taxonID	type	description	language	source
03C987A6FFFC641C769F732BFDEF7F6C.taxon	materials_examined	Type species: Drassus hypocrita Simon, 1878 from France.	en	Fomichev, Alexander A., Marusik, Yuri M. (2021): A survey of East Palaearctic Gnaphosidae (Araneae). 11. New data on Gnaphosidae (Araneae) from Tajikistan. Zootaxa 4966 (4): 443-457, DOI: https://doi.org/10.11646/zootaxa.4966.4.3
03C987A6FFFC641C769F732BFDEF7F6C.taxon	discussion	Comments. So far, ten species distributed from Iberian Peninsula to Turkey are known in the genus (WSC 2021). One undetermined species has been reported from northern Iran (Kashefi et al. 2013) and there are doubtful records of the generotype from Kazakhstan and Uzbekistan (Mikhailov 2013). Our record from Tajikistan is the easternmost in the whole genus range.	en	Fomichev, Alexander A., Marusik, Yuri M. (2021): A survey of East Palaearctic Gnaphosidae (Araneae). 11. New data on Gnaphosidae (Araneae) from Tajikistan. Zootaxa 4966 (4): 443-457, DOI: https://doi.org/10.11646/zootaxa.4966.4.3
03C987A6FFFC641C769F7067FB7C7BCB.taxon	description	Figs 1 – 8, 65 – 66.	en	Fomichev, Alexander A., Marusik, Yuri M. (2021): A survey of East Palaearctic Gnaphosidae (Araneae). 11. New data on Gnaphosidae (Araneae) from Tajikistan. Zootaxa 4966 (4): 443-457, DOI: https://doi.org/10.11646/zootaxa.4966.4.3
03C987A6FFFC641C769F7067FB7C7BCB.taxon	materials_examined	Type. TAJIKISTAN, Khatlon Region: holotype ♂ (ZMMU), Vaksh Karatau Mt. Range, Khodzhamaston Mt., 38 ° 01.192 ’ N, 68 ° 57.144 ’ E, 1040 m, 24.04.2015 (Y. M. Marusik).	en	Fomichev, Alexander A., Marusik, Yuri M. (2021): A survey of East Palaearctic Gnaphosidae (Araneae). 11. New data on Gnaphosidae (Araneae) from Tajikistan. Zootaxa 4966 (4): 443-457, DOI: https://doi.org/10.11646/zootaxa.4966.4.3
03C987A6FFFC641C769F7067FB7C7BCB.taxon	etymology	Etymology. The specific epithet is derived from the country of origin.	en	Fomichev, Alexander A., Marusik, Yuri M. (2021): A survey of East Palaearctic Gnaphosidae (Araneae). 11. New data on Gnaphosidae (Araneae) from Tajikistan. Zootaxa 4966 (4): 443-457, DOI: https://doi.org/10.11646/zootaxa.4966.4.3
03C987A6FFFC641C769F7067FB7C7BCB.taxon	diagnosis	Diagnosis. The new species differs from all congeners by simple embolus (Em) lacking coils (vs. coiled) (cf. Figs 4, 6 – 8 and figs 4, 8 in Hervé et al. 2009).	en	Fomichev, Alexander A., Marusik, Yuri M. (2021): A survey of East Palaearctic Gnaphosidae (Araneae). 11. New data on Gnaphosidae (Araneae) from Tajikistan. Zootaxa 4966 (4): 443-457, DOI: https://doi.org/10.11646/zootaxa.4966.4.3
03C987A6FFFC641C769F7067FB7C7BCB.taxon	description	Description. Male. Habitus as in Fig. 1. Total length 6.05. Carapace 2.8 long, 2.25 wide. Coloration (freshly molted specimen). Prosoma, all limbs and spinnerets light yellow. Abdomen dark yellow with small gray spots dorsally, yellow ventrally. Leg spination: I: Fe d 1 - 1 - 1, p 0 - 1 - 2; Ti p 1 - 0 - 1, v 2 - 2 - 2; Mt v 2 - 2 - 0. II: Fe d 1 - 1 - 1, p 0 - 1 - 1; Ti d 0 - 1 - 0, p 1 - 0 - 1, v 2 - 2 - 2; Mt p 0 - 1 - 0, v 2 - 2 - 0. III: Fe d 1 - 1 - 1, p 0 - 1 - 1, r 0 - 1 - 1; Pt r 1; Ti d 1 - 1 - 0, p 1 - 1 - 1, r 0 - 1 - 1, v 2 - 2 - 2; Mt d 2 - 2 - 0, p 0 - 1 - 0, r 0 - 1 - 0, v 2 - 0 - 2. IV: Fe d 1 - 1 - 1, p 0 - 1 - 1, r 0 - 1 - 1; Pt r 1; Ti d 1 - 1 - 1, p 1 - 1 - 1, r 1 - 1 - 1, v 2 - 2 - 2; Mt d 2 - 2 - 0, p 0 - 1 - 0, r 0 - 1 - 0, v 2 - 2 - 0. Leg measurements: I: 10.08 (2.73, 1.5, 2.5, 2.05, 1.3); II: 8.39 (2.35, 1.25, 1.93, 1.73, 1.13); III: 7.64 (2.13, 1.05, 1.53, 1.9, 1.03); IV: 10.63 (2.8, 1.33, 2.25, 2.95, 1.3). Palp as in Figs 2 – 5, 6 – 8. Tibial apophysis bifurcate, with claw-like superior prong (Sp) and blunt inferior prong (Ip), forming a right angle; median apophysis (Ma) almost straight and pointed; embolus whip-like, without outgrowth; subtegulum (St) rounded. Female. Unknown.	