taxonID	type	description	language	source
03BB87E5FFF7FFD4AE18FC7AFCADFE7C.taxon	materials_examined	Type material examined: Guadeloupe: Trois Rivieres, 1 male, 3 females, detritus vegeteaux, without further data (IRSNB, all syntypes). Additional material examined: Guadeloupe: 1 female without further data (NHMW); St. Vincent: Leeward side, 1 male, 1 female without further data, leg. H. H. Smith (NHMW); St. Thomas: 1 male without further data (NHMW).	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFF7FFD4AE18FC7AFCADFE7C.taxon	diagnosis	Diagnosis: The species is larger than P. nitens and eyes are remarkably smaller. Eyes are only 0.25 times longer than temples, while they are tree times longer than temples in P. nitens. Additionally, punctation on pronotum and elytra is denser and coarser than in P. nitens.	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFF7FFD4AE18FC7AFCADFE7C.taxon	description	Description: Length: 2.4 mm. Colour: dark yellow, antennae and legs lighter yellow. Head: 0.30 mm wide, 0.48 mm long; eyes 0.25 times longer than temples; temples smoothly rounded; disc coarsely and densely punctate; distance between punctures 0.5 times as wide as diameter of punctures; on clypeus punctation slightly sparser; surface shiny, with weak longitudinally reticulate microsculpture. Antennae with 2 nd and 3 rd antennomere equal in length; 4 th antennomere quadrate; the following ones wider than long and increasing in width; 8 th antennomere less wide than 7 th and 9 th antennomere and distinctly shorter. Pronotum: 0.30 mm long, 0.48 mm wide; distinctly emarginate in front of posterior edge; only half as wide as at anterior edge; coarsely and densely punctate; still coarser and denser punctate than on head; distance between punctures on average 0.25 times as wide as diameter of punctures; surface shiny, with weak irregular microsculpture. Elytra: 0.50 mm long, 0.53 mm wide; with dense and deep punctation; distance between punctures similar as on pronotum; surface shiny, with weak irregular microsculpture; microsculpture slightly denser than on pronotum. Abdomen with finer and sparser punctation than on elytra and microsculpture more distinct; thus, surface less shiny than elytra; laterally with long dark setae, tergite of last abdominal segment with distinct central tooth.	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFF0FFD4AECFFD9BFCADF9DB.taxon	materials_examined	Type material examined: Brazil: Brésil, without further data, 1 female, Guadeloupe: Base terre, without further data, 1 female (syntypes, IRSNB). Additional material examined: Peru: Madre de Dios, Pentiacolla Lodge, 5.5 km NW El Mirador Trail, Alto Madre de Dios River (12 ° 30.10 ' S, 71 ° 15.28 ' W), 500 m elevation, 23. - 26. Oct. 2000, 1 female, leg. R. Brooks, (PERU 1 B 00 100), flight intercept trap (KU).	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFF0FFD4AECFFD9BFCADF9DB.taxon	diagnosis	Diagnosis: At present, only 2 species of the genus are known from the Neotropics. Therefore, the diagnosis given in P. crassulus informs about the differences between the two species.	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFF0FFD4AECFFD9BFCADF9DB.taxon	description	Description: Length: 2.0 mm. Colour: yellow, antennae and legs lighter yellow. Head: 0.25 mm wide, 0.35 mm long; eyes large, 3 times longer than temples; temples with distinct angles; punctation distinct, but sparse and moderately fine; distance between punctures on average wider than diameter of punctures; surface shiny with weak and sparse microsculpture. Antennae with 2 nd and 3 rd antennomere equal in length; 4 th to 6 th antennomeres not wider than preceding antennomeres; 4 th and 5 th antennomeres quadrate, 6 th one shorter, wider than long; antennomeres 7 to 11 distinctly wider than preceding antennomeres; 7 th antennomere quadrate, 8 th distinctly shorter and smaller; the penultimate antennomeres wider than long. Pronotum: 0.28 mm long, 0.38 mm wide; distinctly emarginate in front of posterior edge; 0.3 times as wide as at anterior angles; punctation slightly coarser and denser than on head, in particular on both sides of the midline; distance between punctures on average smaller than diameter of punctures; surface shiny and nearly polished; with weak irregular microsculpture. Elytra: 0.43 mm long, 0.45 mm wide; with similar punctation as on pronotum; surface polished. Abdomen with finer and sparser punctation than on elytra and microsculpture more distinct than on fore body; scarcely shiny; laterally with long dark setae, tergite of last abdominal segment with distinct central tooth.	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFF1FFD5AEC7FF54FED6F9F0.taxon	materials_examined	Type material: Holotype, male: Brazil: Amazonas, Manaus, male, floating meadow, # WM 13, 15.4.1981, leg. J. Adis (INPA). Paratypes: floating meadow, 13 specimens, # WM 13, # WM 12, 15.4.1981, 3 specimens, # WM 10, 6.2.1981, 2 specimens, # WM 13, 16.3.1981, 1 specimen, # WM 12, 17.2.1981, 1 specimen, # WM 12, 18.5.1981, 1 specimen, 3.3.1982, leg. J. Adis; 1 specimen, 3.3.1982, tree eclector, leg. J. Adis (INPA, UIC, AMNH).	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFF1FFD5AEC7FF54FED6F9F0.taxon	diagnosis	Diagnosis: The species closely resembles E. titschacki in colour, shape of antennae, and shape of abdominal tergite VIII of male. In E. adisi the two pronotal impressions in the midline are absent, the apical part of the central lobe of the aedeagus is placed in a more or less rectangular angle to the basal part and the paramera are also angulated, whereas they are straight in E. titschacki. Posterior edge of abdominal tergite VIII of male is slightly obtuse in E. adisi, whereas it is of more semicircular shape in E. titschacki.	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFF1FFD5AEC7FF54FED6F9F0.taxon	description	Description: Length: 1.8 – 1.9 mm. Colour: dark yellow, legs lighter yellow. Head: 0.23 mm long, 0.33 mm wide; eyes large and distinctly prominent; temples behind eyes narrowed to the neck without rectangular angle; setiferous punctation fine, but distinct; distance between punctures as wide as diameter of punctures; a midline without or with sparser punctures; setae pointing to the midline; surface with weak and sparse microsculpture, shiny, but not polished. Antennae with 2 nd antennomere short and globular; 3 rd antennomere conical and slightly longer than 2 nd; antennomeres 4 to 6 more or less quadrate; the following antennomeres wider than long and increasing in width except 8 th antennomere being smaller than 7 th and 9 th; 10 th antennomere 1.5 times wider than long. Pronotum: 0.27 mm long, 0.33 mm wide; widest shortly behind anterior angle; slightly narrowed to posterior deep emargination; two long dark marginal setae and one dark seta at anterior edge; sides additionally with several dark shorter setae; setiferous punctation as on head; with irregular smooth midline; discal setae yellow pointing to midline; surface as on head with weak microsculpture and shiny, but not polished. Elytra: 0.35 mm long, 0.40 mm wide; widest in the posterior half; distinctly narrowed to shoulders; base distinctly deeper than disc; setiferous punctation and surface as on pronotum; sides with several long dark setae. Abdomen much finer and sparser punctate than elytra; surface less shiny; abdominal tergite VIII with few long dark setae in a more or less dense pubescence and obtuse posterior edge. Aedeagus with thick central part and nearly rectangular apical part; paramera also angulate, but with obtuse angle with basal and slender apical segment; apical transparent plate circular; diameter of transparent plate approximately half as wide as length of apical segment of paramera.	