identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03A09A04A731FF906EDCFCFE2FE6FBFF.text	03A09A04A731FF906EDCFCFE2FE6FBFF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Blattochaeta Reitter 1910	<div><p>Genus Blattochaeta Reitter, 1910</p> <p>Diagnosis. A member of the subterranean tribe Leptodirini, subtribe Bathysciina, group Théléomorphes, division II sensu Jeannel 1924, defined by: bathysciod body shape, protarsi four segmented in both sexes, protibiae lacking comb (pecten) along the lateral external margin and lacking external apical spurs, pronotum evenly convex, trapezoidal and widest at its base, inner sac (endophallus) of median lobe with sclerotized structures (apical reinforcement bands, medially positioned sclerotized structures, feather-like structures, connection nodules and basal phanera).</p> <p>Distribution map for the genus Blattochaeta is given (Fig. 16).</p> </div>	http://treatment.plazi.org/id/03A09A04A731FF906EDCFCFE2FE6FBFF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Čeplík, Dávid;Lakota, Ján;Čeplík, Jaroslav	Čeplík, Dávid, Lakota, Ján, Čeplík, Jaroslav (2021): Two new species of the genus Blattochaeta Reitter, 1910 (Insecta, Coleoptera Leiodidae, Cholevinae, Leptodirini) from Montenegro. Zootaxa 4969 (1): 86-100, DOI: https://doi.org/10.11646/zootaxa.4969.1.4
03A09A04A731FF976EDCFB6A2902FC6C.text	03A09A04A731FF976EDCFB6A2902FC6C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Blattochaeta brankojalzici D. Ceplik, Lakota & J. Ceplik 2021	<div><p>Blattochaeta brankojalzici D. Čeplík, Lakota &amp; J. Čeplík, sp. nov.</p> <p>(Figs 1–4, 11a–11b)</p> <p>Material studied. Holotype, ♂: two labels: MONTENEGRO „ <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.361973&amp;materialsCitation.latitude=43.07397" title="Search Plazi for locations around (long 19.361973/lat 43.07397)">Montenegro</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.361973&amp;materialsCitation.latitude=43.07397" title="Search Plazi for locations around (long 19.361973/lat 43.07397)">Sinjajevina Mts.</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.361973&amp;materialsCitation.latitude=43.07397" title="Search Plazi for locations around (long 19.361973/lat 43.07397)">Njegovuđa village</a>, Rudanca, Blažova pećina cave, coordinates 43°04‘26.3“N, 19°21‘43.1“E (WGS 84), 27.05.2015, D. Čeplík, J. Lakota, G. Dunay &amp; J. Čeplík lgt.“ (white printed); „ HOLOTYPE Blattochaeta brankojalzici sp. nov., D. Čeplík, Lakota &amp; J. Čeplík det., 2021“ (red printed), (CNHM). Paratypes, 2 ♀♀: same data as for the holotype but collected 27.05.2015 – 14.05.2016 by traps (CDC, CJL). Paratypes are labelled with red printed labels „ PARATYPE Blattochaeta brankojalzici sp. nov., D. Čeplík, Lakota &amp; J. Čeplík det., 2021“.</p> <p>Description. Body medium–sized, bathyscioid, elliptically elongate, convex, weakly pigmented, with fine pubescence and punctures, colour from yellowish–brown to reddish–brown (Fig. 1 – holotype slightly immature). Body length 4.55 mm in male, 4.68–5.00 mm in females. L: 4.05–4.40 mm. Head approximately as long as wide or slightly longer than wide (due to the measuring technique of the head capable of being retracted), HL: 0.74 mm, HW: 0.74 mm, eyes completely absent, with posterior neck region and genae more or less glabrous. Transverse occipital crest present but weakly–defined. Clypeus subhexagonal, clypeofrontal suture distinct, frontoclypeus densely setose, labrum densely setose. Mandibles short, robust, with proximal and subapical incisors and one or two small teeth between. Maxillary ultimate palpomere conical, apically pointed, slender and much shorter than the thick penultimate palpomere. Antennae (AL: 2.27–2.34 mm), finely pubescent (Figs 2a–2b), inserted in the middle of the median third of the head. Pedicel approximately 1.77 times longer than scape. Antennomere 8 as