taxonID	type	description	language	source
B43EBA76FF90EE2049AE4ACDFCE0F94E.taxon	materials_examined	Material examined. PORTUGAL: ALGARVE: 1 ♀: Serra Monchique E Monchique, 37 ° 19 ’ N 8 ° 31 ’ W. 530 m a. s. l. 10.04.2002. H. Meybohm leg. (cA); ESTREMADURA: 4 ♂♂, 1 ♀: Lisboa, Serra do Sintra. 24.05.1992. P. Wunderle leg. (cW); SPAIN: PAIS VASCO: 1 ♀: Irun. 10.02.2005. A. Anichtchenko leg. (cSh); 1 ♀: same data. 17 - 19.03.2005 (cSh); 1 ♂, 3 ♀♀: same data. 09.04.2005 (cSh); 1 ♂, 2 ♀♀: same data. 05.2006 (cSh); 1 ♀: same data. 15.06.2006 (cSh); ARAGÓN: 2 ♂♂: Barbastro, Ariño de Cregenzán. 09.04.1995. D. A. Lott leg. (BMNH); 1 ♀: Villanueva de Sigena. 09.04.1995. D. A. Lott leg. (BMNH); 1 ♂: Guadalavier. 19.05.1994. D. A. Lott leg. (BMNH); 1 ♂: Estrella (BMNH); CASTILLA Y LÉON: 2 ♂♂: Sierra de Gredos, 10 km SSW Burgohondo, 40 ° 21 ’ 15 N 04 ° 49 ’ 10 W. 1250 m a. s. l. 25.03.2007. V. Assing leg. (cA); 2 ♂♂, 1 ♀: Jarandilla, Caseres. 16.06.1969. A. Comellini leg. (MHNG); 1 ♂: Sierra de la Demanda. 2.8.1971. (MHNG); EXTREMADURA: 2 ♂♂: Jarandilla. 29.06.1969. A. Comellini leg. (MHNG); CASTILLA LA MANCHA: 1 ♂: Daimiel. 14.08.1969. A. Comellini leg. (MHNG); CATALUñA: 1 ♂: Santa Perpètua de Gaiá. 08.04.1995. D. A. Lott leg. (BMNH); VALENCIA: 1 ♂: El Toro. 950 m a. s. l. 07.04.1995. D. A. Lott leg. (BMNH); ANDALUSÍA: 1 ♂: Sierra Magina, 37.72 ° N - 3.46 ° W. 2000 m a. s. l. 03 - 04.05.2014. A. V. Shavrin leg. (cSch); 3 ♂♂, 2 ♀♀: Sierra Nevada, Capileira, Rio Nante, 36.99 ° N - 3.35 ° W. 1550 m a. s. l. 08.05.2014. A. V. Shavrin leg. (cSh); 3 ♂♂, 4 ♀♀: Capileira. 1600 m a. s. l. 22.03.1994. V. Assing leg. (cA); 1 ♀: Sierra Nevada, Puerto de la Ragua, 37.09 ° N - 3.02 ° W. 2000 m a. s. l. 08 - 09.05.2014. A. V. Shavrin leg. (cSch); 3 ♂♂, 3 ♀♀: Sierra de Bermeja, Jubrique. 500 m a. s. l. 26.03.1994. V. Assing leg. (cA); 1 ♂, 1 ♀: Sierra Nevada, Guejar Sierra. 1200 m a. s. l. 28.09.1993. P. Wunderle leg. (cW); 1 ♀: San Pedro, Ronda. 14 - 21.03.1990. Mattern leg. (cW); 4 ♂♂: Alpujarra Capileira. 1900 m a. s. l. 22.03.1994. P. Wunderle leg. (cW); 2 ♂♂, 1 ♀: Málaga, Jubrique. 500 m a. s. l. 26.03.1994. P. Wunderle leg. (cW); 1 ♂, 2 ♀♀: Sierra de Cazorla. 1400 m a. s. l. 06.10.1993. P. Wunderle leg. (cW); 1 ♂: Esteperona (MHNG); 1 ♂, 1 ♀: Castille Puerto de Bejar. 800 - 1400 m a. s. l. 05 - 06.1957. G. Fagel leg. (MHNG); 1 ♂: Puerto de la Ragua, Granada E. 1200 m a. s. l. 17.07.1971. A. Comellini leg. <Lesteva fontinalis infrasp. gustavi Det. Herman 2001> (MHNG). Additional material. FRANCE: 1 ♂: ‘ Htes Pyrénées’ (IRSN); MOROCCO: 1 ♂, 1 ♀: Grand Atlas Mts., Oukaimeden env., 31 ° 13 ’ 09.5 ’’ N - 0.7 ° 49 ’ 29.7 ’’ W. 2200 - 2300 m a. s. l. 16 - 17.06.2010. A. Anichtchenko leg. (cSh); 2 ♂♂: Alrene, 55 km S Marrakesh. 5000 ft. 26. iv. 1961. P. N. Lawrence leg. (BMNH); LOCALITY NOT SPECIFIED: 1 ♂: ‘ Fauvel’, ‘ Pyrenäen’ (SF); 1 ♂, 1 ♀: ‘ Pyren. ’, ‘ Fauvel’, ‘ heeri ’ (SMNH).	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF90EE2049AE4ACDFCE0F94E.taxon	discussion	Remarks. Based on a study of material from several collections, L. sicula heeri was recorded by Lohse (1960) from the Pyrenees and central Spain. Horion (1963) recorded it from southern Spain (“ Sierra Nevada, Kiesenwetter leg. Ca. 1850, 6 Ex. Z. S. M. ”). Some studied specimens from southern Spain and Morocco have a paler body, welldeveloped hind wings and relatively wide elytra as in L. sicula sicula Erichson, 1840, a subspecies distributed in Italy and Algeria. Lohse (1960) also noted this morphological similarity for specimens from Guadalupe, South Spain. This is probably morphological variability (from apterous to macropterous forms) of the species at its southern border of the distribution. The aedeagi of both subspecies have similar internal and external structures. A genetic analysis would be required for a clarification of the status and possible synonymy of both taxa. The species is here recorded from Morocco for the first time.	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF90EE2749AE4D1CFDA5FB4D.taxon	description	(Figs. 1, 3 – 5, 8)	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF90EE2749AE4D1CFDA5FB4D.taxon	materials_examined	Type material examined. Paratypes: 1 ♂: ‘ Asturies: Covadonga | (Cangas de Onis) | rio Covadonga | VI. 1965 G. Fagel’ <printed>, ‘ Paratypoid’ <red, handwritten>, ‘ Coll. | G. A. Lohse | MHNG- 1994 ’ <printed>, ‘ PARATYPE | Lesteva | aterrima sp. n. | Det. Lohse 1967 ’ <yellow, printed>, ‘ MHNG | ENTO | 00087306 ’ <printed>, ‘ Lesteva (s. str.) | aterrima Lohse, 1967 | Shavrin A. V. det. 2021 ’ (MHNG); 1 ♀: same labels, but ‘ MHNG | ENTO | 00087307 ’ <printed> (MHNG). Material examined. SPAIN: ASTURIAS: 10 ♂, 8 ♀♀: Puerto de La Cubilla, 1400 m a. s. l. 08.06.1991. P. Wunderle leg. (cA, cW); 3 ♂♂, 1 ♀: same data, but L. Zerche leg. (cW, MHNG); 1 ♂, 1 ♀: Oviedo, Picos de Europa, Covadonga Lago Enol. 1600 m a. s. l. 03.06.1990 (cZan); 1 ♂: Covadonga, Hutte Vega Redonga. 1300 m a. s. l. 07.06.1991. L. Zerche leg. (MHNG).	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF90EE2749AE4D1CFDA5FB4D.taxon	description	Redescription. Measurements (n = 25): HW: 0.68 – 0.75; HL: 0.47 – 0.52; OL: 0.24 – 0.25; LT: 0.07 – 0.08; AL (holotype): 2.07; PL: 0.75 – 0.78; PWMax: 0.83 – 0.87; PWMin: 0.60 – 0.65; EL: 1.58 – 1.62; ESL: 1.49 – 1.52; EW: 1.39 – 1.42; MTbL (holotype): 1.02; MTrL (holotype): 0.57; AW: 1.35 – 0.38; AedL: 0.67 – 0.82; FL: 2.85 – 2.96; TL: 3.50 – 4.55 (holotype: 4.45). Body dark brown, sometimes with paler apical abdominal tergites; antennes and legs yellow-brown; mouthparts and tarsi yellow. Punctation of head very dense, coarse and moderately deep, interspaces between punctures in middle about as long as diameter of one nearest puncture, finer in vertex and finer and denser around eyes; punctation of neck as that of middle portion of head; punctation of pronotum moderately regular, about as that on middle part of head, but slightly deeper, slightly sparser in mediobasal third; scutellum with