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFF1FFD5AEC7FF54FED6F9F0.taxon	etymology	Etymology: The species is named in honour of my passed friend Prof. Joachim Adis, who found this species in the Central Amazon near Manaus during his studies in the Várzea of the Amazon river.	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFF2FFD7AF7BFF54FE54FCDC.taxon	materials_examined	Type material: Holotype, male: Panama: Chiriqui, Finca Palo Santo, 5.3.1959, leg. H. S. Dybas (AMNH). Paratypes: 179 specimens with same date as holotype; NW Volcán de Chiriqui, litter near stream (82 ° 40 ' 32 " W, 8 ° 49 ' 19 " N), 1450 m elevation, 4 males, 6 females, 16.5.2001, 1 female, 17.5.2001, leg. L. Herman; 25.7 km NW Volcán de Chiriqui, litter near stream (82 ° 46 ' 24 " W, 8 ° 50 ' 06 " N), 1150 m elevation, 3 females, 27.12.2001, leg. L. Herman; 11.5 km E Volcán de Chiriqui, Rio Sereno, tree fall litter in forest (82 ° 46 ' 24 " W, 8 ° 50 ' 05 " N), 1150 m elevation, 1 female, 26.12.2001, leg. L. Herman; Panama, road to Cerro Campana, litter near seep (79 ° 54 ' 50 " W, 8 ° 41 ' 20 " N), 645 m elevation, 4 specimens, 1.1.2002, leg. L. Herman; road to Cerro Campana, litter near stream (82 ° 55 ' 12 " W, 8 ° 41 ', 00 " N), 730 m elevation, 3 specimens, 22.12.2001, leg. L. Herman; Costa Rica: Puntarenas, 35 km NE San Vito, nr. Las Alturas (82 ° 50.08 ' W, 8 ° 56.16 ' N), trail to Cerro Echandi, 1950 m elevation, beyond trail marker 30, litter near stream, 78 specimens, 19.3.1991, leg. L. Herman; 7.5 km on trail to Cerro Echandi, Rio Bella Vista, litter near stream, 34 specimens, 21.3.1991, leg. L. Herman; Las Cruces Botanical Garden, near San Vito (82 ° 57.19 W, 8 ° 47.28 ' N), 1066 m elevation, 1 specimen, 27.2.1985, leg. L. Herman; Alajuela, nr. Poasito (84 ° 11.35 ' W, 10 ° 10.09 N), 6.6 km W Jct. Rt. 126 (9) & 120, 2011 m elevation, Volcán Poas, litter near stream, 6 females, 16.3.1991, leg. L. Herman; Cartago, 16.5 km N Capellades (83 ° 47.07 ' W, 9 ° 55.27 ' N), Rd. to Volcán de Turrialba, 2538 m elevation, leaf litter under hummock in pasture, 7 specimens, 14.3.1991, leg. Herman; Heredia, 2 km S Sacramento, Rd. to Volcán Barba, 2011 m elevation, litter near stream, 1 male, 15.3.1991, leg. L. Herman; La Selva Biol. Station, 1 specimen, 18.2.1985, leg. L. Herman; 27 km N Heredia (83 ° 59 ' W, 10 ° 39 ' W), Rt 9, 1828 m elevation, 1 specimen, 9.2.1985, leg. L. Herman; Ecuador: Pichincha, 18 - 20 km NE Alluriquin, old Quito-Sto Domingo rd., 1460 m elevation, litter, 1 specimen, 21.10.1988, leg. L. Herman; 1950 m elevation, litter, 1 specimen, 23.10.1988; 43 km N, Las Palmeras, old Qt Sto. Dgo. rd. km 59, litter, 2 females, 23.10.1988, leg. L. Herman; Napo, 7 km W Baeza, litter, 2400 m elevation, 1 male, 31.10.1988, leg. L. Herman (AMNH, UIC).	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFF2FFD7AF7BFF54FE54FCDC.taxon	diagnosis	Diagnosis: The species resembles E. mexicanus and E. simplex in size and colour. It can be easily differentiated from these two species by the elongate antennae. Whereas in E. mexicanus and E. simplex antennae are only slightly longer than head and pronotum combined, antennae of E. dybasi are as long as head plus pronotum plus half of elytra. Moreover, the shape of the elytra also differs distinctly between E. dybasi and both E. mexicanus and E. simplex, respectively. Width of elytra of E. dybasi is more than 1.5 times as wide as pronotum, whereas it is only 1.3 times as wide in E. mexicanus and E. simplex. In all three species tergite VIII is slightly emarginate at posterior edge. In E. mexicanus the emargination is weak and smooth without distinct acute prominences at outer posterior angles. In E. dybasi a short stylus is placed at posterior angles, while an acute teeth is present in E. simplex.	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFF2FFD7AF7BFF54FE54FCDC.taxon	description	Description: Length: 2.3 mm. Colour: brown; elytra and antennae red; legs yellow. Head: 0.30 mm long, 0.40 mm wide; eyes distinctly prominent; temples half as long as eyes; smoothly rounded to neck; fore-head distinctly narrowed to acute front edge of clypeus; densely and deeply punctate; distance between setiferous punctures on average less than 1 / 2 of diameter of punctures; vertex with impunctate midline; without microsculpture; surface shiny. Antennae longer than head and pronotum combined; 2 nd antennomere short; 3 rd conical antennomere longer than 2 nd; 3 rd to 7 th antennomeres longer than wide; 8 th antennomere slightly narrower and shorter than preceding and following antennomeres. Pronotum: 0.38 mm long, 0.38 mm wide; smoothly rounded in anterior half; distinctly emarginate in posterior half; lateral margin not visible in dorsal aspect; setiferous punctures dense and coarse; with narrow impunctate midline; on average distance between punctures half as wide as diameter of punctures; with small, but deep circular impression in central position of posterior half. Elytra: 0.55 mm long, 0.60 mm wide; sides smoothly rounded; widest in posterior third; setiferous punctation distinctly sparser and weaker than on pronotum; distance between punctures at least as wide as diameter of punctures, but on average distinctly wider. Abdomen with abdominal tergite III densely and deeply punctate; distance between punctures 1 / 2 as wide as diameter of punctures; density of punctation decreasing from tergite III to tergite VII; distance between punctures on tergite VII 2 to 3 times as wide as diameter of punctures; abdominal tergite VIII widely emarginate and with acute angles; on top of each acute angle a short stylus; at base of tergite VIII two setiferous pores within irregular dark spots. Aedeagus with more or less straight central lobe; paramera straight and with small irregular plate at apex.	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFF2FFD7AF7BFF54FE54FCDC.taxon	etymology	Etymology: The specific name refers to the name of its first collector Dr. H. S. Dybas.	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFF3FFD8AEC7FC69FCD0FD27.taxon	materials_examined	Type material examined: Guadeloupe: without further data, 1 male, 4 females (syntypes, IRSNB) Further material examined: Guadeloupe: Gourbeyre, 6 specimens, without further data (AMNH, UIC); Dominica: Long Ditton, females, 20.06.1911 and 21.06.1911 (AMNH).	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFF3FFD8AEC7FC69FCD0FD27.taxon	diagnosis	Diagnosis: The species resembles E. titschacki, E. microphthalmus, E. adisi, E. venezuelanus; E. moratus, E. nevermanni, E. subtilis, and E. pecki in colouration and size. Among these species E. euplectoides is characterised by the two pronotal impressions, a longitudinal one in the anterior half and a circular one in the posterior half. Furthermore, the antennae are longer than head and pronotum combined, whereas they are shorter or only as long as head and pronotum combined in the other yellow species. The aedeagus is similar to E. titschacki, but tergite VIII of male is obtuse, whereas it is elongate in E. titschacki. In comparison to the other yellow species the punctation of the elytra is relatively sparse.	