en	Fomichev, Alexander A., Marusik, Yuri M. (2021): A survey of East Palaearctic Gnaphosidae (Araneae). 11. New data on Gnaphosidae (Araneae) from Tajikistan. Zootaxa 4966 (4): 443-457, DOI: https://doi.org/10.11646/zootaxa.4966.4.3
03C987A6FFFC641C769F7067FB7C7BCB.taxon	distribution	Distribution. Known only from the type locality.	en	Fomichev, Alexander A., Marusik, Yuri M. (2021): A survey of East Palaearctic Gnaphosidae (Araneae). 11. New data on Gnaphosidae (Araneae) from Tajikistan. Zootaxa 4966 (4): 443-457, DOI: https://doi.org/10.11646/zootaxa.4966.4.3
03C987A6FFFC641C769F7067FB7C7BCB.taxon	discussion	Comments. The generic placement is provisional and is due to the bifurcated tibial apophysis. Drassodex tajikistanicus sp. n. lacks a highly coiled or twisted embolus which is typical for Drassodex, and is likely to be transferred to a new genus in future. Furthermore, most of Drassodex species mainly occur in Pyrenees and Alps, mountains of south-western Europe (Hervé et al. 2009). Genus Nomisia Dalmas, 1921 Type species: Pythonissa exornata C. L. Koch, 1839 from Germany. Comments. To date, 38 species distributed from Canary Islands to China and south to South Africa are known in the genus (WSC 2021). Three species has been recorded from Central Asia (Mikhailov 2013).	en	Fomichev, Alexander A., Marusik, Yuri M. (2021): A survey of East Palaearctic Gnaphosidae (Araneae). 11. New data on Gnaphosidae (Araneae) from Tajikistan. Zootaxa 4966 (4): 443-457, DOI: https://doi.org/10.11646/zootaxa.4966.4.3
03C987A6FFFC641A769F74E7FAE77E00.taxon	description	Figs 9 – 21, 65 – 66.	en	Fomichev, Alexander A., Marusik, Yuri M. (2021): A survey of East Palaearctic Gnaphosidae (Araneae). 11. New data on Gnaphosidae (Araneae) from Tajikistan. Zootaxa 4966 (4): 443-457, DOI: https://doi.org/10.11646/zootaxa.4966.4.3
03C987A6FFFC641A769F74E7FAE77E00.taxon	materials_examined	Material examined. TAJIKISTAN, Khatlon Region: 3 ♀ (ZMMU), Khuroson Distr., Gandzhina, 37 ° 57.701 ’ N, 68 ° 33.717 ’ E, 716 m, cliff, dry slopes, under stones, 21.04.2015 (Y. M. Marusik); 1 ♂ 1 ♀ (ZMMU), Dangara Distr., SW slope of Sanglogh (= Sanglok) Mt. Range, 38 ° 13.091 ’ N, 69 ° 14.282 ’ E, 1362 m, 30.04.2015 (Y. M. Marusik); 1 ♀ (ZMMU), Pyandzh Karatau Mt. Ridge, foothills of Astana Mt., 37 ° 23.202 ’ N, 69 ° 14.810 ’ E, 1674 m, 4.05.2015 (S. L. Zonstein); 1 ♂ (ISEA, 001.8557), Shurobdarya River valley, Bal’dzhuvon Village, 38 ° 18.835 ’ N, 69 ° 41.011 ’ E, stony clay hills covered with meadow vegetation, 900 – 1000 m, 15.04.2019 (A. A. Fomichev); 2 ♀ (ISEA, 001.8558), Tabakchi Mt. Range, 3 km ENE from Kalininabad Town, 37 ° 52.918 ’ N, 68 ° 58.719 ’ E, dry stony shiblyak shrubland, 1030 – 1280 m, 18.04.2019 (A. A. Fomichev); 1 ♂ 1 ♀ (ISEA, 001.8559), Aruktau Mt. Range, 7 km SW from Sambuli Village, 37 ° 45.602 ’ N, 68 ° 32.741 ’ E, dry stony shiblyak shrubland with rocks, 660 – 980 m, 09.04.2019 (A. A. Fomichev); Districts of Republican Subordination: 1 ♂ 1 ♀ (ZMMU), 38 th km of Varzob Hwy, Takob Gorge, env. of Dehmalik Vil., 38 ° 50.829 ’ N, 68 ° 54.637 ’ E, 805 m, 8.05.2015 (Y. M. Marusik); 4 ♂ 3 ♀ (ZMMU), 48 th km of Varzob Hwy, 38 ° 55.531 ’ N, 68 ° 48.292 ’ E, 1530 m, S exposed slope with Juglans litter & under stones, 7.05.2015 (Y. M. Marusik & M. Saidov); 4 ♂ 4 ♀ (ZMMU), Hissar Mt. Range, Ramit Res., Darai Holmon (Kholmon) creek gorge, 38 ° 45.364 ’ N, 69 ° 18.286 ’ E, 1370 m, 2.05.2015 (Y. M. Marusik); 6 ♀ (ZMMU), Hissar Mts., Romit Res., Surkhob Gorge, 1.5 km W Soni Vil., 38 ° 50 ’ N, 69 ° 26 ’ E, 1750 m, 5.07.2019 (S. L. Zonstein); 1 ♂ (ISEA, 001.8560), right side of the Pandema River valley (left tributary of Kafirnigan River), Poyonob Village, 38 ° 42.302 ’ N, 69 ° 18.283 ’ E, humid stony shiblyak shrubland with rocks, 1300 – 1370 m, 22.04.2019 (A. A. Fomichev); 1 ♂ 1 ♀ (ISEA, 001.8561), right side of the Kafirnigan River valley, Romit Village, 38 ° 43.225 ’ N, 69 ° 18.387 ’ E, humid stony shiblyak shrubland with rocks, 1200 – 1850 m, 21 – 22.04.2019 (A. A. Fomichev).	en	Fomichev, Alexander A., Marusik, Yuri M. (2021): A survey of East Palaearctic Gnaphosidae (Araneae). 11. New data on Gnaphosidae (Araneae) from Tajikistan. Zootaxa 4966 (4): 443-457, DOI: https://doi.org/10.11646/zootaxa.4966.4.3
03C987A6FFFC641A769F74E7FAE77E00.taxon	diagnosis	Diagnosis. The males of N. palaestina can be easily separated from all congeners by the large S-shaped embolus (Em). The females differ from others Central Asian species in having triangular epigynal fovea (Fo).	en	Fomichev, Alexander A., Marusik, Yuri M. (2021): A survey of East Palaearctic Gnaphosidae (Araneae). 11. New data on Gnaphosidae (Araneae) from Tajikistan. Zootaxa 4966 (4): 443-457, DOI: https://doi.org/10.11646/zootaxa.4966.4.3
03C987A6FFFC641A769F74E7FAE77E00.taxon	description	Description. See Levy (1995).	en	Fomichev, Alexander A., Marusik, Yuri M. (2021): A survey of East Palaearctic Gnaphosidae (Araneae). 11. New data on Gnaphosidae (Araneae) from Tajikistan. Zootaxa 4966 (4): 443-457, DOI: https://doi.org/10.11646/zootaxa.4966.4.3
03C987A6FFFC641A769F74E7FAE77E00.taxon	distribution	Distribution. Previously this species was reported from Greece, Turkey, Syria, Israel and Iran (Chatzaki 2010; Zamani et al. 2020). New records from Tajikistan represent the easternmost localities of this species.	en	Fomichev, Alexander A., Marusik, Yuri M. (2021): A survey of East Palaearctic Gnaphosidae (Araneae). 11. New data on Gnaphosidae (Araneae) from Tajikistan. Zootaxa 4966 (4): 443-457, DOI: https://doi.org/10.11646/zootaxa.4966.4.3
03C987A6FFFC641A769F74E7FAE77E00.taxon	discussion	Comments. Recorded from Tajikistan for the first time; apparently, N. palaestina is common and widespread in south-western regions of the country (Khatlon Region) (Figs 65 – 66). Genus Poecilochroa Westring, 1874 Type species: Pythonissa variana C. L. Koch, 1839 from Germany. Comments. This genus comprises 38 species and has almost worldwide distribution, being absent in North America, Northern Asia and Australia (WSC 2021). Only two species has been recorded from Central Asia.	en	Fomichev, Alexander A., Marusik, Yuri M. (2021): A survey of East Palaearctic Gnaphosidae (Araneae). 11. New data on Gnaphosidae (Araneae) from Tajikistan. Zootaxa 4966 (4): 443-457, DOI: https://doi.org/10.11646/zootaxa.4966.4.3
03C987A6FFFA641A769F71B1FE497A8C.taxon	description	Figs 22 – 29, 65 – 66.	en	Fomichev, Alexander A., Marusik, Yuri M. (2021): A survey of East Palaearctic Gnaphosidae (Araneae). 11. New data on Gnaphosidae (Araneae) from Tajikistan. Zootaxa 4966 (4): 443-457, DOI: https://doi.org/10.11646/zootaxa.4966.4.3