indistinct, very fine and sparse punctation; punctation of elytra similar to that on pronotum, but sparser, slightly finer and denser in parascutellar portion and along suture; abdominal tergites with very fine and dense punctation. Head and neck with very dense isodiametric microsculpture, distinctly coarser in infraorbital ridges; pronotum with fine, transverse microreticulation except for mediobasal portion; scutellum with very dense, fine meshes; abdominal tergites with distinct transverse microsculpture. Habitus as in Fig. 1. Head transverse, 1.4 times as broad as long, distinctly and widely elevated in middle; middle portion separated from lateral portions by wide and deep longitudinal depressions; eyes moderately large, convex, about three times as long as temples (lateral view); distance between ocelli markedly shorter than distance between ocellus and posterior margin of eye. Antennae reaching basal third of elytra when reclined, with moderately elongate antennomeres; basal antennomere disitnctly more than twice as long as wide, 2 slightly narrower than basal antennomere, 3 slightly longer than 2, 4 slightly broader and longer than 3, 5 – 7 slightly longer and broader than 4, 8 – 10 slightly shorter than 7, apical antennomere slightly broader than 10. Pronotum evenly convex, 1.1 times as broad as long, 1.1 – 1.2 times as broad as head, very wide in apical third, sharply narrowed toward elongate subparallel latero-basal margins; middle portion of pronotum with indistinct, oval and moderately wide paired impressions and without or with indistinct to distinct and deep, transverse impression in mediobasal third. Elytra somewhat flattened, transverse, longer than broad, slightly less than twice as long as pronotum, significantly widened apicad. Male. Apical margins of abdominal tergite VIII slightly and sternite VIII concave. Aedeagus with very wide basal portion, significantly narrowed toward very narrow median lobe, with small rounded apex; parameres wide, exceeding apex of median lobe, widened apically, with two long apical and preapical setae; lateral sides of median lobe with several moderately long setae; internal sac without visible sclerotized structures (Fig. 3). Lateral aspect of aedeagus as in Figs. 4 – 5. Female. Apical margins of abdominal tergite VIII and sternite VIII rounded. Comparative notes. Regarding the general shape of the aedeagus and the presence of a deep concavity between the basal portion, median lobe and the narrow, curved ventral process, L. aterrima is similar to L. benicki Lohse, 1958, a species distributed in the Alps, and L. martinae Zanetti, 2008, a species described from central Italy (Zanetti 2008). Lesteva aterrima differs from these species by the smaller size of the aedeagus, a narrower apical part of the median lobe, and different features of its setation (see Fig. 4 in Zanetti (2008 )). Based on the general shape of the body, all three species are similar to L. pubescens, from which they are distinguished by the morphology of the aedeagus.	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF90EE2749AE4D1CFDA5FB4D.taxon	distribution	Distribution. Lesteva aterrima is known from several localities in Cantabrian mountain range, in north-western Spain (Fig. 8). Natural history. Specimens were collected at elevations from 1300 to 1600 m a. s. l. Some specimens were sifted from wet mosses near streams.	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF97EE2449AE4F1EFCF5F855.taxon	description	(Figs. 2, 6 – 7)	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF97EE2449AE4F1EFCF5F855.taxon	materials_examined	Type material examined. Holotype, ♂ [aedeagus glued on the same card under the specimen]: ‘ Mallorca | Balearen’ <handwritten>, ‘ Lesteva [handwritten] | balearica n. sp. [handwritten] | det. Dr. G. A. Lohse’ <printed>, ‘ Holotypus’ <red, handwritten>, ‘ Coll. | G. A. Lohse | MHNG- 1994 ’ <printed>, ‘ HOLOTYPE | Lesteva | balearica sp. n. | Det. Lohse 1967 ’ <red, printed>, ‘ MHNG | ENTO | 00087308 <printed>, ‘ Lesteva (s. str.) | balearica Lohse, 1967 | Shavrin A. V. det. 2021 ’ <printed> (MHNG). Paratypes: 1 ♂ [aedeagus glued on the same card under the specimen]: same labels as the holotype except for: ‘ Para- | typoid’ <red, handwritten>, ‘ PARATYPE | Lesteva | balearica sp. n. | Det. Lohse 1967 ’ <yellow, printed>, ‘ MHNG | ENTO | 00087309 ’ <printed> (MHNG); 1 ♂, 2 ♀♀ [one specimen without head and pronotum]: ‘ Mallorca | IV 65 | Benick’ <two labels, handwritten>, ‘ balearica | Lohse | Paratyp’ <two labels, handwritten>, ‘ Coll. | G. A. Lohse | MHNG- 1994 ’ <printed>, ‘ PARATYPE | Lesteva | balearica sp. n. | Det. Lohse 1967 ’ <three labels, yellow, printed>, ‘ MHNG | ENTO | 00087310 ’ <printed>, ‘ MHNG | ENTO | 00087311 ’ <printed>, ‘ MHNG | ENTO | 00087312 ’ <printed> (MHNG); 3 ♂♂ [dissected, preparation with the aedeagus in Canada balsam on plastic card are pinned under related specimens]: same labels as above, except for: ‘ MHNG | ENTO | 00087313 ’ <printed>, ‘ MHNG | ENTO | 00087314 ’ <printed>, ‘ MHNG | ENTO | 00087315 ’ <printed> (MHNG); 1 ♂ [without right antennomeres 3 - 11], 1 ♀ [without left antennomeres 3 - 11]: same labels, but: ‘ MHNG | ENTO | 00087316 ’ <printed>, ‘ MHNG | ENTO | 00087317 ’ <printed> (MHNG); 1 ♂, 2 ♀♀: ‘ Mallorca | IV. 65 Benick’ <handwritten>, ‘ Lesteva | balearica Lohse | Paratyp. ’ <handwritten>, ‘ PARATYPE | Lesteva | balearica sp. n. | Det. Lohse 1967 ’ <yellow, printed>, ‘ MHNG | ENTO | 00087318 ’ <printed>, ‘ MHNG | ENTO | 00087319 ’ <printed>, ‘ MHNG | ENTO | 00087320 ’ <printed> (MHNG); 2 ♀♀ (one specimen without left antennomeres 3 - 11): same labels, but: ‘ MHNG | ENTO | 00087321 ’ <printed>, ‘ MHNG | ENTO | 00087322 ’ <printed> (MHNG); 2 ♀♀: same data as the previous except for: ‘ MHNG | ENTO | 00087323 ’ <printed>, ‘ MHNG | ENTO | 00087324 ’ <printed> (MHNG); 1 ♂ [dissected]: ‘ Mallorca | Balearen | IV 1965 G. Benick’ <handwritten>, ‘ Lesteva | balearica n. sp. | det. G. A. Lohse’ <printed>, ‘ Paratypus’ <red, printed> (cZan). All paratypes with additional printed labels: ‘ Lesteva (s. str.) | balearica Lohse, 1967 | Shavrin A. V. det. 2021 ’. Material examined. SPAIN: Balearic islands: 1 ♀: Mallorca, Gorg Blau, Casa Nova, 39 ° 49 ‘ N 02 ° 50 E. 14.05.1978. H. Malicky leg. (cA).	