taxonID	type	description	language	source
03E6D8662140FFF9E3A040BBFE9EFCD1.taxon	description	Paralectotypes: 1  „ Whitehead | 68259 | Borneo Kina Balu | Fry Coll. 1905 - 100. “ (BMNH). 1  „ Whitehead; Borneo Kina Balu; Fry Coll. 1905 - 100.; Anomala felicia, Arrow Cotype “ (BMNH). [= Anomala kinabalensis Ohaus, 1910]; 1  „ Whitehead | Borneo Kina Balu | Fry Coll. 1905 - 100. “ (BMNH). [uncertain species identity]. Note: Because the type series is not monospecific a lectotype designation was necessary.	en	Zorn, Carsten (2007): Notes on Anomala felicia A ,, 1910 and its relatives from Borneo with descriptions of six new species (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (1): 81-91, DOI: 10.21248/contrib.entomol.57.1.81-91, URL: https://www.contributions-to-entomology.org/article/view/1693
03E6D8662140FFF9E3A040BBFE9EFCD1.taxon	materials_examined	Additional material: 1 , 1  „ Kinabalu Borneo “ (NHMB). 1  „ BORNEO, Sabah W., Crocker Range, W of Apinapin, ii. 2000, M. Snižek leg. “ (CCZ). 1  „ BORNEO, SABACH W. route RANAU – TAMBUNAN 2.2000 Ltg. Snižek “ (CCZ). 1 , 1  „ BORNEO Sabah Crocker Range Keningau V. 93 LEGRAND leg. “ (CDK).	en	Zorn, Carsten (2007): Notes on Anomala felicia A ,, 1910 and its relatives from Borneo with descriptions of six new species (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (1): 81-91, DOI: 10.21248/contrib.entomol.57.1.81-91, URL: https://www.contributions-to-entomology.org/article/view/1693
03E6D8662140FFF9E3A040BBFE9EFCD1.taxon	description	Redescription: Body shape. Elongate ovoid; length. 9.2 - 10.3 mm; width. 5.1 - 5.5 mm. Color. Dorsal face, abdominal sternites (except the last one in  ), tibiae and tarsi reddish brown with metallic green shine; ventral face including the femora, pygidium and antennae brownish yellow; lateral parts of the pronotum yellow; inner limitation of the yellow lateral band notched in the middle (Fig. 19); laterobasal part of pygidium sometimes darkened. Head. Clypeus trapezoid; clypeus and frons with sometimes confluent, rather coarse punctures; vertex finely and not densely punctate; clavus distinctly shorter than antennomeres 1 - 6 combined but longer than the funiculus. Pronotum. Ca. 1.7 x as broad as long, widest at base; sides subparallel in the posterior half, strongly convergent in the anterior half; anterior angles square; posterior angles obtuse and narrowly rounded; basal bead not interrupted; punctation fine and moderately dense on disc; punctures separated by 1 - 2 (3) diameters. Scutellum. Punctured like the disc of the pronotum. Elytra. With regular, impressed striae; intervals slightly elevated; 2 nd and 3 rd interstice with interrupted and not impressed secondary striae; subsutural interstice irregularly punctured anteriorly, with an indistinct stria in the posterior half. Pygidium. With sparse, rather coarse, ocellated punctation; distal margin with erect setae. Metasternum. With well-separated, ocellated punctures and rather short and sparse setosity in the lateral part. Abdominal sternites. Sparsely punctured in the middle, with denser punctures laterally; with a transverse row of setae. Protibiae. Bidentate, terminal tooth shortly bent outwards; lateral tooth small, square. Metatibiae. Rather short, a little enlarged in the middle and constricted before the apex. Claws. Modified claw of the pro- and mesotarsi apically bifid, protarsi enlarged but with angle or tooth at the inner margin. Aedeagus. Figs 1 - 2.	en	Zorn, Carsten (2007): Notes on Anomala felicia A ,, 1910 and its relatives from Borneo with descriptions of six new species (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (1): 81-91, DOI: 10.21248/contrib.entomol.57.1.81-91, URL: https://www.contributions-to-entomology.org/article/view/1693
