taxonID	type	description	language	source
03AF8785A378276EFF0420BE1F77F867.taxon	description	(Figs 1 – 4, 25)	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
03AF8785A378276EFF0420BE1F77F867.taxon	materials_examined	Type material. Holotype: MALAYSIA (Sabah): ♂, three labels: “ (N. BORNEO) / Headquarter / Mt. Kinabalu / 1. V. 1980 / M. & A. Sakai ” [white, printed], “ Collection / of Ent. Lab. / Ehime Univ. ” [blue, printed], “ HLAVACIELLUS / cincinnalis m. / P. Jałoszyński, 2021 / HOLOTYPUS ” [red, printed] (EUMJ).	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
03AF8785A378276EFF0420BE1F77F867.taxon	diagnosis	Diagnosis. Body large, exceeding 2 mm in length; head in male modified, with median ‘ lock’ of dense setae inserted between pair of tiny tubercles far above level of posterior margins of eyes, and flanked by another pair of similar tubercles situated more anteriorly and separated by distance twice as wide as posterior tubercles, setal ‘ lock’ directed anterad; pronotum distinctly narrower than elytra; each of antennomeres 3 – 10 at least slightly elongate; aedeagus in ventral view stout, slightly less than 3 times as long as broad and distinctly narrowing from base to apex; parameres with conspicuously thick subapical setae.	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
03AF8785A378276EFF0420BE1F77F867.taxon	description	Description. Body of male (Fig. 1) moderately stout, strongly convex, uniformly light brown; setae slightly lighter than cuticle; BL 2.10 mm. Head (Fig. 2) broadest at large, strongly convex and coarsely faceted eyes, HL 0.30 mm, HW 0.51 mm; frons unmodified, only indistinctly convex; supraantennal tubercles prominent; vertex indistinctly convex and modified in its posterior region, far beyond eyes, with two pairs of tiny tubercles forming trapeze, and with slender median tuft of dense setae inserted between posterior tubercles and directed anteriorly. Punctures on frons and vertex fine, inconspicuous; setae moderately dense and long, suberect. Antennae slender, AnL 1.45 mm, BL / AnL 1.45; scape indistinctly elongate, pedicel about as long as broad, antennomeres 3 – 10 each slightly or distinctly elongate, 11 distinctly shorter than 9 – 10 combined, twice as long as broad. Pronotum subtrapezoidal, broadest at base; PL 0.60 mm, PW 0.78 mm, PI 0.77; anterior margin strongly rounded; sides with finely crenulate lateral carinae, indistinctly sinuate; posterior corners slightly sharp-angled and with acute tips; posterior margin shallowly bisinuate; pronotal disc strongly convex, with area at each posterior corner distinctly impressed; antebasal pits equally distant from lateral and posterior margins. Punctures on pronotum fine and inconspicuous; setae long, dense, suberect. Elytra together oval, broadest distinctly in front of middle; EL 1.20 mm, EW 0.95 mm, EI 1.26; subhumeral lines sharply marked, each developed as step-wise border between elevated humeral region and median area, as long as 0.3 EL; sides of elytra strongly rounded; elytral apices separately rounded. Punctures and setae similar to those on pronotum. Legs long and slender, unmodified. Aedeagus (Figs 3 – 4) elongate but not very slender; AeL 0.45 mm; median lobe in ventral view slightly less than 3 times as long as broad, tapering from base to apex; endophallic structures poorly sclerotized, indistinct; parameres with conspicuously thick subapical setae. Female. Unknown.	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
03AF8785A378276EFF0420BE1F77F867.taxon	distribution	Distribution. Northern Borneo (Fig. 25).	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
03AF8785A378276EFF0420BE1F77F867.taxon	etymology	Etymology. The adjective cincinnalis is derived from the Latin noun cincinnus, a lock of hair; the name refers to the median ‘ lock’ of setae on the vertex.	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
03AF8785A378276EFF0420BE1F77F867.taxon	discussion	Remarks. Hlavaciellus cincinnalis belongs to large-bodied species, clearly exceeding 2 mm in length. The conspicuous tuft of setae on the vertex inserted between a pair of small tubercles and flanked by another pair of more anteriorly situated tubercles is a unique apomorphy of this species and allows for unambiguous identification.	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
03AF8785A37A2768FF0423A31F32FBBD.taxon	description	(Figs 5 – 8, 25)	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
