taxonID	type	description	language	source
E101B54FFFF49251FEB74A89FC36FD01.taxon	type_taxon	Type species: Haltica muscorum KOCH, 1803, by monotypy.	en	Nadein, Konstantin S. (2009): Revision of the genus Mniophila S, 1831 (Coleoptera: Chrysomelidae). Beiträge Zur Entomologie = Contributions to Entomology 59 (1): 103-131, DOI: 10.21248/contrib.entomol.59.1.103-131, URL: https://www.contributions-to-entomology.org/article/view/1742
E101B54FFFF49251FEB74A89FC36FD01.taxon	distribution	Distribution: Range of distribution of the genus Mniophila embraces Western and Central Europe, Balkans, Crimea, Caucasus, and Turkey (Figs 8 - 10). Mniophila muscorum has the widest range (Fig. 8), from Pyrenees (Spain) to Carpathians (Ukraine). The northern boundary of this species is in Dania, Norway, and Sweden. The rest of the species have ranges smaller than Mniophila muscorum. It is worthy to mention the high level of endemism among the species of Mniophila. The only ranges of M. muscorum and M. bosnica, and probably, M. caucasica sp. n. and M. transcaucasica sp. n. are partly overlapping (Figs 8 - 10).	en	Nadein, Konstantin S. (2009): Revision of the genus Mniophila S, 1831 (Coleoptera: Chrysomelidae). Beiträge Zur Entomologie = Contributions to Entomology 59 (1): 103-131, DOI: 10.21248/contrib.entomol.59.1.103-131, URL: https://www.contributions-to-entomology.org/article/view/1742
E101B54FFFF49251FEB74A89FC36FD01.taxon	description	Redescription: Body dark, brown to black, shining, with greenish or bronzy metallic luster or without one; sometimes in brown individuals pronotum and elytra with light margin; sometimes partly coloured individuals occur; legs and antennae light to dark brown. Body elliptical and moderately convex to clearly rounded and rather convex (Figs 1 A-E; 3 A-C; 4 A; 5 A, B; 6 A-D; 7 A); head not or barely visible from above. Head (Figs 1 F; 4 H; 5 C; 7 E) large; vertex large, wide, its surface covered with weakly developed smooth shagrination coarse, well developed, grainy shagrination. Ocular sulci not developed. Frontal calli triangular or rhomboidal, small, its surface smoother than that of the vertex and frons; separated from vertex by thin supracallinal sulci. Frontal ridge distinct, convex, triangular and flattened basally. Anterofrontal ridge weakly convex, straight. Antennal grooves large and deep, covered with large and coarse shagrination. Frontal part of head moderately short to long. Antennal sockets situated closely to each other, separated from eye margin by impressed interval in length is almost equal to socket. Eyes small, elliptical, flattened to convex, widely spaced. Antennae (Figs 1 M, 3 D; 4 G; 5 E; 6 K; 7 D) short; three apical segments distinctly widened; remaining segments thinner; eighth antennomere thinner and smaller than seventh; first and second antennomeres clearly longer and thicker than following five segments. Labrum small, with deep frontal notch situated medially. Pronotum (Figs 1 G; 3 E, 4 B; F; 6 E; 7 B) large, widely transverse. Apical and basal edges with rather thin margins; lateral margin wider. Anterofrontal callosity large, thickened, not forming acute denticle; posterolateral callosity smaller, poorly developed. Basal edge of pronotum curved in sides, distinctly rounded medially or widely rounded. Pronotal surface covered with visible, large shagrination. Punctation usually present, not coarse, sparse, and shallow, poorly visible on pronotal microsculpture, rarely impunctate. Elytra slightly elongate with elongate elytral apices to clearly rounded, without elongate apices. Punctation variable: impunctate to punctured densely; usually punctures arranged in regular or partly confused striae, depth and frequency, and size of punctures variable; secondary punctation from undeveloped to well developed, often as large as punctures in striae and dense. Elytral interstices smooth. Hind wings completely reduced. Pro-, meso-, and metathorax strongly shortened; mesothorax shorter than pro- and metathorax. Procoxal cavities open. Legs (1 I-L; 3 F-I; 4 C-F; 5 G-J; 6 F, G, I, J; 7 F-I) not long, usually thick, rarely thinner; first protarsomere of male larger and thicker than in female; hind tibiae and often that of middle and foreleg curved, sometimes fore and middle tibiae straight; hind femora usually wide (Fig. 1 H), rarely thin (Fig. 5 D). Aedeagus (Figs 2, 3 J, K; 4 I, J; 5 K-M; 6 L-N; 7 J, K) rather large, curved basally, apical 2 / 3 almost straight to distinctly curved; apex shape variable: rounded, widened, elongated and narrowing, with or without apical denticle. Spermatheca small, collo large, thick, straight, nodulus comparatively narrow, not very long, duct short, moderately curved, not forming loops, ramus globose, globular. Body length: 1.1 - 1.6 mm, width – 0.8 - 1.2 mm. Preimaginal stages. Egg and first instar larva have been described by COX (1997).	en	Nadein, Konstantin S. (2009): Revision of the genus Mniophila S, 1831 (Coleoptera: Chrysomelidae). Beiträge Zur Entomologie = Contributions to Entomology 59 (1): 103-131, DOI: 10.21248/contrib.entomol.59.1.103-131, URL: https://www.contributions-to-entomology.org/article/view/1742
E101B54FFFF1925EFF0B48FBFDDEFEDA.taxon	description	(Figs 1, 2, 8)	en	Nadein, Konstantin S. (2009): Revision of the genus Mniophila S, 1831 (Coleoptera: Chrysomelidae). Beiträge Zur Entomologie = Contributions to Entomology 59 (1): 103-131, DOI: 10.21248/contrib.entomol.59.1.103-131, URL: https://www.contributions-to-entomology.org/article/view/1742
E101B54FFFF1925EFF0B48FBFDDEFEDA.taxon	materials_examined	Type material: Type material is lost according to DOGUET (1994). To prevent confusion of this taxon I here designate a neotype of Mniophila muscorum in order to have a unique bearer of this name and the standard for its application. A neotype specimen, male, from Bulgaria with labels: 1. BG: Stara Planina, Botev-Massiv, N-Seite, Kar an der Pobita Glava, 1775 m; 2. unter Juniperus, Moss 42 ° 43 ' 36 '' N 24 ° 53 ' 38 '' O 27. V. 2000 leg. Zerche. A neotype is deposited in DEI. Type locality: Stara Planina Mountain Range: Botev-Massiv (Bulgaria). Other material examined: Great Britain: UK: W. Sussex Rewell Hill. 16. ix. 2003 P. G. Booth, 4 specimens (NHML). – Oxford Kent 26.3.70 / in moss, 4 specimens (NHML). – Westerham 27. V. 1923 CET., 18 specimens (NHML). – H. Donisthorpe B. M. 1934 - 4, 2 specimens (NHML). – Chatham Distr. J. J. W., 5 specimens (NHML). – Barcombe 4. - 24, 1 specimen (NHML). – Chatham Kent J. J. W., 6 specimens (NHML). – Westmorland Ambleside ix. 1969 / in moss, 3 specimens (NHML). – Tavertham Kent J. J. W., 2 specimens (NHML). France: Tours Desbroch., 3 specimens (DEI). – France: Htes Pyr. 1856 – 1875 Coll. L. Pandellé, 5 specimens (IRSNB). – France: Htes Pyr. H. Pyrénée 1856 – 1875 Coll. L. Pandellé, 5 specimens (IRSNB). – Tarbes, 3 specimens (LM). – France: Nièvre Vallée de Sardy / leaf litter / P. M. Hammond 8 - 20. viii. 1982 B. M. 1982 - 354, 4 specimens (NHML). – Tarbes, 3 specimens (ZIN). – Vosges. / (Bône) [mislabelled], 4 specimens (ZIN). – 3. M. 746. Gall. M. muscorum, 3 specimens (ZIN). – St. Béat H. - Garonne, 7 specimens (ZIN). – France Chamonix 200 m / 19. VIII. 1974 leg. S. Mahunka, 3 specimens (HNHM). – Moisd., 9 specimens (DEI). – Corsica 19.6.09, 1 specimen (DEI). – Vizzavona Corsica 19.6.09, 1 specimen (DEI). – Corsica Diener, 1 specimen (HNHM). – Corsica, 1 specimen (MTD). Spain: Mniophila muscorum Altos Pirineos col. Perez Arcas, 2 specimens (MNCN). Belgium: Relgie Nemen / Mon-sur-Lesse 19.21 - V- 1977 N. I. V. Krift, 3 specimens (ZMUA). Germany: Thür. Wald gebel 4.6. 