03E6D8662140FFF9E3A040BBFE9EFCD1.taxon	diagnosis	Diagnosis: Anomala feliciafelicia is easily separated from the other members of this species group by the shape of the aedeagus which is simple and not elongate as in the other species. Moreover, the inner limitation of the yellow lateral patch of the pronotum is notched in the middle (Fig. 19), not evenly broad from anterior to posterior angle or extremely narrowed in the posterior half.	en	Zorn, Carsten (2007): Notes on Anomala felicia A ,, 1910 and its relatives from Borneo with descriptions of six new species (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (1): 81-91, DOI: 10.21248/contrib.entomol.57.1.81-91, URL: https://www.contributions-to-entomology.org/article/view/1693
03E6D8662140FFF9E3A040BBFE9EFCD1.taxon	distribution	Distribution: Anomala feliciafelicia is only known from the Kina Balu region in the province of Sabah (Fig. 22).	en	Zorn, Carsten (2007): Notes on Anomala felicia A ,, 1910 and its relatives from Borneo with descriptions of six new species (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (1): 81-91, DOI: 10.21248/contrib.entomol.57.1.81-91, URL: https://www.contributions-to-entomology.org/article/view/1693
03E6D8662143FFF9E21C4092FEE4F988.taxon	materials_examined	Holotype:  „ Museum Leiden Malaysia SABAH: Long Pa Sia: Banks of S. Pa Sia; sec veg. 4 ° 25´N 115 ° 43´E. 1090 m 14 - 28 Oct 1986. J. Huisman | RMNH Leiden Loan 11454 “ (RMNH). Paratypes: 2   „ Museum Leiden Malaysia SABAH: Long Pa Sia: Banks of S. Pa Sia; sec veg. 4 ° 25´N 115 ° 43´E. 1090 m 14 - 28 Oct 1986. J. Huisman | RMNH Leiden Loan 11454 “ (RMNH). 3   „ BORNEO, SABAH W. route RANAU – TAMBUNAN 2.2000 Ltg. Snizek “ (CCZ). 1  „ N. Borneo. Bettotan, NR. SANDAKAN. Aug. 12 th 1937 | Anomala kinabalensis Ohs. “ (NMHB). 1  „ N. Borneo. Bettotan, NR. SANDAKAN. Aug; 12 th 1937. | Anomala kinabalensis Ohs. “ (NMHB).	en	Zorn, Carsten (2007): Notes on Anomala felicia A ,, 1910 and its relatives from Borneo with descriptions of six new species (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (1): 81-91, DOI: 10.21248/contrib.entomol.57.1.81-91, URL: https://www.contributions-to-entomology.org/article/view/1693
03E6D8662143FFF9E21C4092FEE4F988.taxon	description	Description and diagnosis: Length. 9.2 - 10.0 mm; width. 5.1 - 5.6 mm. Anomala snizeki shares most morphological characters with A. feliciafelicia as described above. The inner limitation of the yellow lateral band of the pronotum is not notched but usually straight from anterior to posterior margin (Fig. 20), a character that is shared with A. kaltengensis, A. bifida, A. aequalis, A. kinabalensiskinabalensis and A. mahakamensis. From these five species, A. snizeki can only be distinguished by the shape of the aedeagus. The parameres are strongly elongate whereas the ventral plate shows no extensions (Figs 3 - 4).	en	Zorn, Carsten (2007): Notes on Anomala felicia A ,, 1910 and its relatives from Borneo with descriptions of six new species (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (1): 81-91, DOI: 10.21248/contrib.entomol.57.1.81-91, URL: https://www.contributions-to-entomology.org/article/view/1693
