taxonID	type	description	language	source
76524E00FFDE357FFF1CFDA38FD31D20.taxon	description	(Figs. 1 A – B, 3 A)	en	Reis, Patricia Do Rosario, Gil-Azevedo, Leonardo H., Ferreira-Keppler, Ruth L. (2021): Two new species of Tanytarsus van der Wulp (Diptera: Chironomidae) from Costa Rica. Zootaxa 4966 (1): 84-90, DOI: https://doi.org/10.11646/zootaxa.4966.1.9
76524E00FFDE357FFF1CFDA38FD31D20.taxon	materials_examined	Type material. Holotype: adult male, Costa Rica, Prov. San José, Moravia, Zurquí de Moravia (10 ° 02 ’ 58 ” N – 84 ° 00 ’ 57 ” W), Creek 1 north. 1600 m Canopy Malaise trap. 30. iii to 6. iv. 2013, (IN- BIOCRI-COSTA RICA).	en	Reis, Patricia Do Rosario, Gil-Azevedo, Leonardo H., Ferreira-Keppler, Ruth L. (2021): Two new species of Tanytarsus van der Wulp (Diptera: Chironomidae) from Costa Rica. Zootaxa 4966 (1): 84-90, DOI: https://doi.org/10.11646/zootaxa.4966.1.9
76524E00FFDE357FFF1CFDA38FD31D20.taxon	etymology	Etymology. Named in honor of Dr. Brian V. Brown, the Principal Investigator of ZADBI project.	en	Reis, Patricia Do Rosario, Gil-Azevedo, Leonardo H., Ferreira-Keppler, Ruth L. (2021): Two new species of Tanytarsus van der Wulp (Diptera: Chironomidae) from Costa Rica. Zootaxa 4966 (1): 84-90, DOI: https://doi.org/10.11646/zootaxa.4966.1.9
76524E00FFDE357FFF1CFDA38FD31D20.taxon	diagnosis	Diagnostic characters: The new species can be separated from others by the following combination of characters: Y-shaped anal tergite bands; anal point elongated, with a trifid apex; two pairs of long bars; superior volsella almost sub-rectangular; digitus finger-like, short, pointed, not reaching the margin of the superior volsella; median volsella with simple lamellar setae mounted apically and 4 regular setae; gonostylus short, straight, of almost uniform width from base to apex.	en	Reis, Patricia Do Rosario, Gil-Azevedo, Leonardo H., Ferreira-Keppler, Ruth L. (2021): Two new species of Tanytarsus van der Wulp (Diptera: Chironomidae) from Costa Rica. Zootaxa 4966 (1): 84-90, DOI: https://doi.org/10.11646/zootaxa.4966.1.9
76524E00FFDE357FFF1CFDA38FD31D20.taxon	description	Description. Adult male (n = 1). Small, total length 1.80 mm. Wing length 1.24 mm. Colouration. Head yellowish-brown, flagellum and maxillary palp pale brown. Thorax and legs yellowish. Abdomen yellowish. Head. Eyes bare, with little dorsomedian extension. Frontal tubercles short, 7.5 µm long. Temporal setae 6, uniserial. Antennae missing. Clypeus with 12 setae. Palpomeres 2 – 5, lengths (in µm): 30, 100, 65,160. Thorax. Ac 5, biserial; Pa 1; Scts 6. Halteres with 5 setae on both. Dorsocentral setae (Dc) not observed. Wing (Fig. 3 A). Width 0.39 mm. VR 1.24. Sc and M bare, the remaining wing veins and cells setose. Brachiolum without seta. Legs. Foreleg bearing single tibial spur, 25 μm long. Mid leg tibia bearing two spurs, 30 μm and 40 μm long, and two separated combs. Hind leg tibia bearing two spurs, 27.5 μm and 40 μm long, and two separated combs. Lengths of leg segments and leg ratios as in Table 1. Hypopygium (Figs. 1 A – B). Tergite IX 142 μm long, with 2 shorts distal setae. Y-shaped anal tergite bands “ fused ” on the median part of tergite and reaching anal point, about 34 μm long. Anal point elongated (36 µm long) and slender, lateral margins concave, in the dorsal aspect apex with three rounded portions, ventromedian projection longer than the length of the anal point, lateral margin bearing 3 distal setae; AnPR 3.60. Two pairs of anal point bars, not surpassing the apex of anal point; AnPBR 2.00. Superior volsella sub rectangular, elongated (32 µm long), with 4 setae on dorsal surface and 2 setae on anteromedian margin (one dorsal, one ventral), posteromedian corner elongated and rounded; digitus 11 µm long, pointed, finger-like, not extending beyond margin of superior volsella. Stem of median volsella about 12 μm long, with simple lamellar setae mounted apically and 4 regular setae. Inferior volsella ca. 71 μm long, S-shaped, apex with a prominent triangular swelling. Gonocoxite 67 μm long. Gonostylus short (51 μm long), straight, of almost uniform width from base to apex, rounded tip. HR 1.41.	en	Reis, Patricia Do Rosario, Gil-Azevedo, Leonardo H., Ferreira-Keppler, Ruth L. (2021): Two new species of Tanytarsus van der Wulp (Diptera: Chironomidae) from Costa Rica. Zootaxa 4966 (1): 84-90, DOI: https://doi.org/10.11646/zootaxa.4966.1.9