22 W. Liebmann, 5 specimens (DEI). – Ob. Bayern Umg. Königsee 2.8.21, 2 specimens (DEI); the same, 21.7.21, 3 specimens (DEI); the same, 26.7.21, 2 specimens (DEI); the same, 27.7.21, 1 specimen (DEI); the same, 30.7.21, 1 specimen (DEI). – Ramsau / Obb / 14.6.1962 Willi Lucht, 1 specimen (HNHM). – Grünwald 28.11.49 / München Bühlmann, 1 specimen (DEI). – Germania Rhön 16.5. 21 W. Liebmann, 1 specimen (DEI). – Germ. m., 1 specimen (DEI). – Germ. c., 1 specimen (DEI). – Boppard, 2 specimens (DEI). – Wechley Oldenburg 25. VI. 91, 1 specimen (DEI). – Eifel Boellger, 1 specimen (DEI). – Ems v. Heyden Nassau, 1 specimen (DEI). – aenea Melsheimer Heidelberg Jenisson, 1 specimen (DEI). – Bayern Muscorum Kork., 2 specimens (ZMUC). – Gees Eifel 14. VII. 1972 S. v. Helinsbergen, 1 specimen (ZMUA). – P. Lamy Hanau, 5 specimens (ZIN). – Mniophila muscorum E. H. German. Dohrn, 1 specimen (ZMKU). – Heidelbg. Haag., 4 specimens (HNHM). – Welle, 1 specimen (MTD). – Rein SK x. 92, 3 specimens (MTD). – Heidelberg, 2 specimens (MTD). – Preetz, 2 specimens (MTD). – Ulm Württbg, 226 specimens (MTD). Poland: Silesia, 1 specimen (DEI). – Beskiden Lissahora-Geb. TH. v. Wanka, 5 specimens (DEI). – Westpreussen, 4 specimens (DEI). – Beskiden Reitter, 2 specimens (ZMUA). – Beskiden Lisahora- Geb. TH. v. Wanka, 1 specimen (ZMUA). – Beskiden Lissagebiet TH. v. Wanka, 2 specimens (ZMUA). – Bolechow VIII. 1924, 1 specimen (HNHM). – Polonia Bolechow. VIII. 1926, 1 specimen (HNHM). – Silesia coll. Lichtn., 1 specimen (HNHM). – Tatra, 1 specimen (MTD). – Glatz-Geb., 1 specimen (MTD). – G. Schneebrg, 1 specimen (DEI). – Lysahora, 1 specimen (MTD). Switzerland: Pilatus, 3 specimens (DEI). – Switzerland Crans: Valais moss. VIII. 73, 5 specimens (NHML). – Switzerland near Zurich, Sihlwald 10 year Fagus forest barber trap 27. V. 1999 V. Chumak, 1 specimen (NC). – Helvetia Bürgenstock 880 M. X. 6.52 v. d. Wiel, 1 specimen (ZMUA). Austria: Austria, 1 specimen (DEI). – leodinensis Wesm. Tirol Rosenhauer, 1 specimen (DEI). – Stiermark, 1 specimen (ZMUC). – Miller Austria, 2 specimens (ZMUA). – Wien Reitter, 3 specimens (ZMUA). – Wechselgeb. A. i., Mader, 8 specimens (HNHM). – Umgeb. von Wien, 2 specimens (MTD). – Wien Umgeb. A. Winkler Tullnerbach, 1 specimen (NHMW). – Lunz Austria, 5 specimens (MTD). – Steiermark Stuhleck, 1 specimen (NHMW). – Holdhaus Villach, 2 specimens (NHMW). Italy: Italia. Novara Macugnana Geo. C. Krüger Coll. O. Leonhard, 1 specimen (DEI). – Certosa di Pesio Ligur. Alp. 23. VII. 11, 1 specimen (DEI). – Calabria Sta Eufemia d’Aspromonte 1905, 1 specimen (DEI). – Como-See Villa, 1 specimen (DEI). – Folgaria S. Tirol 11. VII. 14, 1 specimen (DEI). – Bozen Ti. V. Oertzen, 2 specimens (ZMUA). – Südtyrol Bozen 79 Reitter, 1 specimen (HNHM). – Castrozza, 1 specimen (HNHM). – Tauf. Sand. Schneifer, 5 specimens (MTD). Slovenia: Tr. V. Beszédes St. Radegund, 1 specimen (HNHM). Czech Republic: Altvater Letzner, 4 specimens (DEI). – Altvater, 6 specimens (DEI). – Moravia, 3 specimens (ZMUA). – Staudg. Moravia, 1 specimen (ZMUA). – Moravia Reitter, 1 specimen (ZMUA). – Altvater Mähren A. Otto, 1 specimen (ZMUA). – Fleischer Moravia, 1 specimen (ZIN). – Mniophila muscorum Koch. Fleischer Morav., 2 specimens (ZIN). – Moravia Reitter, 3 specimens (HNHM). – Moravia Besciden Reitter, 1 specimen (HNHM). – Altvater Letzner, 6 specimens (MTD). – Gesenke 7.92, 24 specimens (MTD). Rumania: Carpathes Azuga A. L. Montandon, 2 specimens (DEI). – Bullea-See Schuster 95 / Siebenbürg., 3 specimens (DEI). – Banat 1909 Herculesbad leg. M. Hilf Coll. O. Leonhard, 1 specimen (DEI). – Siebenbürg. / Negoi 95 Schuster, 1 specimen (DEI). – Siebenbürg. / Bucsecs 95 Schuster, 1 specimen (DEI). – Siebenbürgen Roterturmpass, 1 specimen (NHMW). – Bucsecs 98 Schuster, 1 specimen (ZMUA). – Transsylvania Strobl., 1 specimen (ZMUA). – Schuler-Geb. Schuster 98, 2 specimens (NHMW). – Schuler- Geb. Schuster 98, 1 specimen (SMF). – Schuler Geb. Flach. VI. 96, 1 specimen (SMF). – Rumänien Sinaia v. Bodemeyer, 1 specimen (SMF). – Deubel Tr. Rosenauer G., 1 specimen (SMF). – Transsilvania Ober Kerz Reitter, 1 specimen (ZIN). – Azuga Walachei, 9 specimens (HNHM). – Azuga Rumänien, 4 specimens (HNHM). – Flinsberg, 1 specimen (HNHM). – Rumänien Fogarasch-Geb. Barcaci 31. V. 1981, 1 specimen (MTD). – Flinsberg D. Ludy, 1 specimen (MTD). – Transsylv, 2 specimens (MTD). – Siebenbg., 6 specimens (MTD). – N. Hagymás Holdhaus, 2 specimens (NHMW). Hungary: Bokony Hungar. Heyden, 2 specimens (DEI). – Mniophila muscorum Ungarn, 1 specimen (ZIN). – Hung. 2 specimens (ZIN). Ukraine: Gadzhyna Çerna-Gora, 2 specimens (LM). – Diana-lak Podk. Rusy VI. Zoufal., 4 specimens (ZMUA). – Ukraine, E Carpathians Zakarpattya, nr. Velyka Ugol’ka Vill., karst 19. VII. 2001 N. N. Yunakov, 3 specimens (NC). – Ukraine, Zakarpatskaya obl., Marmarosh, chrebet Holovatschiu, g. Nenyaska 12.07.2000 N. Yunakov / verkhnyaja chast’ lesnogo poyasa 1800 m, elnik, v mokhovoy podstilke vozle pney, 9 specimens (NC). Croatia: Łlitvica Croatia Heyden, 4 specimens (DEI). Bosnia and Herzegovina: Bosnia 1902 Maklen-Pass O. Leonhard, 2 specimens (DEI). – Bosnia Bjelašnica Pl. O. Leonhard, 1 specimen (DEI). – Bosnia Bjelasnica – Pl. O. Leonhard, 1 specimen (DEI). – Bosnia Rovinaja Planina 1931. VII. 7. leg. Dr. J. Fodor, 1 specimen (HNHM). – Bosnia, Jahorina 1935. X. leg. Dr. J. Fodor, 1 specimen (HNHM). – Bosnien Reitter. Leder, 1 specimen (HNHM). – Trebevic. Bosnia Fodor 1929. V. 11, 1 specimen (HNHM). – Bosnia, Brdo Tresnjevik 1938. VII. 26 leg. Dr. J. Fodor, 4 specimens (HNHM). – Herzegovina Bjelasnica 1901, 2 specimens (DEI). – Herzegovina Ubli 1903 O. Leonhard, 6 specimens (DEI). – Herzegovina, 2 specimens (ZIN). Montenegro: Krivosije Paganetti, 6 specimens (DEI). – Krivosije Paganetti, 2 specimens (ZMUA). – Krivosije Paganetii, 5 specimens (HNHM). – Crna Gora, Durmitor Dolina Susice 1933. VII. 7 - 27. leg. Dr. J. Fodor, 3 specimens (HNHM). – Crna Gora, Biela Gora Trebinje 1929. VII. 25 