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFF3FFD8AEC7FC69FCD0FD27.taxon	description	Description: Length: 1.8 mm. Colour: yellow, antennae and legs lighter yellow. Head: 0.20 mm long, 0.30 mm wide; eyes as large as temples; temples smoothly rounded; punctation dense and distinct; distance between punctures on average half as wide as diameter of punctures; surface shiny with weak and sparse microsculpture. Antennae longer than head and pronotum combined; 2 nd and 3 rd antennomere equal in length; following 4 antennomeres at least scarcely longer than wide and slightly increasing in width; penultimate antennomeres quadrate. Pronotum: 0.30 mm long, 0.33 mm wide; distinctly emarginate in front of posterior edge; 0.3 times smaller than at anterior angles; in the front half of a longitudinal impression in the midline; in the posterior half a circular impression; deeply and densely punctate; distance between punctures on average as wide as diameter of punctures; surface shiny, nearly polished, without microsculpture. Elytra: 0.40 mm long, 0.40 mm wide; with similar deep, but sparser punctation as on pronotum; distance between punctures is on average slightly wider than diameter of punctures; surface shiny, with weak irregular microsculpture. Abdomen densely and deeply punctate; punctation similar as on elytra; without microsculpture; surface shiny; laterad with long dark setae, abdominal tergite VIII of male slightly narrowed distad and smoothly emarginate at posterior edge; with few longer and darker setae among short sparse pubescence. Aedeagus slender with smoothly curved apical part; paramera slightly longer than central lobe and with short transparent plate.	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFFCFFD9AF7BFD11FF5CFD71.taxon	materials_examined	Type material: Holotype, male: Mexico: Chiapas, Union Juárez (15 ° 03 ' N, 92 ° 04 ' W), NE slope of Volcan Tacaná, 2000 m elevation, male, Dec. 21, 1975, leg. H. Frania, # 200 - 75 (AMNH).	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFFCFFD9AF7BFD11FF5CFD71.taxon	diagnosis	Diagnosis: E. franiae is the largest Espeson species of the Neotropical region, actually known. Its darker reddish colouration is similar as in E. dybasi, E. mexicanus, E. hermani, and E. simplex. It resembles E. dybasi in the pronotal and elytral punctation, and in the shape and position of the circular pronotal impression. It can be distinguished from E. dybasi by the larger size and the narrower elytra. Whereas the ratio of pronotal to elytral width is 1.4 in E. franiae, E. mexicanus, E. simplex, and E. hermani, the ratio is 1.6 in E. dybasi. Abdominal tergite VIII of males and the aedeagus resembles those of E. simplex, but paramera in E. franiae are angulate in the middle and a pair of transparent spots is placed at base of tergite VIII.	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFFCFFD9AF7BFD11FF5CFD71.taxon	description	Description: Length: 2.7 mm. Colour: brown, elytra lighter reddish, legs dark yellow. Head: 0.30 mm long, 0.43 mm wide; more or less globular in shape; widest at eyes and with rounded temples; fore-head triangular; setiferous punctation deep and dense; on average distance between punctures less than half as wide as diameter of punctures; with moderately wide impunctate midline; surface without microsculpture and polished. Antennae slightly longer than head and pronotum combined; 2 nd antennomere globular, half as long as conical 3 rd antennomere; 4 th to 6 th antennomere more or less quadrate; following antennomeres slightly wider than long and slightly increasing in width with exception of 8 th antennomere; 8 th antennomere slightly smaller than 7 th and 9 th antennomeres. Pronotum: 0.37 mm long, 0.45 mm wide; widest in anterior half; sides of anterior half rounded; in posterior half deeply emarginate; lateral margin in anterior half present, but not visible in dorsal aspect; setiferous punctation coarse and dense; punctures distinctly deeper and larger than head; distance between punctures less than half as wide as diameter of punctures; surface without microsculpture and polished. Elytra: 0.63 mm long, 0.65 mm wide; sides smoothly rounded from shoulders to posterior angles; anterior edge distinctly emarginate; posterior edge slightly emarginate; setiferous punctation irregular; size of punctures variable; fine micro-punctures between punctures of normal size and few larger and deeper punctures; on average distance between punctures as wide as diameter of punctures; surface without microsculpture and polished. Abdomen on anterior segments III to V coriaceously punctate; density of punctation decreasing to posterior segments; segment VIII twice as long as preceding segments; posterior edge of tergite VIII deeply emarginate; at outer posterior angles with short stylus; with few long and black setae and more dense fine setation; at posterior emargination with transverse lighter band with longitudinal striae; at base of tergite VIII two lighter spots surrounded by darker circle. Aedeagus with apical part of central lobe as long as basal part; apical part smoothly rounded; paramera with small transparent plate at apex; diameter of transparent plate not wider than apical short stylus; paramera with triangular prominence at middle.	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFFCFFD9AF7BFD11FF5CFD71.taxon	etymology	Etymology: The specific name refers to its collector H. Frania.	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFFDFFDAAEC7FCD4FFECFCFE.taxon	materials_examined	Type material: Holotype, male: French Guiana: Eaux Claires, N of Saül, (3 ° 39.44 ' N, 53 ° 13.17 ' W), 160 m elevation, in litter, male, X- 5 - 1995, leg. L. Herman (AMNH). Paratypes: Costa Rica: Puntarenas, 6.5 km W Rincon (83 ° 31 ' W, 8 ° 41 ' N), Fundacion Neotropica, litter near stream # 2531, female, 25.3.1991, leg. L. Herman; Heredia, La Selva, Biol. Station, nr. Puerto Viejo de Sarapiqui, male # 2109, 18.2.1985, leg. L. Herman (AMNH, UIC).	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFFDFFDAAEC7FCD4FFECFCFE.taxon	diagnosis	Diagnosis: E. hermani is one of the darker reddish Neotropical Espeson species. Among these species, it is conspicuous by the slightly lighter red elytra and the longitudinal impression in the anterior half of the pronotum. In the male, the species can be easily identified by the central spine at abdominal tergite VIII, the large triangular shape of the transparent lobe of the paramera, and the short apical part of the central lobe of the aedeagus.	