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF97EE2449AE4F1EFCF5F855.taxon	description	Redescription. Measurements (n = 15): HW: 0.68 – 0.77; HL: 0.49 – 0.52; OL: 0.25 – 0.27; LT: 0.10; AL (holotype): 2.05; PL: 0.73 – 0.80; PWMax: 0.81 – 0.87; PWMin: 0.60 – 0.67; EL: 1.51 – 1.62; ESL: 1.37 – 1.50; EW: 1.35 – 1.42; MTbL (holotype): 0.97; MTrL (holotype): 0.55; AW: 1.21 – 1.35; AedL: 0.65 – 0.85; FL: 2.82 – 3.06; TL: 3.50 – 4.60 (holotype: 4.45). Body yellow-brown to reddish-brown, usually with darker head and large, elongated, yellow to yellow-brown spot on apical half of each elytron, and paler apical abdominal segments; mouthparts, antennomeres and legs yellow to yellow-brown, usually with protibia somewhat darker. Punctation of head moderately dense and fine, sparser in middle and between ocelli, and distinctly denser around eyes; punctation of neck very sparse in middle, finer and coarser on lateral portions; punctation of pronotum about as than on middle portion of head, but slightly sparser, particularly in mediobasal third; scutellum with very fine, irregular punctation; punctation of elytra similar to that on pronotum, but sparser; abdominal tergites with regular, distinct and fine punctation, markedly finer on abdominal tergite VII. Latero-apical portions of head between supra-antennal prominces and apical margins of eyes with coarse and very dense sculpture; mediobasal portion of neck with transverse microreticulation, lateral portions of neck with moderately coarse transverse meshes; some specimens with fine, transverse microsculpture in medioapical third; abdominal tergites with distinct isodiametric sculpture. Habitus as in Fig. 2. Head 1.3 – 1.4 times as broad as long, distinctly elevated in middle; middle portion separated from lateral portions by very deep longitudinal depressions; eyes more than twice as long as temples; distance between ocelli slightly shorter than distance between ocellus and posterior margin of eye. Antennae reaching basal third of elytra when reclined, with moderately elongate antennomeres; basal antennomere more than three times as long as wide, 2 markedly narrower than basal antennomere, 3 slightly longer and narrower than 2, 4 slightly shorter and broader than 3, 5 – 8 slightly broader and longer than 4, 9 slightly broader than 8, 10 slightly shorter than 9. Pronotum slightly broader than long, 1.1 times as broad as head, from their widest anterior third sharply rounded toward elongate, subparallel latero-basal margins; dorsal surface usually without impressions, but some specimens with very indistinct, long, paired impressions in middle and / or transverse impression in mediobasal third. Elytra about as long as broad, 1.8 times as long as pronotum, slightly widened toward hind angles. Abdominal tergites IV and V with paired, transverse and moderately wide tomentose spots. Male. Apical margin of abdominal tergite VIII rounded or slightly concave. Apical margin of abdominal sternite VIII widely and deeply concave. Aedeagus with wide basal part, narrowed toward middle of median lobe, gradually widened toward apical third, with rounded apex; parameres moderately narrow, slightly widened apically, not reaching apex of median lobe, with four moderately short apical setae; internal sac wide and moderately short (Fig. 6). Lateral aspect of aedeagus as in Fig. 7. Female. Apical margin of abdominal tergite VIII straight or rounded. Apical margin of abdominal sternite VIII rounded. Comparative notes. Based on the size and the general shape of the body and antennomeres, length of temples and features of the punctation of the elytra, L. (s. str.) balearica is similar to European L. (s. str.) pubescens, from which it can be distinguished by the paler coloration of the body and the morphology of the aedeagus, without pubescence on the median lobe. Regarding the general shape of the aedeagus, it is somewhat similar to L. (s. str.) fontinalis and the Corsican L. (s. str.) praeses Fauvel, 1900 (Fig. 98 e in Zanetti (1987 )), from which it differs by the paler coloration, longer temples and the shape of the apical part of the median lobe.	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF97EE2449AE4F1EFCF5F855.taxon	distribution	Distribution. Lesteva balearica is an endemic species of Mallorca (Majorca), the largest of the Balearic islands. Natural history. The detailed ecological data are unknown.	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF95EE2B49AE4F45FA8AFADE.taxon	description	(Figs. 9, 14 – 15)	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF95EE2B49AE4F45FA8AFADE.taxon	materials_examined	Type material examined. Lectotype (here designated) of Lesteva fontinalis Kiesenwetter, 1850 ♂ [The specimen is glued sideways on small triangular card; head, pronotum and left legs are missing]: ‘ Kiesenwetter’ <printed>, ‘ fontinalis | M [ons]. Serr [atus Cataloniae]. ’ <handwritten>, ‘ Sammlung | Cl. Müller’ <printed>, ‘ Holotypus | (vermutlich!) [handwritten] | Lesteva [handwritten] | fontinalis Kiesw. [handwritten] | Staatssamml. München’ <red, printed>, ‘ zweifellos die Type, | da nur dies eine Ex. | von mon serrat | St .. [… illegible …] ’ <handwritten>, ‘ Zool. Staatsslg. | München’ <blue, printed>, ‘ LECTOTYPE | Lesteva fontinalis | Kiesenwetter, 1850 | Shavrin A. V. des. 2021 ’ <red, printed> (ZSM). Paratype of Lesteva fontinallis truncata Lohse, 1960 ♂ [aedeagus glued under the specimen]: ‘ Espagne: Estremadura | Guadalupe (scierie) | rio Guadalupejo | V — 1958, G. Fagel’ <printed>, ‘ Lesteva [handwritten] | fontinalis [handwritten] | ssp truncata [handwritten] | det. Dr. G. A. Lohse’ <printed>, ‘ Para- | type’ <red, printed>, ‘ Coll. | G. A. Lohse | MHNG- 