taxonID	type	description	language	source
03FE8550C4244B0DFF6AFC8936ECFADE.taxon	type_taxon	Type species: Micryphantes flavomaculatus C. L. Koch, 1836 (= Euryopis flavomaculata)	en	Rodrigues, Everton Nei Lopes, Marta, Kimberly S., Figueiredo, Cassiano P. (2021): Description of four new species of the spider genus Euryopis Menge, 1868, with new records from Brazil (Araneae, Theridiidae, Hadrotarsinae). Zootaxa 4966 (5): 535-549, DOI: https://doi.org/10.11646/zootaxa.4966.5.3
03FE8550C4244B0DFF6AFC8936ECFADE.taxon	diagnosis	Diagnosis. Euryopis differs from Dipoena, Lasaeola and Phycosoma by the absence of carapace spherical or rounded and by the absence of distinct cervical grooves. Euryopis presents carapace oval (Figs 1 A, 2 A, 3 A, 4 A), not modified. Differs from Guaraniella by the cymbium without deep apical notch. Differs from Audifia by the abdomen without a sclerotized ring around the spinnerets and pedicel and differs from Audifia by the not elongate carapace. Euryopis has an oval (Figs 1 A, 2 A, 4 A, 5 A) or subtriangular abdomen (see Levi 1963, Fig. 9), dorsoventrally flat (Fig. 7 D), sometimes project over the carapace; blackish brown to black (Fig. 1 A), sometimes with silver dots and males with sclerotized scutum on the dorsum (Figs 7 A, D). The females of Euryopis have the epigynum with an oval depression (Fig. 9 C); and males have a distal cymbial projection (Fig. 8 A), however there are some species lacking such projection (Fig. 8 D) or having it little pronounced (Figs 8 B, C).	en	Rodrigues, Everton Nei Lopes, Marta, Kimberly S., Figueiredo, Cassiano P. (2021): Description of four new species of the spider genus Euryopis Menge, 1868, with new records from Brazil (Araneae, Theridiidae, Hadrotarsinae). Zootaxa 4966 (5): 535-549, DOI: https://doi.org/10.11646/zootaxa.4966.5.3
03FE8550C4244B08FF6AFA92344FFBA4.taxon	description	Figs. 1 A – D, 2 A – D, 8 A, 9 A – B, 10 LSIDurn: lsid: zoobank. org: act: C 4 B 11509 - 2611 - 47 E 3 - B 71 A-C 3 BF 3 A 44 DAAD	en	Rodrigues, Everton Nei Lopes, Marta, Kimberly S., Figueiredo, Cassiano P. (2021): Description of four new species of the spider genus Euryopis Menge, 1868, with new records from Brazil (Araneae, Theridiidae, Hadrotarsinae). Zootaxa 4966 (5): 535-549, DOI: https://doi.org/10.11646/zootaxa.4966.5.3
03FE8550C4244B08FF6AFA92344FFBA4.taxon	materials_examined	Type material. Holotype. Brazil: Rio Grande do Sul: Candiota, Miolo farm, 31 ° 23 ’ 00.0 ” S 53 ° 45 ’ 53.7 ” W, 04. X. 2010, J. L. O. Rosado leg., 1 M (MCN 56854). Paratypes. Brazil: Rio Grande do Sul, Candiota, Miolo farm, 31 ° 23 ’ 00.0 ” S, 53 ° 45 ’ 53.7 ” W, 04. X. 2010, J. L. O. Rosado leg., 2 F (MCN 56855); 18. II. 2011, J. L. O. Rosado leg., 5 M (MCN 56856); 18. II. 2011, J. L. O. Rosado leg., 1 F (MCN 56857); 18. II. 2011, J. L. O. Rosado leg., 1 M (MCN 56848); 04. X. 2010, J. L. O. Rosado leg., 2 M, 3 F (MCN 56849); 18. II. 2011, J. L. O. Rosado leg., 4 M, 2 F (MCTP 44010);, 25. VIII. 2011, J. L. O. Rosado leg., 1 M, 1 F (MCTP 44011); 04. X. 2010, J. L. O. Rosado leg., 1 M, 2 F (IBSP 267939); 25. VIII. 2011, J. L. O. Rosado leg., 1 M, 1 F (IBSP 267940); Rio Grande do Sul: Bagé, Malafai farm, 31 ° 16 ’ 00.0 ” S, 54 ° 07 ’ 00.0 ” W, 22. X. 2009, J. L. O. Rosado leg., 1 M (MCTP 44020); 30. V. 2011, J. L. O. Rosado leg., 5 M (MCTP 44021); 22. X. 2009, J. L. O Rosado leg., 1 M (MCN 56850). Rio Grande do Sul, Bagé, Peruzzo farm, 31 ° 15 ’ 05.0 ” S, 54 ° 05 ’ 00.0 ” W, 18. II. 2011, J. L. O. Rosado leg., 2 M (MCTP 44022); 17. V. 2010, J. L. O. Rosado leg., 1 F (MCTP 44023). All collected by pitfall-trap.	en	Rodrigues, Everton Nei Lopes, Marta, Kimberly S., Figueiredo, Cassiano P. (2021): Description of four new species of the spider genus Euryopis Menge, 1868, with new records from Brazil (Araneae, Theridiidae, Hadrotarsinae). Zootaxa 4966 (5): 535-549, DOI: https://doi.org/10.11646/zootaxa.4966.5.3
03FE8550C4244B08FF6AFA92344FFBA4.taxon	etymology	Etymology. The specific name is a noun in apposition, taken from the type locality.	en	Rodrigues, Everton Nei Lopes, Marta, Kimberly S., Figueiredo, Cassiano P. (2021): Description of four new species of the spider genus Euryopis Menge, 1868, with new records from Brazil (Araneae, Theridiidae, Hadrotarsinae). Zootaxa 4966 (5): 535-549, DOI: https://doi.org/10.11646/zootaxa.4966.5.3