long as wide (slightly longer than wide in male L/W: 1.10 and slightly wider than long in females L/W: 0.91). Relative length of antennomeres (male): (1) 0.17, (2) 0.31, (3) 0.17, (4) 0.16, (5) 0.18, (6) 0.16, (7) 0.26, (8) 0.11, (9) 0.22, (10) 0.23 and (11) 0.37. Relative width of antennomeres (male): (1) 0.08, (2) 0.08, (3) 0.07, (4) 0.07, (5) 0.08, (6) 0.09, (7) 0.13, (8) 0.10, (9) 0.13, (10) 0.14 and (11) 0.14. Antennomere length/width ratios (male): (1) 2.12, (2) 3.87, (3) 2.42, (4) 2.28, (5) 2.25, (6) 1.77, (7) 2.00, (8) 1.10, (9) 1.69, (10) 1.64 and (11) 2.64. Relative length of antennomeres (female): (1) 0.18, (2) 0.31, (3) 0.19, (4) 0.15, (5) 0.17, (6) 0.17, (7) 0.26, (8) 0.11, (9) 0.21, (10) 0.19 and (11) 0.33. Relative width of antennomeres (female): (1) 0.10, (2) 0.11, (3) 0.06, (4) 0.07, (5) 0.08, (6) 0.11, (7) 0.14, (8) 0.12, (9) 0.16, (10) 0.18 and (11) 0.15. Antennomere length/width ratios (female): (1) 1.80, (2) 2.81, (3) 3.16, (4) 2.14, (5) 2.12, (6) 1.54, (7) 1.85, (8) 0.91, (9) 1.31, (10) 1.05 and (11) 2.20.</p> <p>Pronotum trapezoidal, strongly convex, wider than long, widest at base (PL/PW: 0.55–0.62), lateral edges evenly rounded, at posterior margin widest, slightly protruding backwards, narrower than elytra. PL: 1.14–1.20 mm. PW: 1.82–2.14 mm. Dorsal surface strongly pubescent. Prosternal ventral surface sparsely pubescent.</p> <p>Elytra elongately oval, pubescent, with maximum width at mid–length, longitudinal parasutural striae absent. EL: 2.91–3.20 mm. EW: 2.00– 2.28 mm. EL/EW: 1.40–1.45. Punctation dense, consisting of approximately 18–20 punctures across the mid–width of each elytron, weak punctuation and pubescence on apical part. Scutellum widely triangular and pubescent.</p> <p>Venter. Mesoventral and metaventral surface sparsely pubescent. Mesocoxal cavities separated, mesoventral process well–developed. Metacoxae separated by bifid posterior metaventral process. Mesoventral carina (Figs 11a– 11b) developed, prolonged posteriorly, not extended over metaventrite, with backwardly oriented setae. Sternites pubescent with short, fine setae, with reticulate and leathery aspect.</p> <p>Protarsi with four slender, undilated segments in both sexes, first protarsomere slightly longer than second and third, fourth longest. Mesotarsi and metatarsi pentamerous in both sexes. Claws wide and well–developed, empodium with one bifurcate seta. Protibiae armed with spines, one very distinct external subapical spine, internal subapical side with one shorter and one longer subapical multi–toothed spurs. Mesotibiae and metatibiae armed with lateral spines, with apical crown of spines (two external spines distinctly longer) of unequal length (Gnaspini et al. 2020) and two internal apical multi–toothed spurs.</p> <p>Genitalia. Aedeagus (Fig. 3) robust, elongate, sclerotised. Inner sac of median lobe elongate and tubular with developed chitinized structures along its length. Aedeagus 1.37 mm long, median lobe maximum width 0.42 mm, basal lamina maximum width 0.59 mm. Median lobe from dorsal aspect sub–parallel, then slightly widened and distinctly narrowed towards rounded apex which is widely beak–shaped and longer than parameres. Parameres elongate, thin, arcuate, curved inward apically, shorter than median lobe, slightly curved before apex club which bears one apical and two lateral short setae. Basal lamina of the median lobe with large posterior expansion. Inner sac of aedeagus (endophallus) consists of apical, medial and basal sections: comparatively short and weakly sclerotized apical reinforcement bands, more sclerotized on its inner margins, medially positioned curved sclerotized structures, feather–like structures, connection nodules and basal phanera; ejaculatory duct is illustrated. Spermatheca with basal, narrow medial and apical regions (Fig. 4). Female ventrite VIII densely pubescent with anterior expansion stout. Each style with four setae, stylus cylindrical, with one long seta.