taxonID	type	description	language	source
C130115AFFC0FFCBFF65FDA8FB8F34D2.taxon	description	The genus Agnoea can be divided into two subgenera Agnoea Walsingham, 1907 and Tubuliferodes Toll, 1956, based on the differences in the gnathos of the male genitalia (Jaeckh, 1972). It is the most speciose group in Lypusinae, currently including 25 species, three of which are known from Korea. Its distribution spans across the Palearctic region but the majority of the congeners occur in Europe. The larvae are associated with the angiosperm genera including Fagus, Carpinus, Populus, Quercus, and Vaccinium (Tokár et al. 2005).	en	Sohn, Jae-Cheon, Lvovsky, Alexander L. (2021): Review of Lypusinae (Lepidoptera: Gelechioidea: Lypusidae) from Korea with a description of a new species of Agnoea Walsingham, 1907. Zootaxa 4966 (3): 385-391, DOI: https://doi.org/10.11646/zootaxa.4966.3.10
C130115AFFC0FFC8FF65FB7EFC383544.taxon	description	(Figs 1, 5)	en	Sohn, Jae-Cheon, Lvovsky, Alexander L. (2021): Review of Lypusinae (Lepidoptera: Gelechioidea: Lypusidae) from Korea with a description of a new species of Agnoea Walsingham, 1907. Zootaxa 4966 (3): 385-391, DOI: https://doi.org/10.11646/zootaxa.4966.3.10
C130115AFFC0FFC8FF65FB7EFC383544.taxon	diagnosis	Diagnosis. This species is closely related to Agnoea (Agnoea) kurentzovi (Lvovsky, 2001) (Fig. 2) in having the asymmetric valvae in the male genitalia (Fig. 6), but differs from the latter in having the disconnected medial bulbs of gnathos, the larger left clasper, and a digitiform process above right sacculus.	en	Sohn, Jae-Cheon, Lvovsky, Alexander L. (2021): Review of Lypusinae (Lepidoptera: Gelechioidea: Lypusidae) from Korea with a description of a new species of Agnoea Walsingham, 1907. Zootaxa 4966 (3): 385-391, DOI: https://doi.org/10.11646/zootaxa.4966.3.10
C130115AFFC0FFC8FF65FB7EFC383544.taxon	description	Description. Head. Vertex and frons dark grayish brown with purplish luster. Antenna 2 / 3 as long as forewing; scape dark grayish brown, intermixed with pale grayish brown scales dorsally; flagellomere dark grayish brown. Labial palpus dark grayish brown. Thorax. Patagium and tegula dark grayish brown; mesonotum dark grayish brown, paler posteriorly, with purplish luster on anterior 1 / 3. Forewing length 6.5 ~ 7.3 mm, narrow, lancelolate apically, dark grayish brown, with purplish luster along costal area; cilia purplish brown. Hindwing grayish brown; cilia purplish brown. Foreleg with coxa and femur grayish brown on outer surface, pale grayish brown on inner surface; tibia dark grayish brown with purplish luster; tarsomeres dark grayish brown, intermixed with pale brown scales ventrally. Midleg with coxa and femur grayish brown; tibia dark grayish brown, with dense purplish-brown, piliform scales; tarsomeres grayish brown dorsally, pale brown ventrally. Hindleg with coxa grayish brown; femur gray, intermixed with purplish gray scales dorsally; tibia with dense, lustrous, yellowish-gray piliform scales; tarsomeres dark yellowish gray dorsally, pale gray ventrally. Abdomen. Terga pale grayish brown, intermixed with gray and purplish gray scales. Sterna dark grayish brown, with purplish luster. Male genitalia (Fig. 5). Uncus deltoidal, dentiform apically. Tegumen subquadrate, as long as uncus. Gnathos as a pair of articulated rami; each ramus stout, bent medially, with large, spinulate bulb at end. Transtilla with digitiform, sparsely-setose lobe laterally. Valva broadened in basal 1 / 3, with linguiform lobe apically, long-setose in distal half; costa very narrow, slightly curved at basal 2 / 5; sacculus broad, narrowed distally, long-setose along upper margin and distal half; clavus in left valva large, drop-shaped, broadened distally, with lying-down spinules along distal margin; clavus in right valva digitiform with subtriangular sclerotized base at distal end of sacculus. Juxta semicircular. Vinculum narrow, converging apically; saccus absent. Sclerotized part of phallus (aedeagus) short, curved in distal half, globular in basal half, obliquely-cut distally, as long as left clavus. Female unknown. Types. Holotype: male, “ HOLOTYPE | Agnoea | danguni | Sohn, 2020 ” [red label], “ KOREA, Gangwon Prov. | Taebaeg, Mt. Taebaegsan | 20 VI 1998 | coll. J. C. Sohn ”. Paratypes: 2 ♂, same data as holotype, [GSN] SJC- 825, 1209. All types are deposited in GJUE.	en	Sohn, Jae-Cheon, Lvovsky, Alexander L. (2021): Review of Lypusinae (Lepidoptera: Gelechioidea: Lypusidae) from Korea with a description of a new species of Agnoea Walsingham, 1907. Zootaxa 4966 (3): 385-391, DOI: https://doi.org/10.11646/zootaxa.4966.3.10