leg. Dr. J. Fodor, 1 specimen (HNHM). – Crna Gora Zabljak 1934. VII. 18. leg. Dr. J. Fodor, 2 specimens (HNHM). – Crna Gora Han Garancic 1938. VII. 19 - 20. leg. Dr. J. Fodor, 4 specimens (HNHM). – Durmitor, Mont. 26. VI. 1958 / Crno Jezero 1400 m / leg. Kaszab & Székessy, 18 specimens (HNHM). – Durmitor, Mont. 3. VII. 1958 / Zmijinje Jezero 1400 m / leg. Kaszab & Székessy, 3 specimens (HNHM). – Durmitor, Mont. 27. VI. 1958 / Crno Jezero cribri ope 1700 m / leg. Kaszab & Székessy, 1 specimen (HNHM). Macedonia: Macedonia Crepolsko Ketsana Stjena, 4 specimens (HNHM). Bulgaria: Bulg. Tschamkorija M. Hilf 1911 Coll. O. Leonhard, 3 specimens (DEI). – BG: Stara Planina, Botev-Massiv, N-Seite, Kar an der Pobita Glava, 1775 m / unter Juniperus, Moos 42 ° 43 ' 36 '' N 24 ° 53 ' 38 '' O 27. V. 2000 leg. Zerche, 2 specimens (DEI). – BG: Ossogovska Planina, O-Gipfel des Ruen-Massivs, N- Hang, 1900 m, unter Steinen am Schneerand gesiebt 42 ° 10 ' 41 '' N 22 ° 33 ' 49 '' O 17. V. 2000 leg. Zerche, 3 specimens (DEI). – BG: Stara Planina, Botev-Massiv, Schaltez-Nordhang S Hütte Pleven, 1705 – 1730 m. Schneefelder in Lawinenrinnen, 16. V. 2001 42 ° 43 ' 43 '' N 24 ° 53 ' 16 '' O leg. Zerche & Behne, 1 specimen (DEI). – BG: Stara Planina, Kom-Massiv, Mali Kom, N-Seite Lawinenrinnen mit Schnee, 1645 – 1750 m, 43 ° 09 ' 52 '' N 23 ° 05 ' 15 '' O 6. V. 2001 leg. Zerche & Behne, 1 specimen (DEI). – BG: Rila-Gebirge, Sitnjakovo S Borovez, 1740 m, Abies, letzte Schneeflecken / 42 ° 14 ' 42 '' N 23 ° 37 ' 37 '' O 28. IV. 2001, leg. Zerche & Behne, 1 specimen (DEI). – Bulgaria. 1928 Mts. Rila. Biro. IX., 5 specimens (HNHM). – Bulgaria 1928. Rila. Biro. IX. 24, 2 specimens (HNHM). Unknown provenance: Muscorum Korch. Pyrenaei, 2 specimens (ZMUC). – Bar. Fux. Haltica muscorum E. H., 1 specimen (ZMUA). – Pyrenäen, 1 specimen (MTD). – Karpaten Scirba / Brancsik, 4 specimens (SMF).	en	Nadein, Konstantin S. (2009): Revision of the genus Mniophila S, 1831 (Coleoptera: Chrysomelidae). Beiträge Zur Entomologie = Contributions to Entomology 59 (1): 103-131, DOI: 10.21248/contrib.entomol.59.1.103-131, URL: https://www.contributions-to-entomology.org/article/view/1742
E101B54FFFF1925EFF0B48FBFDDEFEDA.taxon	distribution	Distribution: Albania, Austria, Belgium, Bosnia, Herzegovina, Bulgaria, Croatia, Czech Republic, Denmark, France, Germany, Hungary, Ireland, Italy, Luxemburg, Macedonia, Montenegro, Netherlands, Norway, Poland, Spain, Rumania, Serbia, Slovakia, Slovenia, Sweden, Switzerland, Ukraine (Carpathians).	en	Nadein, Konstantin S. (2009): Revision of the genus Mniophila S, 1831 (Coleoptera: Chrysomelidae). Beiträge Zur Entomologie = Contributions to Entomology 59 (1): 103-131, DOI: 10.21248/contrib.entomol.59.1.103-131, URL: https://www.contributions-to-entomology.org/article/view/1742
E101B54FFFF1925EFF0B48FBFDDEFEDA.taxon	description	Redescription: Body dark to black, shining, with greenish or rarely bronzy luster. Antennae and legs brown. Body shape rounded with weakly elongated elytral apices to distinctly elliptical. Head large; vertex wide; eyes gently convex to usually clearly convex. Antennae with thick segments. Head surface covered with well developed, large shagrination. Pronotum long with clearly rounded base. Pronotal surface covered with shagrination; punctation large, sparce, shallow, weakly visible. Elytra with greatly variable punctation. Usually punctation well developed, small, dense, striae confused; secondary punctation well developed, often as large as strial. Rarer strial punctation regular and secondary punctation smaller or almost indistinct. Legs thickened; tibiae straight or weakly curved; hind femora wide. First protarsomere of male usually short and moderately narrow. Aedeagus (Fig. 2) ventrally with apical third short with straight margins and nearly straight apex with denticle to elongated, narrow, with narrow apex without denticle. From lateral view apical third clearly gradually narrowed to apex, sometimes wider. Body length – 1.20 - 1.68 mm, width – 0.89 - 1.21 mm.	en	Nadein, Konstantin S. (2009): Revision of the genus Mniophila S, 1831 (Coleoptera: Chrysomelidae). Beiträge Zur Entomologie = Contributions to Entomology 59 (1): 103-131, DOI: 10.21248/contrib.entomol.59.1.103-131, URL: https://www.contributions-to-entomology.org/article/view/1742
E101B54FFFF1925EFF0B48FBFDDEFEDA.taxon	diagnosis	Differential diagnosis: Differs from M. bosnica in structure of aedeagus (Fig. 2): apical third in posterior view elongate with straight margins instead of short with round margins, apex straight or nearly straight with a denticle instead of apex rounded without denticle (Fig. 3 J, K); body more elongate (Fig. 1 A-E); eyes convex (Fig. 1 F); pronotum longer (Fig. 1 G); first metatarsomere thinner and shorter (Fig. 1 K), first protarsomere of male shorter (Fig. 1 L), colouration of the body darker, dark brown to black, usually with weak green luster; elytral punctation usually denser, punctation often confused, secondary punctation well developed, usually as large as punctures in striae. From M. taurica sp. n. differs in structure of aedeagus: apex usually elongate, with narrowing straight margins in posterior view or apical third with straight margins instead of sides toward apex parallel-sided, apical 1 / 4 with straight margins (Fig. 5 K), denticle always present and well developed instead of denticle short or poorly developed; shagrination of the head more developed; eyes more convex; legs thicker; hind femur broader (Figs 1 H, 5 D); punctation of pronotum usually more developed; differs also in the shape of pronotum.	en	Nadein, Konstantin S. (2009): Revision of the genus Mniophila S, 1831 (Coleoptera: Chrysomelidae). Beiträge Zur Entomologie = Contributions to Entomology 59 (1): 103-131, DOI: 10.21248/contrib.entomol.59.1.103-131, URL: https://www.contributions-to-entomology.org/article/view/1742
E101B54FFFFA925DFEB7498FFEA8FDCE.taxon	description	(Figs 3, 9)	en	Nadein, Konstantin S. (2009): Revision of the genus Mniophila S, 1831 (Coleoptera: Chrysomelidae). Beiträge Zur Entomologie = Contributions to Entomology 59 (1): 103-131, DOI: 10.21248/contrib.entomol.59.1.103-131, URL: https://www.contributions-to-entomology.org/article/view/1742