03AF8785A37A2768FF0423A31F32FBBD.taxon	materials_examined	Type material. Holotype: MALAYSIA (Pahang): ♂, two labels: “ W. Malaysia: Pahang / Cameron Highlands / trail 9, 1400 m, 27.3.93 / Löbl & Calame, # 21 ” [white, printed], “ HLAVACIELLUS / carinatus m. / P. Jałoszyński, 2021 / HOLOTYPUS ” [red, printed] (MHNG).	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
03AF8785A37A2768FF0423A31F32FBBD.taxon	diagnosis	Diagnosis. Body large, exceeding 2 mm in length; head in male modified, with broadly V-shaped (anteriorly concave) and sharply marked transverse carina between eyes accompanied anteriorly and posteriorly by glossy and asetose transverse impressions, and with pair of tiny tubercles behind carina; pronotum slightly narrower than elytra; each of antennomeres 3 – 5 as long as broad, and 6 – 10 each slightly elongate; aedeagus in ventral view slender, 3.5 times as long as broad and distinctly narrowing both toward apex and base; parameres with thin subapical setae.	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
03AF8785A37A2768FF0423A31F32FBBD.taxon	description	Description. Body of male (Fig. 5) stout, strongly convex, uniformly dark brown; setae slightly lighter than cuticle; BL 2.10 mm. Head (Fig. 6) broadest at large, strongly convex and coarsely faceted eyes, HL 0.45 mm, HW 0.48 mm; both frons and vertex modified, with broadly V-shaped (anteriorly concave) strongly elevated and thin transverse carina, frons in front of carina distinctly transversely impressed, vertex just behind carina also transversely impressed and with pair of tiny submedian tubercles. Punctures present only behind and in front of impressed areas, fine and inconspicuous; impressions asetose, setae in front of anterior impression sparse, long and suberect, those behind posterior impression distinctly denser and nearly recumbent. Antennae slender, AnL 1.20 mm, BL / AnL 1.75; scape indistinctly elongate, antennomeres 2 – 5 each as long as broad, 6 – 10 each slightly elongate (9 indistinctly so), 11 slightly shorter than 9 – 10 combined, twice as long as broad. Pronotum subrectangular, parallel-sided between base and anterior third; PL 0.63 mm, PW 0.88 mm, PI 0.71; anterior margin weakly rounded; sides with finely crenulate lateral carinae, in posterior 2 / 3 straight; posterior corners right-angled and with blunt tips; posterior margin shallowly bisinuate; pronotal disc strongly convex, with area at each posterior corner distinctly impressed; antebasal pits slightly closer to posterior than to lateral margins. Punctures on pronotum fine and inconspicuous; setae long, dense, suberect. Elytra together oval, broadest distinctly in front of middle; EL 1.03 mm, EW 0.93 mm, EI 1.11; subhumeral lines sharply marked, each developed as step-wise border between elevated humeral region and median area, as long as 0.3 EL; sides of elytra strongly rounded; elytral apices separately rounded. Punctures and setae similar to those on pronotum. Legs long and slender, unmodified. Aedeagus (Figs 7 – 8) elongate and very slender; AeL 0.30 mm; median lobe in ventral view 3.5 times as long as broad, broadest near sub-basal third and with broadly rounded sides; endophallic structures distinct, with three darkly sclerotized elongate and pointed elements in basal half of median lobe; parameres with thin subapical setae. Female. Unknown.	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
03AF8785A37A2768FF0423A31F32FBBD.taxon	distribution	Distribution. Peninsular Malaysia (Fig. 25).	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
03AF8785A37A2768FF0423A31F32FBBD.taxon	etymology	Etymology. The adjective carinatus refers to the conspicuous transverse carina on the head of this species.	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
03AF8785A37A2768FF0423A31F32FBBD.taxon	discussion	Remarks. Hlavaciellus carinatus belongs to large-bodied species, clearly exceeding 2 mm in length. The conspicuous transverse, strongly elevated and thin carina on the vertex accompanied by a pair of tiny posteromedian tubercles and two transverse impressions (anterior and posterior) is a unique apomorphy of this species.	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
03AF8785A37D276AFF04279E1F4AFF1C.taxon	description	(Figs 9 – 12, 25)	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
