3F8E9EFCB482A4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microtrombicula fulgida (Schluger, Grochovskaja, Ngu, Hoe and Tung 1963)	<div><p>Microtrombicula fulgida (Schluger, Grochovskaja, Ngu, Hoe and Tung, 1963)</p> <p>(Figs. 31A, B, 32–34)</p> <p>Trombicula fulgida Schluger, Grochovskaja, Ngu, Hoe and Tung, 1963: 696, figs. 23–29.</p> <p>Microtrombicula fulgida: Kudryashova 2004: 19.</p> <p>Microtrombicula (Microtrombicula) fulgida: Chau et al. 2007: 95, fig. 44.</p> <p>Diagnosis. SIF = 6B-N-2-3111.1000; fsp = 7.7.7; fCx = 1.1.1; fSt = 2.2; fPp = B/B/BbB; fSc: PL&gt; AM&gt; AL; Ip = 547–565; fD = 2H-7-10-5-5-4-4; DS = 31–37; V = 31–42; NDV = 67–74. Sensilla, sternal and coxal setae branched. Standard measurements of type series given in Table 10.</p> <p>Description (larva) [based on lectotype and 3 paralectotypes]. IDIOSOMA (Figs. 31A, B, 32). Eyes 2 + 2; 31–37 barbed dorsal idiosomal setae, including one pair of humeral setae; 4 sternal setae, each with 2 barbs; 31–42 ventral setae; NDV = 67–74.</p> <p>GNATHOSOMA (Fig. 33A, B). Cheliceral blade with tricuspid cap; gnathobase with sparse puncta and 1 pair of branched setae; cheliceral base and palpal femur with sparse puncta; galeala nude; palpal claw with 2 prongs, internal prong longer; setae on palpal femur and genu branched; dorsal and ventral palpal tibial setae branched, lateral palpal tibial seta with 1 branch; palpal tarsus with 6 branched setae and tarsala.</p> <p>SCUTUM (Figs. 31A, 33C). Pentagonal, wider than long, with anterolateral shoulders, with rounded posterior margin, moderately covered with small puncta, with 1 AM, 1 pair of ALs and 1 pair of PLs; AM anterior to level of ALs; all scutal setae uniformly covered with small barbs; PLs longest and ALs shortest scutal setae; sensilla flagelliform, with 8 long branches in distal half; sensillum bases situated close to each other and far anterior to level of PLs (PSB – P-PL = 7–10 µm).</p> <p>LEGS (Fig. 33D–F). All 7-segmented, with 1 pair of claws and claw-like empodium. Leg I: coxa with 1 seta bearing 3 long branches (1B); trochanter 1B; basifemur 1B; telofemur 5B; genu 4B, 3 genualae, microgenuala; tibia 8B, 2 tibialae, microtibiala; tarsus 23B, tarsala, microtarsala distal to tarsala, subterminala, parasubterminala, pretarsala. Leg II: coxa 1B (with 2 short barbs); trochanter 1B; basifemur 2B; telofemur 4B; genu 3B, genuala; tibia 6B, 2 tibialae; tarsus 16B, tarsala, microtarsala at level of tarsala, pretarsala. Leg III: coxa 1B (with 3 long branches); trochanter 1B; basifemur 2B; telofemur 3B; genu 3B, genuala; tibia 6B, tibiala; tarsus 13B, mastitarsala.</p> <p>Host. Rattus tanezumi Temminck [= Rattus flavipectus (Milne-Edwards)]. The host was indicated as a “rat” on the slide labels but was identified as R. flavipectus in the thesis of Grochovskaja (unpublished typescript).</p> <p>Distribution. Vietnam.</p> <p>Type material examined (Fig. 34). Lectotype (here designated): larva ZMMU Tdt-3232 (specimen 5) ex R. tanezumi (labeled as “rat”) No 543, Vietnam, Nghe An Province, Phu Quy, 8 September1956, coll. I.M. Grochovskaja. Paralectotypes: three larvae, ZMMU Tdt-3229 (specimen 1), ZMMU Tdt-3231 (specimen 3), ZMMU Tdt-3280 “ Trombicula vitosa ” (specimen 5), ex R. tanezumi (labeled as “rat”) Nos 571, 568, Vietnam, Nghe An Province, Phu Quy, 10 September 1956, coll. I.M. Grochovskaja.