E101B54FFFFA925DFEB7498FFEA8FDCE.taxon	materials_examined	Type material: Syntypes: Bosnia. Trebević, Apf. / TYPUS, 1 specimen. – Zvijezda / TYPUS, 2 specimens. Type material is deposited in the National Museum of Bosnia and Herzegovina, Sarajevo. Not examined. Type locality: Bosnia (South). Other material examined: Great Britain: Power. Darenth., 8 specimens (NHML). – Surrey. Leith Hill. 16. VI. 1951. J. T. Salmon / in moss, 3 specimens (NHML). – Chatham, Kent. G. C. C., 1 specimen (NHML). – H. Donisthorpe. B. M. 1934 - 4., 3 specimens (NHML). – Oxford Kent 26.3.70 / in moss, 5 specimens (NHML). France: St. Béat H. - Garonne, 7 specimens (ZIN). Germany: Boppard, 3 specimens (DEI). Poland: Riesengebirge Letzner, 3 specimens (DEI). Austria: N-Tirol Reutte leg. Knabl, 1 specimen (DEI). – Zirbitzkogel Diener, 1 specimen (HNHM). – Vorarlb. Allgäu Riezlern 6. - 7.21, 1 specimen (HNHM). – Dobratsch Diener, 2 specimens (HNHM). Czech Republic: Moravia Reitter, 2 specimens (DEI). – Moravia, 1 specimen (ZMUA). Rumania: Ungarn Hatszeg v. Bodemeyer, 1 specimen (DEI). Italy: Italia. Novara Macugnana Geo. C. Krüger Coll. O. Leonhard, 2 specimens (DEI). – Görz 22. VII. 11 Gradisca, 1 specimen (DEI). – Calabria Sta. Eufemia d’Aspromonte 1905 Paganetti, 1 specimen (ZIN). – Calabria Aspromonte Paganetii 1905, 2 specimens (HNHM). – Calabria Aspromonte leg. Paganetti, 1 specimen (HNHM). – Foresta d. Reso (Prov. Pistoia) 5 - 6 - 971 S. Failla, 6 specimens (MZUF). – m. Amiata 24 - 5 - 1969 Castelini-Failla, 1 specimen (MZUF). – S. Vito di Cadore 13 - VIII- 972 S. Failla, 1 specimen (MZUF). – Bozen Ti. V. Oertzen, 3 specimens (ZMUA). – Bozen, 9 specimens (ZMUA). Slovenia: Oestr. Küstenland Fužina 1906 legit. M. Hilf Coll. O. Leonhard, 1 specimen (DEI). Croatia: Capela Croatia Heyden, 10 specimens (DEI). – Brušane Pawel, 10 specimens (HNHM). Bosnia: Bosnia 1909 Kladani leg. M. Hilf Coll. O. Leonhard, 2 specimens (DEI). – Bjelasnica planina Bosnien coll. Leonhard, 2 specimens (DEI). – Bosnia 1902 Maklen-Pass O. Leonhard, 1 specimen (DEI). – Bosnia Bjelasnica-Pl. O. Leonhard, 20 specimens (DEI). – Bosnia, 2 specimens (ZIN). – Bosnia, Sarajevo Bjelasnica Planina 1930. VII. 25. leg. Dr. J. Fodor, 1 specimen (HNHM). – Bosnia Rovinaja Planina 1931. VII. 7 leg. Dr. J. Fodor, 1 specimen (HNHM). – Bosnia, Jajce 1934. XI. 5. leg. Dr. J. Fodor 1929. V. 11, 1 specimen (HNHM). – Bosnia, Brdo Tresnjevik 1938. VII. 26 leg. Dr. J. Fodor, 1 specimen (HNHM). – Trebovic. Bosnia Fodor 1929. V. 11, 1 specimen (HNHM). – Bosn. Herzeg. Ivan Reitter, 1 specimen (HNHM). – Bosnia, Kamenica R. Susica, 1 specimen (HNHM). Herzegovina: Herzegovina Jablanica, 4 specimens (DEI). – Herzegovina, 1 specimen (ZIN). – Herzegovina Vucije bara Jacko. Fodor, 1 specimen (HNHM). Montenegro: Crna Gora, Biela Gora Trebinje 1929. VII. 25, leg. Dr. J. Fodor, 2 specimens (HNHM). – Durm. Mont. 26. VI. 1968 / Crno Jezero 1400 m / leg. Kaszab & Székessy, 1 specimen (HNHM). Bulgaria: BG: Slavianka (Ali-Botusch), S Goleschovo, NW Mt. Gozev Vrach, N-Hang. 1955 m. Schneefeld / einzelne Pinus, 41 ° 22 ' 26 '' N 23 ° 36 ' 28 '' O 6. V. 2000 leg. Zerche, 1 specimen (DEI). – Bulgaria 1912 Trevna V-VI leg. M. Hilf Coll. O. Leonhard, 4 specimens (DEI). – Bulg. Tschamkorija M. Hilf 1911 Coll. O. Leonhard, 4 specimens (DEI). – Bulgaria. 1928. Mt. Rila. Biro. IX., 1 specimen (HNHM). Macedonia: Macedonia Crepolsko Ketsana Stjena, 1 specimen (HNHM). Albania: Ueskueb Albania Dilon, 2 specimens (DEI). Unknown Provenance: Moldavia 1903 - 67, 1 specimen (NHML) [? mislabelled].	en	Nadein, Konstantin S. (2009): Revision of the genus Mniophila S, 1831 (Coleoptera: Chrysomelidae). Beiträge Zur Entomologie = Contributions to Entomology 59 (1): 103-131, DOI: 10.21248/contrib.entomol.59.1.103-131, URL: https://www.contributions-to-entomology.org/article/view/1742
E101B54FFFFA925DFEB7498FFEA8FDCE.taxon	distribution	Distribution: Albania, Austria, Bosnia, Bulgaria, Croatia, Czech Republic, France, Germany, Great Britain, Herzegovina, Italy, Macedonia, Montenegro, Poland, Rumania, Slovenia.	en	Nadein, Konstantin S. (2009): Revision of the genus Mniophila S, 1831 (Coleoptera: Chrysomelidae). Beiträge Zur Entomologie = Contributions to Entomology 59 (1): 103-131, DOI: 10.21248/contrib.entomol.59.1.103-131, URL: https://www.contributions-to-entomology.org/article/view/1742
E101B54FFFFA925DFEB7498FFEA8FDCE.taxon	description	Redescription: Body brown with bronzy lustre or without, sometimes dark-brown, margins of pronotum and elytra sometimes lighter; legs brown. Body distinctly rounded, wide, elytral apices not elongated or weakly elongated. Head large, vertex wide, eyes flattened, rarely weakly convex. Surface of a head with well developed shagrination. Antennae (especially in males) clearly thickened. Pronotum short with very wide and weakly rounded base. Pronotal surface covered with well developed, large shagrination; punctation large, shallow, usually poorly visible. Elytra with punctation arranged in regular striae, sometimes partly confused, secondary punctation smaller than strial, not dense, rarely striae more confused and secondary punctation large. Legs distinctly thickened, wide (especially in males). First protarsomere of male large, long, and wide. Tibiae straight or weakly curved. Hind femora wide. Aedeagus (Fig. 3 J, K) ventrally with apical 1 / 3 with clear, often rounded margins and rounded apex without distinct denticle, medial third usually narrower than apical, from lateral view apical third wide, sharply narrowed to apex. Body length – 1.28 - 1.61 mm, width – 0.95 - 1.19 mm.	en	Nadein, Konstantin S. (2009): Revision of the genus Mniophila S, 1831 (Coleoptera: Chrysomelidae). Beiträge Zur Entomologie = Contributions to Entomology 59 (1): 103-131, DOI: 10.21248/contrib.entomol.59.1.103-131, URL: https://www.contributions-to-entomology.org/article/view/1742
E101B54FFFFA925DFEB7498FFEA8FDCE.taxon	diagnosis	Differential diagnosis: Differs from M. muscorum: structure of aedeagus (Fig. 3 J, K) with apex short in posterior view rounded, apex laterally rounded, apically without denticle instead of apex elongated or short with straight margins and well developed denticle (Fig. 2); body more rounded (Fig. A-C); eyes flattened; prothorax shorter (Fig. 3 E); first protarsomere of male thicker and longer (Fig. 3 G); first metatarsomere of male longer (Fig. 3 F); colouration of the body somewhat lighter, brown to dark brownish with bronzy lustre; elytral punctures usually sparser, usually arranged in regular striae, secondary punctation smaller, usually weakly developed. From M. turcica differs: in structure of aedeagus: apex short at view from behind with rounded sided, apex rounded, without denticle instead of aedeagus ventrally with apical third gradually narrowed to apex; apex almost straight with large denticle; punctated pronotum, tibiae usually thinner and less curved, segments of antennae wider. From M. taurica sp. n. differs: structure of aedeagus with apex short in posterior view, laterally rounded, apex rounded, without denticle instead of aedeagus ventrally toward apex parallel-sided, wide, apical 1 / 4 with straight, narrowing sides and with distinct and straight apex with rather short and wide denticle or with poorly developed one; coloration of the body (without green reflection); legs and antennae thicker, head larger and of another structure, shagrination of the head more developed, punctation of elytra usually more regular and larger; differs also in the shape of pronotum.	en	Nadein, Konstantin S. (2009): Revision of the genus Mniophila S, 1831 (Coleoptera: Chrysomelidae). Beiträge Zur Entomologie = Contributions to Entomology 59 (1): 103-131, DOI: 10.21248/contrib.entomol.59.1.103-131, URL: https://www.contributions-to-entomology.org/article/view/1742