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFFDFFDAAEC7FCD4FFECFCFE.taxon	description	Description: Length: 2.0 mm. Colour: reddish; pronotum and elytra lighter red than head and abdomen; legs yellow. Head: 0.25 mm long, 0.35 mm wide; eyes large and prominent; nearly four times longer than short temples; labrum more or less triangular with short acute apex; setiferous punctation dense and deep; distance between punctures less than half as wide as diameter of punctures; a wide midline impunctate; surface with extremely weak remains of microsculpture between punctures; midline without microsculpture and more or less polished. Antennae distinctly longer than head and pronotum combined; 2 nd antennomere more or less globular; conical 3 rd antennomere slightly longer than 2 nd; 4 th to 5 th antennomeres more or less quadrate; 6 th slightly wider than long; following antennomeres slightly increasing in width and wider than long except smaller 8 th antennomere. Pronotum: 0.30 mm long, 0.35 mm wide; with sides rounded in anterior 2 / 3 rd and emargination in posterior 1 / 3 rd; fine lateral margin visible in anterior half in dorsal aspect; setiferous punctation similar as on head; distance between punctures irregular; laterad on average half as wide as diameter of punctures; impunctate midline in anterior half and there with longitudinal impression; surface with weak remains of microsculpture; midline without microsculpture; shiny; at posterior edge a transverse band with coriaceous ground-sculpture. Elytra: 0.45 mm long, 0.50 mm wide; with sides smoothly rounded and with emargination at both anterior and posterior edge; setiferous punctation distinctly wider than on pronotum; distance between punctures on average as wide as diameter of punctures; a sparse micro-punctation between normal punctures. Abdomen finely and sparsely punctate; density of setiferous punctation decreasing posteriad; on tergites II to V transverse impressions with coriaceous ground sculpture; tergite VIII of male deeply emarginate at posterior edge; emargination divided by acute spine; a stripe of fine longitudinal striae at posterior emargination. Aedeagus with short straight apical part and two times longer basal part; paramera slightly longer than central lobe and with large more or less triangular transparent plate; diameter of transparent plate more than half as long as length of paramera.	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFFDFFDAAEC7FCD4FFECFCFE.taxon	etymology	Etymology: The specific name refers to its collector Lee Herman.	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFFEFFDBAF7BFC4AFFABFC82.taxon	materials_examined	Type material examined: Mexico: 1 female, without further data (holotype, IRSNB) Additional material examined: Mexico: Prov. Veracruz, 1 mi. S Huatusco (96 ° 57 ' W, 19 ° 8 ' N), Berlese extraction, 1344 m elevation, 45 specimens, 2.8.1969, leg. S. & J. Peck (AMNH, UIC).	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFFEFFDBAF7BFC4AFFABFC82.taxon	diagnosis	Diagnosis: The species resembles E. simplex, E. franiae, E. dybasi, and E. hermani in the darker reddish colouration and it size of at least 2.0 mm. It can be differentiated from the similarly large E. hermani by the position of the pronotal impression. In E. mexicanus the longitudinal impression is placed in the posterior half and drop-shaped, whereas in E. hermani the longitudinal impression is placed in the anterior half. Moreover, the elytra in E. hermani are lighter red than head and pronotum, whereas no different colouration between head, pronotum, and elytra is found in E. mexicanus. In males, tergite VIII and aedeagus distinctly differ between the two species. E. simplex, E. dybasi, and, in particular, E. franiae are larger with at least 2.3 mm length. In E. simplex a pronotal impression is absent, in E. dybasi and E. franiae the pronotal impression is placed also in the posterior half, but with a circular shape.	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFFEFFDBAF7BFC4AFFABFC82.taxon	description	Description: Length: 2.0 mm. Colour: dark reddish, elytra dark yellow, antennae and legs yellow. Head: 0.28 mm long, 0.36 mm wide; eyes 0.3 times longer than temples; temples smoothly rounded; punctation dense and deep; distance between punctures half as wide as diameter of punctures; a space in the middle of disc without punctures; without microsculpture; surface polished. Antennae as long as head and pronotum combined; 2 nd and 3 rd antennomeres equal in length; following antennomeres quadrate and slightly increasing in width; 8 th antennomere slightly smaller than preceding and following antennomeres. Pronotum: 0.30 mm long, 0.38 mm wide; distinctly emarginate in front of posterior edge; sides in front of emargination smoothly rounded; widest in anterior third; 0.3 times wider than at posterior angles; punctation deep and dense; on average distance between punctures half as wide as diameter of punctures; at posterior edge with coriaceous punctation; a longitudinal impression in the anterior half; with sparse and weak microsculpture; surface shiny. Elytra: 0.48 mm long, 0.53 mm wide; punctation similar as on pronotum; without microsculpture; surface polished. Abdomen with sparser and finer punctation as on elytra, but microsculpture more distinct; surface less shiny than on elytra; abdominal tergite VIII slightly narrowed posteriad and with undulate straight edge; with few longer setae among sparse pubescence. Aedeagus with slender, more or less straight apical part placed in obtuse angle to basal part of central lobe; paramera thick and long with large apical transparent plate; diameter of apical plate nearly as long as length of apical segment of paramera.	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFFFFFDCAEC7FCA6FE01FE3B.taxon	materials_examined	Type material examined: French Guyana: 1 female, without further data (holotype, IRSNB)	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFFFFFDCAEC7FCA6FE01FE3B.taxon	diagnosis	Diagnosis: Compared to the other yellow species, E. microphthalmus can be differentiated by the overall parallel shape of the body that resembles a species of the genus Pseudespeson; but abdominal segment III is slightly narrowed and tergite VIII is without central spine. The pronotal emargination is deep and its sides are nearly parallel with a distinct lateral margin, whereas they are concave in the other yellow Espeson species and lateral margins are absent or extremely narrow. A dorsal impression is absent as in E. adisi, but in E. adisi the shape of the pronotal emargination is concave and abdominal segment III and IV are narrowed at base. Unfortunately, only the type female is known.	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFFFFFDCAEC7FCA6FE01FE3B.taxon	description	Description: Length: 1.9 mm. Colour: yellow, antennae and legs lighter yellow. Head: 0.25 mm long, 0.30 mm wide; eyes as large as temples; temples smoothly rounded; punctation dense and deep; distance between punctures half as wide as diameter of punctures; without microsculpture; surface polished. Antennae shorter than head and pronotum combined; 2 nd globular, 3 rd conical, both antennomeres equal in length; following 3 antennomeres at least slightly wider than long and not wider than preceding antennomeres; antennomeres 7 to 11 distinctly wider than preceding ones and distinctly wider than long; 8 th antennomere slightly smaller than preceding and following antennomere and three times wider than long; penultimate antennomeres twice as wide as long. Pronotum: 0.28 mm long, 0.38 mm wide; distinctly emarginate in front of posterior edge; with deep impression in the emargination; widest in the middle and slightly narrowed to the anterior angles; deeply and densely punctate on the disc, punctation partly coriaceous; without microsculpture; surface polished. Elytra: 0.45 mm long, 0.43 mm wide; punctation irregularly dense; distance between punctures as wide as or smaller than diameter of punctures; without microsculpture; surface polished. Abdomen with similar punctation as on elytra and weak microsculpture; surface shiny, but less shiny than elytra; tergite of last abdominal segment rounded.	