</p> <p>Remarks. Microtrombicula fulgida is similar to M. munda (Gater, 1932) and differs from it in the greater number of idiosomal setae (fD = 2H-7-10-5-5-4-4, DS = 31–36, V = 32–42, and NDV = 67–74 vs. 2H-6-6-4-4-2-2, 26, 32, and 58, respectively), all coxal and sternal setae branched vs. nude (except coxal setae I), and by longer legs (Ip = 547–565 vs. 488) (Vercammen-Grandjean 1965a).</p> <p>This species is also close to M. nadchatrami Vercammen-Grandjean, 1965 and differs from it in a shorter scutum (ASB = 20–21, PSB = 19–21, SD = 39–43, and AP = 19–22, vs. 24, 25, 49, and 25, respectively), shorter setae (AM = 24–25, AL = 20–22, PL = 29–33, D min = 18–24, D max = 25–26, V min = 9–16, V max = 22–24, vs. 31, 28, 38, 27, 32, 21, and 28, respectively), shorter legs (Ip = 547–565 vs. 644), and less branched sternal and coxal setae (sternal setae with 2 vs. about 6 barbs; coxal setae I and III with 3 vs. about 6 long branches; coxal setae II with 2 vs. multiple short barbs) (Vercammen-Grandjean 1965a).</p> </div>	http://treatment.plazi.org/id/8545EB45EC27A03EFF3F8E9EFCB482A4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Antonovskaia, Anastasia A.;Stekolnikov, Alexandr A.	Antonovskaia, Anastasia A., Stekolnikov, Alexandr A. (2021): Redescriptions of ten chigger mite species (Acariformes: Trombiculidae) from Vietnam. Zootaxa 4969 (1): 1-53, DOI: https://doi.org/10.11646/zootaxa.4969.1.1
8545EB45EC23A022FF3F885CFB5E8166.text	8545EB45EC23A022FF3F885CFB5E8166.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microtrombicula vitosa (Schluger, Grochovskaja, Ngu, Hoe and Tung 1963)	<div><p>Microtrombicula vitosa (Schluger, Grochovskaja, Ngu, Hoe and Tung, 1963)</p> <p>(Figs. 31C, D, 35–37)</p> <p>Trombicula vitosa Schluger, Grochovskaja, Ngu, Hoe and Tung, 1963: 694, figs. 16–22.</p> <p>Microtrombicula (Microtrombicula) vitosa: Wen 1984: 315; Li et al. 1997: 277, fig. 2-10-11; Chau et al. 2007: 98, fig. 46.</p> <p>Diagnosis. SIF = 4B1b1N-N-2-3111.1000; fsp = 7.7.7; fCx = 1.1.1; fSt = 2.2; fPp = B/B/BNB; fSc: PL&gt; AM = AL; Ip = 585–636; fD = 2H-6-6-4-5-5; DS = 24–28; V = 25–31; NDV = 49–58. Sensilla branched; sternal setae nude; coxal setae I with 1 short cilium; coxal setae II and III nude. Standard measurements of type series given in Table 11.</p> <p>Description (larva) [based on lectotype and 7 paralectotypes]. IDIOSOMA (Figs. 31C, D, 35). Eyes 2 + 2; 24–28 barbed dorsal idiosomal setae, including one pair of humeral setae; 4 nude sternal setae; 25–31 ventral setae, nude or having few inconspicuous cilia; NDV = 49–58.</p> <p>GNATHOSOMA (Fig. 36A, B). Cheliceral blade with tricuspid cap; gnathobase with sparse puncta and 1 pair of branched setae; cheliceral base with sparse puncta; galeala nude; palpal claw with 2 prongs, internal prong longer; setae on palpal femur and genu with few short barbs, dorsal palpal tibial seta with few short barbs, lateral palpal tibial seta nude, ventral palpal tibial setae with few branches; palpal tarsus with 4 branched setae, 1 seta with single branch, 1 nude seta and tarsala.