1994 ’ <printed>, ‘ PARATYPE | Lesteva | fontinalis | truncata ssp. n. | Det Lohse 1960 ’ <yellow, printed>, ‘ PARATYPE | Lesteva fontinalis | gustavi nom. n. | Det. Herman 2001 ’ <printed>, ‘ MHNG | ENTO | 00087335 ’ <printed> (MHNG). Paratype ♀, same labels except for ‘ MHNG | ENTO | 00087336 ’ (MHNG). Paratype ♂: ‘ Espagne: Estremadura | Guadalupe | Fuente del Trincho | V — 1958, G. Fagel’ <printed>, ‘ fontinalis [handwritten] | ssp. truncata [hadwritten] | det. Dr. G. A. Lohse’ <printed>, ‘ Coll. | G. A. Lohse | MHNG- 1994 ’ <printed>, ‘ Lesteva fontinalis | infrasp. gustavi nom. n. | Det. Herman 2001 ’ <printed>, ‘ MHNG | ENTO | 00087337 ’ <printed> (MHNG). All paratypes with additional printed label: ‘ Lesteva (str.) fontinalis Kiesenwetter, 1850 | Shavrin A. V. det. 2021 ’. Material examined. SPAIN: ASTURIAS: 2 ♂♂: Puerto Pajares. A. Kricheldorff leg. (NMPC); PAIS VASCO: 1 ♂: Bilbao (MHNG); CATALUñA: 1 ♂: Col d`Ares (Spain Pyren.). 31.05.1971. G. Lohse leg. (MHNG); 2 ♂♂: Barcelona, Monserrat. 22.06.2006. L. Fancello leg. (cZan); 1 ♂: Sierra de Montseny. 13.05.1981. P. Zwick leg. (MHNG); 2 ♂♂, 2 ♀♀: Seo de Urgel (Arfa), de la Coma. 05 - 06.1962. G. Fagel leg. (MHNG); 1 ♀: Seo de Urgel (graviers du Segre). 05 - 06.1962. G. Fagel leg. (MHNG); ARAGÓN: 1 ♂: Ardisa (MHNG); 3 ♂♂, 2 ♀♀: Ordesa, Huesca. 31.05.1965. A. Comellini leg. (MHNG); 12 ♂♂, 2 ♀♀: Teruel, Albarracin. 1200 m a. s. l. 23.05.1993. A. Matern (cA, cW, cSh); 1 ♂, 1 ♀: Mezquita de Jarque. 18.06.1970. A. Comellini leg. (MHNG); CASTILLA Y LÉON: 5 ♂♂, 5 ♀♀: Sierra de la Demanda. H. Meybohm leg. (MHNG); 1 ♂: Santa Colomba de Somoza, Astorga. 31.05.1970. A. Comellini leg. (MHNG); MADRID: 1 ♂: Escorial (MHNG); CASTILLA LA MANCHA: 3 ♂♂: Las Torcas. 1200 m a. s. l. 23.05.1962. H. Freude leg. (MHNG, ZSM); ANDALUSÍA: 3 ♂♂: Sierra de Cazorla, sources of Guadalquivir. 25.06.1975. M. Daccordi leg. (cZan); 1 ♀: Sierra de Cazorla. 23.06.1975. M. Daccordi leg. (cZan); 2 ♂♂: same locality. 27.05.2009. T. Struyve leg. (cZan); 1 ♀: Sierra de Segura, Guadalquivir, 37 ° 50 ’ N - 2 ° 59 ’ W. 2276 ft. 21.05.2002. Starke leg. (cA); 4 ♂♂, 2 ♀♀: Sierra Magina, 37 ° 72 ’ N - 3 ° 46 ’ W. 2000 m a. s. l. 03 - 04.05.2014. A. V. Shavrin leg. (cSh); 1 ♂, 2 ♀♀: ‘ Kiesenwetter’, ‘ Jaen (Sierra) ’, ‘ Sammlung Cl. Müller’, ‘ fontinalis Kiesw’ (ZSM); 1 ♂: ‘ Jaen’, ‘ Sammlung Cl. Müller’, ‘ fontinalis Kiesw’ (ZSM); 1 ♂: Jaen (MHNG); 1 ♂: Jaen, Jaen-Valdepenas Rd., 18 km S Jaen. 1000 m a. s. l. 25.05.1967. M. E. Bacchus & D. Levey (MHNG); 1 ♀, 1 ex.: ‘ S. d. Jaen’, ‘ Sammlung Cl. Müller, ‘ fontinalis? ’ (ZSM); 1 ♂: Cazorla. 800 - 2000 m a. s. l. 05.1943. G. Frey & C. Koch (MHNG); 1 ♂: ‘ fontinalis Ksw type. M … [illegible] ’, ‘ … [illegible] prés Granada’, ‘ Andalousie’ (IRSN); 1 ♂: ‘ Spanien, 1935 Sierra Nevada 2000 m, 16 / 6 O. Lundblad’, ‘ Lesteva fontinalis Kiesw. Det. Scheepeltz 1964 ’ (SMNH); 1 ♂, 1 ♀: ‘ Süd. Spanien’, ‘ Andalusien’ (NMPC); LOCALITY NOT SPECIFIED: 2 ♂♂: ‘ Spanien Scriba’, ‘ Coll. K. Neumann’ (SF). Additional material. FRANCE: 1 ♀: ‘ St. Jean de Luz Basse-Pyrenees’ (NMPC); MOROCCO: 1 ♂, 1 ♀: ‘ Rif Ketama’ (MHNH); LOCALITY NOT SPECIFIED: 1 ♂, ‘ Pyrén [ees]. ’, ‘ fontinalis ’, ‘ L. pandellei J. Bohač det. 1980 ’ (NMPC); 1 ♂: Pyrenees (MHNG).	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF95EE2B49AE4F45FA8AFADE.taxon	discussion	Remarks. Lesteva fontinalis was originally described based on an unspecified number of syntypes from “ Mons serratus [Montserrat] Cataloniae ”. During the study of Lesteva from ZSM, the first author found a strongly damaged male, which undoubtedly belongs to the type series. Consequently, this specimen is designated as the lectotype. Lohse (1960) described L. fontinalis truncata as “ kleine, kurzflügelige und kleinäugige Rasse ” from Central Spain. Herman (2001 a) replaced the preoccupied name with L. fontinalis gustavi Herman, 2001. Based on the study of the type and additional material from Spain and other countries, we did not find pronounced morphological differences between them. Body size, coloration, and the proportions of temples and elytra of L. fontinalis are variable. Thus, the new synonymy is proposed here. Habitus as in Fig. 9. Aedeagus as in Figs. 14 – 15. Lesteva fontinalis is known from Switzerland, France, Spain, and North Africa (Schülke & Smetana 2015). In North Africa it was recorded from Morocco, Algeria, and Tunisia (Fauvel 1871 [= 1872], 1878, 1886, 1902, Champion 1898), but all these records require confirmation. It was recorded from Spain by Fauvel (1871 a [= 1872]) and Lohse (1955 a, 1960).	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF9BEE2849AE4F8EFCFEFE7D.taxon	description	(Figs. 10, 16 – 17)	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF9BEE2849AE4F8EFCFEFE7D.taxon	materials_examined	Material examined. PORTUGAL: CASTELO BRANCO: 1 ♂: Serra da Estrela, S Manteigas, 40 ° 21 ‘ 20 N 7 ° 33 ‘ 39 W. 1073 m a. s. l., bushes, under stones. 18.03.2002. A. Lompe leg. (cA); SPAIN: ARAGÓN: 1 ♂: Pirineos, Monte Perdido, Collata Añisolo. 2400 m a. s. l. 25.08.2005. A. Anichtchenko leg. (cSh); CATALUñA: 1 ♂, 2 ♀♀: Seo de Urgel (Arfa), arr. de la Coma. 05 - 06.1962. G. Fagel leg. (MHNG); ANDALUSÍA: 1 ♂: ‘ Jaen (Espagne) ’ (IRSN); 9 exs.: Jaen-Vaidepenas Rd., 18 km S Jaén. 1000 m a. s. l. 25.05.1967 (BMNH); 2 ♀♀: ‘ Jaen (Sierra) ’, ‘ Sammlung Cl- Müller’, ‘ hanseni sp. n. Lohse det. ’ (ZSM); 1 ♂: Sierra de Cazorla, Torre del Vinagre. 14.05.1975. H. Fülscher leg. (MHNG); 1 ♂: Granada, Sierra de la Sagra, rio Bravate. 1100 m a. s. l. 30.04.1991. M. Tronquet leg. (cZan); 2 ♀♀: Ronda, Puente de la Ventilla. 8 - 20.05.1956. G. Fagel leg. (MHNG); 4 ♂♂, 1 ♀: ‘ Ronda Dr. Cameron’ (BMNH); 1 ♂: Tarifa. 04.1992. P. Poot leg. (cW); 1 ♀: ‘ Andalusien’, ‘ Sammlung M. v. Pfaundler’ (ZSM); 1 ♀: ‘ Prov. Andalus. 20.03.1974 ’ (MHNG); LOCALITY NOT SPECIFIED: 1 ♀: ‘ Hispan. 6 Sz. ’, ‘ Samml. v. Seidlitz’, ‘ hanseni Lohse Lohse det. ’ (ZSM); 1 ♂: ‘ Hispan. ’ (MHNG). Additional material. FRANCE: 1 ♂: Carnus, Eaux Bonnes. 