03E6D8662143FFF9E21C4092FEE4F988.taxon	distribution	Distribution: Anomala snizekisnizeki is known from various localities in Sabah (Fig. 22).	en	Zorn, Carsten (2007): Notes on Anomala felicia A ,, 1910 and its relatives from Borneo with descriptions of six new species (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (1): 81-91, DOI: 10.21248/contrib.entomol.57.1.81-91, URL: https://www.contributions-to-entomology.org/article/view/1693
03E6D8662143FFF9E21C4092FEE4F988.taxon	etymology	Etymology: The species is named after Miroslav Snížek (České Budějovice) who collected some of the paratypes.	en	Zorn, Carsten (2007): Notes on Anomala felicia A ,, 1910 and its relatives from Borneo with descriptions of six new species (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (1): 81-91, DOI: 10.21248/contrib.entomol.57.1.81-91, URL: https://www.contributions-to-entomology.org/article/view/1693
03E6D8662145FFFFE21C402DFBD2FB5E.taxon	materials_examined	Holotype:  „ Puak, Sarawak. G. E. Bryant. 5. V. 14 | G. Bryant Coll. B. M. 1926 - 86. “ (BMNH).	en	Zorn, Carsten (2007): Notes on Anomala felicia A ,, 1910 and its relatives from Borneo with descriptions of six new species (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (1): 81-91, DOI: 10.21248/contrib.entomol.57.1.81-91, URL: https://www.contributions-to-entomology.org/article/view/1693
03E6D8662145FFFFE21C402DFBD2FB5E.taxon	description	Description and diagnosis: Length. 9.2 mm; width. 5.3 mm. Anomala bifida shares most morphological characters with A. feliciafelicia as described above. The inner limitation of the yellow lateral band of the pronotum is not notched but usually straight from anterior to posterior margin (Fig. 20), a character that is shared with A. snizeki, A. kaltengensis, A. aequalis, A. kinabalensiskinabalensis and A. mahakamensis. From these five species, A. kaltengensis can only be distinguished by the shape of the aedeagus. Both, parameres and ventral plate are strongly elongate, the parameres are apically pointed (Figs 7 - 8).	en	Zorn, Carsten (2007): Notes on Anomala felicia A ,, 1910 and its relatives from Borneo with descriptions of six new species (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (1): 81-91, DOI: 10.21248/contrib.entomol.57.1.81-91, URL: https://www.contributions-to-entomology.org/article/view/1693
03E6D8662145FFFFE21C402DFBD2FB5E.taxon	distribution	Distribution: Anomala bifida is only known from the type locality Puak in the province of Sarawak (Fig. 22).	en	Zorn, Carsten (2007): Notes on Anomala felicia A ,, 1910 and its relatives from Borneo with descriptions of six new species (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (1): 81-91, DOI: 10.21248/contrib.entomol.57.1.81-91, URL: https://www.contributions-to-entomology.org/article/view/1693
03E6D8662145FFFFE21C402DFBD2FB5E.taxon	etymology	Etymology: The epithet is referring to the deeply incised parameres found in this species.	en	Zorn, Carsten (2007): Notes on Anomala felicia A ,, 1910 and its relatives from Borneo with descriptions of six new species (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (1): 81-91, DOI: 10.21248/contrib.entomol.57.1.81-91, URL: https://www.contributions-to-entomology.org/article/view/1693
03E6D8662145FFFFE21C43ABFD52FD45.taxon	materials_examined	Holotype:  „ Indonesia Kalimantan Tengah, Tewah 23. II. 96, secondary forest, leg. A. Kallies & C. Zorn “ (CCZ). Paratypes: 1 , 1  - same data as holotype (CCZ).	en	Zorn, Carsten (2007): Notes on Anomala felicia A ,, 1910 and its relatives from Borneo with descriptions of six new species (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (1): 81-91, DOI: 10.21248/contrib.entomol.57.1.81-91, URL: https://www.contributions-to-entomology.org/article/view/1693
