taxonID	type	description	language	source
039787FAFF83211AFF68FF402404800F.taxon	materials_examined	Holotype.  “ SABAH: Tawai Plat. 1300 ft, 8 m. S. Telupid 8. ix. 1977 | At light | M. E. Bacchus B. M. 1978 - 48 | HOLOTYPUS Anomala parastasioides sp. n. det. Zorn, 2007 ” (BMNH).	en	Zorn, Carsten (2007): Three new species of the genus Anomala S, 1819 from Borneo of the fulvofusca-species group (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (2): 277-283, DOI: 10.21248/contrib.entomol.57.2.277-283, URL: https://www.contributions-to-entomology.org/article/view/1704
039787FAFF83211AFF68FF402404800F.taxon	description	Description: Body shape. Length: 10.3 mm, width: 6.0 mm; oval, stout, and convex. Color. Head dark reddish; pronotum, scutellum and elytra yellowish brown with two separated longitudinal black discal patches on the pronotum, the latter not reaching the anterior or posterior margin; elytral pattern as follows: humeral protuberance and the complete anterior margin black; a broad black longitudinal patch along the suture which is dilated before but not reaching the apex; sutural margin itself narrowly reddish; propygidium and a transverse triangular patch at the base of the pygidium dark reddish to black; remaining part of the pygidium yellowish brown; underside including antennae and femora yellowish brown; metasternum somewhat blackened; tibiae and tarsi dark reddish brown; abdominal sternites black with a reddish distal margin. Head. Clypeus subrectangular, strongly reflexed; anterior angles broadly rounded; surface with fine punctures densely covered; frons with larger, somewhat umbilicate punctures; punctures smaller and less dense towards vertex. Pronotum. Width 1.61 times length, broadest somewhat behind the middle; sides evenly curved and strongly convergent towards the acute anterior angles; posterior angles broadly rounded; basal stria not interrupted; punctures moderately fine and rather sparse on the disc, gradually larger and more dense towards the lateral margins. Elytra. With deeply impressed, coarse and somewhat umbilicate punctures arranged in regular striae mixed with sparse micropunctation; primary costae not elevated; subsutural, second and third interstice (the intervals between primary costae) with one secondary row of punctures, the one of the subsutural interstice irregularly duplicated behind the scutellum; the three inner primary striae weakly sulcate at the apex; humeral protuberance prominent and smooth. Pygidium. Convex in profile; punctures umbilicate and deeply impressed, moderately fine at the base, gradually coarser and denser towards the hind margin. Metasternum. Disc smooth and shining, weakly sulcate; lateral parts gradually more densely and coarsely punctured, with rather short and not dense yellowish setae. Abdominal sternites. Densely punctured with some irregular smooth areas; punctures moderately coarse, sometimes transversely confluent; with a transverse series of yellowish semierect setae; distal margin of the last sternite deeply emarginate. Protibia. Bidentate, lateral and terminal teeth rather short and blunt, but apparently somewhat worn out; terminal articulated spine very short, about 1.5 times as long as wide. Metatibia. Moderately slender, 3.1 times as long as wide, slightly fusiform. Claws. External middle claw and internal fore claw cleft; the latter with a very small spine-shaped upper branch; lower branch much broader and longer, with an obtuse angle at its inner margin and excised before the strongly pointed apex (Fig. 14). Aedeagus. See figs 4, 5.	en	Zorn, Carsten (2007): Three new species of the genus Anomala S, 1819 from Borneo of the fulvofusca-species group (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (2): 277-283, DOI: 10.21248/contrib.entomol.57.2.277-283, URL: https://www.contributions-to-entomology.org/article/view/1704
039787FAFF83211AFF68FF402404800F.taxon	diagnosis	Diagnosis: Anomala parastasioidesparastasioides sp. n. is easily distinguishable from its relatives, A. fulvofusca, A. matangensis, A. pumilispumilis and A. pumiloides, by the remarkable coloration pattern with distinct black patches on pronotum and elytra and the different structure of the aedeagus with hirsute parameres. Moreover, A. parastasioidesparastasioides sp. n. is distinctly bigger than the other species.	en	Zorn, Carsten (2007): Three new species of the genus Anomala S, 1819 from Borneo of the fulvofusca-species group (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (2): 277-283, DOI: 10.21248/contrib.entomol.57.2.277-283, URL: https://www.contributions-to-entomology.org/article/view/1704