03FE8550C4244B08FF6AFA92344FFBA4.taxon	diagnosis	Diagnosis. Males of Euryopis candiota n. sp. resemble those of Euryopis talaveraensis González, 1991 (see González 1991, figs 7 – 9), but differ from the latter species by the basal position of the spermatic ducts on the tegulum (Fig. 8 A), while the spermatic duct is positioned apically in E. talaveraensis; the embolus with well sclerotized apex (Fig. 8 A) and conductor distally pointed not trespassing the edge of the cymbium (Fig. 8 A), whereas E. talaveraensis has a non-sclerotized embolus and conductor distally rounded; the cymbium apex of E. candiota n. sp. has a thin projection (Fig. 8 A), absent in E. talaveraensis, and differ by the shape of carapace, pear shaped (Fig. 1 A), while the later present a rounded shape. Females of E. candiota n. sp. also resemble those of E. talaveraensis González, 1991 (see González 1991, figs 2 – 4), but differ by the small copulatory opening, medially positioned (Fig. 9 A), copulatory opening is large and basally positioned in E. talaveraensis and the spermathecae, closer to each other (Fig. 9 B), but further apart in the later species.	en	Rodrigues, Everton Nei Lopes, Marta, Kimberly S., Figueiredo, Cassiano P. (2021): Description of four new species of the spider genus Euryopis Menge, 1868, with new records from Brazil (Araneae, Theridiidae, Hadrotarsinae). Zootaxa 4966 (5): 535-549, DOI: https://doi.org/10.11646/zootaxa.4966.5.3
03FE8550C4244B08FF6AFA92344FFBA4.taxon	description	Description. Male holotype (MCN 56854). Total length 2.43. Carapace length 0.94, width 0.71, height 0.37. Clypeus height 0.25. Sternum length 0.44, width 0.42. Abdomen length 1.17, width 0.81, height 0.63. Leg formula IV / I / II / III; segment length (I / II / III / IV): femora 0.48 / 0.42 / 0.42 / 0.56; patellae + tibiae 0.67 / 0.60 / 0.56 / 0.81; metatarsi + tarsi 0.72 / 0.60 / 0.62 / 0.79; total 1.87 / 1.62 / 1.60 / 2.16. Carapace pear-shaped (Fig. 1 A), reddish brown (Fig. 1 A, D). Chelicerae lighter than carapace. Ocular region slightly pronounced, eyes with black margins. Eye diameter and interdistances: AME 0.04, PME 0.04, ALE 0.06, PLE 0.06, AME – AME 0.04, AME – ALE 0.02, PME – PME 0.04, PME – PLE 0.04. Sternum (Fig. 1 B) paler than the carapace, with blackened edges. Reddish brown elongated pedicel with two longitudinal blackish bands (Fig. 1 A, B). Abdomen dorsally oval (Fig. 1 A), darker. Spinnerets pale yellow. Cymbium oval in dorsal view; with thin projection apically; embolus and conductor sclerotized, both distally pointed (Fig. 8 A); the spermatic ducts basally in the tegulum, form loops ventrally next to the subtegulum. Female paratype (MCN 56855). Total length 2.49. Carapace length 0.94, width 0.71. Clypeus height 0.21. Sternum length 0.56, width 0.48. Abdomen length 1.42, width 0.98, height 0.90. Leg formula IV / I / II / III; segment length (I / II / III / IV): femora 0.48 / 0.42 / 0.42 / 0.56; patellae + tibiae 0.67 / 0.60 / 0.56 / 0.81; metatarsi + tarsi 0.72 / 0.60 / 0.62 / 0.79; total 1.87 / 1.62 / 1.60 / 2.16. Carapace (Fig. 2 A) yellowish brown, lighter than male, with elevated cephalic region (Fig. 2 D). Chelicerae yellowish brown, same coloration as the labium. Ocular region slightly pronounced, eyes with black edges (Fig. 2 C). Eye diameter and interdistances: AME 0.06, PME 0.06, ALE 0.08, PLE 0.04, AME – AME 0.06, AME – ALE 0.02, PME – PME 0.06, PME – PLE 0.04. Sternum darker than the carapace, with blackened edges (Fig. 2 B). Pedicel yellowish brown elongated with two longitudinal blackish bands (Fig. 2 A). In dorsal view, abdomen oval (Fig. 2 A). In ventral view (Fig. 2 B), slightly yellowish brown, laterally blackened. Spinnerets pale yellow. Epigynum with a slightly sclerotized plate, longer than wider (Fig. 9 A); with one oval copulatory opening positioned medially on the plate (Fig. 9 A). Internal genitalia with two pairs of round spermathecae, close together (Fig. 9 B); uncoiled copulatory ducts (Fig. 9 B) and short and uncoiled fertilization ducts (Fig. 9 B). Natural history. It is already known that spiders of the genus Euryopis are associated with myrmecophagy, which was suggested by several authors (Berland 1933; Levi 1954; Carico 1978; Gertsch 1979; Porter & Eastmond 1982; Forster et al. 1990; Cushing 2012; Líznarová & Pekár 2019). The area where the material was collected is rich in myrmecofauna (Rosado et al. 2013) which corroborates the idea proposed by Foster et al. (1990) that Hadrotarsinae would be myrmecophages. This species is abundantly found in the soil of the vineyards in Southern Brazil.	en	Rodrigues, Everton Nei Lopes, Marta, Kimberly S., Figueiredo, Cassiano P. (2021): Description of four new species of the spider genus Euryopis Menge, 1868, with new records from Brazil (Araneae, Theridiidae, Hadrotarsinae). Zootaxa 4966 (5): 535-549, DOI: https://doi.org/10.11646/zootaxa.4966.5.3