E101B54FFFF99243FF0B4E57FE16FE40.taxon	description	(Figs 4, 10)	en	Nadein, Konstantin S. (2009): Revision of the genus Mniophila S, 1831 (Coleoptera: Chrysomelidae). Beiträge Zur Entomologie = Contributions to Entomology 59 (1): 103-131, DOI: 10.21248/contrib.entomol.59.1.103-131, URL: https://www.contributions-to-entomology.org/article/view/1742
E101B54FFFF99243FF0B4E57FE16FE40.taxon	materials_examined	Type material: Holotype: Turkey: Between Yol-Üstü and Rize, 600 m., 15. V. 1967, C. Besuchet leg., male. – Paratypes: the same label as holotype, 8 specimens. – Hopa-Arhavi, 14. V. 1967, C. Besuchet leg., 3 specimens. – deposited in the Museum of Natural History of Geneva. Two paratype specimens studied (labels numbered): 1. Turquie Rize yol üstu / Rize 600 m, 14. V. 67 Cl. Besuchet; 2. Paratypus Mniophila muscorum turcica L. Medv. – male and female (LM). Type locality: Turkey: Rize.	en	Nadein, Konstantin S. (2009): Revision of the genus Mniophila S, 1831 (Coleoptera: Chrysomelidae). Beiträge Zur Entomologie = Contributions to Entomology 59 (1): 103-131, DOI: 10.21248/contrib.entomol.59.1.103-131, URL: https://www.contributions-to-entomology.org/article/view/1742
E101B54FFFF99243FF0B4E57FE16FE40.taxon	distribution	Distribution: Turkey (Rize, Artvin).	en	Nadein, Konstantin S. (2009): Revision of the genus Mniophila S, 1831 (Coleoptera: Chrysomelidae). Beiträge Zur Entomologie = Contributions to Entomology 59 (1): 103-131, DOI: 10.21248/contrib.entomol.59.1.103-131, URL: https://www.contributions-to-entomology.org/article/view/1742
E101B54FFFF99243FF0B4E57FE16FE40.taxon	description	Redescription: Body dark brown with weak bronzy luster; legs and antennae brown. Body almost rounded with weakly elongated elytral apices. Head large, long, eyes moderately convex; frons long, frontal ridge strongly convex; antennal grooves rather deep. Head surface covered with shagrination. Antennae not thick, segments short. Pronotum moderately long with wide and widely rounded base. Its surface covered with well developed shagrination, impunctate. Elytra smooth, punctation weakly developed, punctures large, poorly visible, shallow; in apical third invisible. Punctures arranged in partly confused striae; secondary punctation not developed. Notch between metathoracic cavities concave. Legs rather thick, clearly curved. First protarsomere of male moderately wide, not very long; metafemora wide. Aedeagus ventrally with apical third gradually narrowed to apex; apex almost straight with large denticle; basal 2 / 3 gradually widened toward apical third; from lateral view apical 2 / 3 almost straight, slightly curved apically. Body length – 1.2 - 1.5 mm, width – 0.98 mm.	en	Nadein, Konstantin S. (2009): Revision of the genus Mniophila S, 1831 (Coleoptera: Chrysomelidae). Beiträge Zur Entomologie = Contributions to Entomology 59 (1): 103-131, DOI: 10.21248/contrib.entomol.59.1.103-131, URL: https://www.contributions-to-entomology.org/article/view/1742
E101B54FFFF99243FF0B4E57FE16FE40.taxon	diagnosis	Differential diagnosis: From M. transcaucasica sp. n. differs: in structure of aedeagus (Fig. 4 I, J), apex almost straight instead of apex obtuse; structure of head (Fig. 4 H), particularly in more convex eyes, frontal ridge more convex and more developed, antennal grooves deeper, frontal calli more developed; notch between metathoracic cavities concave; first metatarsomere longer and narrower (Fig. 4 C); dorsal punctation poorly developed. From M. caucasica sp. n. differs: in structure of aedeagus, apex almost straight instead of aedeagal apex rather wide, straight, apical 1 / 4 in posterior view more or less sharply narrowed to apex; apical segments of antennae shorter; tibiae thicker and more curved; first protarsomere (Fig. 4 D) and that of mesotarsomere of males shorter and wider; dorsal punctation poorly developed and shagrination well developed. From M. bosnica differs: in structure of aedeagus, ventrally with apical third gradually narrowed to apex, the latter almost straight with large denticle instead of aedeagal apex with rounded sides and apex, the latter without denticle; pronotum impunctate; tibiae usually more curved (Fig. 4 E, F), antennal segments thinner (Fig. 4 G).	en	Nadein, Konstantin S. (2009): Revision of the genus Mniophila S, 1831 (Coleoptera: Chrysomelidae). Beiträge Zur Entomologie = Contributions to Entomology 59 (1): 103-131, DOI: 10.21248/contrib.entomol.59.1.103-131, URL: https://www.contributions-to-entomology.org/article/view/1742
E101B54FFFE79241FF0B4925FC0FFCEA.taxon	description	(Figs 5, 10)	en	Nadein, Konstantin S. (2009): Revision of the genus Mniophila S, 1831 (Coleoptera: Chrysomelidae). Beiträge Zur Entomologie = Contributions to Entomology 59 (1): 103-131, DOI: 10.21248/contrib.entomol.59.1.103-131, URL: https://www.contributions-to-entomology.org/article/view/1742
E101B54FFFE79241FF0B4925FC0FFCEA.taxon	materials_examined	Type material: Holotype (labels numbered): 1. Ukraine, Crimea, ascent to Chatyr-Dagh mid plateau from Sosnovka Vill. 18. VI. 2003 Nadein K.; 2. Quercus-Fraxinus-Fagus forest, on Fagus stem and in moss, male (ZIN). – Paratypes: the same label as holotype, 4 females (NC). – Ukraine, Crimea, ascent to Chatyr-Dagh mid plateau from Sosnovka Vill. 18. VI. 2003 Yunakov N., 1 male, 1 female (NC). – Crimea, Chatyr-Dagh nr. Bin-Bash-Koba Cave mid plateau, on Fagus stem in forest, 100 m, 22. VI. 2003 Nadein K., 1 female (ZIN). – 1. Crimea Bakhchisarayskiy Distr. Bol’shoy Kan’yon Gorge, 5.05.1999 N. Yunakov; 2. in forest, on moss among stones, 5 females (ZIN), 4 females (DEI). – 1. Crimea Bol’shoy Kan’yon Gorge, reserved area, 05.05.1999 A. Drogvalenko; 2. in moss on stones, 12 females (KUMN). – 1. Ukraine, Crimea, 1 km S Pereval’noe Vill., left bank of Angara Riv., 1. VII. 2008 Nadein K. leg.; 2. Quercus- Fagus-Carpinus forest, in moss at base of Quercus stem, 2 females (DEI). – Ukraine, Crimea, Dolgorukovskaya Yajla, upper stream of Burul’cha Riv., env. Kolan-Bair Mt., h = 800 m, 29. VI. 2008 Nadein K. leg.; 2. Fagus-Quercus-Carpinus forest, in moss on stones, 2 males 3 females (SIZK), 3 males, 3 females (NHML).	en	Nadein, Konstantin S. (2009): Revision of the genus Mniophila S, 1831 (Coleoptera: Chrysomelidae). Beiträge Zur Entomologie = Contributions to Entomology 59 (1): 103-131, DOI: 10.21248/contrib.entomol.59.1.103-131, URL: https://www.contributions-to-entomology.org/article/view/1742