03C987A6FFFA641A769F71B1FE497A8C.taxon	materials_examined	Material examined. TAJIKISTAN, Khatlon Region: 1 ♂ (ZMMU), Dangara Distr, SW slope of Sanglogh (= Sanglok) Mt. Range, 38 ° 13.091 ’ N, 69 ° 14.282 ’ E, 1362 m, 30.04.2015 (Y. M. Marusik).	en	Fomichev, Alexander A., Marusik, Yuri M. (2021): A survey of East Palaearctic Gnaphosidae (Araneae). 11. New data on Gnaphosidae (Araneae) from Tajikistan. Zootaxa 4966 (4): 443-457, DOI: https://doi.org/10.11646/zootaxa.4966.4.3
03C987A6FFFA641A769F71B1FE497A8C.taxon	diagnosis	Diagnosis. The species is similar to the European species P. furcata Simon, 1914, from which it can be separated by the ventral branch of the tibial apophysis (Vb) directed forward (vs. directed downward), short prolateral fold (Pf) of conductor (Cn) and by having a projection (Pb) on the base of embolus (Eb) (cf. Figs 22 – 23, 26 and figs 3 – 4 in Di Franco (2001 )).	en	Fomichev, Alexander A., Marusik, Yuri M. (2021): A survey of East Palaearctic Gnaphosidae (Araneae). 11. New data on Gnaphosidae (Araneae) from Tajikistan. Zootaxa 4966 (4): 443-457, DOI: https://doi.org/10.11646/zootaxa.4966.4.3
03C987A6FFFA641A769F71B1FE497A8C.taxon	discussion	Comment. This species was treated in three taxonomic publications and previously known from original figures only.	en	Fomichev, Alexander A., Marusik, Yuri M. (2021): A survey of East Palaearctic Gnaphosidae (Araneae). 11. New data on Gnaphosidae (Araneae) from Tajikistan. Zootaxa 4966 (4): 443-457, DOI: https://doi.org/10.11646/zootaxa.4966.4.3
03C987A6FFFA641A769F71B1FE497A8C.taxon	description	Description. Male. Habitus as in Fig. 29. Total length 6.2. Carapace: 2.8 long, 2.05 wide. Coloration. Carapace, sternum, labium, maxillae and chelicerae brown. Legs and palps brown, legs III and IV lighter. Abdomen beigegray with brown scutum. Spinnerets gray. Leg spination: I: Fe: d 1 - 1 - 1, p 0 - 0 - 1; Ti v 0 - 0 - 2; Mt v 2 - 0 - 0. II: Fe d 1 - 1 - 1, p 0 - 1 - 1; Ti v 0 - 0 - 2; Mt v 2 - 0 - 0. III: Fe d 1 - 1 - 1, p 0 - 1 - 1, r 0 - 0 - 1; Pt r 1; Ti d 1 - 0 - 0, p 1 - 1 - 1, r 1 - 0 - 1, v 1 - 1 - 2; Mt d 0 - 1 - 0, p 0 - 1 - 0, r 1 - 0 - 0, v 0 - 2 - 0; IV: Fe d 1 - 1 - 0, p 0 - 0 - 1, r 0 - 0 - 2; Ti p 2 - 0 - 1, r 0 - 0 - 1, v 3 - 1 - 2; Mt d 0 - 2 - 0, p 2 - 1 - 0, r 2 - 1 - 0, v 3 - 2 - 0. Leg measurements. I: 6.41 (1.95, 1.15, 1.4, 1.18, 0.73); II: 6.31 (1.9, 1.1, 1.38, 1.2, 0.73); III: 5.54 (1.8, 0.93, 0.71, 1.35, 0.75); IV: 7.59 (2.1, 1.05, 1.63, 1.98, 0.83). Palp as in Figs 22 – 27, 28. Tibial apophysis splits into short, triangular dorsal branch (Db) and long, digitiform and straight ventral branch; tegulum ovate, with an indistinct constriction in the middle part; sperm duct (Sd) forming one loop; petal-shaped, membranous conductor (Cn) envelopes embolus prolaterally; embolus sharply pointed, with a wide square base (Eb). Female. Unknown.	en	Fomichev, Alexander A., Marusik, Yuri M. (2021): A survey of East Palaearctic Gnaphosidae (Araneae). 11. New data on Gnaphosidae (Araneae) from Tajikistan. Zootaxa 4966 (4): 443-457, DOI: https://doi.org/10.11646/zootaxa.4966.4.3
03C987A6FFFA641A769F71B1FE497A8C.taxon	distribution	Distribution. Uzbekistan and southwestern Tajikistan (new record).	en	Fomichev, Alexander A., Marusik, Yuri M. (2021): A survey of East Palaearctic Gnaphosidae (Araneae). 11. New data on Gnaphosidae (Araneae) from Tajikistan. Zootaxa 4966 (4): 443-457, DOI: https://doi.org/10.11646/zootaxa.4966.4.3