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFF8FFDDAECFFDD8FC3EFB12.taxon	materials_examined	Type material examined: St. Thomas: female without further data (holotype of E. moratus, ZMHU); 1 male and 3 females (paratypes, ZMHU); Brazil: São Paulo, Barueri, 3 females, in nest of the ant Camponotus rufipes F., 3.6.1967, leg. Lenko; (holotype and paratype of E. lenkoi, MNHW). Further material examined: Cuba: Havana, Sierra Bouilla, 18.11.1928, male, leg A. Bierig; Soledad (80 ° 16 ' W, 22 ° 09 N), Santa Clara, female, 1.8.1932, leg. B. B. Leavitt (AMNH); Puerto Rico: El Yunque (PUE) male, May 1938, leg. Darlington (AMNH); St. Thomas: 1 female, without further data, 1 female with information: ex coll. Staudinger & Bang-Haas (ZMHU); 4 females without further data (NHMW); 5 males, 5 females, leg. v. Eggers (NHMW); 3 males, 4 females, col. Eppelsheim Steind. (NHMW, AMNH, UIC); Guadeloupe: 1 female without further data (NHMW); St. Vincent: 2 females without further data (NHMW, BMNH); Grenada: Mount Gay East (Leeward side), 2 females without further date, leg. H. H. Smith (NHMW); 1 female, Leeward side, without further data (BMNH); Trinidad: 5 mi SE Rio Claro (61 ° 10 ' 16 W, 10 ° 18 ' 17 N), female, 3.1.1935, leg. N. A. Weber (AMNH); Ecuador: Pichincha, W. of Alluriquin, Tinalandia, 790 - 850 m elevation, litter nr. stream # 2724 - 2725, female, 19. - 20.5.1993, leg. L. Herman (AMNH); Limoncocha, 40 km E Puerto Francisco Orellana, Rio Napo, 1 male, 2 females, 21. - 27.9.1979, leg. Balogh (HNMB); Brazil: Guanabara (43 ° 07 ' W, 22 ° 46 ' S), Represa Rio Grande, female, March-April 1972, leg. F. M. Oliveira (AMNH); Paraguay: Puerto President Stroessner (Cuidad del Este: 54 ° 61.67 ' W, 25 ° 51.67 ' S), Hungarian, Soil. Zool. Exped., 1 male, 6 females, 5.1.1966, leg Balogh et Mahunka (HNMB).	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFF8FFDDAECFFDD8FC3EFB12.taxon	diagnosis	Diagnosis: The species is certainly closely related to E. titschacki and E. adisi concerning the yellow colour, shape of elytra, and structure of aedeagus. The species can be easily distinguished from the two species by the pair of weak pronotal impressions on both sides of the midline in front of posterior edge. Furthermore, it is differing from E. adisi by the slightly smaller size and shorter paramera. The shape of abdominal tergite VIII in males is very similar in both species, but posterior edge in E. moratus is slightly emarginate, whereas it is more obtuse in E. adisi and semicircular in E. titschacki.	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFF8FFDDAECFFDD8FC3EFB12.taxon	description	Description: Length: 1.4 – 1.6 mm. Colour: yellow. Head: 0.20 mm long, 0.27 mm wide; eyes not prominent, but large, as long as temples; temples smoothly rounded behind eyes to distinctly separated neck; setiferous punctation distinct, moderately fine; distance between punctures on average as wide as diameter of punctures; on disc sparser than laterally; a wide midline without punctures; surface without microsculpture and polished. Antennae as long as head and pronotum combined; 2 nd antennomere globular, 3 rd conical, but not longer than 2 nd; 4 th antennomere wider than long, but not wider than 3 rd; the following antennomeres increasing in width; the penultimate antennomere 2 times wider than long. Pronotum: 0.20 mm long, 0.23 mm wide; widest in the anterior half; smoothly rounded to the front edge, anterior angles scarcely present; distad with deep emargination; setiferous punctation similar as on head, but slightly coarser; distance between punctures half as wide as diameter of punctures; with pair of weak longitudinal impressions on each side of the midline in the posterior half; surface without microsculpture and polished. Elytra: 0.30 mm long, 0.33 mm wide; widest in the middle; slightly narrowed to the shoulders and the posterior edge; setiferous punctation distinctly sparser and finer than on pronotum; distance between punctures on average two times wider than diameter of punctures; surface without microsculpture, polished. Abdomen with setiferous punctation still finer and sparser than on elytra; setae pointing posteriad; surface with weak microsculpture, less shiny than elytra; abdominal tergite VIII of male differing from female tergite; posterior edge of male tergite widely emarginate and with indistinct lateral prominence that is expressed by two thicker setae; small midline without setae. Aedeagus with thick and large basal part in central lobe and sharply angulate short apical part; paramera also short, but slightly longer than central lobe; apical transparent plate circular.	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFF8FFDDAECFFDD8FC3EFB12.taxon	discussion	Remarks: Unfortunately the Brazilian specimens described as E. lenkoi by Scheerpeltz (1970) are all females. A significant difference between the West Indian E. moratus specimens could not be found. Scheerpeltz (1970) also described the high similarity of the two species. He focused on the wide antennomeres of E. lenkoi compared to E. moratus. However, also E. moratus is characterised by extremely wide antennomeres. Therefore, although hesitating, I synonymise the two species.	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFF9FFDEAEC7FB36F91CFD46.taxon	materials_examined	Type material examined: Holotype, female: Costa Rica: Hamburgfarm, Ebene Limon, unter loser Rinde, female, 4.10.1928, leg. P. Nevermann (FMNH).	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFF9FFDEAEC7FB36F91CFD46.taxon	diagnosis	Diagnosis: Unfortunately only the female holotype is known. The species resembles E. microphthalmus, E. adisi, and E. simplex in the absence of pronotal impressions. Compared to E. simplex, E. nevermanni is smaller in size and as long as E. adisi and E. microphthalmus. It can be differentiated from E. microphthalmus by the wider elytra and from E. adisi by the more rectangular temples.	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFF9FFDEAEC7FB36F91CFD46.taxon	description	Description: Length: 1.8 mm. Colour: yellow. Head: 0.20 mm long, 0.33 mm wide; with eyes not prominent and temples nearly as long as eyes; fore-head nearly parallel in front of eyes; clypeus triangular; labrum with short spine in middle; punctation dense and deep; distance between punctures irregular; on average slightly smaller than diameter of punctures; neck with dense coriaceous punctation; with small impunctate midline; surface without microsculpture and shiny. Antennae nearly as long as head and pronotum combined; 2 nd antennomere globular and nearly as long as conical 3 rd antennomere; 4 th to 6 th antennomeres more or less quadrate; 7 th to 10 th antennomeres increasing in width and wider than long; except smaller 8 th antennomere. Pronotum: 0.25 mm long, 0.34 mm wide; rounded in anterior two third; in posterior third emarginate; with small margin in posterior half in front of emargination; anterior half not margined; setiferous punctation more regular; as deep as on head, but less dense; on average distance between punctures as wide as diameter of punctures; without impression on disc; surface without microsculpture and shiny. Elytra: 0.31 mm long, 0.38 mm wide; with smoothly rounded sides, but not distinctly wider than pronotum; punctation less dense and deep than on pronotum and surface still more shiny. Abdomen only with segment III with deep transverse impression at base; following segments without transverse impression; setiferous punctures distinctly finer than on fore-body.	