14.06.1991. P. Wunderle leg. (cW); 1 ♀: ‘ Col de Aras Pyren. or. ’ (MHNG); MOROCCO: 7 exs.: 11 km SE Asni, 50 km S Marrakesh 22. iv. 1961. P. N. Lawrence (BMNH); 1 ♂: Tizi-n-Test Pass. 2100 m a. s. l. 10.04.1989. (ZMUC); 1 ♂: Haut Atlas, NE Tizi-n-Test, 30 ° 54 ’ 12 N 08 ° 18 ’ 39 W. 1710 m a. s. l., edge of stream. 26.12.2002. V. Assing leg. (cA); LOCALITY NOT SPECIFIED: 1 ♂: ‘ Atlas [… illegible ...] ’ (BMNH).	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF9BEE2849AE4F8EFCFEFE7D.taxon	discussion	Remarks. Lesteva hanseni is widely distributed in Europe (Schülke & Smetana 2015). It was recorded from Morocco (Coiffait 1973) and southern Spain (Lohse 1960). Habitus as in Fig. 10. The shape of the apical portion of the median lobe is somewhat variable (Fig. 16 and Fig. 92 c in Zanetti (1987 )). Lateral aspect of aedeagus as in Fig. 17. The species is here recorded from Portugal for the first time.	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF98EE2949AE4855FB54FEC9.taxon	materials_examined	Material examined. SPAIN: PAIS VASCO: 1 ♂, 1 ♀: Torre. 2000 m a. s. l .. 15.11.1997. P. Poot leg. (cW); ARAGÓN: 1 ♂: Monte Perdido, Collata Añisolo. 2400 m a. s. l. 25.08.2005. A. Anichtchenko leg. (cSh); CASTILLA Y LÉON: 1 ♂: Astorga. Paganetti leg. (FMNH); 1 ♂: Palencia. Paganetti leg. (FMNH); MADRID: 1 ♂, 1 ♀: Cercedilla (MNCN); 50 ♂♂, 42 ♀♀: same loality. 05.1909. Bolivar leg. (MNCN); 1 ♂: Madrid. 06.1909 (MNCN); 1 ♂: Madrid. 1926. Lindberg leg. (MZHF); 1 ♀: Navacerrada. Martin leg. (IRSN); 1 ♂: Guadarrama, Eppelsheim leg. (SF); 1 ♂: ‘ Serra Guadarrama Escorial’ (NHMB); 2 ♀♀: same locality. J. Ildefonso leg. (NMPC); EXTREMADURA: 1 ♂, 1 ♀: La Granja (IRSN, BMNH); ANDALUSÍA: 2 ♂♂, 1 ♀, 13 exs.: Granada, Mt. Mulhacen. 2000 m a. s. l. 21.05.1967. M. E. Bacchus & B. Levey leg. (BMNH, MHNG); 1 ♂: Capileira. 1500 m a. s. l. 28.09.1993. P. Wunderle leg. (cW); 1 ♀: Sierra Nevada, Barranco de Cauchiles. 2000 m a. s. l. 23.07.1971. (cW); 1 ex.: ‘ Andalusia’ (ZIN); 2 ♂♂, 1 ♀: ‘ Kraatz’, ‘ Andalusien’, ‘ Coll. K. Neumann’ (SF); 1 ♂: ‘ Sierra Nevada nivicola Kr. ’ (IRSN); LOCALITY NOT SPECIFIED: 1 ♂: ‘ Spanien’ (NHMB). Additional material. FRANCE: 1 ♀: Grottes de Bétharram (IRSN); 10 exs: Col de Peyresourde. 5,000 - 6,500 ft, in wet moss by stream. 04.09.1968 (BMNH); 1 ♂, 1 ♀: ‘ Lesteva pubescens Mnh. H. Pyrenees’ (MNCN); MOROCCO: 1 ♂: Sidi Ali Lake, 50 km S Ifrane. 6700 ft, island in lake, flood debris. 11.05.1961 (BMNH); ALGERIA: 1 ♂: ‘ mt. Babor Algérie’ (IRSN); LOCALITY NOT SPECIFIED: 3 exs.: ‘ Pyrenaei’ (ZIN).	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF98EE2949AE4855FB54FEC9.taxon	discussion	Remarks. Lesteva l. longoelytrata is a West Palaearctic subspecies, distributed in Orkney Islands, Faroe Islands, Iceland and Britain Isles through North Africa, Europe, Caucasus, Turkey, Lebanon, Cyprus, Turkmenistan and Iran (Shavrin 2014), replaced by L. l. maura Erichson, 1840 in Corsica and Sardinia, and by L. l. cretica Lohse & Steel, 1961 in Crete (Zanetti 1987). The status of these subspecies requires confirmation. In North Africa L. l. longoelytrata is known from Morocco (Koch 1941, Coiffait 1973) and Algeria (Fauvel 1878, 1886, 1902). This species is very common in the Iberian Peninsula (Gamarra & Outerelo 2008) and known from both Portugal and Spain (Kiesenwetter 1851, Heyden 1870, Quedenfeldt 1884, Champion & Chapman 1905, Lindberg & Bernhauer 1931, Koch 1940, Jeannel & Jarrige 1949, Jarrige 1954, Lohse 1960, Outerelo et al. 1998, etc.).	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF99EE2F49AE4B81FC5BFD21.taxon	description	(Figs. 8, 11, 18 – 19)	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF99EE2F49AE4B81FC5BFD21.taxon	materials_examined	Type material examined. Holotype, ♂ [apical part of the aedeagus slightly damaged; aedeagus contained in a small glass vial with a cork pinned under the card with the beetle]: ‘ Covilhã | 20.5. [19] 12 ’ <handwritten in black>, ‘ Samml. | K. Daniel’ <printed>, ‘ Typus [handwritten in blue] | Lesteva | lusitana n. sp. [handwritten in blue] | G. A. Lohse det. ’ <printed>, ‘ Holotypus [printed] ♂ | Lesteva | lusitana Lohse | 1955 | Staatssamml. München [printed] ’ <red label, handwritten in black>, ‘ Zool. Staatsslg. | München’ <blue label, printed> (ZSM). Cotype, 1 ♂: ‘ Covilhã | 20.5. [19] 12 ’ <handwritten>, ‘ Sammlg. | K. Daniel’ <handwritten>, [red full label], ‘ cotypus’ <handwritten>, ‘ Lesteva | lusitana n. sp. [handwritten] ’ <printed>, ‘ G. A. Lohse det. ’ <printed>, ‘ Coll. | G. A. Lohse | MHNG- 1994 ’ <printed>, ‘ COTYPE | Lesteva | lusitana sp. n. | Det. Lohse 1955 ’ <red, printed>, ‘ MHNG | ENTO | 00087329 ’ <printed>, ‘ Lesteva (s. str.) | lusitana Lohse, 1955 | Shavrin A. V. det. 2021 ’ <printed> (MHNG). Material examined. PORTUGAL: ALTO TÂMEGA: 3 ♂♂, 2 ♀♀: Serra do Gêres. 800 m a. s. l. 26.05.1992. P. Wunderle leg. (cW); CASTELO BRANCO: 2 ♂♂: Sierra de Estrela. H. Franz leg. (NHMW); 1 ♂, 1 ♀: Estrela. 1950 m a. s. l. 10.07.1979. G. Osella leg. (cZan); SPAIN: GALICIA: 1 ♂: Lugo, Valle de Florenza. H. Franz leg. (NHMW); 1 ♂: Sierra de Queixa. 1300 - 1780 m a. s. l. 27 - 28.07.1992. M. Tronquet leg. (cZan); CASTILLA Y LEÓN: 1 ♂: Cordillera Cantabrica, NE Puerto de Ventana, 43 ° 04 ’ N - 6 ° 00 ’ W. 1585 m a. s. l. 13.06.2000. D. W. Wrase leg. (cSch); CANTABRIA: 1 ♀: Reinosa, Campoo Cebernigua, 15 km NW Reinosa. 06.06.1991. P. Wunderle leg. (cW); ASTURIAS: 1 ♂: Picos de Europa, Covadonga, Lago Enol. 1200 m a. s. l. 02.06.1990. (cZan); 1 ♀: same data, Cornion. 2000 m a. s. l. 04.07.1994. T. Assmann leg. (cA).	