</p> <p>Sexual dimorphism: Female slightly larger than male.Antennal length 2.34 mm in male and 2.27 mm in females. Antennomere 8, ratio L/W: 1.10 in male and 0.91 in females. Shape of mesoventral carina variable.</p> <p>Differential diagnosis and remarks. Blattochaeta brankojalzici sp. nov. differs from other species by antennomere 8 approximately as long as wide (slightly longer than wide in male L/W: 1.10 and slightly wider than long in females L/W: 0.91), versus antennomere 8 longer than wide (L/W: 1.22 in males, 1.09 in females, Blattochaeta peterhlavaci sp. nov.), or antennomere 8 much longer than wide (L/W: 1.72 in male, Blattochaeta remyi; L/W: 1.66 in male, Blattochaeta montenegrina; L/W: 1.80–2.02, Blattochaeta marianii kusijanovici; L/W: 1.50–2.50, studied Blattochaeta marianii ssps.; L/W: approximately 2.00, Blattochaeta matchai, according Jeannel, 1924). Related available species of the genus can be separated from Blattochaeta brankojalzici sp. nov. further as follows:</p> <p>– Blattochaeta peterhlavaci sp. nov. by: denser punctation on elytra, which is approximately 20–25 punctures across the mid–width of each elytron (versus approximately 18–20 punctures in Blattochaeta brankojalzici sp. nov.); basal lamina maximum width 0.49 mm (versus basal lamina maximum width 0.59 in Blattochaeta brankojalzici sp. nov.); spermatheca with only slightly narrow medial region (versus spermatheca with distinctly narrow medial region in Blattochaeta brankojalzici sp. nov.); other differences in aedeagi and spermathecae of both species are shown in the accompanying figures.</p> <p>– Blattochaeta remyi by: total body length approximately 4.30 mm in male, length approximately 3.80 mm in male (versus total body length 4.55–5.00 mm, length 4.05–4.40 mm in new species); EW: 1.94 mm in male (versus EW: 2.00– 2.28 mm in new species); aedeagus length 1.10 mm (versus aedeagus length 1.37 mm in new species); basal lamina maximum width 0.41 mm (versus basal lamina maximum width 0.59 mm in new species); median lobe maximum width 0.34 mm (versus median lobe maximum width 0.42 mm in new species); parameres slightly longer than median lobe (versus parameres shorter than median lobe in new species); other differences in aedeagi of both species are shown in the accompanying figures.</p> <p>– Blattochaeta montenegrina by: antennal length 2.85 mm in male (versus antennal length 2.27–2.34 mm in new species); EW: 2.34 in male (versus EW: 2.00– 2.28 mm in new species); sparse punctation on elytra, which is approximately 15 punctures across the mid–width of each elytron (versus approximately 18–20 punctures in new species); other differences in aedeagi of both species are shown in the accompanying figures.</p> <p>– Blattochaeta marianii ssps. and B. matchai by: antennal length 2.50–3.20 mm (versus antennal length 2.27– 2.34 mm in new species); basal lamina maximum width 0.46–0.50 mm (versus basal lamina maximum width 0.59 mm in new species); total body length 4.85–5.80 mm, length 4.20–5.00 mm (versus total body length 4.55–5.00 mm, length 4.05–4.40 mm in new species); EW: 2.30–2.70 mm (versus EW: 2.00– 2.28 mm in new species).</p> <p>Etymology. Dedicated to our dear friend biospeleologist Branko Jalžić (Zagreb, Croatia), Croatiaʼs authority on speleobiology, speleology and speleodiving.</p> <p>Distribution and natural history. The new species is known only from the type locality, the Blažova pećina cave (Fig. 15), coordinates 43°04‘26.3“N, 19°21‘43.1“E (WGS 84), situated at the elevation of 1451 m a.s.l., and located near the villages Njegovuđa and Rudanca in northern Montenegro, Sinjajevina Mts. The holotype was found on the cave wall, about one meter above surface and approximately 50 meters from the cave entrance. Two aditional females were attracted to the pitfall trap. The bait used was a combination of rotten fish and blue cheese, with a saturated solution of vinegar and salt as preservative. Topography of the Blažova pećina cave was provided in Njunjić et al. 2015.