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFFAFFDFAF7BFCE3FF4AFC71.taxon	materials_examined	Type material: Holotype, male: Mexico: Cordoba, Prov. Veracruz, tropical evergreen forest, Berlese, male, 7.8.1969, leg. J. Peck (AMNH). Paratypes: 65 specimens with same data as holotype; 385 specimens with same data as holotype, but 4.8.1969; Cordoba, Paraje Nueve Nacimiento, tropical evergreen forest, Berlese, 26 specimens, 8.1969, leg. S & J. Peck; San Blas, junction San Blas and Tepic roads, 9 specimens, 4.8.1960, without collector information; Nuevo, Monterrey León, Chipinque Mesa (115 ° 15.49 ' W, 25 ° 35.14 ' N), forest leaf litter, 1 female, 22.6.1969, leg. S. & J. Peck; Tamaulipas, Gomez Farias, Rancho del Cielo, 1100 m elevation, 1 female, 4.7.1969, leg. S. & J. Peck; Veracruz, Fortin (96 ° 59.59 ' W, 18 ° 53.49 ' W) 1 male, 15 females, 5.8.1969, leg. S. & J. Peck (AMNH, UIC).	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFFAFFDFAF7BFCE3FF4AFC71.taxon	diagnosis	Diagnosis: Compared with the small yellow species, E. titschacki, and E. adisi, E. pecki is easily to identify by the extremely small eyes. The species is as small as E. titschacki and both species are slightly smaller than E. adisi. Antennae are slightly shorter than in E. titschacki. In E. titschacki antennae are as long as head and pronotum combined, whereas they are slightly shorter than pronotum in E. pecki. Both species, E. pecki and E. titschacki, can be distinguished from E. adisi by the presence of two deep impressions on the pronotum, whereas E. adisi has no impression on pronotum. E. venezuelanus that is also similar to these three species is slightly darker. In males, the deep emargination of abdominal tergite VIII resembles that in E. simplex, but E. simplex is much larger, and punctation of tergite VIII is denser. Furthermore, the semicircular large apical plates of paramera are unique in the Neotropical Espeson species.	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFFAFFDFAF7BFCE3FF4AFC71.taxon	description	Description: Length: 1.6 mm. Colour: yellow. Head: 0.20 mm long, 0.25 mm wide; eyes extremely small; only four or five ocellae; nearly invisible in dorsal aspect; temples twice as long as eyes and convexly prominent; fore-head straightly narrowed to more or less acute front edge of clypeus; setiferous punctation moderately dense and fine; on clypeus finer and denser than on lateral part of vertex; disc with impunctate midline; surface shiny; laterad with remains of weak microsculpture; disc polished. Antennae nearly as long as head and pronotum combined; 1 st and 2 nd antennomere oblong, 3 rd conical; antennomeres 1 to 3 nearly equal in length; antennomere 4 and 5 more or less quadrate; following antennomeres wider than long; 8 th antennomere narrower and smaller than antennomeres 7 and 9; 10 th antennomere nearly twice as wide as long. Pronotum: 0.23 mm long, 0.27 mm wide; with weak lateral margin; only visible in posterior half in dorsal aspect; deeply emarginate in posterior half; posterior edge only half as wide as width in anterior half; setiferous punctation deeper and coarser than on head; distance between punctures on average only half as wide as diameter of punctures; in anterior and posterior half each a small, but deep impression; surface shiny; laterad with remains of microsculpture, but surface mostly polished. Elytra: 0.26 mm long, 0.33 mm wide; with setiferous punctation slightly weaker and sparser than on pronotum; with remains of microsculpture, but surface mostly polished and shiny. Abdomen with coarser punctation on anterior segments than on distal segments; microsculpture slightly more distinct than on elytra and, thus, surface less shiny; tergite VIII of male deeply emarginate in middle and with long acute teeth at outer angles; sparsely punctate also at lateral margin. Aedeagus with long and thick basal part; apical part more or less straight and only half as long as basal part; paramera with large apical plate of semicircular shape.	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFFAFFDFAF7BFCE3FF4AFC71.taxon	etymology	Etymology: The specific name refers to its collector James Peck.	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFFBFFC0AEC7FBD4FFA4FB05.taxon	materials_examined	Type material: Holotype, male: Brazil: Nova Teutonia, without further data, leg. F. Plaumann (ZMHU), (52.23 ' W, 27.11 ' S). Paratypes: Nova Teutonia, 300 - 500 m elevation, without further data, 1 female, leg. F. Plaumann (ZMHU), 4 males, 3 females, VI. 1960, leg. F. Plaumann (ZMHU, UIC), 2 females, VII. 1959, leg. F. Plaumann (ZMHU), 1 male, XI. 1955, leg. F. Plaumann (ZMHU); Paraguay: Puerto Presidente Stroessner (Cuidad del Este: 54 ° 61.67 ' W, 25 ° 51.67 ' S), Hungarian Zoological Expedition, 1 male from ethylene-glycol soil traps, 27.12.1965 - 7.1.1966, leg. A. Zicssi (HNMB).	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFFBFFC0AEC7FBD4FFA4FB05.taxon	diagnosis	Diagnosis: The impressions on the pronotum are absent in this species and in this respect it resembles E. adisi and E. microphthalmus. But in size and colour it is more similar to E. mexicanus. Compared to E. microphthalmus, elytra of E. simplex are wider and sides are curved, whereas sides are more or less parallel in E. microphthalmus. The species is unique by the shape of the abdominal tergite VIII of male that laterally carries a distinct tooth. The two specimens without date were already labelled as holotype and paratype by Wendeler, but never described.	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFFBFFC0AEC7FBD4FFA4FB05.taxon	description	Description: Length: 2.3 mm. Colour: dark red, head brown, leg and antennae lighter red. Head: 0.25 mm long, 0.33 mm wide; eyes not prominent and small; temples well developed, at least 0.3 times longer than eyes; temples behind eyes shortly parallel, then rounded in a smooth curve; neck distinctly separated from the head; setiferous punctation coarse and dense; distance between punctures less than half as wide as diameter of punctures; on the disc with distinct impunctate midline widened to the front; clypeus sparsely punctate; surface without micro- sculpture and polished. Antennae as long as head and pronotum combined; 2 nd antennomere globular, 3 rd conical, but not longer than 2 nd, antennomeres 4 and 5 quadrate, the following antennomeres slightly wider than long; 8 th antennomere slightly smaller than preceding and following antennomeres. Pronotum: 0.30 mm long, 0.37 mm wide; widest in the anterior third, rounded in a smooth curve to the front edge; without distinct front angles; posteriad with emargination, forming a nearly rectangular angle; setiferous punctation coarse and dense, puncture still deeper and larger than on head; distance between punctures less than 1 / 4 as wide as diameter of punctures; surface without microsculpture and polished. Elytra: 0.43 mm long, 0.50 mm wide; widest in the posterior half; posterior angles rounded in a smooth curve; shoulders indistinct, too; setiferous punctation as large as on pronotum, but slightly sparser; between setiferous coarse punctation a finer and sparser micro-punctation; distance between coarser punctures irregular, on average half as wide as diameter of punctures. Abdomen with anterior tergites as coarsely and densely punctate as on head, posterior tergite sparser and finer punctate; abdominal tergite VIII in male emarginate at posterior edge and with lateral tooth; two long setae inserted anteriad and with relatively long and dense pubescence, but with a wide midline without setae. Aedeagus slender with slightly angulate apical part; basal segment of paramera thick, apical segment slender; apical transparent plate circular; placed on a short stylus.	