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF99EE2F49AE4B81FC5BFD21.taxon	description	Redescription. Measurements (n = 15): HW: 0.67 – 0.70; HL: 0.45 – 0.50; OL: 0.20 – 0.25; LT: 0.08 – 0.10; AL (holotype): 1.69; PL: 0.62 – 0.70; PWMax: 0.72 – 0.80; PWMin: 0.60 – 0.66; EL: 1.50 – 1.63; ESL: 1.35 – 1.55; EW: 1.35 – 1.40; MTbL (holotype): 0.92; MTrL (holotype): 0.50; AW: 1.18 – 1.38; AedL: 0.82 – 0.86; FL: 2.62 – 3.18; TL: 3.20 – 4.00 (holotype: 3.90). Body brown to reddish-brown, sometimes with slightly paler elytra; antennae and legs yellow-brown; mouthparts and tarsi yellow to yellow-brown. Punctation of head moderately large, sparser in middle and denser between longitudinal depressions and eyes; punctation of pronotum dense, slightly deeper than that of head. Habitus as in Fig. 11. Head 1.4 times as broad as long, with distinctly elevated middle portion, indistinct transverse impression at level of middle length of eyes and separated from lateral portions by very deep longitudinal depressions; eyes more than twice as long as temples. Antennae reaching one third of elytra when reclined, with slightly thickened antennomeres 7 – 10; antennomeres 2 – 7 approximately equal in length, 8 – 9 slightly shorter than 7, 10 about as long as and slightly broader than 9. Pronotum slightly broader than long, about as wide as head, widest in anterior third portion, sharply narrowed toward subparallel latero-basal margins, with obtuse or subacute hind angles; mediobasal third of pronotum with distinct transverse impression. Elytra about as broad as long, more than twice as long as pronotum. Metatarsus less than twice as long as metatibia, with basal metatarsomere slightly shorter than combined length of tarsomeres 2 – 3. Abdomen with paired transverse tomentose spots in the middle of abdominal tergite V. Male. Apical margin of abdominal tergite VIII rounded. Apical margin of abdominal sternite VIII widely concaved. Aedeagus with very wide basal portion, strongly narrowed toward acute apex; parameres narrow, widened apically, distinctly exceeding apex of median lobe, with four long apical and preapical setae; internal sac moderately short and narrow (Fig. 18). Lateral aspect of aedeagus as in Fig. 19. Female. Apical margins of abdominal tergites and sternites VIII rounded. Comparative notes. Regarding the coloration, large and deep punctation of the pronotum, the shape of the apical margins of the elytra, and the length of the apical metarsomere, L. lusitana is similar to L. monticola and L. osellai sp. n. Regarding the shape of antennomeres it is more similar to L. monticola, from which it is distinguished by smaller body size and broader pronotum. From both these species, it differs by the shape of narrow median lobe, significantly longer parameres, and the different structure of the internal sac.	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF99EE2F49AE4B81FC5BFD21.taxon	distribution	Distribution. The species is known from several localities in central Portugal and the north-western provinces of Spain (Fig. 8). Natural history. Specimens were collected at elevations from 450 to 1585 m a. s. l. Detailed ecological data are unknown.	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF99EE2F49AE4B81FC5BFD21.taxon	discussion	Remarks. The species is here recorded from Spain for the first time.	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF9FEE2D49AE48E9FCDDFE59.taxon	description	(Figs. 8, 12, 20 – 22)	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF9FEE2D49AE48E9FCDDFE59.taxon	materials_examined	Type material examined. Holotype, ♂ [The aedeagus glued on the triangle card pinned under the card with specimen]: ‘ Barçges H [autes]. Pyr [énées]. VIII. [19] 49 P. Marié’ <handwritten>, ‘ Bovr’ <handwritten>, ‘ TYPE’ <red, printed>, ‘ Holotypus Lesteva mariei Jarrige’ <handwritten> (MHNH). Paratypes: 1 ♂, 1 ♀: ‘ Bagn [ères]. de Bigorre H [autes]. P [yrénées]. P. Nadar’ (MHNH); 1 ♂: ‘ Lac Bleu H [autes]. Pyr [énées]. Ch. F [agniez]. ’ (MHNH); 1 ♀: ‘ [... illegible ..] du Laurentli Carlitte N 2200 m’ (MHNH); 1 ♂: ‘ St Engrâce B [asses-]. P [yrénées] [… illegible …] ’ (MHNH). All paratypes with printed in red label: ‘ PARATYPE’. Material examined. SPAIN: GALICIA: 1 ♂: Orense, Pena Trevinca. 1800 m a. s. l .. 03.06.1995. W. Starke leg. (cA); CANTABRIA: 1 ♀: Espinama, 27.07.1972 (MHNG); 1 ♂, 4 ♀♀: same locality, Valle de Salvoron. 30.07.1972. (MHNG); 1 ♂ Picos de Europa, Covadonga, Lago Enol. 1600 m a. s. l. 03.06.1990. (cZan); ASTURIAS: 1 ♂ Picos de Europa, Covadonga, Lago Enol. 1600 m a. s. l. 03.06.1990. (cZan); 1 ♂, 1 ♀: Oviedo, Puerto de Cubilla. 1400 m a. s. l. 08.06.1991. L. Zerche leg. (MHNG); 1 ♂: Puerto de La Cubilla, 1400 m a. s. l. 08.06.1991. P. Wunderle leg. (cW); 1 ♂, 1 ♀: Oviedo, Puerto de Cubilla. 1400 m a. s. l. 08.06.1991. L. Zerche leg. (MHNG). Additional material. FRANCE: 1 ♂, 3 ♀♀: Eaux-Bonnes. 14.06.1991. P. Wunderle leg. (cA, cW); 1 ♀: Gavarnie, Gédre-Tromouse. 22.07.1972. (cA); 1 ♂: Gedre. 22.07.1972. (MHNG); 1 ♂: El Portelet. 1400 m a. s. l. 12.06.1991. P. Wunderle leg. (cW); 1 ♀: Aucun, Vallée de Arrens. 15.06.1991. P. Wunderle leg. (cW); 1 ♂: Le Tech Vallée de Arrens. 1200 m. 14.06.1991. L. Zerche leg. (cW); 1 ♂: Gavarnie, Vallée d`Ossoue. 23.07.1972. (MHNG); 1 ♂: Casteil (Canigou). 1960 - 2020 m a. s. l. 15.07.2003. M. Tronquet leg. (cT); 1 ♂, 1 ♀: same locality. 06.1958. L. Levasseur leg. (MHNH); 1 ♀: between Lac de Cap-de-Long and d`Oredon. 1900 - 2200 m a. s. l, in wet moss by streams. 05.09.1968. P. Hammond leg. (MHNG); 1 ♂: Ariège, Vallèes Villages Mountagnes. 08.1958. (MHNG); 1 ♂: Forges d`Abel. 28.08.1938. B. P. Giraud leg. (MHNG); ANDORRA: 1 ♂, 1 ♀: Puerto de Envalira. 2200 - 2300 m a. s. l. 30.05.1994. M. Schülke & B. Grünberg leg. (cZan).	