</p> <p>Associated subterranean Coleoptera from the type locality: Anthroherpon sinjajevina Njunjić et al. 2015.</p> </div>	http://treatment.plazi.org/id/03A09A04A731FF976EDCFB6A2902FC6C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Čeplík, Dávid;Lakota, Ján;Čeplík, Jaroslav	Čeplík, Dávid, Lakota, Ján, Čeplík, Jaroslav (2021): Two new species of the genus Blattochaeta Reitter, 1910 (Insecta, Coleoptera Leiodidae, Cholevinae, Leptodirini) from Montenegro. Zootaxa 4969 (1): 86-100, DOI: https://doi.org/10.11646/zootaxa.4969.1.4
03A09A04A736FF976EDCFBA72F60FA40.text	03A09A04A736FF976EDCFBA72F60FA40.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Blattochaeta hawelkai Knirsch 1929	<div><p>Blattochaeta hawelkai Knirsch, 1929</p> <p>Remarks: Blattochaeta hawelkai was described from its type locality (according the catalogue) located in Ledenice Mts., Montenegro. Despite our effort to study Knirsh‘s type material deposited in the collection in The Field Museum of Natural History (Chicago, Illinois, USA), we were not successful due to Covid–19 restrictions. Ledenice Mts. is considerably distant from other localities of Blattochaeta. It is approximately 50 km from the type locality of the Blattochaeta brankojalzici sp. nov. and 100 km from the type locality of the Blattochaeta peterhlavaci sp. nov. Large geographical distances have been shown to represent natural barriers to the possibility of spreading a particular species of subterranean insect. Consequently, in most cases, the particular geographical populations represent separate species. With this in mind, we here describe two new species of Blattochaeta without their comparison with Blattochaeta hawelkai Knirsch, 1929.</p> </div>	http://treatment.plazi.org/id/03A09A04A736FF976EDCFBA72F60FA40	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Čeplík, Dávid;Lakota, Ján;Čeplík, Jaroslav	Čeplík, Dávid, Lakota, Ján, Čeplík, Jaroslav (2021): Two new species of the genus Blattochaeta Reitter, 1910 (Insecta, Coleoptera Leiodidae, Cholevinae, Leptodirini) from Montenegro. Zootaxa 4969 (1): 86-100, DOI: https://doi.org/10.11646/zootaxa.4969.1.4
03A09A04A736FF976EDCF98B29F5F884.text	03A09A04A736FF976EDCF98B29F5F884.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Blattochaeta marianii Reitter 1910	<div><p>Blattochaeta marianii Reitter, 1910</p> <p>Material studied. 2 ♂♂, 2 ♀♀: Montenegro, Krivošije Mts., Minos jama pit, VIII.2005, E. Quéinnec lgt.; 2 ♂♂, 2 ♀♀: Montenegro, Krivošije Mts., Ericova jama pit, 09.10.2019, B. Jalžić lgt.; 1 ♂: Montenegro, Orjen Mts., an unknown cave, 1913, K. Absolon lgt.; 1 ♂: Montenegro, Krivošije Mts., pećina Na Pode cave, K. Absolon lgt.; 3 ♂♂, 2 ♀♀: Montenegro, Dragaljsko polje, Bucovica Mts., Moss pit, 12.09.2004 – 11.05.2005, R. Mlejnek lgt.; 2 ♂♂, 2 ♀♀: Bosnia and Herzegovina, Orjen Mts., Mijatova Torina pećina, 19.07.2008, D. Čeplík lgt. (all in CDC).</p> </div>	http://treatment.plazi.org/id/03A09A04A736FF976EDCF98B29F5F884	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Čeplík, Dávid;Lakota, Ján;Čeplík, Jaroslav	Čeplík, Dávid, Lakota, Ján, Čeplík, Jaroslav (2021): Two new species of the genus Blattochaeta Reitter, 1910 (Insecta, Coleoptera Leiodidae, Cholevinae, Leptodirini) from Montenegro. Zootaxa 4969 (1): 86-100, DOI: https://doi.org/10.11646/zootaxa.4969.1.4