E101B54FFFE79241FF0B4925FC0FFCEA.taxon	etymology	Etymology: The specific epithet refers to geographical distribution of the new species that is endemic to Crimea.	en	Nadein, Konstantin S. (2009): Revision of the genus Mniophila S, 1831 (Coleoptera: Chrysomelidae). Beiträge Zur Entomologie = Contributions to Entomology 59 (1): 103-131, DOI: 10.21248/contrib.entomol.59.1.103-131, URL: https://www.contributions-to-entomology.org/article/view/1742
E101B54FFFE79241FF0B4925FC0FFCEA.taxon	distribution	Distribution: Ukraine: Crimean Mountains.	en	Nadein, Konstantin S. (2009): Revision of the genus Mniophila S, 1831 (Coleoptera: Chrysomelidae). Beiträge Zur Entomologie = Contributions to Entomology 59 (1): 103-131, DOI: 10.21248/contrib.entomol.59.1.103-131, URL: https://www.contributions-to-entomology.org/article/view/1742
E101B54FFFE79241FF0B4925FC0FFCEA.taxon	description	Description: Body black, shining, often with greenish luster; legs and antennae yellow-reddish to light brown. Body almost rounded, with weakly elongated elytral apices. Head small, vertex moderately wide; eyes convex. Head surface covered with poorly developed, smoothed shagrination, sometimes vertex almost smooth. Antennae thin. Pronotum comparatively long, with distinctly rounded base, its surface covered with fine shagrination, punctation small, weakly visible among surface’s microsculpture. Elytra with small, dense punctation, striae usually confused; secondary punctation usually well developed, size as large as in striae or nearly so; rarely striae regular. Legs thin; first protarsomere of male almost not widened or weakly widened; tibiae straight or slightly curved, metafemora narrow. Aedeagus (Fig. 5 K-M) ventrally toward apex parallel-sided, wide, apical 1 / 4 with straight, narrowing sides and with distinct and straight apex with rather short, wide denticle or denticle poorly developed; from lateral view apical half gradually narrowed to apex. Body length – 1.34 - 1.59 mm, width – 0.95 - 1.21 mm.	en	Nadein, Konstantin S. (2009): Revision of the genus Mniophila S, 1831 (Coleoptera: Chrysomelidae). Beiträge Zur Entomologie = Contributions to Entomology 59 (1): 103-131, DOI: 10.21248/contrib.entomol.59.1.103-131, URL: https://www.contributions-to-entomology.org/article/view/1742
E101B54FFFE79241FF0B4925FC0FFCEA.taxon	diagnosis	Differential diagnosis: From M. caucasica sp. n. differs: in structure of aedeagus (Fig. 5 K-M) with poorly developed denticle or with wider and shorter one, from lateral view narrower; flattened eyes (Fig. 5 C); apical antennal segments shorter (Fig. 5 E); shape of pronotum with more rounded and more elongated base (Fig. 5 F); head more elongated with vertex narrower, hind femora narrower (Fig. 5 D). From M. transcaucasica sp. n. differs: in structure of aedeagus, wide, ventrally toward apex parallelsided, apical 1 / 4 with straight, narrowing sides and with distinct and straight apex with rather short, wide denticle or poorly developed instead of aedeagus ventrally with apical third gradually narrowed to apex, the latter obtuse with well developed denticle; head longer, shagrination of head less developed, frontal calli more developed; pronotum longer with more convex base; tibiae less curved and usually straight (Fig. 5 G, H); hind femora narrower; notch between metathoracic cavities concave; first protarsomere of male narrower. From M. turcica differs: in structure of aedeagus, wide, ventrally toward apex parallel-sided, apical 1 / 4 with straight, narrowing laterally, apex distinct and straight with rather short, wide denticle or poorly developed compared to aedeagus ventrally with apical third gradually narrowed to apex, the latter almost straight with large denticle; tibiae nearly straight and usually less curved; shagrination of head less developed; hind femora narrower; pronotum punctate; tibiae thicker; eyes more flattened.	en	Nadein, Konstantin S. (2009): Revision of the genus Mniophila S, 1831 (Coleoptera: Chrysomelidae). Beiträge Zur Entomologie = Contributions to Entomology 59 (1): 103-131, DOI: 10.21248/contrib.entomol.59.1.103-131, URL: https://www.contributions-to-entomology.org/article/view/1742
E101B54FFFE59244FF0B4FFFFED0FBFD.taxon	description	(Figs 6, 10)	en	Nadein, Konstantin S. (2009): Revision of the genus Mniophila S, 1831 (Coleoptera: Chrysomelidae). Beiträge Zur Entomologie = Contributions to Entomology 59 (1): 103-131, DOI: 10.21248/contrib.entomol.59.1.103-131, URL: https://www.contributions-to-entomology.org/article/view/1742
E101B54FFFE59244FF0B4FFFFED0FBFD.taxon	materials_examined	Type material: Holotype (labels numbered): Georgia: 1. Manglis 1879, 2. k. G. Siversa, male (ZIN). – Paratypes: Russia: Krasnodarsk. Kray Severskiy r-n Ubinskoe lesn. Belov 12. VII. 1975, 3 females (DEMU). – The same label, 24. VII. 1975, 1 female (DEMU). – The same label, 18. V. 1975, 1 male (DEMU). – Sev. Osetia, 1300 m, Buron yuzh. Alagir 16. VIII. 1979 Kurbatov, 1 female (DEMU). – Kavkazskiy zapov. Yuzhnoe lesn. Kordon Pslukh Nikitskiy 15. VII. 1976, 1 male, 1 female (DEMU). – The same label, 14. VII. 1976, 1 male, 1 female (DEMU). Abkhazia: Abkhazia Myusserskiy zapov. 11 - 16. III. 1988 leg. V. Belov, 1 female (BMPU). – Abkhazia, okrestn. Pitsundy, Myussera 4 - 12. X. 1986 leg. V. Belov, 1 female (BMPU). – 13.04.1990 Abkhazia Gumistinskiy zapov. Tsimur. V podstilke, S. Kurbatov, 1 female (LM). – W Caucasus Abkhazia Pskhu valley, Bzyb’ River, mouth of Reshevie Riv., 591 m N 43 ° 20 ' 19.23 " E 40 ° 50 ' 27.64 " 28. VI. 2007 K. Nadein leg. / lower terrace, in moss on Carpinus stems and on stones in forest, 1 male, 7 females (DEI). – W Caucasus Abkhazia W Gumista Riv., 1 km above mouth Gumiripsha Riv. 450 m N 43 ° 14 ' 45.80 " E 40 ° 54 ' 40.26 " 21. VI. 2007 K. Nadein leg. / Fagus-Carpinus forest, slopes in gorge, in moss and in litter, 2 males, 6 females, (NC). – W Caucasus Abkhazia W Gumista Riv., 1 km above mouth Gumiripsha Riv. 450 m N 43 ° 14 ' 45.80 " E 40 ° 54 ' 40.26 " 20. VI. 2007 K. Nadein leg. / Fagus-Carpinus forest, slopes to river, in moss on Fagus stem, 4 males (ZIN). – W Caucasus Abkhazia Bagri-Yashta MtR, nr Anchkho Pass, h = 1800 - 2000 m N 43 ° 28 ' 39.64 " E 40 ° 42 ' 28.49 " 2. VII. 2007 K. Nadein leg. / sweeping in Fagus - Carpinus forest and edge of meadows, 6 males, 1 females (KUMN). – W Caucasus Abkhazia middle course of Ghegha Riv. Gheghskoe Gorge, h = 500 m N 43 ° 25 ' 43.11 " E 40 ° 26 ' 51.04 " 13. VII. 2007 K. Nadein leg. / Picea - Abies - Fagus forest in moss, lower and middle terrases along river, 2 males, 4 females (HNHM). – W Caucasus Abkhazia Psheghishkhva Mt., nr Malaya Ritsa Lake, h = 1200 m N 43 ° 28 ' 25.73 " E 40 ° 30 ' 04.47 " 8 – 9. VII. 2007 K. Nadein leg. / Abies-Fagus forest, in