C130115AFFC0FFC8FF65FB7EFC383544.taxon	distribution	Distribution. Korea (endemic).	en	Sohn, Jae-Cheon, Lvovsky, Alexander L. (2021): Review of Lypusinae (Lepidoptera: Gelechioidea: Lypusidae) from Korea with a description of a new species of Agnoea Walsingham, 1907. Zootaxa 4966 (3): 385-391, DOI: https://doi.org/10.11646/zootaxa.4966.3.10
C130115AFFC0FFC8FF65FB7EFC383544.taxon	discussion	Remarks. This is the first record of the subgenus Agnoea from Korea. Agnoea danguni represents the easternmost species of the subgenus in Eurasia.	en	Sohn, Jae-Cheon, Lvovsky, Alexander L. (2021): Review of Lypusinae (Lepidoptera: Gelechioidea: Lypusidae) from Korea with a description of a new species of Agnoea Walsingham, 1907. Zootaxa 4966 (3): 385-391, DOI: https://doi.org/10.11646/zootaxa.4966.3.10
C130115AFFC0FFC8FF65FB7EFC383544.taxon	etymology	Etymology. This species is named after “ Dangun ”, the legendary founder of Korea. There is an altar memorizing the ancestor in the Mountain Taebaegsan, the type locality of the species.	en	Sohn, Jae-Cheon, Lvovsky, Alexander L. (2021): Review of Lypusinae (Lepidoptera: Gelechioidea: Lypusidae) from Korea with a description of a new species of Agnoea Walsingham, 1907. Zootaxa 4966 (3): 385-391, DOI: https://doi.org/10.11646/zootaxa.4966.3.10
C130115AFFC3FFC9FF65F980FE693544.taxon	description	(Figs 3, 7, 9)	en	Sohn, Jae-Cheon, Lvovsky, Alexander L. (2021): Review of Lypusinae (Lepidoptera: Gelechioidea: Lypusidae) from Korea with a description of a new species of Agnoea Walsingham, 1907. Zootaxa 4966 (3): 385-391, DOI: https://doi.org/10.11646/zootaxa.4966.3.10
C130115AFFC3FFC9FF65F980FE693544.taxon	diagnosis	Diagnosis. Kim & Lee (2020) distinguished this species from Agnoea josephinae, based on differences in the external appearance and the male genitalia, and the divergence in the COI sequences. The female genitalia of Agnoea digitiella differ from ones of A. josephinae in having the longer ductus bursae: ca. 5 × longer than corpus bursae in A. digitiella but 4 × in A. josephinae (Fig. 9 vs. 10).	en	Sohn, Jae-Cheon, Lvovsky, Alexander L. (2021): Review of Lypusinae (Lepidoptera: Gelechioidea: Lypusidae) from Korea with a description of a new species of Agnoea Walsingham, 1907. Zootaxa 4966 (3): 385-391, DOI: https://doi.org/10.11646/zootaxa.4966.3.10
C130115AFFC3FFC9FF65F980FE693544.taxon	description	Description. Forewing length 6.3 – 7.9 mm (average 7.08 mm, n = 10). External appearance of females same as that of males. Female genitalia (Fig. 9). Papillae anales small, oblique distally, narrowed apically, setose. Apophysis posterioris 2 × longer than apophysis anterioris; apophysis anterioris curved at basal 1 / 3. Ostium bursae small. Ductus bursae ca. 5 × longer than corpus bursae, narrow; antrum columnar, strongly sclerotized laterally, 1 / 4 as long as apophysis anterioris. Corpus bursae obovate, with signum at caudal 2 / 5; signum subrhomboidal, scobinate, less sclerotized longitudinally at middle.	en	Sohn, Jae-Cheon, Lvovsky, Alexander L. (2021): Review of Lypusinae (Lepidoptera: Gelechioidea: Lypusidae) from Korea with a description of a new species of Agnoea Walsingham, 1907. Zootaxa 4966 (3): 385-391, DOI: https://doi.org/10.11646/zootaxa.4966.3.10