03FE8550C4244B08FF6AFA92344FFBA4.taxon	distribution	Distribution. Brazil (state of Rio Grande do Sul, Fig. 10).	en	Rodrigues, Everton Nei Lopes, Marta, Kimberly S., Figueiredo, Cassiano P. (2021): Description of four new species of the spider genus Euryopis Menge, 1868, with new records from Brazil (Araneae, Theridiidae, Hadrotarsinae). Zootaxa 4966 (5): 535-549, DOI: https://doi.org/10.11646/zootaxa.4966.5.3
03FE8550C4244B08FF6AFA92344FFBA4.taxon	description	Variation. Carapace in males and females range from reddish brown to yellowish brown. Pigmentation in dorsal and ventral view of abdomen and sternum of males and females range from reddish brown to yellow. Additional material examined. Brazil. Rio Grande do Sul: Candiota (Miolo farm, 31 ° 23 ’ 44.8 ” S, 53 ° 45 ’ 53.7 ” W), 26. II. 2010, J. L. O Rosado leg., 1 M (MZUnisinos Ara 00863); 1 M, 1 F (MZUnisinos Ara 00880); 1 F (MZUnisinos Ara 00885); 1 M (MZUnisinos Ara 00897); 1 M (MZUnisinos Ara 00891); 1 M (MZUnisinos Ara 00908); 17. V. 2010, 1 M (MZUnisinos Ara 00862); 1 M (MZUnisinos Ara 00893); 1 M (MZUnisinos Ara 00902); 2 M (MZUnisinos Ara 00843); 1 M (MZUnisinos Ara 00859); 1 M (MZUnisinos Ara 00864); 1 M (MZUnisinos Ara 00872); 1 M (MZUnisinos Ara 00879); 1 M (MZUnisinos Ara 00881); 06. VIII. 2010, 2 M (MZUnisinos Ara 00854); 1 M (MZUnisinos Ara 00855); 1 F (MZUnisinos Ara 00884); 1 M (MZUnisinos Ara 00894); 1 F (MZUnisinos Ara 00901); 2 M (MZUnisinos Ara 00903); 1 M (MZUnisinos Ara 00909); 1 M (MZUnisinos Ara 00910); 04. X. 2010, 1 M (MZUnisinos Ara 00856); 1 M (MZUnisinos Ara 00858); 1 M (MZUnisinos Ara 00860); 2 M (MZUnisinos Ara 00861); 3 M (MZUnisinos Ara 00865); 1 M (MZUnisinos Ara 00866); 2 M (MZUnisinos Ara 00867); 1 M (MZUnisinos Ara 00868); 3 M (MZUnisinos Ara 00869); 7 M (MZUnisinos Ara 00870); 1 M (MZUnisinos Ara 00871); 18. II. 2011, 2 M (MZUnisinos Ara 00836); 1 M (MZUnisinos Ara 00837); 1 M (MZUnisinos Ara 00838); 1 M (MZUnisinos Ara 00839); 2 M (MZUnisinos Ara 00840); 1 M (MZUnisinos Ara 00841); 2 M (MZUnisinos Ara 00842); 1 M (MZUnisinos Ara 00845); 1 M (MZUnisinos Ara 00846); 1 M (MZUnisinos Ara 00847); 1 M (MZUnisinos Ara 00848); 6 M (MZUnisinos Ara 00849); 2 M (MZUnisinos Ara 00850); 1 M (MZUnisinos Ara 00851); 1 M (MZUnisinos Ara 00852); 1 M (MZUnisinos Ara 00853); 1 M (MZUnisinos Ara 00883); 1 M (MZUnisinos Ara 00888); 1 M (MZUnisinos Ara 00895); 4 M (MZUnisinos Ara 00962); 2 M (MZUnisinos Ara 00963); 30. V. 2011, 1 M (MZUnisinos Ara 00857); 1 M (MZUnisinos Ara 00886); 1 M (MZUnisinos Ara 00887); 1 M (MZUnisinos Ara 00890); 1 M (MZUnisinos Ara 00896); 1 F (MZUnisinos Ara 00898); 2 M (MZUnisinos Ara 00899); 2 M (MZUnisinos Ara 00900); 2 M (MZUnisinos Ara 00951); 1 M (MZUnisinos Ara 00952); 2 M (MZUnisinos Ara 00953); 1 M (MZUnisinos Ara 00954); 1 M (MZUnisinos Ara 00955); 1 M (MZUnisinos Ara 00956); 1 M (MZUnisinos Ara 00957); 1 M (MZUnisinos Ara 00958); 1 M (MZUnisinos Ara 00959); 1 F (MZUnisinos Ara 00960); 1 F (MZUnisinos Ara 00961); 25. VIII. 2008, 1 M (MZUnisinos Ara 00873); 1 F (MZUnisinos Ara 00874); 1 M (MZUnisinos Ara 00875); 1 M (MZUnisinos Ara 00876); 3 M (MZUnisinos Ara 00877); 1 M (MZUnisinos Ara 00878); 1 M (MZUnisinos Ara 00882); 1 M (MZUnisinos Ara 00964); Rio Grande do Sul: Bagé (Peruzzo farm, 31 ° 15 ’ 05.0 ” S, 54 ° 05 ’ 00.0 ” W), 04. X. 2010, J. L. O Rosado leg., 1 M (MZUnisinos Ara 00973); 1 M (MZUnisinos Ara 00974); 17. V. 2010, 1 M (MZUnisinos Ara 00981); 18. II. 2011, 1 M (MZUnisinos Ara 00965); 1 M (MZUnisinos Ara 00968); 1 M (MZUnisinos Ara 00971); 25. VIII. 2011, 1 M (MZUnisinos Ara 00972); 1 M (MZUnisinos Ara 00982); 1 M (MZUnisinos Ara 00967); 30. V. 2011, 4 M (MZUnisinos Ara 00969); 1 M (MZUnisinos Ara 00970); 1 M (MZUnisinos Ara 00975); 1 M (MZUnisinos Ara 00977); 1 M (MZUnisinos Ara 00978); 1 M (MZUnisinos Ara 00979); 1 M (MZUnisinos Ara 00980). All collected by pitfall-trap.	en	Rodrigues, Everton Nei Lopes, Marta, Kimberly S., Figueiredo, Cassiano P. (2021): Description of four new species of the spider genus Euryopis Menge, 1868, with new records from Brazil (Araneae, Theridiidae, Hadrotarsinae). Zootaxa 4966 (5): 535-549, DOI: https://doi.org/10.11646/zootaxa.4966.5.3