039787FAFF83211AFF68FF402404800F.taxon	etymology	Etymology: The species name is referring to the superficial resemblance of A. parastasioides to some small species of the Ruteline genus Parastasia WESTWOOD, 1842.	en	Zorn, Carsten (2007): Three new species of the genus Anomala S, 1819 from Borneo of the fulvofusca-species group (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (2): 277-283, DOI: 10.21248/contrib.entomol.57.2.277-283, URL: https://www.contributions-to-entomology.org/article/view/1704
039787FAFF802118FED4FC9E2785878A.taxon	materials_examined	Holotype.  “ Za 21 | Mt. Matanga, W. Sarawak, G. E. Bryant. 16. II. 14. | G. Bryant Coll. B. M. 1926 - 86. | HOLOTYPUS Anomala matangensis sp. n. det. Zorn, 2007 ” (BMNH).	en	Zorn, Carsten (2007): Three new species of the genus Anomala S, 1819 from Borneo of the fulvofusca-species group (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (2): 277-283, DOI: 10.21248/contrib.entomol.57.2.277-283, URL: https://www.contributions-to-entomology.org/article/view/1704
039787FAFF802118FED4FC9E2785878A.taxon	description	Description: Body shape. Length: 7.8 mm, width: 4.3 mm; elongate oval, dilated behind; strongly convex. Color. Head reddish brown; pronotum reddish brown on the disc, yellowish brown towards the lateral margins; scutellum yellowish; elytra dark brown, with the apical part broadly pale yellow; sutural margin reddish; underside including femora and antennae pale yellowish; propygidium, pygidium, abdominal sternites, tibiae and tarsi reddish brown. Head. Clypeus trapezoid, anterior angles rounded; anterior margin reflexed; punctures moderately fine; frons transversely punctured, somewhat rugose; punctures more fine and sparse towards vertex. Pronotum. Width 1.56 times length; in dorsal view broadest at its base; lateral margins straight and subparallel posteriorly, curved and convergent in the anterior third; anterior angles acute; posterior angles rounded; basal stria complete; whole surface with fine and rather sparse punctation. Elytra. With impressed, coarse and somewhat umbilicate punctures, arranged in regular striae; sparse micropunctation intermixed; primary costae not elevated; subsutural, second and third interstice with one secondary row of punctures, the one of the subsutural interstice irregularly duplicated behind the scutellum; the three inner primary striae weakly sulcate at the apex; humeral and apical protuberance distinct, but not very prominent. Pygidium. Moderately fine punctured; punctures somewhat oblong near the distal margin. Metasternum. Disc sulcate, with minute sparse punctures; towards the lateral margin with very dense, umbilicate punctures and short yellowish setae. Abdominal sternites. Densely, sometimes confluently punctured; with a transverse series of short yellowish setae. Protibia. With a long, acute lateral tooth and a long pointed terminal tooth which is strongly curved outwards; terminal articulated spine short, about 2 times as long as wide. Metatibia. 2.5 times as long as wide, slightly fusiform. Claws. External middle claw and internal front claw cleft; the upper branch of the latter very small and spine-shaped; the lower branch much broader and longer, with an obtuse angle at its inner margin and distinctly excised before the strongly pointed apex. Aedeagus. See figs 6, 7.	en	Zorn, Carsten (2007): Three new species of the genus Anomala S, 1819 from Borneo of the fulvofusca-species group (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (2): 277-283, DOI: 10.21248/contrib.entomol.57.2.277-283, URL: https://www.contributions-to-entomology.org/article/view/1704
039787FAFF802118FED4FC9E2785878A.taxon	diagnosis	Diagnosis: Anomala matangensismatangensis is very similar to A. fulvofusca but can be distinguished by the different aedeagus (Figs 5, 6) and the coloration as described above. The degree of variation of the coloration pattern, however, is uncertain as long as no further material is available.	en	Zorn, Carsten (2007): Three new species of the genus Anomala S, 1819 from Borneo of the fulvofusca-species group (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (2): 277-283, DOI: 10.21248/contrib.entomol.57.2.277-283, URL: https://www.contributions-to-entomology.org/article/view/1704