03C987A6FFFA641A769F71B1FE497A8C.taxon	discussion	Comments. The new combination is proposed based on the large retrolateral tibial apophysis divided into branches and short apical embolus partly entwined by membranous conductor, which is similar to those known in Poecilochroa Westring, 1874.	en	Fomichev, Alexander A., Marusik, Yuri M. (2021): A survey of East Palaearctic Gnaphosidae (Araneae). 11. New data on Gnaphosidae (Araneae) from Tajikistan. Zootaxa 4966 (4): 443-457, DOI: https://doi.org/10.11646/zootaxa.4966.4.3
03C987A6FFF86417769F7490FEC87C10.taxon	materials_examined	Type species. Melanophora subterranea C. L. Koch, 1833 from Germany.	en	Fomichev, Alexander A., Marusik, Yuri M. (2021): A survey of East Palaearctic Gnaphosidae (Araneae). 11. New data on Gnaphosidae (Araneae) from Tajikistan. Zootaxa 4966 (4): 443-457, DOI: https://doi.org/10.11646/zootaxa.4966.4.3
03C987A6FFF86417769F7490FEC87C10.taxon	discussion	Comments. With 392 valid species (WSC 2021), Zelotes is the most speciose genus of the family. Although it has never been the subject of a global revision, during the last 50 years over 200 species previously classified in this genus have been transferred to other genera, including new ones: Aponetius Kamura, 2020, Civizelotes Senglet, 2012, Heser Tuneva 2004, and Marinarozelotes Ponomarev, 2020. Still, the genus remains paraphyletic, judging by the shape of the copulatory organs. One of the distinct species groups that doesn’t depict much similarity to the generotype is the puritanus group, which accommodates at least ten species known primarily from the Palaearctic (one species has a Holarctic range). We consider the following species in this group: Z. anatolyi sp. n.; Z. gussakovskyi Charitonov, 1951; Z. khatlonicus sp. n.; Z. mikhailovi Marusik, 1995; Z. parascrutatus Levy, 1998; Z. potanini Schenkel, 1963; Z. puritanus Chamberlin, 1922; Z. scrutatus O. Pickard-Cambridge, 1872; Z. zin Levy, 1998 and possibly Z. hummeli Schenkel, 1963. This group is characterized by solid, anteriorly rounded wide epigynal pockets (= hoods), elongate epigynal fovea rounded posteriorly, blunt tibial apophysis, and embolus being covered by “ intercalary ” sclerite (= part of embolic division).	en	Fomichev, Alexander A., Marusik, Yuri M. (2021): A survey of East Palaearctic Gnaphosidae (Araneae). 11. New data on Gnaphosidae (Araneae) from Tajikistan. Zootaxa 4966 (4): 443-457, DOI: https://doi.org/10.11646/zootaxa.4966.4.3
03C987A6FFF76416769F73BAFCEE7C6B.taxon	description	Figs 30 – 31, 33 – 35, 39 – 41, 45, 47 – 49, 65 – 66.	en	Fomichev, Alexander A., Marusik, Yuri M. (2021): A survey of East Palaearctic Gnaphosidae (Araneae). 11. New data on Gnaphosidae (Araneae) from Tajikistan. Zootaxa 4966 (4): 443-457, DOI: https://doi.org/10.11646/zootaxa.4966.4.3
03C987A6FFF76416769F73BAFCEE7C6B.taxon	materials_examined	Type. TAJIKISTAN, Khatlon Region: holotype ♂ (ZMMU), Tigrovaya Balka Res., 37 ° 10.459 ’ N, 68 ° 23.047 ’ E, 316 m, tugai forest with thick litter, 6.07.2015 (Y. M. Marusik & M. Saidov). Additional material examined. TAJIKISTAN, Khatlon Region: ♀ (ZMMU), Shaartuz Distr., Khushody Vil., 37 ° 09.132 ’ N 68 ° 04.195 ’ E, 378 m, sandy desert, chiefly under shrubs, 19.04.2015 (Y. M. Marusik).	en	Fomichev, Alexander A., Marusik, Yuri M. (2021): A survey of East Palaearctic Gnaphosidae (Araneae). 11. New data on Gnaphosidae (Araneae) from Tajikistan. Zootaxa 4966 (4): 443-457, DOI: https://doi.org/10.11646/zootaxa.4966.4.3