03FE8550C4214B0AFF6AFB043620FA7C.taxon	description	Figs. 3 A – D, 4 A – D, 8 B, 9 C, D, 10 LSIDurn: lsid: zoobank. org: act: 7 A 1 C 6 E 5 A- 90 BE- 48 A 3 - 919 E- 12 CCC 33 BF 8 DC	en	Rodrigues, Everton Nei Lopes, Marta, Kimberly S., Figueiredo, Cassiano P. (2021): Description of four new species of the spider genus Euryopis Menge, 1868, with new records from Brazil (Araneae, Theridiidae, Hadrotarsinae). Zootaxa 4966 (5): 535-549, DOI: https://doi.org/10.11646/zootaxa.4966.5.3
03FE8550C4214B0AFF6AFB043620FA7C.taxon	materials_examined	Type material. Holotype. Brazil: Paraná: Palmas, Refugio de Vida Silvestre dos Campos de Palmas [Campos de Palmas Wildlife Refuge], 26 ° 32 ’ 22.50 ” S, 51 ° 39 ’ 11.93 ” W, 22 – 29. X. 2018, G. Galeti et al. leg., 1 M (IBSP 269920). Paratypes. Brazil: Paraná: Palmas, Refugio de Vida Silvestre dos Campos de Palmas, 26 ° 32 ’ 22.50 ” S, 51 ° 39 ’ 11.93 ” W, 22 – 29. X. 2018, G. Galeti et al. leg., 1 F (IBSP 269921); 22 – 29. X. 2018, G. Galeti et al. leg., 1 F (IBSP 269922); 22 – 29. X. 2018, G. Galeti et al. leg., 1 M (IBSP 269923); 22 – 29. X. 2018, G. Galeti et al. leg., 1 M (MCN 56851); 22 – 29. X. 2018, G. Galeti et al. leg., 1 M (MCTP 44012); 22 – 29. X. 2018, G. Galeti et al. leg., 1 M (IBSP 267941). All collected by pitfall-trap.	en	Rodrigues, Everton Nei Lopes, Marta, Kimberly S., Figueiredo, Cassiano P. (2021): Description of four new species of the spider genus Euryopis Menge, 1868, with new records from Brazil (Araneae, Theridiidae, Hadrotarsinae). Zootaxa 4966 (5): 535-549, DOI: https://doi.org/10.11646/zootaxa.4966.5.3
03FE8550C4214B0AFF6AFB043620FA7C.taxon	etymology	Etymology. The specific name is a reference to the somatic characteristic from the male of this species, which has a nose-like projection in the clypeus.	en	Rodrigues, Everton Nei Lopes, Marta, Kimberly S., Figueiredo, Cassiano P. (2021): Description of four new species of the spider genus Euryopis Menge, 1868, with new records from Brazil (Araneae, Theridiidae, Hadrotarsinae). Zootaxa 4966 (5): 535-549, DOI: https://doi.org/10.11646/zootaxa.4966.5.3
03FE8550C4214B0AFF6AFB043620FA7C.taxon	diagnosis	Diagnosis. Males of Euryopis nasuta n. sp. resemble those of Euryopis camis Levi, 1963 (see Levi, 1963, fig. 19) but E. nasuta n. sp. differs from E. camis by the sclerotized and elongated embolus (Fig. 8 B), while the embolus is much less sclerotized and rounded in the later; by the different course of the ducts present in the tegulum in ventral view (Fig. 8 B), and by the presence of nose-like projection in the clypeus (Fig. 3 D), absent in E. camis. Females of E. nasuta n. sp. also resemble those of E. camis (see Levi, 1963, figs 17, 18) but differ from E. camis in having the copulatory opening positioned medially on the epigynal plate (Fig. 9 C), while the latter species is posteriorly positioned.	en	Rodrigues, Everton Nei Lopes, Marta, Kimberly S., Figueiredo, Cassiano P. (2021): Description of four new species of the spider genus Euryopis Menge, 1868, with new records from Brazil (Araneae, Theridiidae, Hadrotarsinae). Zootaxa 4966 (5): 535-549, DOI: https://doi.org/10.11646/zootaxa.4966.5.3
03FE8550C4214B0AFF6AFB043620FA7C.taxon	description	Description. Male holotype (IBSP 269920). Total length 2.43. Carapace length 0.96, width 0.71. Clypeus height 0.31. Sternum length 0.60, width 0.52. Abdomen length 1.17, width 0.81, height 0.79. Leg formula IV / I / II / III; segment length (I / II / III / IV): femora 0.48 / 0.42 / 0.42 / 0.56; patellae + tibiae 0.67 / 0.60 / 0.56 / 0.81; metatarsi + tarsi 0.72 / 0.60 / 0.62 / 0.79; total 1.87 / 1.62 / 1.60 / 2.16. Carapace oval (Fig. 3 A), reddish brown, with blackened stripes. Chelicerae (Fig. 3 B) pale orange. Eye diameter and interdistances: AME 0.04, PME 0.04, ALE 0.06, PLE 0.04, AME – AME 0.04, AME – ALE 0.02, PME – PME 0.06, PME – PLE 0.04. Clypeus present a nose-like projection (Fig. 3 D). Sternum (Fig. 3 B) lighter than the carapace, with blackened edges. Reddish brown elongated pedicel with two longitudinal blackish bands (Fig. 3 A, B, D). Abdomen oval (Fig. 3 A), darker than carapace. Spinnerets pale yellow. Cymbium oval in dorsal view; embolus sclerotized and distally pointed; conductor enlarged, and loop-like duct in the median apophysis (Fig. 8 B). Female paratype (IBSP 269921). Total length 2.49. Carapace length 0.94, width 0.71. Clypeus height 0.21. Sternum length 0.58, width 