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF9FEE2D49AE48E9FCDDFE59.taxon	description	Redescription. Measurements (n = 30): HW: 0.68 – 0.81; HL: 0.46 – 0.68; OL: 0.17 – 0.23; LT: 0.12 – 0.16; AL (holotype): 1.80; PL: 0.65 – 0.83; PWMax: 0.85 – 1.02; PWMin: 0.56 – 0.77; EL: 1.33 – 1.58; ESL: 1.16 – 1.41; EW: 1.38 – 1.61; MTbL (holotype): 0.90; MTrL (holotype): 0.47; AW: 1.28 – 1.49; AedL: 0.70 – 0.85; FL: 2.58 – 3.27; TL: 3.75 – 4.65 (holotype: 4.00). Body brown or reddish-brown, sometimes with somewhat paler elytra and apical abdominal tergites; antennomeres and legs yellow-brown to brown (sometimes antennes paler); mouthparts and tarsi yellow. Punctation of head dense, fine and moderately deep, sparser in middle and between ocelli, finer and denser around eyes; punctation of pronotum similar to that in middle portion of head, but markedly denser, with interspaces between punctures in middle as wide as diameters of nearest one-two punctures, sparser in mediobasal portion; punctation of elytra regular, sparse, larger and deeper than that in pronotum, finer in prescutellar and sometimes along suture; abdominal tergites without distinct punctures. Lateral portions of head around eyes sometimes with indistinct fine meshes; pronotum usually without microreticulation, but some specimens with very fine and indistinct transverse meshes in medioapical third; abdominal tergites with distinct isodiametric microsculpture. Habitus as in Fig. 12. Head 1.1 – 1.4 times as broad as long, widely elevated in middle; middle portion separated from lateral portions by very deep and moderately wide longitudinal depressions; eyes about one and a half times as long as temples. Antennae with elongate antennomeres; basal antennomere disitnctly more than twice as long as broad, 2 significantly narrower and shorter than basal antennomere, 3 slightly longer than 2, 4 slightly broader than 3, 5 – 8 slightly narrower and about as long as 4, 9 – 10 slightly shorter and wider than 8, apical antennomere about 1.4 times as long as 10, from about middle shaprly narrowed toward acute apex. Pronotum 1.2 – 1.3 times as broad as long, 1.2 times as broad as head, from their widest apical third sharply narrowed toward subparallel latero-basal margins; dorsal surface sometimes with paired oval impressions in about middle and without or with indistinct, transverse mediobasal impression. Elytra slightly broader than long, 1.6 – 1.7 times as long as pronotum, significantly widened toward hind angles. Male. Apical margin of abdominal tergite VIII rounded. Apical margin of abdominal sternite VIII deeply concave. Aedeagus with wide moderately median lobe, from apical third significantly narrowed toward small, rounded apex; parameres slightly exceeding apex of median lobe; internal sac wide and long (Figs. 20, 22). Lateral aspect of aedeagus as in Fig. 21. Female. Apical margins of abdominal tergite VIII and sternite VIII rounded. Comparative notes. Regarding the general shapes of the body and antennomeres, body size and coloration, L. (s. str.) mariei is similar to the Iberian L. (s. str.) mateui, but differs by shorter elytra, the shape of the median lobe and length of the parameres (for details see the key below).	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF9FEE2D49AE48E9FCDDFE59.taxon	distribution	Distribution. Lesteva mariei is widely distributed in the Pyrenees and Cordillera Cantabrica (Fig. 8). Natural history. Specimens were collected at elevations from 1200 to 2300 m a. s. l. Detailed ecological data are unknown, but some specimens from Spain were collected by sifting wet mosses near streams.	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF9FEE2D49AE48E9FCDDFE59.taxon	discussion	Remarks. Lesteva mariei was originally described from several localities of the French Pyrenees. The species is here recorded from Spain for the first time.	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF9DEE3249AE4831FAA0FB61.taxon	description	(Figs. 8, 13, 23 – 28)	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF9DEE3249AE4831FAA0FB61.taxon	materials_examined	Type material examined. Paratypes of Lesteva mateui Jarrige, 1954, 2 ♀♀: ‘ Corral del Veleta Sierra Nevada 3200 m J. Mateu’ <handwritten>, ‘ PARATYPE’ <printed in red> (MHNH). Holotype of Lesteva cobosi Jarrige, 1954, ♂ [The aedeagus is glued on the separate card]: ‘ ♂ ’ <handwritten>, ‘ F [uen] te. Bermejo. Jaén Iruela S [ierra]. Cazorla Mateau-Cobos coll. ’ <handwritten>, ‘ Holotype’ <red, printed>, ‘ Lesteva (s. str.) cobosi Coiff. H. Coiffait det 1970 ’ <handwritten> (MHNH). Paratype, 1 ♀: same data as the holotype except for red printed label ‘ Paratype’ (MHNH). Holotype of Lesteva cazorlana Lohse, 1987, ♂ [without right antennomeres 6 - 11]: ‘ ESPAGNE JAEN | Sierra de Cazorla | Guadalquivir | (source) 25 - VII- 71 | A. SENGLET’ <printed>, ‘ Holotypus’ <red, handwritten>, ‘ Lesteva | cazorlana | n. sp. ’ <handwritten>, ‘ Coll. | G. A. Lohse | MHNG- 1994 ’ <printed>, ‘ HOLOTYPE | Lesteva | cazorlana sp. n. | Det. Lohse 1987 ’ <red, printed>, ‘ MHNG | ENTO | 00087325 ’ <printed>, ‘ Lesteva (s. str.) | mateui Jarrige, 1954 | Shavrin A. V. det. 2021 ’ <printed> (MHNG) Material examined. SPAIN: ANDALUSÍA: 1 ♂: Albacete, Mundo River, Chorros del Rio Mundo. 01.06.2002. I. Ribeira & A. Cieslak leg. (cA); 1 ♂, 1 ♀: Jaén, Iznatoraf, Sima los Tejos. 30.10.2005. (cA); 3 ♂♂: Cazorla Barranco de Teatinos. 1300 m a. s. l. 25.06.1975. (cZan); 5 ♂♂, 2 ♀♀: Sierra de Cazorla. 1400 m a. s. l. 06.10.1993. P. Wunderle leg. (cA, cW, cSh); 2 ♂, 5 ♀♀: same data. 20.03.1974 (MHNG); 3 ♂♂, 1 ♀: same Mts., Torre del Vinagre. 14.05.1975. H. Fülscher leg. (MHNG); 2 ♂♂, 1 ♀: Guadalquivir (source), 25.07.1971. A. Senglet leg. (MHNG); 9 ♂♂, 4 ♀♀: Sierra Nevada, Capileira, Rio Nante, 36 ° 99 ’ N - 3 ° 35 ’ W. 1550 m a. s. l. 08 - 09.05.2014. A. V. Shavrin leg. (cSh); 1 ♂, 4 ♀♀: same locality. 1600 m a. s. l. 22.03.1994. V. Assing leg. (cA); 1 ♂: same data. 1900 m a. s. l. 23.03.1994. (cA); 6 ♂♂, 1 ♀: same data. 1500 m a. s. l. 28.09.1993. P. Wunderle leg. (cA, cW, MHNG); 1 ♀: Sierra Nevada, Laguna de las Yeguas. 2900 m a. s. l. 18.05.2005. A. Anichtchenko leg. (cSh); 1 ♀, 6 exs.: same locality. 17.08.1966 (MHNH); 2 ♂♂, 2 ♀♀: Bano San Juan. 2900 m a. s. l. 18.08.1966. G. Ledoux leg. (MHNH); 1 ♂, 2 ♀♀: same data. 2600 m a. s. l. 08.1967. G. Ledoux leg. (MHNH); 1 ♂: ‘ Sierra Nevada 17.7. [18] 79. D. S. ’, ‘ Sharp Coll. 1905 - 303. ’ (MHNG); 1 ♂, 1 ♀: ‘ Sa Nevada Hisp. VI. 50 G. P. ’ (MHNH); 2 ♂♂: Güejar-Sierra, Loma de la Cuna de los Cuartos. 