C130115AFFC3FFC9FF65F980FE693544.taxon	materials_examined	Material examined. 4 ♂, Gyonggi Prov., Dongducheon, Mt. Soyosan, 13 VII 1998 (JC Sohn), [GSN] SJC- 770; 1 ♂, Gangwon Prov., Taebaeg, Mt. Maebongsan, 11 VII 1998 (JC Sohn), [GSN] SJC- 577; 3 ♂ 2 ♀, Gangwon Prov., Inje, Mt. Jeombongsan, 7 VII 2006 (JC Sohn), [GSN] SJC- 1199 (♀); 1 ♀, Jeonnam Prov., Gurae, Sandong, Mt. Jirisan, Nogodan (N 35 ° 17 ′ 35.4 ″ E 127 ° 31 ′ 41.1 ″, alt. 1344 m), 20 VII 2004 (JC Sohn), [GSN] SJC- 547.	en	Sohn, Jae-Cheon, Lvovsky, Alexander L. (2021): Review of Lypusinae (Lepidoptera: Gelechioidea: Lypusidae) from Korea with a description of a new species of Agnoea Walsingham, 1907. Zootaxa 4966 (3): 385-391, DOI: https://doi.org/10.11646/zootaxa.4966.3.10
C130115AFFC3FFC9FF65F980FE693544.taxon	distribution	Distribution. Korea (endemic).	en	Sohn, Jae-Cheon, Lvovsky, Alexander L. (2021): Review of Lypusinae (Lepidoptera: Gelechioidea: Lypusidae) from Korea with a description of a new species of Agnoea Walsingham, 1907. Zootaxa 4966 (3): 385-391, DOI: https://doi.org/10.11646/zootaxa.4966.3.10
C130115AFFC3FFC9FF65F980FE693544.taxon	discussion	Remarks. This species was described on the basis of only one male specimen. Female characters are described for the first time in the present study. Additional collecting localities are provided for Agnoea digitiella, expanding its distributional range.	en	Sohn, Jae-Cheon, Lvovsky, Alexander L. (2021): Review of Lypusinae (Lepidoptera: Gelechioidea: Lypusidae) from Korea with a description of a new species of Agnoea Walsingham, 1907. Zootaxa 4966 (3): 385-391, DOI: https://doi.org/10.11646/zootaxa.4966.3.10
C130115AFFC2FFCFFF65F8F9FCA036F4.taxon	description	(Figs 4, 8, 10)	en	Sohn, Jae-Cheon, Lvovsky, Alexander L. (2021): Review of Lypusinae (Lepidoptera: Gelechioidea: Lypusidae) from Korea with a description of a new species of Agnoea Walsingham, 1907. Zootaxa 4966 (3): 385-391, DOI: https://doi.org/10.11646/zootaxa.4966.3.10
C130115AFFC2FFCFFF65F8F9FCA036F4.taxon	diagnosis	Diagnosis. This species is similar to Agnoea digitiella in external appearance but differs from the latter in having the larger size (forewing length 8.5 – 9.6 mm) and the longer saccular extension in the male genitalia (Fig. 8).	en	Sohn, Jae-Cheon, Lvovsky, Alexander L. (2021): Review of Lypusinae (Lepidoptera: Gelechioidea: Lypusidae) from Korea with a description of a new species of Agnoea Walsingham, 1907. Zootaxa 4966 (3): 385-391, DOI: https://doi.org/10.11646/zootaxa.4966.3.10
C130115AFFC2FFCFFF65F8F9FCA036F4.taxon	materials_examined	Material examined. 4 ♂, 1 ♀, Chungbuk Prov., Jecheon, Hansu, Songgyeri, Mt. Weolagsan (N 36 ° 51 ′ 15.6 ″ E 128 ° 04 ′ 53.8 ″), 28 V 2013 (JC Sohn), [GSN] SJC- 1200 (♂), 1201 (♀).	en	Sohn, Jae-Cheon, Lvovsky, Alexander L. (2021): Review of Lypusinae (Lepidoptera: Gelechioidea: Lypusidae) from Korea with a description of a new species of Agnoea Walsingham, 1907. Zootaxa 4966 (3): 385-391, DOI: https://doi.org/10.11646/zootaxa.4966.3.10
C130115AFFC2FFCFFF65F8F9FCA036F4.taxon	distribution	Distribution. Korea, Japan (Hokkaido), Russia, Europe. Bionomics. The larvae feed on the dead leaves of Carpinus, Fagus, Populus, Quercus, and Vaccinium (Toll, 1964; Harper et al., 2002). They carry a case composed of the dead leaves. It is known in Europe that the larval stage lasts two years (Jäckh 1959).	en	Sohn, Jae-Cheon, Lvovsky, Alexander L. (2021): Review of Lypusinae (Lepidoptera: Gelechioidea: Lypusidae) from Korea with a description of a new species of Agnoea Walsingham, 1907. Zootaxa 4966 (3): 385-391, DOI: https://doi.org/10.11646/zootaxa.4966.3.10
C130115AFFC2FFCFFF65F8F9FCA036F4.taxon	discussion	Remarks. Koo et al. (2021) reported this species for the first time from Korea. Our specimens of A. josephinae show some differences in the male genitalia from the ones illustrated in Koo et al. (2021). Systematic value of those differences needs to be critically reviewed with COI barcodes.	en	Sohn, Jae-Cheon, Lvovsky, Alexander L. (2021): Review of Lypusinae (Lepidoptera: Gelechioidea: Lypusidae) from Korea with a description of a new species of Agnoea Walsingham, 1907. Zootaxa 4966 (3): 385-391, DOI: https://doi.org/10.11646/zootaxa.4966.3.10