03A09A04A736FF966EDCF8CF2EE8FD98.text	03A09A04A736FF966EDCF8CF2EE8FD98.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Blattochaeta marianii subsp. kusijanovici Polak & Jalzic 2009	<div><p>Blattochaeta marianii kusijanovici Polak &amp; Jalžić, 2009</p> <p>Material studied. Paratype, 1 ♀: Croatia, Sniježnica Mts., Glogova jama pit, 24.08.2005, B. Jalžić lgt., (CDC).</p> <p>Remarks: Blattochaeta marianii with its subspecies is recorded e.g., Pretner (1977), Polak &amp; Jalžić (2009) and were collected with additional recent finds in many caves and pits, approximately more than twenty speleological localities in Orjen Mts. and Krivošije Mts. in southwestern Montenegro and several localities in southeastern Bosnia and Herzegovina, except only B. marianii kusijanovici which is currently known only from its type locality in Croatia. Specimens are deposited in different museums and private collections. The type locality of some Blattochaeta marianii ssps., and of Blattochaeta matchai Jeannel, 1924 are not exact, since the collectors, for example Leo Weirather, for unknown personal reasons, in the past often did not provide to the species authors the correct and real names of type localities (caves and pits). This has already been mentioned by Pretner (1977) and Polak &amp; Jalžić (2009). Based on the above, it would be necessary to collect as much fresh material as possible from most localities from Orjen and Krivošije Mts., for detailed study of all morphological external and internal characters of species to resolve the status and variability of these taxa. Currently, due to the lack of sufficient quantity of material we are not able to resolve here this taxonomic problem.</p> </div>	http://treatment.plazi.org/id/03A09A04A736FF966EDCF8CF2EE8FD98	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Čeplík, Dávid;Lakota, Ján;Čeplík, Jaroslav	Čeplík, Dávid, Lakota, Ján, Čeplík, Jaroslav (2021): Two new species of the genus Blattochaeta Reitter, 1910 (Insecta, Coleoptera Leiodidae, Cholevinae, Leptodirini) from Montenegro. Zootaxa 4969 (1): 86-100, DOI: https://doi.org/10.11646/zootaxa.4969.1.4
03A09A04A737FF966EDCFDF32936FD70.text	03A09A04A737FF966EDCFDF32936FD70.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Blattochaeta montenegrina Jeannel 1930	<div><p>Blattochaeta montenegrina Jeannel, 1930</p> <p>(Fig. 14)</p> <p>Material studied. 1 ♂: Montenegro, Lovćen Mts., Boljanovica jama pit, 26.06.2001, D. Čeplík lgt., (CDC).</p> </div>	http://treatment.plazi.org/id/03A09A04A737FF966EDCFDF32936FD70	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Čeplík, Dávid;Lakota, Ján;Čeplík, Jaroslav	Čeplík, Dávid, Lakota, Ján, Čeplík, Jaroslav (2021): Two new species of the genus Blattochaeta Reitter, 1910 (Insecta, Coleoptera Leiodidae, Cholevinae, Leptodirini) from Montenegro. Zootaxa 4969 (1): 86-100, DOI: https://doi.org/10.11646/zootaxa.4969.1.4
03A09A04A737FF9D6EDCFCDB2810FBD8.text	03A09A04A737FF9D6EDCFCDB2810FBD8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Blattochaeta peterhlavaci D. Ceplik, Lakota & J. Ceplik 2021	<div><p>Blattochaeta peterhlavaci D. Čeplík, Lakota &amp; J. Čeplík, sp. nov.</p> <p>(Figs 5–10, 12a–12b)</p> <p>Material studied. Holotype, ♂: MONTENEGRO, two labels: „ Montenegro, Rumija Mts., Phoenix pit, 06.08.2005, R. Mlejnek lgt.“ (white printed); „ HOLOTYPE Blattochaeta peterhlavaci sp. nov., D. Čeplík, Lakota &amp; J. Čeplík det., 2021“ (red printed), (CNHM). Paratypes: 11 ♂♂, 4 ♀♀, 41 specimens sex not determined, same data as for the holotype; 7 ♂♂, 5 ♀♀: „Montenegro, Rumija Mts., White Virgin pit, 10.08.2005; 15 ♂♂, 9 ♀♀, 37 specimens sex not determined: „ Montenegro, Rumija Mts., Ice Virgin pit, 19.05– 01.08.2005, 07.08.2005 – 27.05.2006 traps; 1 ♂, labelled: „ Montenegro, Rumija Mts., Little Virgin pit, 05.08.2005; all R. Mlejnek lgt.“ (CDC, CJL, CPH, CJC). Paratypes are labelled with red printed labels „ PARATYPE Blattochaeta peterhlavaci sp. nov., D. Čeplík, Lakota &amp; J. Čeplík det., 2021“.