03AF8785A37D276AFF04279E1F4AFF1C.taxon	materials_examined	Type material. Holotype: MALAYSIA (Pahang): ♂, two labels: “ W. Malaysia: Pahang / Cameron Highlands / trail 9, 1400 m, 27.3.93 / Löbl & Calame, # 21 ” [white, printed], “ HLAVACIELLUS / microtuberculatus m. / P. Jałoszyński, 2021 / HOLOTYPUS ” [red, printed] (MHNG). Paratypes (2 exx.): 1 ♀, same data as for holotype; 1 ♀, “ W. Malaysia: Pahang # 18 b / Cameron Highls, 1550 m / Gunung Jasar, trail 11 / Löbl & Calame, 24.3.93 ”; paratypes with yellow “ PARATYPUS ” labels (MHNG, cPJ).	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
03AF8785A37D276AFF04279E1F4AFF1C.taxon	diagnosis	Diagnosis. Body small, not reaching 2 mm in length and conspicuously stout, with elytra about as long as broad; head in males with unmodified frons and vertex, but in both sexes with pair of tiny, barely discernible submedian tubercles on vertex; pronotum slightly narrower than elytra; each of antennomeres 3 – 4 and 7 – 10 about as long as broad, and 5 – 6 each indistinctly elongate; aedeagus in ventral view stout, 3 times as long as broad and distinctly narrowing both toward apex and base; parameres with thin subapical setae.	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
03AF8785A37D276AFF04279E1F4AFF1C.taxon	description	Description. Body of male (Fig. 9) stout, strongly convex, uniformly dark brown; setae distinctly lighter than cuticle; BL 1.78 mm. Head (Fig. 10) broadest at large, strongly convex and coarsely faceted eyes, HL 0.33 mm, HW 0.45 mm; frons and vertex unmodified, weakly convex, vertex with pair of tiny submedian tubercles (barely discernible; observations at various angles are necessary to notice tubercles). Punctures on frons and vertex fine and inconspicuous; setae long, moderately dense and suberect. Antennae slender, AnL 1.23 mm, BL / AnL 1.45; scape indistinctly elongate, antennomeres 3 – 4 and 7 – 10 each about as long as broad, and 5 – 6 each indistinctly elongate, 11 distinctly shorter than 9 – 10 combined, twice as long as broad. Pronotum subrectangular, broadest near anterior third but indistinctly narrowing toward base; PL 0.58 mm, PW 0.85 mm, PI 0.68; anterior margin weakly rounded; sides with finely crenulate lateral carinae, in posterior 2 / 3 straight; posterior corners nearly right-angled and with blunt tips; posterior margin shallowly bisinuate; pronotal disc strongly convex, with area at each posterior corner distinctly impressed; antebasal pits slightly closer to posterior than to lateral margins. Punctures on pronotum fine and inconspicuous; setae long, dense, suberect. Elytra together oval, broadest distinctly in front of middle; EL 0.88 mm, EW 0.93 mm, EI 0.95; subhumeral lines sharply marked, each developed as step-wise border between elevated humeral region and median area, slightly exceeding 0.3 EL; sides of elytra strongly rounded; elytral apices separately rounded. Punctures and setae similar to those on pronotum. Legs long and slender, unmodified. Aedeagus (Figs 11 – 12) elongate but not very slender; AeL 0.30 mm; median lobe in ventral view 3 times as long as broad, broadest near sub-basal third and with broadly rounded sides; endophallic structures distinct, in ventral view appearing as single elongate subtriangular element, but in lateral view two separate tubular structures can be seen, proximal one bent at sharp angle near base of median lobe; parameres with thin subapical setae. Female. Externally similar to male but with clearly shorter antennae in relation to body. BL 1.78 – 1.85 mm; HL 0.30 – 0.40 mm, HW 0.45 – 0.48 mm, BL / AnL 1.82 – 1.95; PL 0.55 – 0.58 mm, PW 0.85 – 0.90 mm, PI 0.64 – 0.65; EL 0.90 mm, EW 0.90 – 0.93 mm, EI 0.97 – 1.00.	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
03AF8785A37D276AFF04279E1F4AFF1C.taxon	distribution	Distribution. Peninsular Malaysia (Fig. 25).	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
03AF8785A37D276AFF04279E1F4AFF1C.taxon	etymology	Etymology. The adjective microtuberculatus refers to the pair of tiny tubercles on the vertex in both sexes.	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
03AF8785A37D276AFF04279E1F4AFF1C.taxon	discussion	Remarks. Hlavaciellus microtuberculatus belongs to small-bodied species, clearly below 2 mm in length. The most conspicuous feature of this otherwise unremarkable species is a very stout body, with elytra about as long as broad, or even indistinctly broader than long; in all remaining species the elytra are at least slightly elongate.	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