03E6D8662145FFFFE21C43ABFD52FD45.taxon	description	Description and diagnosis: Length. 9.2 - 10.3 mm, width. 5.1 - 5.5 mm. Anomala kaltengensiskaltengensis shares most morphological characters with A. felicia as described above. The inner limitation of the yellow lateral band of the pronotum is not notched but usually straight from anterior to posterior margin (Fig. 20), a character that is shared with A. bifida, A. snizeki, A. aequalis, A. kinabalensis and A. mahakamensis. From these five species, A. kaltengensis can only be distinguished by the shape of the aedeagus. The parameres and ventral plate are strongly elongate, the parameres are somewhat thickened apicad (Figs 5 - 6).	en	Zorn, Carsten (2007): Notes on Anomala felicia A ,, 1910 and its relatives from Borneo with descriptions of six new species (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (1): 81-91, DOI: 10.21248/contrib.entomol.57.1.81-91, URL: https://www.contributions-to-entomology.org/article/view/1693
03E6D8662145FFFFE21C43ABFD52FD45.taxon	distribution	Distribution: Anomala kaltengensis is only known from the type locality Tewah in Central Kalimantan (Fig. 22).	en	Zorn, Carsten (2007): Notes on Anomala felicia A ,, 1910 and its relatives from Borneo with descriptions of six new species (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (1): 81-91, DOI: 10.21248/contrib.entomol.57.1.81-91, URL: https://www.contributions-to-entomology.org/article/view/1693
03E6D8662145FFFFE21C43ABFD52FD45.taxon	etymology	Etymology: The species is named after the abbreviation “ Kalteng ” for the type locality, the province Kalimantan Tengah (Central Kalimantan).	en	Zorn, Carsten (2007): Notes on Anomala felicia A ,, 1910 and its relatives from Borneo with descriptions of six new species (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (1): 81-91, DOI: 10.21248/contrib.entomol.57.1.81-91, URL: https://www.contributions-to-entomology.org/article/view/1693
03E6D8662145FFFEE21C4626FBB9F9EB.taxon	materials_examined	Holotype:  „ Ouest Borneo Deby | Coll. et determ. J. La Fontaine. | Coll. Kraatz | Ohaus det. “ (DEI).	en	Zorn, Carsten (2007): Notes on Anomala felicia A ,, 1910 and its relatives from Borneo with descriptions of six new species (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (1): 81-91, DOI: 10.21248/contrib.entomol.57.1.81-91, URL: https://www.contributions-to-entomology.org/article/view/1693
03E6D8662145FFFEE21C4626FBB9F9EB.taxon	description	Description and diagnosis: Length. 9.6 mm; width. 5.6 mm. Anomala aequalisaequalis shares most morphological characters with A. feliciafelicia as described above. The inner limitation of the yellow lateral band of the pronotum is not notched but usually straight from anterior to posterior margin (Fig. 20), a character that is shared with A. snizeki, A. kaltengensis, A. bifida, A. kinabalensiskinabalensis and A. mahakamensis. From these five species, A. aequalisaequalis can only be distinguished by the shape of the aedeagus. Both, parameres and ventral plate are strongly elongate. The parameres are apically not pointed nor angled (Figs 9 - 10).	en	Zorn, Carsten (2007): Notes on Anomala felicia A ,, 1910 and its relatives from Borneo with descriptions of six new species (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (1): 81-91, DOI: 10.21248/contrib.entomol.57.1.81-91, URL: https://www.contributions-to-entomology.org/article/view/1693