</p> <p>Description. Body medium–sized, bathyscioid, elliptically elongate, convex, weakly pigmented, with fine pubescence and punctures, colour from yellowish–brown to reddish–brown. Body length 4.45–4.67 mm in males, 4.50–4.85 mm in females. L: 3.83–4.22 mm. Head (Fig. 10) approximately as long as wide or slightly longer than wide (due to the measuring technique of the head capable of being retracted), HL: 0.74 mm, HW: 0.74 mm, eyes completely absent, with posterior neck region and genae more or less glabrous. Transverse occipital crest present but weakly–defined. Clypeus subhexagonal, clypeofrontal suture distinct, frontoclypeus densely setose, labrum densely setose. Mandibles short, robust, with proximal and subapical incisors and one or two small teeth between. Maxillary ultimate palpomere conical, apically pointed, slender and much shorter than the thick penultimate palpomere. Antennae (AL: 2.00– 2.28 mm), finely pubescent (Figs 8a–8b), inserted in the middle of the median third of the head. Pedicel approximately 1.60 times longer than scape. Antennomere 8 slightly longer than wide (L/W: 1.22 in males, 1.09 in females). Relative length of antennomeres (male): (1) 0.17, (2) 0.25, (3) 0.18, (4) 0.15, (5) 0.18, (6) 0.13, (7) 0.27, (8) 0.11, (9) 0.23, (10) 0.24 and (11) 0.37. Relative width of antennomeres (male): (1) 0.08, (2) 0.08, (3) 0.07, (4) 0.07, (5) 0.08, (6) 0.08, (7) 0.13, (8) 0.09, (9) 0.14, (10) 0.14 and (11) 0.13. Antennomere length/width ratios (male): (1) 2.12, (2) 3.12, (3) 2.57, (4) 2.14, (5) 2.25, (6) 1.62, (7) 2.07, (8) 1.22, (9) 1.64, (10) 1.71 and (11) 2.84. Relative length of antennomeres (female): (1) 0.17, (2) 0.29, (3) 0.18, (4) 0.15, (5) 0.18, (6) 0.14, (7) 0.26, (8) 0.12, (9) 0.20, (10) 0.20 and (11) 0.33. Relative width of antennomeres (female): (1) 0.09, (2) 0.10, (3) 0.08, (4) 0.08, (5) 0.09, (6) 0.10, (7) 0.12, (8) 0.11, (9) 0.13, (10) 0.13 and (11) 0.14. Antennomere length/width ratios (female): (1) 1.88, (2) 2.90, (3) 2.25, (4) 1.87, (5) 2.00, (6) 1.40, (7) 2.16, (8) 1.09, (9) 1.53, (10) 1.53 and (11) 2.35.</p> <p>Pronotum (Fig. 6) trapezoidal, strongly convex, wider than long, widest at base (PL/PW: 0.52–0.60), lateral edges evenly rounded, at posterior margin widest, slightly protruding backwards, narrower than elytra. PL: 1.03–1.20 mm. PW: 1.96–2.00 mm. Dorsal surface strongly pubescent. Prosternal ventral surface sparsely pubescent.</p> <p>Elytra (Fig. 7) elongately oval, pubescent, with maximum width at mid–length, longitudinal parasutural striae absent. EL: 2.80–3.02 mm. EW: 2.11–2.22 mm. EL/EW: 1.32–1.43. Punctation very dense, consists approximately of 20–25 punctures across the mid–width of each elytron, weak punctuation and pubescence on apical part. Scutellum widely triangular and pubescent.</p> <p>Venter. Mesoventral and metaventral surface sparsely pubescent. Mesocoxal cavities separated, mesoventral process well–developed. Metacoxae separated by bifid posterior metaventral process. Mesoventral carina (Figs 12a–12b) developed, prolonged posteriorly, not extended over metaventrite, with backwardly oriented setae. Sternites pubescent with short, fine setae, with reticulate and leathery aspect.</p> <p>Protarsi with four slender, undilated segments in both sexes, first protarsomere slightly longer than second and third, fourth longest. Mesotarsi and metatarsi pentamerous in both sexes. Claws wide and well–developed, empodium with one bifurcate seta. Protibiae armed with spines, one very distinct external subapical spine, internal subapical side with one shorter and one longer subapical multi–toothed spurs. Mesotibiae and metatibiae armed with lateral spines, with apical crown of spines (two external spines distinctly longer) of unequal length (Gnaspini et al. 2020) and two internal apical multi–toothed spurs.