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFFBFFC0AEC7FBD4FFA4FB05.taxon	etymology	Etymology: The specific name was adopted from the preliminary name given by Wendeler. The name derived from the same Latin word simplex meaning simply and presumably refers to the simple surface of the pronotum without impressions.	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFE4FFC1AF7BFB27FFF6FC2B.taxon	materials_examined	Type material examined: Mexico: Veracruz, Cordoba, without further data, leg. Fenyes (holotype in FMNH); Cordoba, without further data, 2 males, 2 females (paratypes, NHMW, BMNH, NMPC). Additional material examined: Brazil: Paranà, Rio Negro, 3 females without further data, leg. Reichensperger (NHMW); Minas Gerais, Passa Quatro, 1 female without further data (NHMW).	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFE4FFC1AF7BFB27FFF6FC2B.taxon	diagnosis	Diagnosis: The species resembles E. mexicanus in punctation of the fore-body and the impressions of the pronotum. But E. mexicanus is larger than E. subtilis and the impression in the anterior half of the pronotum is longitudinal, whereas it is circular in E. subtilis.	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFE4FFC1AF7BFB27FFF6FC2B.taxon	description	Description: Length: 1.8 mm. Colour: red, head darker red, legs and antennae dark yellow. Head: 0.20 mm long, 0.30 mm wide; eyes scarcely prominent; temples continuously narrowed to the neck; only half as long as eyes; setiferous punctation coarse and deep; distance between punctures only half as wide as diameter of punctures; a wide smooth midline impunctate; surface without microsculpture and polished. Antennae as long as head and pronotum combined; with 2 nd antennomere quadrate, 3 rd conical and not longer than 2 nd, the following antennomeres more or less quadrate and increasing in width; penultimate antennomeres only slightly wider than long. Pronotum: 0.27 mm long, 0.30 mm wide; widest in the middle, narrowed in a smooth curve to front angle, emargination in front of the posterior angle forming an obtuse angle; setiferous punctation still deeper and larger than on head; distance between punctures half as wide as diameter of punctures; a wide midline without punctures; within midline a short longitudinal impression in anterior half and a circular impression in posterior half; surface without microsculpture and polished. Elytra: 0.37 mm long, 0.40 mm wide; widest in the posterior half, with setiferous punctation differing in size; coarser punctation sparser than on pronotum, distance between punctures on average as wide as or wider than diameter of punctures, between coarser punctation a sparse finer punctation; surface without microsculpture and polished. Abdomen with punctation as coarse and dense as on pronotum, on each abdominal tergite a wide impunctate midline; posterior edge of tergite VIII in male smoothly rounded; without emargination. Aedaegus with thick basal part of central lobe and apical part placed in rectangular angle to basal part. The paramera not found in the single male specimen.	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFE5FFC3AEC7FBFEFDE5FEBD.taxon	materials_examined	Type material examined: Peru: Ayacucho, Sivia (73 ° 51.09 ' W, 12 ° 30.45 ' S) (male holotype FMNH); 3 females with same data as holotype (paratypes, FMNH, NHMW); from the same location, leaf litter; female 22.5.1936, leg. Titschack (paratypes, FMNH). Further material examined: Mexico: Nayarit (104 ° 50.42 ' W, 21 ° 45.07 ' S), junction San Blas and Tepec roads 300 m elevation, 64 specimens, 4.8.1960, without information of the collector (AMNH, UIC); Venezuela: Bolivar, Carabobo (61.24 ' W, 6.18 ' N), Via Palmichal, 750 - 850 m elevation, 3 males, 9 females, Nov. 2005, leg. Brachat (UIC, VAC), 900 m elevation; 2 females, 22.11.2005, leg. Brachat (UIC, VAC); Ecuador: Pichincha, Santo Domingo de los Colorados (79 ° 10 ' W, 0 ° 14 ' S), 17 km SE, Tinalandia, 900 m elevation, 1 male, 1 female, 16. - 21.10.1988, 1 female 19. - 20.5.1998, leg. L. Herman (AMNH); Pastaza, 2.6 km S of Santa Clara, (01 ° 17 ' 20 " S, 77 ° 53 ' 20 " W), 790 m elevation, secondary lowland forest with sparse understory vegetation, shaded, sifting leaf litter between tabular roots, 2 females, 15. - 16.11.2006, leg. Fikacek & Skuhrovec (NMPC); Alluriquin, 43 km N, Las Palmeras, old Qto-Sto. Dgo. rd. km 59, litter, 23.10.1988, leg. L. Herman (AMNH); Limoncocha, 40 km E Puerto Francisco Orellana, Rio Napo, 2 females, 21. - 27.9.1979, leg. Balogh (HNMB); Napo, 35.5 km NE El Chaco, Cascadas San Rafael (00 ° 06 ' 00 " S, 77 ° 34 ' 51 " W), 1200 m elevation, low primary forest on summit of river gorge, sparse understory vegetation, lot of leaf litter, sifting leaf litter, 2 males, 1 female, 29.11.2006, leg. Fikacek & Skuhrovec (NMPC); Peru: Cuzco, Madre de Dios, Cuzco Amazonica, secondary forest (69 ° 02 ' 06 W, 12 ° 36 ' 48 S), 300 m elevation, 5 males, 16 females; 17.5.1995, leg. D. Agosti (AMNH, UIC); Junin, San Ramón de Pangoa (75 ° 19 ' 60 W, 11 ° 07 ' 60 S), 40 km SE Satipo, 750 m elevation, male, 25.3.1972, leg. R. T. Schuh (UIC); Huanuco, Panguana (74 ° 56 ' W, 9 ° 37 ' S), Chocha, female, 15.3.1976, leg. W. Hanagarth (UIC); Brazil: Santa Catarina, Nova Teutonia, Aug. 1953, female, leg. F. Plaumann (AMNH); Mato Grosso, Nossa Sinhora de Livramento (56 ° 36.24 ' W, 16 ° 15.24 ' S), Pirizal, Faz. Retiro Novo, from V. divergens tree collected by fogging, female, 22.02.2000, leg. M. Marques (UFMT). Paraguay: Puerto Presidente Stroessner (Ciudad del Este: 54 ° 61.67 ' W, 25 ° 51.67 S) Hungarian Soil-Zoology Exp., 1 females, 5.1.1966, leg. Balogh et Mahunka (HNMB); Puerto Presidente Stroessner (Ciudad del Este: 54 ° 61.67 ' W, 25 ° 51.67 ' S) Hungarian Soil-Zoology Exp., Acaray waterfall, 1 female, 2.1.1966, leg. Loksa (HNMB); Argentina: Salta, Aguas Blancas-Yaculica (64 ° 22.25 ' W, 22 ° 43.44 ' S), 460 m elevation, yungas forest, leaf litter, female, 25.10.1994, leg. J. Carpenter & D. Agosa (AMNH).	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFE5FFC3AEC7FBFEFDE5FEBD.taxon	diagnosis	Diagnosis: The species is very similar to E. pecki, E. adisi and E. moratus in size and colouration. It differs from E. adisi in the presence of the two pronotal impressions. Females are very similar to E. pecki, because both species have similar pronotal impressions in the midline. It is distinguished from E. pecki by the longer antennae and in the male by the different shape of tergite VIII that is elongate and semicircular at apex in E. titschacki, but short and emarginate in E. pecki.	