03E6D8662145FFFEE21C4626FBB9F9EB.taxon	distribution	Distribution: Anomala aequalisaequalis is only known from the imprecise type locality, West Borneo (Fig. 22).	en	Zorn, Carsten (2007): Notes on Anomala felicia A ,, 1910 and its relatives from Borneo with descriptions of six new species (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (1): 81-91, DOI: 10.21248/contrib.entomol.57.1.81-91, URL: https://www.contributions-to-entomology.org/article/view/1693
03E6D8662145FFFEE21C4626FBB9F9EB.taxon	etymology	Etymology: The species name was chosen because A. aequalis cannot be distinguished by external characters from most of its relatives but can only be differentiated by the shape of the aedeagus.	en	Zorn, Carsten (2007): Notes on Anomala felicia A ,, 1910 and its relatives from Borneo with descriptions of six new species (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (1): 81-91, DOI: 10.21248/contrib.entomol.57.1.81-91, URL: https://www.contributions-to-entomology.org/article/view/1693
03E6D8662147FFFDE21C43B4FEE2FB28.taxon	description	Paralectotype: 1  “ N. BORNEO Kina Balu Waterstraat | Anomala kinabalensis Ohs. Cotype ” (NMHB). Note: Because it is not entirely certain that the type series is monospecific a lectotype designation was necessary.	en	Zorn, Carsten (2007): Notes on Anomala felicia A ,, 1910 and its relatives from Borneo with descriptions of six new species (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (1): 81-91, DOI: 10.21248/contrib.entomol.57.1.81-91, URL: https://www.contributions-to-entomology.org/article/view/1693
03E6D8662147FFFDE21C43B4FEE2FB28.taxon	materials_examined	Additional material: 1  “ BORNEO Grubauer | Anomala kinabalensis Ohs. Cotype ” (NMHB). 1  “ Limbang 12 - 2 - 10 | Anomala sarawakensis Ohs. Cotype ” (NMHB). 10 , 3  “ N. Borneo Brunei Waterstradt S. ” (ZMHB, CCZ). 1  “ Nord-Borneo Kinabalu III. 1969 | Nord-Borneo Kinabalu III. 1969 | Anomala kinabaluensis Ohs. det. G. Frey, 1967 / 68 ” (NHMB). 1  “ Borneo | Ohaus determin. Anomala kinabalensis Ohs. ” (NHMB). 1 , 3  “ Sabah: Sook, 1500 ft. 17 m. SW Keningau, 15. viii. 1977 | At light | M. E. Bacchus B. M. 1978 - 48 ” (BMNH). 1  “ SABAH: Mt. Kinabalu Tenom Keningau 4 - 8. iii. 1964. J. Smart. Royal Soc. Exped. B. M. 1964 - 250 ” (BMNH). 3  “ Borneo | Museum Leiden verz. F. T. Valck Lucassen | Anomala kinabalensis Ohs. det. Ohaus 1928 ” (RMNH). 1 , 1  “ Kinabalu, Borneo, 1500 m H. Rolle, Berlin S. W. 1 l | Coll. Kraatz | Ohaus det. ” (DEI). 1  “ Kinabalu, Borneo, 1500 m H. Rolle, Berlin S. W. 1 l | Coll. Kraatz | Anomala kinabalensis Ohs. | Ohaus det. ” (DEI). 1  “ N. BORNEO: Bukit Kretam area. J. D. H. Hedley. B. M. 1952 - 463. ” (BMNH). 1  “ N. BORNEO: Bukit Kretam area. J. D. H. Hedley. B. M. 1952 - 463. ” (BMNH).	en	Zorn, Carsten (2007): Notes on Anomala felicia A ,, 1910 and its relatives from Borneo with descriptions of six new species (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (1): 81-91, DOI: 10.21248/contrib.entomol.57.1.81-91, URL: https://www.contributions-to-entomology.org/article/view/1693