</p> <p>Genitalia. Aedeagus (Fig. 9) robust, elongate, sclerotised. Inner sac of median lobe elongate and tubular with developed chitinized structures along its length. Aedeagus 1.34–1.37 mm long, median lobe maximum width 0.44 mm, basal lamina maximum width 0.49 mm. Median lobe from dorsal aspect sub–parallel, then gradually narrower and distinctly narrowed towards pointed apex which is beak–shaped and longer than parameres. Pointed, more or less sinuate and more or less curved beak–shaped apex of the median lobe is well–visible from lateral aspect. Parameres elongate, thin, arcuate, curved inward apically, shorter than median lobe, curved before apex club which bears one apical and two lateral short setae. Basal lamina of the median lobe with large posterior expansion. Inner sac of aedeagus (endophallus) consists of apical, medial and basal sections: comparatively long and weakly sclerotized apical reinforcement bands, more sclerotized on its curved inner margins, medially positioned curved sclerotized structures, feather–like structures, connection nodules and basal phanera; ejaculatory duct is illustrated. Spermatheca with basal, only slightly narrow medial and apical regions (Fig. 5). Female ventrite VIII densely pubescent with anterior expansion stout. Each style with four setae, stylus cylindrical, with one long seta.</p> <p>Sexual dimorphism:Female slightly larger than male.Antennal length 2.22–2.28 mm in males and 2.00– 2.22 mm in females. Antennomere 8, ratio L/W: 1.22 in males and 1.09 in females. Shape of mesoventral carina variable.</p> <p>Intra–species morphological variability (inside the population from the type locality, between and inside populations from all recorded localities): slightly variable shape of the elytra; more or less variable shape of the mesoventral carina (visible from lateral aspect); slightly variable structures of inner sac of the median lobe; slightly variable shape of posterior expansion of the aedeagus; variable shape of more or less sinuate–more or less curved beak–shaped apex of the median lobe (visible from lateral aspect); slightly variable shape of the spermatheca.</p> <p>Differential diagnosis and remarks. Blattochaeta peterhlavaci sp. nov. differs from other species by antennomere 8 slightly longer than wide (L/W: 1.22 in males, 1.09 in females) versus antennomere 8 approximately as long as wide (in male L/W: 1.10, in females L/W: 0.91, Blattochaeta brankojalzici sp. nov.), or antennomere 8 much longer than wide (L/W: 1.72 in male, Blattochaeta remyi; L/W: 1.66 in male, Blattochaeta montenegrina; L/W: 1.80–2.02, Blattochaeta marianii kusijanovici; L/W: 1.50–2.50, other studied specimens of Blattochaeta marianii ssps.; L/W: approximately 2.00, Blattochaeta matchai, from Jeannel, 1924). Related available species of the genus can be separated from Blattochaeta peterhlavaci sp. nov. further as follows:</p> <p>– Blattochaeta brankojalzici sp. nov. by: sparse punctation on elytra, which is approximately 18–20 punctures across the mid–width of each elytron (versus dense, approximately 20–25 punctures in Blattochaeta peterhlavaci sp. nov.); basal lamina maximum width 0.59 mm (versus basal lamina maximum width 0.49 in Blattochaeta peterhlavaci sp. nov.); spermatheca with distinctly narrow medial region (versus spermatheca with only slightly narrow medial region in Blattochaeta peterhlavaci sp. nov.); other differences in aedeagi and spermathecae of both species are shown in the accompanying figures.</p> <p>– Blattochaeta remyi by: total body length approximately 4.30 mm in male, length approximately 3.80 mm in male (versus total body length 4.45–4.85 mm, length 3.83–4.22 mm in new species); EW: 1.94 mm in male (versus EW: 2.11–2.22 mm in new species); sparse punctation on elytra, which is approximately 18–20 punctures across the mid–width of each elytron (versus dense, approximately 20–25 punctures in new species); antennal length 2.40 mm in male (versus antennal length 2.00– 2.28 mm in new species); aedeagus length 1.10 mm (versus aedeagus length 1.34–1.37 mm in new species); basal lamina maximum width 0.41 mm (versus basal lamina maximum width 0.49 mm in new species); median lobe maximum width 0.34 mm (versus median lobe maximum width 0.44 mm in new species); parameres slightly longer than median lobe (versus parameres shorter than median lobe in new species); other differences on aedeagi of both species are shown in the accompanying figures.