0.46. Abdomen length 1.42, width 0.98, height 0.79. Leg formula IV / I / II / III; segment length (I / II / III / IV): femora 0.48 / 0.42 / 0.42 / 0.56; patellae + tibiae 0.67 / 0.60 / 0.56 / 0.81; metatarsi + tarsi 0.72 / 0.60 / 0.62 / 0.79; total 1.87 / 1.62 / 1.60 / 2.16. Carapace oval (Fig. 4 A), yellowish brown with darkened edges. Eye diameter and interdistances: AME 0.04, PME 0.04, ALE 0.06, PLE 0.06, AME – AME 0.06, AME – ALE 0.04, PME – PME 0.04 PME – PLE 0.04. Sternum scutiform (Fig. 4 B), lighter than the carapace with blackened edges. Pedicel yellowish, shorter than in males (Fig. 4 A, B). In dorsal view, abdomen oval (Fig. 4 A), dark brown with irregular median yellow pattern. In ventral view, predominantly yellow with two black spots on the posterior laterals (Fig. 4 B). Epigynum with a slightly sclerotized plate, wider than long. Copulatory opening oval, with well-developed ridge (Fig. 9 C), located at the median-anterior area of epigynum (Fig. 9 C) and internally with two pairs of round spermathecae, first pair of spermathecae larger (Figs 9 C, D), short and coiled fertilization ducts (Fig. 9 D) and uncoiled copulatory ducts (Fig. 9 D). Natural history. This species was collected at three different areas in the Atlantic forest (Campos de Palmas Wildlife Refuge), at two locations with riparian forest and at one location without riparian forest. A total of eight specimens were collected in an area with riparian forest and only one in a region without riparian forest.	en	Rodrigues, Everton Nei Lopes, Marta, Kimberly S., Figueiredo, Cassiano P. (2021): Description of four new species of the spider genus Euryopis Menge, 1868, with new records from Brazil (Araneae, Theridiidae, Hadrotarsinae). Zootaxa 4966 (5): 535-549, DOI: https://doi.org/10.11646/zootaxa.4966.5.3
03FE8550C4214B0AFF6AFB043620FA7C.taxon	distribution	Distribution. Brazil (state of Paraná, Fig. 10). Additional material examined. Brazil. Paraná: Palmas (Campos de Palmas Wildlife Refuge 26 ° 32 ’ 37.2 ” S, 51 ° 38 ’ 40.1 ” W), 22 – 29. X. 2018, G. Galeti et al. leg., 1 M (MZUnisinos Ara 00911); 22 – 29. X. 2018, G. Galeti et al. leg., 1 M (MZUnisinos Ara 00912). All collected by pitfall trap.	en	Rodrigues, Everton Nei Lopes, Marta, Kimberly S., Figueiredo, Cassiano P. (2021): Description of four new species of the spider genus Euryopis Menge, 1868, with new records from Brazil (Araneae, Theridiidae, Hadrotarsinae). Zootaxa 4966 (5): 535-549, DOI: https://doi.org/10.11646/zootaxa.4966.5.3
03FE8550C4234B04FF6AFA3D37F0F82E.taxon	description	Figs. 5 A – D, 6 A – D, 8 C, 9 E, F, 10 LSIDurn: lsid: zoobank. org: act: FAD 9973 - 09 A 8 - 4 D 7 F-AAA 6 - A 9 BE 81 FCFB 70	en	Rodrigues, Everton Nei Lopes, Marta, Kimberly S., Figueiredo, Cassiano P. (2021): Description of four new species of the spider genus Euryopis Menge, 1868, with new records from Brazil (Araneae, Theridiidae, Hadrotarsinae). Zootaxa 4966 (5): 535-549, DOI: https://doi.org/10.11646/zootaxa.4966.5.3
03FE8550C4234B04FF6AFA3D37F0F82E.taxon	materials_examined	Type material. Holotype. Bolivia: La Paz Department, Murillo (Mecapaca 16 ° 39 ’ 51.5 ” S, 68 ° 00 ’ 56.6 ” W), III. 1980, 1 M (MCN 25528). Paratype: La Paz Department, Murillo (Mecapaca 16 ° 39 ’ 51.5 ” S, 68 ° 00 ’ 56.6 ” W), III. 1980, 1 F (MCN 25528).	en	Rodrigues, Everton Nei Lopes, Marta, Kimberly S., Figueiredo, Cassiano P. (2021): Description of four new species of the spider genus Euryopis Menge, 1868, with new records from Brazil (Araneae, Theridiidae, Hadrotarsinae). Zootaxa 4966 (5): 535-549, DOI: https://doi.org/10.11646/zootaxa.4966.5.3
03FE8550C4234B04FF6AFA3D37F0F82E.taxon	etymology	Etymology. The specific name is derived from the type locality.	en	Rodrigues, Everton Nei Lopes, Marta, Kimberly S., Figueiredo, Cassiano P. (2021): Description of four new species of the spider genus Euryopis Menge, 1868, with new records from Brazil (Araneae, Theridiidae, Hadrotarsinae). Zootaxa 4966 (5): 535-549, DOI: https://doi.org/10.11646/zootaxa.4966.5.3