03E6D8662147FFFDE21C43B4FEE2FB28.taxon	diagnosis	Diagnosis: Length. 9.1 - 10.5 mm; width. 5.2 - 6.2 mm. Anomala kinabalensiskinabalensis shares most morphological characters with A. feliciafelicia as described above. The inner limitation of the yellow lateral band of the pronotum is not notched but usually straight from anterior to posterior margin (Fig. 20), a character that is shared with A. snizeki, A. kaltengensis, A. bifida, A. aequalisaequalis and A. mahakamensis. From these five species, A. kinabalensis can only be distinguished by the shape of the aedeagus. Both, parameres and ventral plate are strongly elongate. The parameres are angled apically and bent downwards and are very similar to those of A. mahakamensis. The extension of the ventral plate is apically broader than in A. mahakamensis (Figs 11 - 12).	en	Zorn, Carsten (2007): Notes on Anomala felicia A ,, 1910 and its relatives from Borneo with descriptions of six new species (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (1): 81-91, DOI: 10.21248/contrib.entomol.57.1.81-91, URL: https://www.contributions-to-entomology.org/article/view/1693
03E6D8662147FFFDE21C43B4FEE2FB28.taxon	distribution	Distribution: Anomala kinabalensiskinabalensis is only known from the Kinabalu region in the province of Sabah (Fig. 22).	en	Zorn, Carsten (2007): Notes on Anomala felicia A ,, 1910 and its relatives from Borneo with descriptions of six new species (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (1): 81-91, DOI: 10.21248/contrib.entomol.57.1.81-91, URL: https://www.contributions-to-entomology.org/article/view/1693
03E6D8662147FFF3E21C47C9FB9AFEA3.taxon	materials_examined	Holotype:  “ Oost-BORNEO Sangasanga-dalem | coll. Zoölogisch Museum acq. 1938 ” (ZMAN).	en	Zorn, Carsten (2007): Notes on Anomala felicia A ,, 1910 and its relatives from Borneo with descriptions of six new species (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (1): 81-91, DOI: 10.21248/contrib.entomol.57.1.81-91, URL: https://www.contributions-to-entomology.org/article/view/1693
03E6D8662147FFF3E21C47C9FB9AFEA3.taxon	description	Description and Diagnosis: Length. 11.1 mm; width. 6.9 mm. Anomala mahakamensismahakamensis shares most morphological characters with A. felicia as described above. With A. snizeki, A. kaltengensis, A. bifida, A. kinabalensis and A. aequalisaequalis it has in common a yellow lateral marking of the pronotum of which the inner limitation is not notched but usually straight from anterior to posterior margin (Fig. 20). From these five species, A. mahakamensis can only be distinguished by the shape of the aedeagus which is extremely similar to that of A. kinabalensis. Both, parameres and ventral plate are strongly elongate. The parameres are angled apically and bent downwards. The extension of the ventral plate is apically narrower than in A. kinabalensis (Figs 13 - 14).	en	Zorn, Carsten (2007): Notes on Anomala felicia A ,, 1910 and its relatives from Borneo with descriptions of six new species (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (1): 81-91, DOI: 10.21248/contrib.entomol.57.1.81-91, URL: https://www.contributions-to-entomology.org/article/view/1693
03E6D8662147FFF3E21C47C9FB9AFEA3.taxon	distribution	Distribution: Anomala mahakamensismahakamensis is only known from Sangasanga-dalam in the province of Kalimantan Timur (East Kalimantan) (Fig. 22).	en	Zorn, Carsten (2007): Notes on Anomala felicia A ,, 1910 and its relatives from Borneo with descriptions of six new species (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (1): 81-91, DOI: 10.21248/contrib.entomol.57.1.81-91, URL: https://www.contributions-to-entomology.org/article/view/1693