03AF8785A37F276BFF04246F1E59F9ED.taxon	description	(Figs 13 – 16, 25)	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
03AF8785A37F276BFF04246F1E59F9ED.taxon	materials_examined	Type material. Holotype: MALAYSIA (Pahang): ♂, two labels: “ E. Malaysia: Sarawak / confl. Sun Oyan and / Mujong riv., E. Kapit / 50 m, 18. V. 1994, # 5 a / Löbl & Burckhardt ” [white, printed], “ HLAVACIELLUS / diversipilosus m. / P. Jałoszyński, 2021 / HOLOTYPUS ” [red, printed] (MHNG). Paratype: 1 ♂, same data as for holotype (cPJ).	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
03AF8785A37F276BFF04246F1E59F9ED.taxon	diagnosis	Diagnosis. Body very small, reaching only 1.6 mm in length; head in males with pair of transverse, shallow and indistinct postocular impressions and several setae on median frontal region divergent laterad along anterior margins of impressions; each of antennomeres 1 – 10 about as long as broad; aedeagus in ventral view moderately stout, 3.2 times as long as broad and distinctly narrowing both toward apex and base; parameres with thin subapical setae.	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
03AF8785A37F276BFF04246F1E59F9ED.taxon	description	Description. Body of male (Fig. 13) stout, strongly convex, uniformly dark brown; setae distinctly lighter than cuticle; BL 1.58 – 1.60 mm. Head (Fig. 14) broadest at large, strongly convex and coarsely faceted eyes, HL 0.25 mm, HW 0.43 mm; frons and vertex weakly convex at middle, vertex behind each eye with shallow transverse impression. Punctures on frons and vertex fine and inconspicuous; setae long, moderately dense and suberect, a few setae inserted on median frontal region are strongly divergent laterad along anterior margins of postocular impressions. Antennae slender, AnL 1.05 mm, BL / AnL 1.50 – 1.52; antennomeres 1 – 10 each about as long as broad, 11 distinctly shorter than 9 – 10 combined, twice as long as broad. Pronotum subrectangular, broadest at base but indistinctly narrowing anterad; PL 0.45 – 0.48 mm, PW 0.73 mm, PI 0.62 – 0.66; anterior margin weakly rounded; sides with finely crenulate lateral carinae, in posterior 2 / 3 straight; posterior corners sharp-angled and with acute tips; posterior margin shallowly bisinuate; pronotal disc strongly convex, with area at each posterior corner distinctly impressed; antebasal pits slightly closer to posterior than to lateral margins. Punctures on pronotum fine and inconspicuous; setae long, dense, suberect. Elytra together oval, broadest distinctly in front of middle; EL 0.88 mm, EW 0.80 mm, EI 1.09; subhumeral lines sharply marked, each developed as step-wise border between elevated humeral region and median area, slightly exceeding 0.3 EL; sides of elytra strongly rounded; elytral apices separately rounded. Punctures and setae similar to those on pronotum. Legs long and slender, unmodified. Aedeagus (Figs 15 – 16) elongate but not very slender; AeL 0.33 – 0.35 mm; median lobe in ventral view 3.2 times as long as broad, broadest near sub-basal third and with broadly rounded sides; endophallic structures distinct, in ventral view appearing as two elongate elements, also in lateral view two separate tubular structures can be seen, proximal one bent at sharp angle near base of median lobe, distal one with cup-like broadening situated near middle of median lobe; parameres with thin subapical setae. Female. Unknown.	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
03AF8785A37F276BFF04246F1E59F9ED.taxon	distribution	Distribution. Borneo (Fig. 25).	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
03AF8785A37F276BFF04246F1E59F9ED.taxon	etymology	Etymology. The adjective diversipilosus refers to the setal pattern on the head, with some setae directed laterad along postocular impressions.	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
03AF8785A37F276BFF04246F1E59F9ED.taxon	discussion	Remarks. Among SE Asian species, Hlavaciellus diversipilosus has the smallest adults, with the body length not exceeding 1.6 mm. The small size, body shape, transverse postocular impressions on the head, and the endophallic structures of the aedeagus in lateral view with a cup-like broadened distal portion of the distal sclerite is a unique combination of characters that allows for unambiguous identification of this species.	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