moss on stones and on Abies stem, 2 males, 6 females (DEI). – W Caucasus Abkhazia Bagri- Yashta MtR., Anchkho Mt. nr Auatkhara loc., h = 1500 m N 43 ° 29 ' 38 " E 40 ° 40 ' 26 " 5. VII. 2007 K. Nadein leg. / Abies-Fagus forest, in moss on stones, 4 males, 4 females (NHML). – W Caucasus Abkhazia Aguripsta Riv., 4 km NW of Pskhu Vill., nr. Svyataya Mt. N 43 ° 25 ' 36.80 " E 40 ° 49 ' 08.40 " 29. VI. 2007 K. Nadein leg. / Fagus-Carpinus forest, in moss on Carpinus stem, h = 800 m, 3 males, 7 females (IRSNB). – W Caucasus Abkhazia Bzybskiy Mount. Range, S slope of Dou Mt., h = 1200 m N 43 ° 17 ' 12.00 " E 40 ° 53 ' 39.21 " 23. VI. 2007 K. Nadein leg. / Fagus-Carpinus- Quercus - Castanea forest, in moss on stony slopes along road under Rhododendron bushes, 4 males, 9 females (NHMW). Georgia: The same labels as holotype, 1 male (ZIN). – Manglis [illegible words in a second line] 1881, 2 females (ZIN). – Manglis 1880 / G. Siversa, 6 females, 2 males (ZIN). – Caucasus Swanetien Leder. Reitter / M. muscorum Koch. Coll. Reitter, 1 female (HNHM). – Gruzia okr. Borzhomi 28 - 29. VII. 1977 v lesu pod kamnem Belov, 1 male, 1 female (DEMU). – Caucasus Tbatani 79 Leder (Reitter), 1 female (ZMUA). – Kaukas Leder / M. muscorum v. wroblewskii Wk. Coll. Reitter, 1 female (HNHM). – Kaukas Leder / M. muscorum Koch. Coll. Reitter, 2 females (HNHM). – Caucasus Leder. Reitter / M. muscorum v. wroblewskii Wk. Coll. Reitter, 1 female (HNHM). – Kaukas Leder / Mn. muscorum v. caucasica Hktg., 1 female (DEI). – Kaukas. Schneider / Mn. muscorum v. caucasica Hktg., 1 female (DEI). Azerbaijan: Azerbaijan: Caucasus Belokany Distr., 9 km N Vill. Katekh, Zagatala Res. Katekhchay Riv. valley 26. VI. 2004 K. Nadein leg.; 2. N 41 ° 45 ' 02 '' E 46 ° 37 ' 01 '' left bank, E slope of Fidjugel’ MtR, 700 - 800 m Fagus forest, 1 male, 2 females (NC). – Caucasus Helenendorf Reitter, 1 female (ZMUA).	en	Nadein, Konstantin S. (2009): Revision of the genus Mniophila S, 1831 (Coleoptera: Chrysomelidae). Beiträge Zur Entomologie = Contributions to Entomology 59 (1): 103-131, DOI: 10.21248/contrib.entomol.59.1.103-131, URL: https://www.contributions-to-entomology.org/article/view/1742
E101B54FFFE59244FF0B4FFFFED0FBFD.taxon	etymology	Etymology: The specific epithet refers to geographical distribution of this new species that is widespread in Caucasus.	en	Nadein, Konstantin S. (2009): Revision of the genus Mniophila S, 1831 (Coleoptera: Chrysomelidae). Beiträge Zur Entomologie = Contributions to Entomology 59 (1): 103-131, DOI: 10.21248/contrib.entomol.59.1.103-131, URL: https://www.contributions-to-entomology.org/article/view/1742
E101B54FFFE59244FF0B4FFFFED0FBFD.taxon	distribution	Distribution: Russia (North Caucasus: Krasnodarskiy Terr.), Abkhazia, Georgia, Azerbaijan.	en	Nadein, Konstantin S. (2009): Revision of the genus Mniophila S, 1831 (Coleoptera: Chrysomelidae). Beiträge Zur Entomologie = Contributions to Entomology 59 (1): 103-131, DOI: 10.21248/contrib.entomol.59.1.103-131, URL: https://www.contributions-to-entomology.org/article/view/1742
E101B54FFFE59244FF0B4FFFFED0FBFD.taxon	description	Description: Body dark to black, shining, often with greenish luster; rarely brown with bronze luster in partly coloured specimens; legs brown. Body roundellipticall. Head large, short; eyes convex; frontal ridge moderately convex; frontal calli well developed, convex; supracallinal sulcus visible. Head surface covered with large, well developed shagrination. Antennal grooves not very deep. Antennae with moderately thick and comparatively long segments. Pronotum long, quite apparently not widely rounded, its surface covered with distinct shagrination, rarely smooth; punctation developed, not large and dense, weakly visible among surface’s microsculpture. Elytra with variable states of punctuation, usually punctation well developed, sometimes elytra almost impunctate; elytral striae usually confused, not large; secondary punctation well developed, dense, size often as large as strial punctures; interstices smooth. Notch between metathoracic cavities concave (Fig. 6 H). Legs not very thick; tibiae curved; metafemora wide; first protarsomere of male not wide. Aedeagus (Fig. 6 L-N) ventrally with apical 1 / 4 more or less sharply narrowed to apex; apex rather wide, straight, with denticle; from lateral view apical 2 / 3 almost straight, wide, clearly narrowed to apex. Body length – 1.14 - 1.58 mm, width – 0.91 - 1.12 mm.	en	Nadein, Konstantin S. (2009): Revision of the genus Mniophila S, 1831 (Coleoptera: Chrysomelidae). Beiträge Zur Entomologie = Contributions to Entomology 59 (1): 103-131, DOI: 10.21248/contrib.entomol.59.1.103-131, URL: https://www.contributions-to-entomology.org/article/view/1742
E101B54FFFE59244FF0B4FFFFED0FBFD.taxon	diagnosis	Differential diagnosis: From M. transcaucasica sp. n. differs: in structure of aedeagus (Fig. 6 L-N), ventrally with apical 1 / 4 more or less sharply narrowed to apex, the latter rather wide and straight instead of aedeagus ventrally with apical third gradually narrowed to apex; less curved tibiae (Fig. 6 I, J); first protarsomere of male longer and thinner (Fig. 6 F); eyes more convex; ocular sulci and frontal calli more developed, apical antennal segments longer (Fig. 6 K); pronotum longer (Fig. 6 E), notch between metathoracic cavities concave (Fig. 6 H); last metatarsomere thicker (Fig. 6 G). From M. turcica differs: in structure of aedeagus, ventrally with apical 1 / 4 more or less sharply narrowed to apex, the latter rather wide, straight instead of aedeagus ventrally with apical third gradually narrowed to apex, the latter almost straight; head shorter with deep antennal grooves; apical antennal segments longer; tibiae thinner and less curved; usually well developed punctation of pronotum and usually poorer developed shagrination. From M. taurica sp. n. differs: in structure of aedeagus, ventrally with apical 1 / 4 more or less sharply narrowed to apex, the latter rather wide, straight, denticle well developed instead of aedeagus ventrally toward apex parallelsided, wide, apical 1 / 4 with straight, narrowing sides and with distinct and straight apex with rather short, wide denticle or poorly developed; head shorter and wider, eyes more convex, apical antennal segments longer; hind femora thicker; differs also in the shape of pronotum with less rounded base.	en	Nadein, Konstantin S. (2009): Revision of the genus Mniophila S, 1831 (Coleoptera: Chrysomelidae). Beiträge Zur Entomologie = Contributions to Entomology 59 (1): 103-131, DOI: 10.21248/contrib.entomol.59.1.103-131, URL: https://www.contributions-to-entomology.org/article/view/1742