03E6D8662147FFF3E21C47C9FB9AFEA3.taxon	etymology	Etymology: The new species is named after the Mahakam river in East Kalimantan.	en	Zorn, Carsten (2007): Notes on Anomala felicia A ,, 1910 and its relatives from Borneo with descriptions of six new species (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (1): 81-91, DOI: 10.21248/contrib.entomol.57.1.81-91, URL: https://www.contributions-to-entomology.org/article/view/1693
03E6D8662149FFF3E21C424CFCD3FB12.taxon	materials_examined	Holotype:  „ At light | SARAWAK: Gungong Mulu Nat. Oark [sic!] R. G. S. Exped. 1977 - 8 J. D. Holloway et al. B. M. 1978 - 206 | Site 1. January Camp 4, Mulu 1790 m. 452463 Lower montane (moss) forest, MV – canopy “ (BMNH). Paratypes: 1  „ SARAWAK: Gunong Mulu Nat. Park R. G. S. Exped. 1977 - 8 J. D. Holloway et al. B. M. 1978 - 206 | Site 19. March W. Melinau Gorge 100 m. 457567 Alluvial forest. Acl-understorey “ (CCZ). 1  „ Old Secondary forest. | Native collected | SARAWAK: Foot of Mt. Dulit. Junction of rivers Tinjar & Lejok. 6. x. 1932. | Oxford Univ. Exp. B. M. Hobby & A. W. Moore. B. M. 1933 - 254 | Ohaus determ. Anomala felicia Arr.?  “ (BMNH). 1 , 1  „ Dr. J. Bosscha Sambas Borneo occ. | Mus. Leiden | Anomala kinabalensis Ohs. Cotype “ (NMHB, CCZ). 1  „ Sarawak Baram R. | Anomala sarawakensis Cotype Ohs. “ (NMHB). 2 , 5  „ Dr. J. Bosscha Sambas Borneo occ. | Museum Leiden ANOMALA (A.) SARAWAKENSIS OHS. “ (CCZ, RMNH). 1  „ Dr. J. Bosscha Sambas Borneo occ. | Ohaus determ. Anomala sarawakensis Ohs. “ (RMNH).	en	Zorn, Carsten (2007): Notes on Anomala felicia A ,, 1910 and its relatives from Borneo with descriptions of six new species (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (1): 81-91, DOI: 10.21248/contrib.entomol.57.1.81-91, URL: https://www.contributions-to-entomology.org/article/view/1693
03E6D8662149FFF3E21C424CFCD3FB12.taxon	description	Description and diagnosis: Length. 9.5 - 10.4 mm; width. 5.5 - 6.1 mm. Anomala manseri shares most morphological characters with A. felicia as described above. The inner limitation of the yellow lateral band of the pronotum is strongly narrowed towards the posterior angle, a character that is only shared with A. sarawakensissarawakensis (Fig. 21). Moreover, A. manserimanseri and A. sarawakensissarawakensis appear to be slightly larger than the other species of this complex, but too few specimens are available for final conclusions. From A. sarawakensissarawakensis the new species can only be distinguished by the shape of the aedeagus. The parameres are much more elongated and pointed than in A. sarawakensissarawakensis (Figs 15 - 16).	en	Zorn, Carsten (2007): Notes on Anomala felicia A ,, 1910 and its relatives from Borneo with descriptions of six new species (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (1): 81-91, DOI: 10.21248/contrib.entomol.57.1.81-91, URL: https://www.contributions-to-entomology.org/article/view/1693
03E6D8662149FFF3E21C424CFCD3FB12.taxon	distribution	Distribution: Anomala manserimanseri is known from the Baram River region in east Sarawak and Sambas in West Kalimantan (Fig. 22).	en	Zorn, Carsten (2007): Notes on Anomala felicia A ,, 1910 and its relatives from Borneo with descriptions of six new species (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (1): 81-91, DOI: 10.21248/contrib.entomol.57.1.81-91, URL: https://www.contributions-to-entomology.org/article/view/1693