03FE8550C4234B04FF6AFA3D37F0F82E.taxon	diagnosis	Diagnosis. Males of Euryopis boliviensis n. sp. resemble those of Euryopis nasuta, but E. boliviensis n. sp. differs from E. nasuta in having the apical portion of the conductor with a thin hook-like projection (Fig. 8 C), absent in E. nasuta, and by the absence of nose-like projection in the clypeus, present in the latter. Females of the E. boliviensis n. sp. resemble those of E. nasuta, but differ by having the second spermathecae further apart (Figs 9 E, F), and closer to each other in the E. nasuta, and by a larger and more sclerotized copulatory opening (Fig. 9 E) than in E. nasuta.	en	Rodrigues, Everton Nei Lopes, Marta, Kimberly S., Figueiredo, Cassiano P. (2021): Description of four new species of the spider genus Euryopis Menge, 1868, with new records from Brazil (Araneae, Theridiidae, Hadrotarsinae). Zootaxa 4966 (5): 535-549, DOI: https://doi.org/10.11646/zootaxa.4966.5.3
03FE8550C4234B04FF6AFA3D37F0F82E.taxon	description	Description. Male holotype (MCN 25528). Total length 2.33. Carapace length 0.94, width 0.77, clypeus height 0.33. Sternum length 0.50, width 0.46. Abdomen length 1.19, width 0.86, height 0.58. Leg formula IV / I / II / III; segment length (I / II / III / IV): femora 0.63 / 0.60 / 0.56 / 0.73; patellae + tibiae 0.79 / 0.73 / 0.69 / 0.88; metatarsi + tarsi 0.84 / 0.79 / 0.79 / 0.94; total 2.26 / 2.12 / 2.04 / 2.55. Carapace oval (Fig. 5 A), orange brown. Ocular region slightly pronounced, eyes with black margins (Fig. 5 C). Eye diameter and interdistances: AME 0.02, PME 0.04, ALE 0.04, PLE 0.06, AME – AME 0.04, AME – ALE 0.02, PME – PME 0.06, PME – PLE 0.04. Fovea marked dark brown (Fig. 5 A). Sternum scutiform (Fig. 5 B), yellow, lighter than the carapace. Legs yellow with blackish pigment on the sides, except on the coxae and trochanter. Pedicel elongated with two longitudinal blackened bands (Fig. 5 A, 5 B, 5 D). Abdomen oval (Fig. 5 A, 5 B), dorsally brown with irregular yellow pattern. In ventral view, predominantly yellow with a blackened spot in median-anterior area and two blackened spots on the posterior laterals. Spinnerets yellow. Palp with cymbium oval in dorsal view; the conductor distally hook-shaped and embolus apically sclerotized (Fig. 8 C); conductor enlarged, and median apophysis with loop-like duct (Fig. 8 C). Female paratype (MCN 25528). Total length 2.54. Carapace length 0.96, width 0.79. Clypeus height 0.14. Sternum length 0.56, width 0.44. Abdomen length 1.65, width 1.05, height 0.94. Leg formula IV / I / III / II; segment length (I / II / III / IV): femora 0.73 / 0.65 / 0.60 / 0.71; patellae + tibiae 0.88 / 0.79 / 0.81 / 1.05; metatarsi + tarsi 0.92 / 0.73 / 0.77 / 1.00; total 2.53 / 2.17 / 2.18 / 2.76. Carapace (Fig. 6 A) yellow, oval, with blackish edges. Ocular region slightly bulged, eyes with black margins (Fig. 6 C). Eye diameter and interdistances: AME 0.06, PME 0.06, ALE 0.06, PLE 0.06, AME – AME 0.04, AME – ALE 0.02, PME – PME 0.06, PME – PLE 0.04. Sternum scutiform (Fig. 6 B), yellow with blackened edges. Pedicel yellow with two longitudinal blackened bands (Fig. 6 A). In dorsal view, abdomen longer than wide (Fig. 6 A, 6 B), sparse setae, predominantly yellow with irregular pattern of dark brown spots. In ventral view abdomen yellow with two black spots on the posterior laterals (Fig. 6 B). Spinnerets yellow. Epigynum with a slightly sclerotized plate, longer than wide (Fig. 9 E). Internal genitalia with two pairs of round spermathecae with the similar size, first pair of spermathecae slightly larger (Fig. 9 F), short and coiled fertilization ducts (Fig. 9 F) and uncoiled copulatory ducts (Fig. 9 F). Natural history. Unknown.	en	Rodrigues, Everton Nei Lopes, Marta, Kimberly S., Figueiredo, Cassiano P. (2021): Description of four new species of the spider genus Euryopis Menge, 1868, with new records from Brazil (Araneae, Theridiidae, Hadrotarsinae). Zootaxa 4966 (5): 535-549, DOI: https://doi.org/10.11646/zootaxa.4966.5.3
03FE8550C4234B04FF6AFA3D37F0F82E.taxon	distribution	Distribution. Bolivia (La Paz, Fig. 10).	en	Rodrigues, Everton Nei Lopes, Marta, Kimberly S., Figueiredo, Cassiano P. (2021): Description of four new species of the spider genus Euryopis Menge, 1868, with new records from Brazil (Araneae, Theridiidae, Hadrotarsinae). Zootaxa 4966 (5): 535-549, DOI: https://doi.org/10.11646/zootaxa.4966.5.3
03FE8550C42C4B07FF6AFF5C36EEF829.taxon	description	Figs. 7 A – D, 8 D, 10 LSIDurn: lsid: zoobank. org: act: EA 3 AF 009 - 3083 - 41 A 7 - A 1 B 0 - 0 FE 4 D 86 FE 034	en	Rodrigues, Everton Nei Lopes, Marta, Kimberly S., Figueiredo, Cassiano P. (2021): Description of four new species of the spider genus Euryopis Menge, 1868, with new records from Brazil (Araneae, Theridiidae, Hadrotarsinae). Zootaxa 4966 (5): 535-549, DOI: https://doi.org/10.11646/zootaxa.4966.5.3