76524E00FFDE357FFF1CFDA38FD31D20.taxon	discussion	Remarks. The anal point and the gonostylus are the main characters that differentiate the male T. briani sp. nov. from other species. In the identification key for the males of Tanytarus ortoni species group, the shape of the anal point of T. briani is similar to the species Tanytarsus yara and Tanytarsus castelnaui. The trifid apex, in dorsal aspect, of the anal point is similar to what is observed in T. yara; the ventromedian projection of the anal point is similar to that of T. castelnaui. However, T. briani can be easily distinguished from the two above mentioned species by the presence of two pairs of long anal bars in contrast to a pair of short anal bars found in T. yara and T. castelnaui. The small gonostylus, of almost uniform width, does not correspond to any species of Tanytarus ortoni species group, this format apparently being unique among the Neotropical species of Tanytarsus.	en	Reis, Patricia Do Rosario, Gil-Azevedo, Leonardo H., Ferreira-Keppler, Ruth L. (2021): Two new species of Tanytarsus van der Wulp (Diptera: Chironomidae) from Costa Rica. Zootaxa 4966 (1): 84-90, DOI: https://doi.org/10.11646/zootaxa.4966.1.9
76524E00FFDE357FFF1CFDA38FD31D20.taxon	distribution	Distribution and ecological notes. Tanytarsus briani was collected at an elevation of about 1,600 meters, demonstrating the wide adaptive range of chironomids of the Tanytarsini tribe, as almost all species of the Tanytarsus ortoni species group have been recorded from lowland areas of tropical climate.	en	Reis, Patricia Do Rosario, Gil-Azevedo, Leonardo H., Ferreira-Keppler, Ruth L. (2021): Two new species of Tanytarsus van der Wulp (Diptera: Chironomidae) from Costa Rica. Zootaxa 4966 (1): 84-90, DOI: https://doi.org/10.11646/zootaxa.4966.1.9
76524E00FFDC3579FF1CFDA98E431A90.taxon	description	(Figs. 2 A – C, 3 B)	en	Reis, Patricia Do Rosario, Gil-Azevedo, Leonardo H., Ferreira-Keppler, Ruth L. (2021): Two new species of Tanytarsus van der Wulp (Diptera: Chironomidae) from Costa Rica. Zootaxa 4966 (1): 84-90, DOI: https://doi.org/10.11646/zootaxa.4966.1.9
76524E00FFDC3579FF1CFDA98E431A90.taxon	materials_examined	Type material. Holotype: adult male, Costa Rica, Prov. San José, Moravia, Zurquí de Moravia (10 ° 02 ’ 58 ” N – 84 ° 00 ’ 57 ” W), Creek 1 north. 1600 m. Canopy Malaise trap. 12 – 19. iv. 2013 (INBIOCRI-COSTA RICA).	en	Reis, Patricia Do Rosario, Gil-Azevedo, Leonardo H., Ferreira-Keppler, Ruth L. (2021): Two new species of Tanytarsus van der Wulp (Diptera: Chironomidae) from Costa Rica. Zootaxa 4966 (1): 84-90, DOI: https://doi.org/10.11646/zootaxa.4966.1.9
76524E00FFDC3579FF1CFDA98E431A90.taxon	etymology	Etymology. Named after the country where the species was found. Noun in apposition.	en	Reis, Patricia Do Rosario, Gil-Azevedo, Leonardo H., Ferreira-Keppler, Ruth L. (2021): Two new species of Tanytarsus van der Wulp (Diptera: Chironomidae) from Costa Rica. Zootaxa 4966 (1): 84-90, DOI: https://doi.org/10.11646/zootaxa.4966.1.9
76524E00FFDC3579FF1CFDA98E431A90.taxon	diagnosis	Diagnosis. Tanytarsus costarica can be separated from other species of the genus by the following combination of characters: head without frontal tubercles; Y-shaped anal tergite bands; anal point elongated, wide, divided into three portions, with two pairs of long bars; superior volsella with posteromedian lobe strongly produced; digitus finger-like; median volsella with simple lamellar setae mounted apically and 4 regular setae.	en	Reis, Patricia Do Rosario, Gil-Azevedo, Leonardo H., Ferreira-Keppler, Ruth L. (2021): Two new species of Tanytarsus van der Wulp (Diptera: Chironomidae) from Costa Rica. Zootaxa 4966 (1): 84-90, DOI: https://doi.org/10.11646/zootaxa.4966.1.9