03AF8785A37E2765FF04256E1B27FD15.taxon	description	(Figs 17 – 20, 25)	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
03AF8785A37E2765FF04256E1B27FD15.taxon	materials_examined	Type material. Holotype: INDONESIA (Sumatra): ♂, two labels: “ SUMATRA: Jambi / Mt Kerinci, 1750 - / 1850 m, 11 - 12. XI. / 1989, Agosti, Löbl / Burckhardt # 11 ” [white, printed], “ HLAVACIELLUS / concavus m. / P. Jałoszyński, 2021 / HOLOTYPUS ” [red, printed] (MHNG).	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
03AF8785A37E2765FF04256E1B27FD15.taxon	diagnosis	Diagnosis. Body very large, exceeding 2.2 mm in length; head in males with large and deep median impression between eyes filled with short and dense setae directed toward its center; each of antennomeres 3 – 10 elongate; aedeagus in ventral view moderately stout, 3 times as long as broad and distinctly narrowing both toward apex and base; parameres with thin subapical setae and conspicuously broadened and then rapidly narrowed apical portions.	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
03AF8785A37E2765FF04256E1B27FD15.taxon	description	Description. Body of male (Fig. 17) stout, strongly convex, uniformly light brown; setae distinctly lighter than cuticle; BL 2.23 mm. Head (Fig. 18) broadest at large, strongly convex and coarsely faceted eyes, HL 0.30 mm, HW 0.60 mm; frons and vertex with large and deep circular median impression. Punctures on frons in front of impression fine and inconspicuous; setae between supraantennal tubercles and on sides of impression long, moderately dense and suberect, impression densely filled with short, recumbent setae directed toward its center. Antennae slender, AnL 1.63 mm, BL / AnL 1.37; scape indistinctly elongate, pedicel as long as broad, antennomeres 3 – 10 each elongate (4 and 5 weakly so), 11 slightly shorter than 9 – 10 combined, 2.2 × as long as broad. Pronotum subrectangular, equally broad at base and at anterior third; PL 0.68 mm, PW 0.93 mm, PI 0.73; anterior margin weakly rounded; sides with finely crenulate lateral carinae, in posterior 2 / 3 distinctly concave; posterior corners sharp-angled but with blunt tips; posterior margin shallowly bisinuate; pronotal disc strongly convex, with area at each posterior corner distinctly impressed; antebasal pits slightly closer to posterior than to lateral margins. Punctures on pronotum fine and inconspicuous; setae long, dense, suberect. Elytra together oval, broadest distinctly in front of middle; EL 1.25 mm, EW 1.08 mm, EI 1.16; subhumeral lines sharply marked, each developed as step-wise border between elevated humeral region and median area, as long as 0.3 EL; sides of elytra strongly rounded; elytral apices separately rounded. Punctures and setae similar to those on pronotum. Legs long and slender, unmodified. Aedeagus (Figs 19 – 20) elongate but not very slender; AeL 0.45 mm; median lobe in ventral view 3 times as long as broad, broadest near sub-basal fourth and with broadly rounded sides near base, but lateral margins straight in distal 2 / 3; endophallic structures distinct, in ventral view appearing as two elongate elements, but in lateral view one tubular structure can be seen bent at sharp angle near base of median lobe; parameres with thin subapical setae, each with broadened and then rapidly narrowed distal region. Female. Unknown.	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
03AF8785A37E2765FF04256E1B27FD15.taxon	distribution	Distribution. Sumatra (Fig. 25).	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
03AF8785A37E2765FF04256E1B27FD15.taxon	etymology	Etymology. Concavus is a Latin adjective meaning “ hollow, concave ” and refers to the large median concavity on the head.	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
03AF8785A37E2765FF04256E1B27FD15.taxon	discussion	Remarks. The conspicuous round impression densely filled with short setae on the head is a unique apomorphy of this species; in addition, H. concavus has the largest adults of all known members of this genus, exceeding 2.2 mm in length.	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
03AF8785A3702767FF0420761821FF31.taxon	description	(Figs 21 – 25)	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