039787FAFF802118FED4FC9E2785878A.taxon	etymology	Etymology: The species epithet is referring to the type locality, Mt. Matang in western Sarawak, Borneo.	en	Zorn, Carsten (2007): Three new species of the genus Anomala S, 1819 from Borneo of the fulvofusca-species group (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (2): 277-283, DOI: 10.21248/contrib.entomol.57.2.277-283, URL: https://www.contributions-to-entomology.org/article/view/1704
039787FAFF812118FF68FDAC27E8802A.taxon	materials_examined	Holotype.  “ BORNEO-SABAH Banjaran Maitland Sapulut 22. - 24.5.1995 Ivo Jeniš leg. | HOLOTYPUS Anomala pumiloides sp. n. det. Zorn, 2007 ” (CCZ). Paratypes. 1 , 1  “ BORNEO-SABAH Banjaran Maitland Sapulut 22. - 24.5.1995 Ivo Jeniš leg. | PARATYPUS Anomala pumiloides sp. n. det. Zorn, 2007 ” (CCZ).	en	Zorn, Carsten (2007): Three new species of the genus Anomala S, 1819 from Borneo of the fulvofusca-species group (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (2): 277-283, DOI: 10.21248/contrib.entomol.57.2.277-283, URL: https://www.contributions-to-entomology.org/article/view/1704
039787FAFF812118FF68FDAC27E8802A.taxon	description	Description: Body shape. Length: 7.0 - 7.5 mm, width: 3.3 - 3.7 mm; elongate oval; subparallel posteriorly; strongly convex. Color. Entirely yellowish brown; head between the eyes, some indistinct transverse patches on the disc of the pronotum and the entire abdomen in the male sex darker brown. Head. Clypeus trapezoid; anterior angles rounded; anterior margin reflexed; punctures moderately fine; frons densely and sometimes confluently punctured; punctures gradually finer and more sparse towards the vertex. Pronotum. Width 1.64 times length; in dorsal view lateral margin evenly curved and strongly converging towards the acute anterior angles; broadest somewhat behind the middle; posterior angles rounded; basal stria complete; the whole surface covered with moderately sized, umbilicate punctures. Elytra. With rather coarse and umbilicate punctures arranged in regular striae and intermixed irregular, sparse micropunctation; primary costae not elevated; subsutural, second and third interstice with one secondary row of punctures, the one of the subsutural interstice irregularly duplicated at its base; the three inner primary striae weakly sulcate at the apex; humeral protuberance indistinct. Pygidium. Densely covered with coarse, umbilicate punctures. Metasternum. Disc smooth with a longitudinal impression; lateral part with dense umbilicate punctures and indistinct, short setosity. Abdominal sternites. Densely, sometimes confluently punctured; with a transverse series of sparse short yellowish setae. Protibia. With a long acute lateral tooth and a long pointed terminal tooth which is strongly curved outwards; terminal articulated spine about 5 times as long as wide. Metatibia. 2.8 times as long as wide, weakly fusiform. Claws. External middle claw and internal front claw cleft; upper branch of the latter very small and spine-shaped; the lower branch much more broad and long, somewhat dilated at its inner margin; with a strongly pointed apex. Aedeagus. See figs 8, 9.	en	Zorn, Carsten (2007): Three new species of the genus Anomala S, 1819 from Borneo of the fulvofusca-species group (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (2): 277-283, DOI: 10.21248/contrib.entomol.57.2.277-283, URL: https://www.contributions-to-entomology.org/article/view/1704
039787FAFF812118FF68FDAC27E8802A.taxon	diagnosis	Diagnosis: Anomala pumiloidespumiloides is very similar to A. pumilis but can easily be distinguished by the larger elongated body shape, the strongly rounded sides and the more coarse punctation of the pronotum and the less reflexed clypeus. Moreover, the hind tibiae are more fusiform in A. pumilispumilis and the parameres are distinctly longer (Figs 10, 11).	en	Zorn, Carsten (2007): Three new species of the genus Anomala S, 1819 from Borneo of the fulvofusca-species group (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (2): 277-283, DOI: 10.21248/contrib.entomol.57.2.277-283, URL: https://www.contributions-to-entomology.org/article/view/1704
039787FAFF812118FF68FDAC27E8802A.taxon	etymology	Etymology: The species name is referring to the notable resemblance of the new species to A. pumilis.	en	Zorn, Carsten (2007): Three new species of the genus Anomala S, 1819 from Borneo of the fulvofusca-species group (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (2): 277-283, DOI: 10.21248/contrib.entomol.57.2.277-283, URL: https://www.contributions-to-entomology.org/article/view/1704