03C987A6FFF76416769F73BAFCEE7C6B.taxon	etymology	Etymology. The specific epithet is derived from Khatlon Region.	en	Fomichev, Alexander A., Marusik, Yuri M. (2021): A survey of East Palaearctic Gnaphosidae (Araneae). 11. New data on Gnaphosidae (Araneae) from Tajikistan. Zootaxa 4966 (4): 443-457, DOI: https://doi.org/10.11646/zootaxa.4966.4.3
03C987A6FFF76416769F73BAFCEE7C6B.taxon	diagnosis	Diagnosis. The male of the new species can be diagnosed from all other Central Asian species of puritanus group by large circular intercalary sclerite (Is) (vs. triangular, square, ovate or oblong) (cf. Figs 34 – 40 and 54, 57, 60, 63). The female of Zelotes khatlonicus sp. n. differs from those of Z. mikhailovi Marusik, 1995 by converging anterior parts of lateral margins (Lm) (vs. parallel or divergent). Also, from Z. potanini Schenkel, 1963 it can be distinguished by () – shaped fovea (Fo) and lateral margins twice longer than receptacles (Re) (vs. bowl-shaped fovea and lateral margins as long as receptacles). From Z. puritanus Chamberlin, 1922 it can be distinguished by () – shaped epigynal fovea and the anterior epigynal hood (Ah) being twice wider than receptacle (vs. bowl-shaped fovea and the anterior hood as wide as receptacle) (cf. Figs 47 – 48 and 50 – 53).	en	Fomichev, Alexander A., Marusik, Yuri M. (2021): A survey of East Palaearctic Gnaphosidae (Araneae). 11. New data on Gnaphosidae (Araneae) from Tajikistan. Zootaxa 4966 (4): 443-457, DOI: https://doi.org/10.11646/zootaxa.4966.4.3
03C987A6FFF76416769F73BAFCEE7C6B.taxon	description	Description. Male. Habitus as in Fig. 31. Total length 4.25. Carapace 2.1 long, 1.65 wide. Coloration. Carapace brown with dark gray edges. Chelicerae brown. Maxillae and labium yellow-brown. Sternum yellowish orange. Palps yellowish gray. Legs: I – III — Fe, Pt, Ti — gray with yellow spots, Mt, Ta — yellow; IV — Fe, Pt, Ti, Mt — gray, Ta — yellow. Coxae yellow. Abdomen dark gray, almost black dorsally, yellow-gray ventrally. Scutum brown-gray. Spinnerets dark gray, almost black. Leg spination: I: Fe d 1 - 1 - 0, p 0 - 0 - 1; II: Fe d 1 - 1 - 0, p 0 - 0 - 1; III: Fe d 1 - 1 - 0, p 0 - 1 - 1, r 0 - 1 - 1; Pt r 1; Ti p 1 - 1 - 1, r 0 - 1 - 1, v 2 - 2 - 2; Mt d 0 - 1 - 0, p 1 - 1 - 0, r 1 - 1 - 0, v 2 - 0 - 0. IV: Fe d 1 - 1 - 0, p 0 - 1 - 1, r 0 - 0 - 1; Ti p 1 - 0 - 1, r 1 - 1 - 1, v 2 - 2 - 2; Mt d 0 - 2 - 0, p 1 - 1 - 0, r 1 - 1 - 0, v 2 - 2 - 0. Leg measurements. I: 5.79 (1.6, 0.98, 1.25, 1.08, 0.88); II: 5.04 (1.38, 0.83, 1.05, 0.95, 0.83); III: 4.16 (1.18, 0.65, 0.78, 0.9, 0.65); IV: 6.34 (1.73, 0.93, 1.35, 1.5, 0.83). Palp as in Figs 33 – 35, 39 – 41, 45. Tibial apophysis straight, broad with blunt tip, as long as tibia, and as wide as ½ of tibia; bulb almost 2 times (1.8) longer than wide; intercalary sclerite (Is) circular; terminal apophysis (Ta) triangular, as wide as intercalary sclerite; embolar projection (Ep) pointed, as thick as terminal apophysis; median apophysis (Ma) tooth-like, directed upwards; embolus (Em) long, almost as longs as width of bulb, almost straight, with anteriorly bent tip. Female. Habitus as in Fig. 30. Total length 6.2. Carapace: 2.95 long, 2.3 wide. Carapace brown with dark gray edges. Chelicerae brown. Labium and maxillae light brown. Sternum yellow-orange. Palps yellow. Legs: I, II, IV — Fe, Pt, Ti — yellow-gray, Mt, Ta — yellow; III — Fe, Pt — yellow-gray, Ti, Mt, Ta — yellow. Coxae light yellow. Abdomen dark gray, almost black dorsally, yellow-gray ventrally. Spinnerets dark gray, almost black. Leg spination: I: Fe d 1 - 1 - 0, p 0 - 0 - 1. II: Fe d 1 - 1 - 0, p 0 - 0 - 1. III: Fe d 1 - 1 - 0, p 1 - 0 - 1, r 0 - 1 - 1; Pt r 1; Ti p 2 - 0 - 1, r 0 - 1 - 1, v 2 - 2 - 2; Mt d 0 - 1 - 0, p 1 - 1 - 0, r 1 - 1 - 0, v 2 - 2 - 0. IV: Fe d 1 - 1 - 0, p 1 - 0 - 1, r 0 - 0 - 1; Ti p 1 - 0 - 1, r 2 - 1 - 1, v 2 - 2 - 2; Mt d 0 - 2 - 0, p 1 - 1 - 0, r 1 - 1 - 0, v 2 - 0 - 2. Leg measurements. I: 6.97 (2.05, 1.28, 1.48, 1.18, 0.98); II: 6.25 (1.78, 1.13, 1.28, 1.13, 0.93); III: 5.51 (1.58, 0.88, 1.0, 1.2, 0.85); IV: 8.03 (2.25, 1.23, 1.65, 1.9, 1.0). Epigyne as in Figs 47 – 49. Epigyne twice longer than wide; fovea elongated oval, twice longer than wide; anterior hood 1.3 times wider than fovea; receptacles large (diameter equal to fovea’s width), closely spaced; copulatory ducts (Cd) thin (in ventral view), forming 3 loops and bear hemispherical glands (Gl) located on a halfway between hood and posterior edge of fovea.	en	Fomichev, Alexander A., Marusik, Yuri M. (2021): A survey of East Palaearctic Gnaphosidae (Araneae). 11. New data on Gnaphosidae (Araneae) from Tajikistan. Zootaxa 4966 (4): 443-457, DOI: https://doi.org/10.11646/zootaxa.4966.4.3
03C987A6FFF76416769F73BAFCEE7C6B.taxon	distribution	Distribution. Known only from the type locality.	en	Fomichev, Alexander A., Marusik, Yuri M. (2021): A survey of East Palaearctic Gnaphosidae (Araneae). 11. New data on Gnaphosidae (Araneae) from Tajikistan. Zootaxa 4966 (4): 443-457, DOI: https://doi.org/10.11646/zootaxa.4966.4.3
03C987A6FFF76416769F73BAFCEE7C6B.taxon	discussion	Comments. The holotype male and the female were collected in different localities. Distance between Tigrovaya Balka Reserve and Khushody Village is about 30 – 40 km, which is relatively close. Moreover, both specimens belong to puritanus species group and have similar habitus. For these reasons, we considered them as conspecific; this hypothesis shall be confirmed or rejected in future when both sexes are collected together. Another species, Z. anatolyi sp. n., which also belongs to the puritanus group and represented by the holotype male has bright colouration, very different from those in Z. khatlonicus sp. n.	en	Fomichev, Alexander A., Marusik, Yuri M. (2021): A survey of East Palaearctic Gnaphosidae (Araneae). 11. New data on Gnaphosidae (Araneae) from Tajikistan. Zootaxa 4966 (4): 443-457, DOI: https://doi.org/10.11646/zootaxa.4966.4.3
03C987A6FFF36412769F729BFE4B7C13.taxon	description	Figs 32, 36 – 38, 42 – 44, 46, 65 – 66.	en	Fomichev, Alexander A., Marusik, Yuri M. (2021): A survey of East Palaearctic Gnaphosidae (Araneae). 11. New data on Gnaphosidae (Araneae) from Tajikistan. Zootaxa 4966 (4): 443-457, DOI: https://doi.org/10.11646/zootaxa.4966.4.3
03C987A6FFF36412769F729BFE4B7C13.taxon	materials_examined	Type. TAJIKISTAN, Khatlon Region: holotype ♂ (ISEA, 001.8556), E foothill of Aruktau Mt. Range, between Khumdon and Sambuli Villages, 37 ° 50.210 ’ N, 68 ° 35.872 ’ E, stony steppe, 420 m, 05.04.2019 (A. A. Fomichev).	en	Fomichev, Alexander A., Marusik, Yuri M. (2021): A survey of East Palaearctic Gnaphosidae (Araneae). 11. New data on Gnaphosidae (Araneae) from Tajikistan. Zootaxa 4966 (4): 443-457, DOI: https://doi.org/10.11646/zootaxa.4966.4.3