03E6D8662149FFF3E21C424CFCD3FB12.taxon	etymology	Etymology: This species is dedicated to Bruno Manser, a Swiss rain forest and human rights activist who is missing in Sarawak since 2000.	en	Zorn, Carsten (2007): Notes on Anomala felicia A ,, 1910 and its relatives from Borneo with descriptions of six new species (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (1): 81-91, DOI: 10.21248/contrib.entomol.57.1.81-91, URL: https://www.contributions-to-entomology.org/article/view/1693
03E6D8662149FFF2E21C47D7FD78F9D3.taxon	description	Syntypes: 1  “ Sarawak Mt. Penrissen 3300 ft. V. 1900 | Anomala sar- awakensis Ohs. Type ” (NMHB). 1  “ Penrissen May 1899 |  | Typus | A. ovatula Ohs. sbsp. sarawakensis Ohs. ” (NMHB). 1  “ Penrissen May 1899 |  | Typus! | A. ovatula Ohs. sbsp. sarawakensis Ohs. ” (NMHB). Note: In the collection of the NMHB are additional specimens labeled as “ Cotypen ” which are not mentioned in the original description and belong in fact to A. manseri. Fig. 20, Anomala snizeki; Fig. 21: Anomala manseri.	en	Zorn, Carsten (2007): Notes on Anomala felicia A ,, 1910 and its relatives from Borneo with descriptions of six new species (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (1): 81-91, DOI: 10.21248/contrib.entomol.57.1.81-91, URL: https://www.contributions-to-entomology.org/article/view/1693
03E6D8662149FFF2E21C47D7FD78F9D3.taxon	materials_examined	Additional material: 1  “ Mt. Matang, W. Sarawak. G. E. Bryant. 2. XII. 13. | G. Bryant Coll. B. M. 1926 - 86. ” (BMNH). 1  “ Mt. Matang, W. Sarawak. G. E. Bryant. 12. 11. 14. | Anomala felicia Arrow [sic!] ” (NHMB).	en	Zorn, Carsten (2007): Notes on Anomala felicia A ,, 1910 and its relatives from Borneo with descriptions of six new species (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (1): 81-91, DOI: 10.21248/contrib.entomol.57.1.81-91, URL: https://www.contributions-to-entomology.org/article/view/1693
03E6D8662149FFF2E21C47D7FD78F9D3.taxon	diagnosis	Diagnosis: Length. 10.5 - 11.5 mm; width. 5.9 - 6.5 mm. Anomala sarawakensissarawakensis shares most morphological characters with A. feliciafelicia as described above. The inner limitation of the yellow lateral band of the pronotum is strongly narrowed towards the posterior angle, a character that is only shared with A. manseri (Fig. 21). Moreover, A. manserimanseri and A. sarawakensissarawakensis appear to be slightly larger in mean than the other species of this complex, but too few specimens are available for final conclusions. Anomala sarawakensisawakensis can only be distinguished from A. manserimanseri by the shape of the aedeagus. The parameres are distinctly shorter than in A. manseri. The apex is somewhat bulged and not pointed (Figs 17 - 18).	en	Zorn, Carsten (2007): Notes on Anomala felicia A ,, 1910 and its relatives from Borneo with descriptions of six new species (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (1): 81-91, DOI: 10.21248/contrib.entomol.57.1.81-91, URL: https://www.contributions-to-entomology.org/article/view/1693
03E6D8662149FFF2E21C47D7FD78F9D3.taxon	distribution	Distribution: Anomala sarawakensissarawakensis is only known from west Sarawak (Gunung Penrissen, Gunung Matang) (Fig. 22). One female from Mt. Lingga cannot be identified with certainty and belongs to A. sarawakensis, A. manseri or a yet unknown species.	en	Zorn, Carsten (2007): Notes on Anomala felicia A ,, 1910 and its relatives from Borneo with descriptions of six new species (Coleoptera: Scarabaeidae: Rutelinae). Beiträge Zur Entomologie = Contributions to Entomology 57 (1): 81-91, DOI: 10.21248/contrib.entomol.57.1.81-91, URL: https://www.contributions-to-entomology.org/article/view/1693