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFE5FFC3AEC7FBFEFDE5FEBD.taxon	description	Description: Length: 1.6 mm. Colour: yellow. Head: 0.20 mm long, 0.25 mm wide; eyes as long as temples, with more than 12 ocellae and distinctly visible in dorsal aspect; temples more or less parallel; fore-head straightly narrowed to more or less acute front edge of clypeus; setiferous punctation laterad denser than on disc; setae pointing to middle; with impunctate midline on vertex; surface laterad with remains of microsculpture; on disc without microsculpture; surface polished and shiny. Antennae as long as head and pronotum combined; 1 st antennomere thick; 2 nd globular and as thick as conical 3 rd antennomere; following four antennomeres more or less quadrate; 8 th antennomere only slightly narrower and smaller than 7 th and 9 th antennomere; 9 th and 10 th antennomere slightly wider than long. Pronotum: 0.25 mm long, 0.30 mm wide; with fine lateral margin visible in its total length in dorsal aspect; deeply emarginate in posterior half; anterior edge approximately 1.5 times as wide as posterior edge; densely and deeply punctate; distance between setiferous punctures on average half as wide as diameter of punctures; in midline with impressions in both anterior and posterior half; surface with remains of microsculpture, but mostly polished and shiny. Elytra: 0.30 mm long, 0.35 mm wide; with similar deep and dense setiferous punctation as pronotum; surface with remains of microsculpture, but moderately polished and shiny. Abdomen on tergites III to VI more weakly, but as densely punctate as elytra; density of punctation decreasing posteriad; microsculpture more distinct than on fore-body; surface less shiny; tergite VIII of male straightly narrowed to rounded apex; sparsely punctate. Aedeagus slender with apical part nearly as long as basal part; paramera slightly longer than central lobe and with small apical transparent plate.	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFE5FFC3AEC7FBFEFDE5FEBD.taxon	discussion	Remarks: No differences could be found between the Mexican specimens and the South American specimens, which let suppose that the species might also occur in other Central American countries. 4 Discussion The remarkable structure of the aedeagus, in particular, the structure of the paramera with the suction-cup-like apical appendix, at least in the genus Espeson, is unique in the family Staphylinidae. Moreover, it seems that the paramera are divided into two parts, similar to the subfamily Aleocharinae. Although the overall habit of the fore-body resembles that in the genus Glyptoma Erichson, 1839, the structure of the aedeagus suggests that both genera are not closely related to Glyptoma as already emphasised by Irmler (2010). Lecoq (1994) mentioned the smaller 8 th antennomere as characteristic to differentiate Pseudespeson from Espeson, but as the comparison of all Espeson species showed the size of the 8 th antennomere varies in this genus. In several species, e. g. E. moratus, the 8 th antennomere equals 7 th and 9 th antennomere in size, but in other species, e. g. E. microphthalmus and E. nevermanni, it is distinctly smaller. In some species, e. g. E. simplex and E. pecki, 8 th antennomere is only slightly smaller than 7 th or 9 th antennomeres. Thus, this character is not appropriate to differentiate the two genera. Furthermore, Lecoq (1994) also noted the shape of the last abdominal tergite as differentiating character between the two genera. Indeed, in the two Neotropical Pseudespeson species the last abdominal tergite is of triangular shape with short apical spine. In Espeson, a triangular shape of the last tergite without spine is found in E. titschacki, a central spine at an emarginate posterior edge is found in E. hermani. The distinct sexual dimorphism in Espeson with simple triangular shape of last abdominal tergites, i. e. E. titschacki, and various shapes in males indicates that the variance in the shape of the last abdominal tergite might be high and provide no valuable differentiating character, too. The geographic distributions of the species are still vague due to the rare records of the species. The genus Pseudespeson was recorded from extreme distant locations, i. e. Guadeloupe and southern Brazil, which indicates that species might occur also in other Neotropical countries. In particular, P. nitens that was recorded from The Caribbean, Brazil and Peru shows a wide distribution, but seem to be rare. In contrast, the species of the genus Espeson are widely distributed from Mexico to Argentina. Some, e. g. E. moratus and E. titschacki, seem to be distributed over the whole Neotropical region or at least in a wide area of South America and the Caribbean. Others, i. e. E. dybasi, were found from several localities of southern Central America and the northern Andean region of South America. Thus, similar geographical distributions can be assumed as in other osorine genera (Irmler 2007, 2010). Concerning the ecology of the species, our knowledge is similarly poor as for the geographical distribution. Referring to the information given on the labels by the collectors, many specimens were found in litter of rain forests. Nevertheless, also other habitats are recorded, e. g. E. adisi was also found by tree eclectors, E. nevermanni was found under bark, and E. moratus even in nests of ants. These records indicate that a variety of habitats may be inhabited by the species. 5 References Bernhauer, M. 1910: Staphylinidae (Coleoptera). – Beiträge zur Fauna Perus 1: 277 - 293. Bernhauer, M. 1942: Neue Staphyliniden aus Kostarika. – Zoologischer Anzeiger 138: 1 - 27. Fauvel, A. 1902: Staphylinides exotiques nouveaux – Revue d‘Entomologie 21: 8 - 37. Hadley, A. 2006: CombineZ 5.3. http: // www. hadleyweb. pwp. blueyonder. co. uk / CZ 5 / combinez 5. htm Herman, L. 2001: Catalogue of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. II. Oxyteline group. – Bulletin of the American Museum of Natural History 265: 1067 - 1806. Irmler, U. 2007: The Tannea (Coleoptera: Staphylinidae: Osoriinae) species of Costa Rica – new species and records, a key, ecological, and geographical remarks. – Brenesia 68: 69 - 85. Irmler, U. 2009: New species and records of the genus Lispinus with a key to the species from Peru (Coleoptera: Staphylinidae: Osoriinae). – Zootaxa 2263: 42 - 58. Irmler, U. 2010: A new genus of Osoriinae in the Neotropical region with a cladistic analysis of the tribe Thoracophorini (Insecta: Coleoptera: Staphylinidae). – Arthropod Systematics and Phylogeny 68: 229 - 237. Lecoq, J. C. 1994: Un nouveau genre et une nouvelle espèce d’Osoriinae de Sierra Leone: Pseudespeson rossi (Coleoptera, Staphylinidae). – Ricerche biologiche in Sierra Leone; 1 Academia Nazionale die Lincei, Rom, 267: 299 - 306. Schaufuss, L. W. 1882: Descriptions de coléptères nouveaux. – Annales de la Société Entomologique de France, 3. Série 62: 43 - 48. Scheerpeltz, O. 1970: Studien über die Arten der Gattungen Espeson Schaufuss und Parespeson Bernhauer (Coleoptera, Staphylinidae, Subfamilie Piestinae, Tribus Thoracophorini). – Mitteilungen der Münchner Entomologischen Gesellschaft 59: 115 - 129. Author´s address: Subject editor: Prof. Dr. Ulrich Irmler Dr. L. Zerche Institut für Ökosystemforschung Abt. Angewandte Ökologie, Universität Olshausenstrasse 40, 24098 Kiel, Germany e-mail: uirmler @ ecology. uni-kiel. de	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
03BB87E5FFE5FFC3AEC7FBFEFDE5FEBD.taxon	discussion	Remarks: No differences could be found between the Mexican specimens and the South American specimens, which let suppose that the species might also occur in other Central American countries.	en	Irmler, U. (2012): The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae). Beiträge Zur Entomologie = Contributions to Entomology 62 (2): 331-360, DOI: 10.21248/contrib.entomol.62.2.331-360