03C987A6FFF36412769F729BFE4B7C13.taxon	etymology	Etymology. The specific epithet is dedicated to Anatoly A. Fomichev (Novosibirsk, Russia), the father of the first author.	en	Fomichev, Alexander A., Marusik, Yuri M. (2021): A survey of East Palaearctic Gnaphosidae (Araneae). 11. New data on Gnaphosidae (Araneae) from Tajikistan. Zootaxa 4966 (4): 443-457, DOI: https://doi.org/10.11646/zootaxa.4966.4.3
03C987A6FFF36412769F729BFE4B7C13.taxon	diagnosis	Diagnosis. The new species belongs to puritanus species group. Zelotes anatolyi sp. n. differs from all Central Asian species of this group, except for Z. gussakovskyi, by square-shaped intercalary sclerite (Is) (vs. triangular, circular, ovate or oblong). From Z. gussakovskyi it can be distinguished by ovate terminal apophysis (Ta) (vs. triangular) (cf. Figs 37, 43 and 54, 57, 60, 63).	en	Fomichev, Alexander A., Marusik, Yuri M. (2021): A survey of East Palaearctic Gnaphosidae (Araneae). 11. New data on Gnaphosidae (Araneae) from Tajikistan. Zootaxa 4966 (4): 443-457, DOI: https://doi.org/10.11646/zootaxa.4966.4.3
03C987A6FFF36412769F729BFE4B7C13.taxon	description	Description. Male. Habitus as in Fig. 32. Total length 5.5. Carapace 2.33 long, 1.75 wide. Coloration. Chelicerae, labium, maxillae and sternum orange. Carapace orange with dark gray edges. Palps yellowish gray. Legs: I, II — Fe, Pt, Ti — gray, Mt, Ta — yellow; III — Fe, Pt — gray, Ti, Mt, Ta — yellow; IV — Fe, Pt, Ti, Mt — gray, Ta — yellow. Coxae yellowish gray. Abdomen dark gray dorsally, yellowish gray ventrally. Scutum brownish gray. Spinnerets dark gray. Leg spination: I: Fe d 1 - 1 - 0, p 0 - 0 - 1. II: Fe d 1 - 1 - 0, p 0 - 0 - 1. III: Fe d 1 - 1 - 0, p 0 - 1 - 1, r 0 - 1 - 1; Pt r 1; Ti p 1 - 1 - 1, r 0 - 1 - 1, v 2 - 2 - 2; Mt d 0 - 1 - 0, p 1 - 1 - 0, r 1 - 1 - 0, v 2 - 0 - 0; IV: Fe d 1 - 1 - 0, r 0 - 0 - 1; Ti p 1 - 1 - 0, r 1 - 1 - 1, v 2 - 2 - 2; Mt d 0 - 1 - 0, p 0 - 1 - 0, r 1 - 1 - 0, v 2 - 0 - 2. Leg measurements. I: 6.33 (1.75, 1.05, 1.35, 1.2, 0.98); II: 5.62 (1.55, 0.93, 1.13, 1.08, 0.93); III: 4.86 (1.35, 0.73, 0.88, 1.05, 0.85); IV: 7.21 (1.93, 1.03, 1.45, 1.75, 1.05). Palp as in Figs 36 – 38, 42 – 44, 46. Tibial apophysis straight and pointed, as long as tibia, about 3 times longer than wide; intercalary sclerite square-shaped; terminal apophysis lanceolate, as long as intercalary sclerite; embolar projection blunt, twice thicker than terminal apophysis, bearing a small tooth (Sm); median apophysis claw-like, directed upwards; embolus straight, shifted retrolaterally. Female. Unknown.	en	Fomichev, Alexander A., Marusik, Yuri M. (2021): A survey of East Palaearctic Gnaphosidae (Araneae). 11. New data on Gnaphosidae (Araneae) from Tajikistan. Zootaxa 4966 (4): 443-457, DOI: https://doi.org/10.11646/zootaxa.4966.4.3
03C987A6FFF36412769F729BFE4B7C13.taxon	distribution	Distribution. Known only from the type locality.	en	Fomichev, Alexander A., Marusik, Yuri M. (2021): A survey of East Palaearctic Gnaphosidae (Araneae). 11. New data on Gnaphosidae (Araneae) from Tajikistan. Zootaxa 4966 (4): 443-457, DOI: https://doi.org/10.11646/zootaxa.4966.4.3
03C987A6FFF36412769F729BFE4B7C13.taxon	discussion	Comments. Zelotes anatolyi sp. n. has an unusually bright coloration: orange carapace, chelicerae and sternum and a partially yellowish legs.	en	Fomichev, Alexander A., Marusik, Yuri M. (2021): A survey of East Palaearctic Gnaphosidae (Araneae). 11. New data on Gnaphosidae (Araneae) from Tajikistan. Zootaxa 4966 (4): 443-457, DOI: https://doi.org/10.11646/zootaxa.4966.4.3