</p> <p>SCUTUM (Figs. 31C, 36C). Pentagonal, wider than long, with anterolateral shoulders, with rounded posterior margin, densely covered with small puncta, with 1 AM, 1 pair of ALs and 1 pair of PLs; AM anterior to level of ALs; all scutal setae uniformly barbed; PLs longest scutal setae, ALs and AM of about equal length; sensilla flagelliform, slightly expanded, with 8 long branches in distal half; sensillum bases situated close to each other and far anterior to level of PLs (PSB – P-PL = 9–14 µm).</p> <p>LEGS (Fig. 36D–F). All 7-segmented, with 1 pair of claws and claw-like empodium. Leg I: coxa with 1 seta bearing 1 short cilium (1b); trochanter with 1 branched seta (1B); basifemur 1B; telofemur 5B; genu 4B, 3 genualae, microgenuala; tibia 8B, 2 tibialae, microtibiala; tarsus 22B, tarsala, microtarsala at level of tarsala, subterminala, parasubterminala, pretarsala. Leg II: coxa with 1 nude seta (1N); trochanter 1B; basifemur 2B; telofemur 4B; genu 3B, genuala; tibia 6B, 2 tibialae; tarsus 16B, tarsala, microtarsala at level of tarsala, pretarsala. Leg III: coxa 1N; trochanter 1B; basifemur 2B; telofemur 3B; genu 3B, genuala; tibia 6B, tibiala; tarsus 13B, mastitarsala.</p> <p>Hosts. Rattus tanezumi Temminck [= Rattus flavipectus (Milne-Edwards)]. The host was indicated as “rat” on the slide labels but was identified as R. flavipectus in the thesis of Grochovskaja (unpublished typescript).</p> <p>Distribution. China (Wen 1984; Li et al. 1997), Vietnam.</p> <p>Type material examined (Fig. 37). Lectotype (here designated): larva ZMMU Tdt-3280 “ Trombicula vitosa -5” (specimen 2) ex R. tanezumi (labeled as “rat”) No 568, Vietnam, Nghe An Province, Phu Quy, 10 September 1956, coll. I.M. Grochovskaja. Paralectotypes: four larvae, ZMMU Tdt-3280 “ Trombicula vitosa ” (specimen 2), ZMMU Tdt-3280 “ Trombicula vitosa -2” (specimen 1), ZMMU Tdt-3280 “ Trombicula vitosa -3” (specimen 5), ZMMU Tdt-3280 “ Trombicula vitosa -4” (specimen 1), same collection data as for lectotype; three larvae, ZMMU Tdt- 3281 “ Trombicula vitosa ” (specimens 1, 2), ZMMU Tdt-3281 “ Trombicula vitosa -2” (specimen 3) ex R. tanezumi (labeled as “rat”) No 577, Vietnam, Nghe An Province, Phu Quy, 11 September 1956, coll. I.M. Grochovskaja.</p> <p>Remarks. This species is similar to M. spicea (Gater, 1932) by the presence of slightly expanded sensilla, but differs by the nude lateral palpal tibial seta (fPp = B/B/BNB vs. B/B/BBB), 7-segmented legs (fsp = 7.7.7 vs. 7.6.6), longer scutum (SD = 46–48 vs. 41–42), and longer legs (Ip = 585–636 vs. 516–534) (Vercammen-Grandjean 1965a).</p> <p>As Vercammen-Grandjean (1965a) noted for M. spicea, the presence of slightly expanded sensilla relates this species to the genus Ascoschoengastia from the tribe Schoengastiini. Probably, Kudryashova (2004) did not list M. vitosa in the catalogue of types because of the uncertainty of the taxonomic position of this species.</p> </div>	http://treatment.plazi.org/id/8545EB45EC23A022FF3F885CFB5E8166	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Antonovskaia, Anastasia A.;Stekolnikov, Alexandr A.	Antonovskaia, Anastasia A., Stekolnikov, Alexandr A. (2021): Redescriptions of ten chigger mite species (Acariformes: Trombiculidae) from Vietnam. Zootaxa 4969 (1): 1-53, DOI: https://doi.org/10.11646/zootaxa.4969.1.1