03FE8550C42C4B07FF6AFF5C36EEF829.taxon	materials_examined	Type material. Holotype. Brazil: Santa Catarina (Garopada 28 ° 01 ’ 36.2 ” S, 48 ° 36 ’ 48.0 ” W), 12 – 15. III. 2001, M. P. Barros leg., 1 M (MCN 33781). Collected by pitfall-trap.	en	Rodrigues, Everton Nei Lopes, Marta, Kimberly S., Figueiredo, Cassiano P. (2021): Description of four new species of the spider genus Euryopis Menge, 1868, with new records from Brazil (Araneae, Theridiidae, Hadrotarsinae). Zootaxa 4966 (5): 535-549, DOI: https://doi.org/10.11646/zootaxa.4966.5.3
03FE8550C42C4B07FF6AFF5C36EEF829.taxon	etymology	Etymology. The specific name is derived from the type locality.	en	Rodrigues, Everton Nei Lopes, Marta, Kimberly S., Figueiredo, Cassiano P. (2021): Description of four new species of the spider genus Euryopis Menge, 1868, with new records from Brazil (Araneae, Theridiidae, Hadrotarsinae). Zootaxa 4966 (5): 535-549, DOI: https://doi.org/10.11646/zootaxa.4966.5.3
03FE8550C42C4B07FF6AFF5C36EEF829.taxon	diagnosis	Diagnosis. The male of Euryopis catarinensis n. sp. is similar to that of Euryopis camis (see Levi, 1963, fig. 19) but differs from this latter species by having the median apophysis more rounded with the posterior margin sclerotized, while the median apophysis is elongated in the later; by the different course of the ducts in the tegulum, medially positioned (Fig. 8 D), while the spermatic duct is positioned basally in the E. camis (see Levi, 1963, fig. 19).	en	Rodrigues, Everton Nei Lopes, Marta, Kimberly S., Figueiredo, Cassiano P. (2021): Description of four new species of the spider genus Euryopis Menge, 1868, with new records from Brazil (Araneae, Theridiidae, Hadrotarsinae). Zootaxa 4966 (5): 535-549, DOI: https://doi.org/10.11646/zootaxa.4966.5.3
03FE8550C42C4B07FF6AFF5C36EEF829.taxon	description	Description. Male holotype (MCN 33781). Total length 1.99. Carapace length 0.92, width 0.73, clypeus height 0.31. Sternum length 0.42, width 0.42. Abdomen length 1.07, width 0.86, height 0.58. Leg formula IV / I / II / III; segment length (I / II / III / IV): femora 0.56 / 0.52 / 0.48 / 0.63; patellae + tibiae 0.63 / 0.56 / 0.58 / 0.79; metatarsi + tarsi 0.63 / 0.56 / 0.54 / 0.73; total 1.82 / 1.64 / 1.60 / 2.15. Carapace oval (Fig. 7 A), dark reddish brown. Chelicerae (Fig. 7 C) reddish brown. Ocular region (Fig. 7 C) slightly pronounced, eyes with black margins. Eye diameter and interdistances: AME 0.04, PME 0.04, ALE 0.06, PLE 0.06, AME – AME 0.06, AME – ALE 0.02, PME – PME 0.06, PME – PLE 0.04. Sternum (Fig. 7 B) lighter than the carapace, with darkened edges. Abdomen oval (Figs 7 A, B), dorsal view orange brown, slightly lighter than the carapace, and darkened edges (Figs 7 A, D). In ventral view, the abdomen has a uniform orange-brown color, with dark pigmentation on the laterals. Spinnerets yellow. Palp with embolus (Fig. 8 D) not sclerotized; median apophysis rounded with spermatic duct 8 - shaped, cymbium oval in dorsal view without apical projection; ventrally, duct in the tegulum as an inverted “ S ” (Fig. 8 D). Female. Unknown. Natural history. Unknown.	en	Rodrigues, Everton Nei Lopes, Marta, Kimberly S., Figueiredo, Cassiano P. (2021): Description of four new species of the spider genus Euryopis Menge, 1868, with new records from Brazil (Araneae, Theridiidae, Hadrotarsinae). Zootaxa 4966 (5): 535-549, DOI: https://doi.org/10.11646/zootaxa.4966.5.3
03FE8550C42C4B07FF6AFF5C36EEF829.taxon	distribution	Distribution. Brazil (state of Santa Catarina, Garopaba, Fig. 10).	en	Rodrigues, Everton Nei Lopes, Marta, Kimberly S., Figueiredo, Cassiano P. (2021): Description of four new species of the spider genus Euryopis Menge, 1868, with new records from Brazil (Araneae, Theridiidae, Hadrotarsinae). Zootaxa 4966 (5): 535-549, DOI: https://doi.org/10.11646/zootaxa.4966.5.3
03FE8550C4294B01FF6AFA5635B1FDC0.taxon	description	Fig. 10	en	Rodrigues, Everton Nei Lopes, Marta, Kimberly S., Figueiredo, Cassiano P. (2021): Description of four new species of the spider genus Euryopis Menge, 1868, with new records from Brazil (Araneae, Theridiidae, Hadrotarsinae). Zootaxa 4966 (5): 535-549, DOI: https://doi.org/10.11646/zootaxa.4966.5.3