76524E00FFDC3579FF1CFDA98E431A90.taxon	description	Description. Adult male (n = 1). Small, total length 2.00 mm. Wing length 1.26 mm. Colouration. Head yellowish-brown; flagellum and maxillary palp pale brown. Thorax and legs yellowish. Abdomen yellowish. Head. Eyes bare, without dorsomedial extension. Frontal tubercles absent. Temporal setae 5 + 5. Antenna with 13 flagellomeres, 680 µm long. AR. 0.38. Length of palpomeres 2 – 5 (in µm): 25, 50, 90, 100. Thorax. Ac 5; Pa 1; Scts 2. Halteres with 6 setae on both. Dorsocentral setae (Dc) not observed. Wing (Fig. 3 B). Width 0.39 mm. VR 1.26. Sc, M, and RM bare, remaining wing veins and cells with setae. Brachiolum with one seta. Legs. Foreleg bearing single tibial spur, 27.5 μm long. Mid leg tibia bearing two spurs, 17.5 μm and 27.5 μm long, and two separated combs. Hind leg tibia bearing two spurs, 17.5 μm and 30 μm long, and two separated combs. Lengths of leg segments and leg ratios as in Table 2. Hypopygium (Figs. 2 A – C). Tergite IX 169 μm long, with 2 distal setae. Y-shaped anal tergite bands “ fused ” on median part of tergite and reaching anal point, about 24 μm long. Anal point elongated (40 µm long), wide (17 µm width), divided into three very distinct portions (Fig. A, C), lateral margin with 2 pairs of setae in distal part; AnPR 2.35. Two pairs of anal point bars elongated (28 μm long), each bar tip split into two apices and reaching beyond apex of anal point; AnPBR 0.86. Superior volsella subrectangular, with 5 setae on dorsal surface, 3 setae on anteromedian margin (2 dorsal, 1 ventral), posteromedian corner and strongly projecting; digitus 13 µm long, pointed, finger-like, extending beyond margin of superior volsella. Stem of median volsella short, about 7 μm long, with simple lamellar setae and 4 regular setae. Inferior volsella ca. 61 μm long. Gonocoxite 67 μm long. Gonostylus 68 μm long, broader at mid-length, rounded tip. HR 0.99.	en	Reis, Patricia Do Rosario, Gil-Azevedo, Leonardo H., Ferreira-Keppler, Ruth L. (2021): Two new species of Tanytarsus van der Wulp (Diptera: Chironomidae) from Costa Rica. Zootaxa 4966 (1): 84-90, DOI: https://doi.org/10.11646/zootaxa.4966.1.9
76524E00FFDC3579FF1CFDA98E431A90.taxon	discussion	Remarks: The superior volsella of T. costarica sp. nov. is superficially similar to those of T. canine, T. comunis and T. mulleri but has a slightly more elongated posteromedian corner. The digitus, on the other hand, is distinct from these species, since it exceeds the lateral margin of the superior volsella, while in the above mentioned three species, the digit does not exceed the superior volsella. However, the best diagnostic character for the male of T. costarica is the anal point, which is distinctly trifid, with the lateral portions showing a pointed tip and the thumblike middle portion, a design unique among the species of the Tanytarsus ortoni group.	en	Reis, Patricia Do Rosario, Gil-Azevedo, Leonardo H., Ferreira-Keppler, Ruth L. (2021): Two new species of Tanytarsus van der Wulp (Diptera: Chironomidae) from Costa Rica. Zootaxa 4966 (1): 84-90, DOI: https://doi.org/10.11646/zootaxa.4966.1.9
76524E00FFDC3579FF1CFDA98E431A90.taxon	distribution	Distribution and ecological notes. T. costarica was also collected from the same study site and with the same type of trap as T. briani (Zurquí de Moravia, Costa Rica), about 1,600 meters above sea level.	en	Reis, Patricia Do Rosario, Gil-Azevedo, Leonardo H., Ferreira-Keppler, Ruth L. (2021): Two new species of Tanytarsus van der Wulp (Diptera: Chironomidae) from Costa Rica. Zootaxa 4966 (1): 84-90, DOI: https://doi.org/10.11646/zootaxa.4966.1.9