</p> <p>– Blattochaeta montenegrina by: total body length approximately 5.00 mm in male, length approximately 4.30 mm in male (versus total body length 4.45–4.85 mm, length 3.83–4.22 mm in new species); EW: 2.34 in male (versus EW: 2.11–2.22 mm in new species); antennal length 2.85 mm in male (versus antennal length 2.00– 2.28 mm in new species); sparse punctation on elytra, approximately 15 punctures across the mid–width of each elytron (versus denser punctation on elytra, approximately 20–25 punctures in new species); basal lamina maximum width 0.58 mm (versus basal lamina maximum width 0.49 mm in new species); other differences on aedeagi of both species are shown in the accompanying figures.</p> <p>– Blattochaeta marianii ssps. and B. matchai by: sparse punctation on elytra, approximately 18–20 punctures across the mid–width of each elytron (versus denser punctation on elytra, approximately 20–25 punctures in new species); antennal length 2.50–3.20 mm (versus antennal length 2.00– 2.28 mm in new species); total body length 4.85–5.80 mm, length 4.20–5.00 mm (versus total body length 4.45–4.85 mm, length 3.83–4.22 mm in new species); EW: 2.30–2.70 mm (versus EW: 2.11–2.22 mm in new species).</p> <p>Etymology. Dedicated to our dear friend Peter Hlaváč (Prague, Czech Republic), a specialist for selected groups of Staphylinidae and Curculionidae and member of Biospeleologica Slovaca.</p> <p>Distribution and data. The new species is known from its type locality, the Phoenix pit (– 70 m), at the elevation of approximately 1100 m a.s.l., and from the nearby White Virgin pit, Ice Virgin pit and Little Virgin pit in Rumija Mts., southeastern Montenegro. The Phoenix pit was described in Lohaj et al. 2016. The new species probably inhabits more speleological localities situated in Rumija Mts. Here are stated translated data from Pretner (1977: 127, 139, 15): “a small female of Blattochaeta, perhaps new species, from a little pit, snow cave near Miligrepa, Mali Mikulići, Rumija Mts, 05.07.1960, Dušan Gavrilovićˮ. This specimen was not studied for this work.</p> <p>Associated subterranean Coleoptera from the type locality: Adriaphaenops rumijaensis Lohaj et al. 2016, Neotrechus suturalis ssp. Schaufuss, 1864, Laemostenus (Antisphodrus) cavicola ssp. Schaum, 1858, Anthroherpon matulici Reitter, 1903, and Anthroherpon taxi ssp. Müller, 1913, from Lohaj et al. 2016.</p> </div>	http://treatment.plazi.org/id/03A09A04A737FF9D6EDCFCDB2810FBD8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Čeplík, Dávid;Lakota, Ján;Čeplík, Jaroslav	Čeplík, Dávid, Lakota, Ján, Čeplík, Jaroslav (2021): Two new species of the genus Blattochaeta Reitter, 1910 (Insecta, Coleoptera Leiodidae, Cholevinae, Leptodirini) from Montenegro. Zootaxa 4969 (1): 86-100, DOI: https://doi.org/10.11646/zootaxa.4969.1.4
03A09A04A73CFF9D6EDCFB0629ECFAA7.text	03A09A04A73CFF9D6EDCFB0629ECFAA7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Blattochaeta remyi Jeannel 1931	<div><p>Blattochaeta remyi Jeannel, 1931</p> <p>(Fig. 13)</p> <p>Material studied. 1 ♂: Montenegro, Lubnice, Berane, Županska pećina cave, 14.07.2008, D. Čeplík lgt., (CDC).</p> </div>	http://treatment.plazi.org/id/03A09A04A73CFF9D6EDCFB0629ECFAA7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Čeplík, Dávid;Lakota, Ján;Čeplík, Jaroslav	Čeplík, Dávid, Lakota, Ján, Čeplík, Jaroslav (2021): Two new species of the genus Blattochaeta Reitter, 1910 (Insecta, Coleoptera Leiodidae, Cholevinae, Leptodirini) from Montenegro. Zootaxa 4969 (1): 86-100, DOI: https://doi.org/10.11646/zootaxa.4969.1.4