03FE8550C4294B01FF6AFA5635B1FDC0.taxon	materials_examined	New records. Brazil: Paraná, Palmas, Refúgio de Vida Silvestre dos Campos de Palmas [Campos de Palmas Wildlife Refuge] (26 ° 32 ’ 37.77 ” S, 51 ° 38 ’ 39.71 ” W), 22 – 29. X. 2008, G. Galeti et al. leg., 1 F (MZUnisinos Ara 0915); 1 M (MZUnisinos Ara 0923). Santa Catarina, Chapecó, Floresta Nacional de Chapecó (27 ° 06 ’ 11.3 ” S, 52 ° 46 ’ 46.8 ”), 26. X. 2018, H. M. Gasparetto leg., 1 M (MZUnisinos Ara 00914); Floresta Nacional de Chapecó (27 ° 03 ’ 24.3 ” S, 52 ° 41 ’ 40.3 ” W), 25. I. 2019, H. M. Gasparetto leg., 1 F (MZUnisinos Ara 00916); 1 M (MZUnisinos Ara 00917); 1 F (MZUnisinos Ara 00918); 1 M (MZUnisinos Ara 00919); 1 M (MZUnisinos Ara 00920); 26. X. 2018, H. M. Gasparetto leg., 1 M (MZUnisinos Ara 00914); 1 M (MZUnisinos Ara 00926); 1 M, 2 F (MZUnisinos Ara 00927); 1 M (MZUnisinos Ara 00928); 2 M, 1 F (MZUnisinos Ara 00929); 1 M (MZUnisinos Ara 00930); 2 M (MZUnisinos Ara 00931); 1 M, 1 F (MZUnisinos Ara 00932); 1 M, 1 F (MZUnisinos Ara 00933); Floresta Nacional de Chapecó (27 ° 03 ’ 37.5 ” S, 52 ° 42 ’ 35.6 ” W), 26. X. 2018, H. M. Gasparetto leg., 2 M (MZUnisinos Ara 00934); 1 M (MZUnisinos Ara 00935); Floresta Nacional de Chapecó (27 ° 04 ’ 48.9 ” S, 52 ° 43 ’ 18.3 ” W), 26. X. 2018, H. M. Gasparetto leg., 2 M (MZUnisinos Ara 00936); 1 M (MZUnisinos Ara 00938); 1 M (MZUnisinos Ara 00939); 1 M (MZUnisinos Ara 00940); 1 F (MZUnisinos Ara 00941); 1 M (MZUnisinos Ara 00942); 1 M (MZUnisinos Ara 00943); 1 F (MZUnisinos Ara 00944); 1 M (MZUnisinos Ara 00945); 1 M (MZUnisinos Ara 00946); 1 M (MZUnisinos Ara 00947); 1 M (MZUnisinos Ara 00948); 1 M (MZUnisinos Ara 00949); Floresta Nacional de Chapecó (27 ° 04 ’ 13.1 ” S, 52 ° 42 ’ 36.8 ” W), 26. X. 2018, H. M. Gasparetto leg., 1 M (MZUnisinos Ara 00937); Floresta Nacional de Chapecó (27 ° 6 ’ 6.44 ” S, 52 ° 47 ’ 24.58 ” W), 07 – 14. I. 2019, G. Galeti et al. leg., 1 F (MZUnisinos Ara 00921); Floresta Nacional de Chapecó (27 ° 6 ’ 21.63 ” S, 52 ° 47 ’ 9.37 ” W), 07 – 14. I. 2019, G. Galeti et al. leg., 1 M (MZUnisinos Ara 00922); Floresta Nacional de Chapecó (27 ° 5 ’ 16.07 ” S, 52 ° 47 ’ 3.28 ” W); Floresta Nacional de Chapecó (27 ° 6 ’ 6.42 ” S, 52 ° 47 ’ 22.91 ” W), 07 – 14. I. 2019, G. Galeti et al. leg., 1 M (MZUnisinos Ara 00925); all collected by pitfall-trap. Rio Grande do Sul, Triunfo, Parque Copesul (29 ° 51 ’ 57.9 ” S, 51 ° 21 ’ 55.4 ” W), 22. X. 2002, R. Ott leg., 3 M, 3 F (MCN 34868); 12. VI. 1991, A. B. Bonaldo leg., 25 M, 21 F (MCN 21189).	en	Rodrigues, Everton Nei Lopes, Marta, Kimberly S., Figueiredo, Cassiano P. (2021): Description of four new species of the spider genus Euryopis Menge, 1868, with new records from Brazil (Araneae, Theridiidae, Hadrotarsinae). Zootaxa 4966 (5): 535-549, DOI: https://doi.org/10.11646/zootaxa.4966.5.3
03FE8550C4294B01FF6AFA5635B1FDC0.taxon	distribution	Distribution. Southern Brazil (Paraná, Santa Catarina and Rio Grande do Sul see Buckup et al. 2010, Fig. 10).	en	Rodrigues, Everton Nei Lopes, Marta, Kimberly S., Figueiredo, Cassiano P. (2021): Description of four new species of the spider genus Euryopis Menge, 1868, with new records from Brazil (Araneae, Theridiidae, Hadrotarsinae). Zootaxa 4966 (5): 535-549, DOI: https://doi.org/10.11646/zootaxa.4966.5.3
03FE8550C4284B01FF6AFDA03190FC58.taxon	description	Fig. 10	en	Rodrigues, Everton Nei Lopes, Marta, Kimberly S., Figueiredo, Cassiano P. (2021): Description of four new species of the spider genus Euryopis Menge, 1868, with new records from Brazil (Araneae, Theridiidae, Hadrotarsinae). Zootaxa 4966 (5): 535-549, DOI: https://doi.org/10.11646/zootaxa.4966.5.3
03FE8550C4284B01FF6AFDA03190FC58.taxon	materials_examined	New record. Brazil: Santa Catarina, Chapecó, Floresta Nacional de Chapecó (27 ° 03 ’ 46.0 ” S 52 ° 41 ’ 05.4 ” W), 25. I. 2019, H. M. Gasparetto leg., 1 F (MZUnisinos Ara 0950).	en	Rodrigues, Everton Nei Lopes, Marta, Kimberly S., Figueiredo, Cassiano P. (2021): Description of four new species of the spider genus Euryopis Menge, 1868, with new records from Brazil (Araneae, Theridiidae, Hadrotarsinae). Zootaxa 4966 (5): 535-549, DOI: https://doi.org/10.11646/zootaxa.4966.5.3
03FE8550C4284B01FF6AFDA03190FC58.taxon	distribution	Distribution. Argentina (Provinces of Buenos Aires, Missiones and San Luis, see González, A. (1990 )). Brazil (Santa Catarina, Chapecó, Floresta Nacional, Fig. 10; Rio Grande do Sul, see Buckup et al. 2010).	en	Rodrigues, Everton Nei Lopes, Marta, Kimberly S., Figueiredo, Cassiano P. (2021): Description of four new species of the spider genus Euryopis Menge, 1868, with new records from Brazil (Araneae, Theridiidae, Hadrotarsinae). Zootaxa 4966 (5): 535-549, DOI: https://doi.org/10.11646/zootaxa.4966.5.3