03AF8785A3702767FF0420761821FF31.taxon	materials_examined	Type material. Holotype: INDONESIA (Sumatra): ♂, two labels: “ SUMATRA: Jambi / Mt Kerinci, 3300 m, 12. XI. 1989 / Agosti, Löbl / Burckhardt # 12 a ” [white, printed], “ HLAVACIELLUS / sumatranus m. / P. Jałoszyński, 2021 / HOLOTYPUS ” [red, printed] (MHNG).	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
03AF8785A3702767FF0420761821FF31.taxon	diagnosis	Diagnosis. Body large, exceeding 2 mm in length; head in males modified: posteromedian region of vertex with group of dense setae forming loose tuft, vertex with shallow transverse postocular impression, and posterior margin of frons behind each supraantennal tubercle with arcuate (posteriorly convex) ridge; each of antennomeres 3 – 9 elongate, 10 about as long as broad; aedeagus in ventral view slender, nearly 4 times as long as broad and distinctly narrowing both toward apex and base; parameres with thin subapical setae and rapidly narrowed apical portions.	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
03AF8785A3702767FF0420761821FF31.taxon	description	Description. Body of male (Fig. 21) stout, strongly convex, uniformly dark brown; setae distinctly lighter than cuticle; BL 2.18 mm. Head (Fig. 22) broadest at large, strongly convex and coarsely faceted eyes, HL 0.38 mm, HW 0.58 mm; posteromedian region of vertex with group of dense setae forming loose tuft, vertex with shallow transverse postocular impression, and posterior margin of frons behind each supraantennal tubercle with arcuate (posteriorly convex) ridge. Frons between supraantennal tubercles with uneven, slightly coarse surface; punctures on remaining areas fine and inconspicuous; setae long, moderately dense and suberect, those on posteromedian region of vertex dense and erect, postocular impression asetose. Antennae slender, AnL 1.38 mm, BL / AnL 1.58; scape indistinctly elongate, pedicel as long as broad, antennomeres 3 – 7 each strongly elongate, 8 and 9 each slightly elongate, 10 about as long as broad, 11 indistinctly shorter than 9 – 10 combined, twice as long as broad. Pronotum subrectangular, equally broad between base and anterior third; PL 0.68 mm, PW 1.00 mm, PI 0.68; anterior margin weakly rounded; sides with finely crenulate lateral carinae, in posterior 2 / 3 straight; posterior corners right-angled and with blunt tips; posterior margin shallowly bisinuate; pronotal disc strongly convex, with area at each posterior corner distinctly impressed; antebasal pits slightly closer to posterior than to lateral margins. Punctures on pronotum fine and inconspicuous; setae long, dense, suberect. Elytra together oval, broadest distinctly in front of middle; EL 1.13 mm, EW 0.98 mm, EI 1.15; subhumeral lines sharply marked, each developed as step-wise border between elevated humeral region and median area, as long as 0.3 EL; sides of elytra strongly rounded; elytral apices separately rounded. Punctures and setae similar to those on pronotum. Legs long and slender, unmodified. Aedeagus (Figs 23 – 24) elongate and slender; AeL 0.40 mm; median lobe in ventral view nearly 4 times as long as broad, broadest near sub-basal third and with broadly rounded sides; endophallic structures in the only known specimen erected, darkly sclerotized and tubular; parameres with thin subapical setae, each with parallel-sided and then rapidly narrowed distal region. Female. Unknown.	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
03AF8785A3702767FF0420761821FF31.taxon	distribution	Distribution. Sumatra (Fig. 25).	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
03AF8785A3702767FF0420761821FF31.taxon	etymology	Etymology. After terra typica, i. e., Sumatra.	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
03AF8785A3702767FF0420761821FF31.taxon	discussion	Remarks. Apart from unique modifications of the head, the only known male of H. sumatranus has the pronotum distinctly broader than elytra, a feature not known in any other congener described so far. Although the aedeagus of the holotype is in erected condition, with endophallic structures projecting distally from the ostium, the shape of parameres in lateral view can be used as one of diagnostic characters. The apical region of each paramere resembles that of the sympatric H. concavus, but in the latter species the parameres are weakly, and in H. sumatranus strongly curved near distal third.	en	Jałoszyński, Paweł (2021): Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae). Zootaxa 4966 (1): 1-15, DOI: https://doi.org/10.11646/zootaxa.4966.1.1