E101B54FFFE0924BFEB74ED7FF29FB4D.taxon	description	(Figs 7, 10)	en	Nadein, Konstantin S. (2009): Revision of the genus Mniophila S, 1831 (Coleoptera: Chrysomelidae). Beiträge Zur Entomologie = Contributions to Entomology 59 (1): 103-131, DOI: 10.21248/contrib.entomol.59.1.103-131, URL: https://www.contributions-to-entomology.org/article/view/1742
E101B54FFFE0924BFEB74ED7FF29FB4D.taxon	materials_examined	Type material: Holotype: Armenia: Kaukasus Dshelal ogly, male (ZIN). – Paratypes: Georgia: Kaukas Leder / M. muscorum Koch. Coll. Reitter, 2 males (HNHM). Caucasus. Mniophila muscorum Caucas / k. Rybakova, male (ZIN).	en	Nadein, Konstantin S. (2009): Revision of the genus Mniophila S, 1831 (Coleoptera: Chrysomelidae). Beiträge Zur Entomologie = Contributions to Entomology 59 (1): 103-131, DOI: 10.21248/contrib.entomol.59.1.103-131, URL: https://www.contributions-to-entomology.org/article/view/1742
E101B54FFFE0924BFEB74ED7FF29FB4D.taxon	etymology	Etymology: The specific epithet refers to geographical distribution of the new species that is distributed in Transcaucasus.	en	Nadein, Konstantin S. (2009): Revision of the genus Mniophila S, 1831 (Coleoptera: Chrysomelidae). Beiträge Zur Entomologie = Contributions to Entomology 59 (1): 103-131, DOI: 10.21248/contrib.entomol.59.1.103-131, URL: https://www.contributions-to-entomology.org/article/view/1742
E101B54FFFE0924BFEB74ED7FF29FB4D.taxon	distribution	Distribution: Armenia, Georgia.	en	Nadein, Konstantin S. (2009): Revision of the genus Mniophila S, 1831 (Coleoptera: Chrysomelidae). Beiträge Zur Entomologie = Contributions to Entomology 59 (1): 103-131, DOI: 10.21248/contrib.entomol.59.1.103-131, URL: https://www.contributions-to-entomology.org/article/view/1742
E101B54FFFE0924BFEB74ED7FF29FB4D.taxon	description	Description: Body brown to dark-brown with weak bronze luster or without one; legs reddish-brownish. Body nearly elongated to clearly rounded. Head large, short; vertex covered with well developed, large shagrination; frontal calli almost not convex; supracallinal sulci poorly visible; frontal ridge short, weakly convex; eyes flattened. Antennae with segments short and thick. Pronotum short, widely transversal, with very widely rounded base, its surface usually covered with well developed, large shagrination; punctures large, sparse, poorly visible. Elytra impunctate to well developed, dense punctation, punctures not large; striae partly confused, secondary punctation well developed. Notch between metathoracic cavities straight (Fig. 7 C). Legs not very thick, tibiae moderately curved; metafemora wide; first protarsomere of male wide and thick. Aedeagus (Fig. 7 J, K) ventrally with apical third gradually narrowed to apex; apex obtuse with well developed denticle; from lateral view nearly gradually curved from basal third, weakly and gradually narrowed toward apex. Body length – 1.34 - 1.47 mm, width – 0.91 - 1.05 mm.	en	Nadein, Konstantin S. (2009): Revision of the genus Mniophila S, 1831 (Coleoptera: Chrysomelidae). Beiträge Zur Entomologie = Contributions to Entomology 59 (1): 103-131, DOI: 10.21248/contrib.entomol.59.1.103-131, URL: https://www.contributions-to-entomology.org/article/view/1742
E101B54FFFE0924BFEB74ED7FF29FB4D.taxon	diagnosis	Differential diagnosis: From M. caucasica sp. n. differs: in structure of aedeagus (Fig. 7 J, K), ventrally with apical third gradually narrowed to apex; apex obtuse instead of aedeagus ventrally with apical 1 / 4 more or less sharply narrowed to apex, the latter rather wide, straight; hind tibiae distinctly curved (Fig. 7 F), first tarsomeres of male shorter and thicker (Fig. 7 H, I); eyes more flattened; ocular sulci and frontal calli less developed (Fig. 7 E); apical antennal segments shorter and thicker (Fig. 7 D), differs in pronotum shape (Fig. 7 B); notch between metathoracic cavities straight (Fig. 7 C); last metatarsomere thinner (Fig. 7 H). From M. taurica sp. n. differs: in structure of aedeagus, ventrally with apical third gradually narrowed to apex; apex obtuse with well developed denticle instead of aedeagus ventrally toward apex parallel-sided, wide, apical 1 / 4 with straight, narrowing sides and with distinct and straight apex with rather short, wide denticle or poorly developed; head shorter, frontal calli less developed, shagrination of head more developed; pronotum shorter with less elongated base; hind femora thicker; tibiae thicker and more curved; notch between metathoracic cavities straight; first protarsomere of male wider. From M. turcica differs: in structure of aedeagus, apex obtuse instead of apex almost straight; head shorter, frontal ridge less convex, antennal grooves shallower; notch between metathoracic cavities straight; eyes more flattened; tibiae thinner; punctation of pronotum well developed; first metatarsomere shorter and wider.	en	Nadein, Konstantin S. (2009): Revision of the genus Mniophila S, 1831 (Coleoptera: Chrysomelidae). Beiträge Zur Entomologie = Contributions to Entomology 59 (1): 103-131, DOI: 10.21248/contrib.entomol.59.1.103-131, URL: https://www.contributions-to-entomology.org/article/view/1742
E101B54FFFE0924BFEB74ED7FF29FB4D.taxon	discussion	Remarks: The materal labelled “ Kaukas Leder ” possibly orignates from Georgia according to LEDER (1878). Acknowledgments I thank S. Shute (NHML), I. Izquierdo (Spain), O. Jäger (MTD), L. Bartolozzi (MZUF), L. Zerche (DEI), O. Merkl (HNHM), P. Limbourg (IRSNB), B. Brugge (ZMUA), O. Martin (ZMUC), H. Schoenmann (NHMW), A. Drogvalenko (KUMN), N. Yunakov (ZIN), N. Nikitsky (ZMMU), K. Makarov (BDMP), Yu. Savitskii (DEMU), and L. Medvedev (Moscow, Russia) for the opportunity to study material in their care. I am thankful to D. Kulijer (National Museum of Bosnia and Herzegovina, Sarajevo) for the information about the type materail of Mniophila bosnica. I sincerely gratitude to V. Virtshenko (Kholodnyi Institute of Botany, Kiev, Ukraine) for the determination of mosses. I thank A. Samuelson (Honoloulu, Hawaii) for the information on Mniophila exulans. I am indebted to D. Furth (Smithsinian Institution, USA) for his linguistic help and valuable suggestions. This study has been partly supported by Ernst Mayr Foundation travel grant in 2007.	en	Nadein, Konstantin S. (2009): Revision of the genus Mniophila S, 1831 (Coleoptera: Chrysomelidae). Beiträge Zur Entomologie = Contributions to Entomology 59 (1): 103-131, DOI: 10.21248/contrib.entomol.59.1.103-131, URL: https://www.contributions-to-entomology.org/article/view/1742
E101B54FFFE0924BFEB74ED7FF29FB4D.taxon	discussion	Remarks: The materal labelled “ Kaukas Leder ” possibly orignates from Georgia according to LEDER (1878).	en	Nadein, Konstantin S. (2009): Revision of the genus Mniophila S, 1831 (Coleoptera: Chrysomelidae). Beiträge Zur Entomologie = Contributions to Entomology 59 (1): 103-131, DOI: 10.21248/contrib.entomol.59.1.103-131, URL: https://www.contributions-to-entomology.org/article/view/1742