1900 - 2100 m a. s. l. 14.06.1991 D. Wrase leg. (cZan); 1 ♂: Málaga, Las Navas de San Luis. M. Cobos leg. (MHNH); LOCALITY NOT SPECIFIED: 1 ♂ [identified by G. A. Lohse]: ‘ Spain. ’ (BMNH).	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF9DEE3249AE4831FAA0FB61.taxon	description	Redescription. Measurements (n = 15): HW: 0.70 – 0.92; HL: 0.50 – 0.62; OL: 0.20 – 0.26; LT: 0.11 – 0.15; AL (mean): 2.30; PL: 0.63 – 0.85; PWMax: 0.84 – 0.96; PWMin: 0.59 – 0.70; EL: 1.38 – 1.65; ESL: 1.11 – 1.45; EW: 1.23 – 1.53; MTbL (averaged): 1.00; MTrL (averaged): 0.60; AW: 1.14 – 1.44; AedL: 0.67 – 0.75; FL: 2.87 – 3.05; TL: 3.00 – 4.80 (holotype of L. mateui: 4.20). Body yellow-brown to brown, with slightly paler elytra; antennae and legs yellow to yellow-brown, with darker apical portion of femora and sometimes tibia; mouthparts and tarsi yellow. Punctation of head dense and fine, slightly larger in middle and denser between longitudinal depressions and eyes, distance between punctures in middle about as broad as nearest puncture; neck with punctation similar to that of middle portion of head; punctation of pronotum denser and sometimes finer than that of head, slightly sparser in medioapical and mediobasal portions; scutellum with very fine and moderately dense, irregular punctation; punctation of elytra distinctly sparser and coarser than that of pronotum. Habitus as in Fig. 13. Head with markedly convex middle portion and very deep longitudinal depressions; eyes small, about twice as long as temples; distance between ocelli markedly shorter than distance between ocellus and posterior margin of eye. Antennae reaching anterior third or middle length of elytra when reclined, with long antennomeres 4 – 10; antennomeres 3 – 7 of approximately equal length, 8 – 10 slightly shorter and broader than 7. Pronotum 1.1 – 1.3 times as broad as long, markedly broader than head, usually very wide in anterior portion. Elytra distinctly broader than long, about twice as long as pronotum, significantly widened apicad. Basal metatarsomere slightly longer than combined length of tarsomeres 2 – 3 and distinctly longer than apical metatarsomere. Male. Apical margin of abdominal tergite VIII convex. Apical margin of abdominal tergite VIII concave. Aedeagus with moderately wide basal portion, gradually narrowing toward median lobe and slightly widened in apical third, with sharply narrowed apical portion and subacute apex; parameres nearly reaching apex of median lobe; internal sac long and moderately wide (Figs. 23, 25, 27). Lateral aspect of aedeagus as in Figs. 24, 26, 28. Female. Apical margins of abdominal tergite VIII and sternite VIII round. Comparative notes. Regarding body, size, coloration, the punctation of the forebody, the proportions of the antennomeres, and the shape of the elytra, L. mateui is similar to L. mariei (see above), from which it is distinguished by slightly longer elytra, a narrower median lobe with a differently shaped apical portion, the length of the parameres, and a different structure of the internal sac.	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF9DEE3249AE4831FAA0FB61.taxon	distribution	Distribution. The species is distributed in South Spain (Fig. 8). Natural history. Specimens were collected at elevations from 1300 to 2900 m a. s. l. The specimens from Rio Nante (Sierra Nevada) were taken from wet mosses near the confluence of a stream and a river.	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF9DEE3249AE4831FAA0FB61.taxon	discussion	Remarks. Lesteva mateui was originally described from several localities of Sierra Nevada. Lesteva cobosi was described from Sierra de Cazorla (Coiffait 1970) and later synonymized with L. mateui by Lohse (1987). In the same article, Lohse presented a new species without description: L. cazorlana Lohse, 1987, referring to specimens with somewhat larger eyes and a longer median lobe, which had been identified by Coiffait as L. mateui and illustrated by him in Figs. 3 g-i (Coiffait 1970). Lohse designated the specimen from his collection (see above) as the holotype of L. cazorlana. A study of material of this morphologically variable species revealed no significant differences between specimens from different localities. The proportions of the longitudinal length of the eyes in relation to the length of the temples, the width of the pronotum, and the apical portion of the median lobe may vary even in material from the same locality (Figs. 23, 25). Thus, L. cazorlana is synonymized with L. mateui.	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF82EE3249AE4F29FDA5F87D.taxon	materials_examined	Material examined. SPAIN: CANTABRIA: 1 ♂: Ruberdiago. 11.06.1990. D. A. Lott leg. (BMNH); ASTURIAS: 4 spec: De Pajares (BMNH); MADRID: 1 ♂: Escorial (IRSN); 1 ♀: same locality. 30.07.1879. L. Bleuse leg. (NMPC); 2 ♀♀: Parque Nacional de Guadarrama, San Ildefonso - Majada Hambrienta. 21.06 - 08.07.2016. J. L. Lencina & J. C. Otero González leg. (cSh); ARAGÓN: 1 ♀: Estrella (IRSN); ANDALUSÍA: 1 ♂: Granada (IRSN); 1 ex.: ‘ Andalucia’ (ZIN); LOCALITY NOT SPECIFIED: 1 ♀: ‘ Spanien’, ‘ L. pandellei J. Bohač det. 80 ’ (NMPC); 1 ♀: ‘ Pyrén. ’, ‘ fontinalis ’, ‘ L. pandellei J. Bohač det. 80 ’ (NMPC).	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF82EE3249AE4F29FDA5F87D.taxon	discussion	Remarks. Lesteva monticola is widespread in Europe and also known from Turkey (Schülke & Smetana 2015). It was recorded from Spain as L. pandellei (Fauvel 1876, Quedenfeldt 1884, Wagner 1927). The record from Algeria (Fauvel 1902) requires confirmation.	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF80EE3149AE4A85FEC8F9FD.taxon	description	(Figs. 8, 29 – 30, 32 – 37)	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF80EE3149AE4A85FEC8F9FD.taxon	materials_examined	Type material examined. Holotype, ♂: ‘ Portugal | Estrela m 1950 | 10. vii. [19] 79 | Osella’, ‘ HOLOTYPE | Lesteva (s. str.) | osellai sp. n. | Zanetti A. & Shavrin A. V. des. 2020 ’ <red, printed> (MCSNV). Paratypes: 23 ♂♂ (three specimens dissected): ‘ Portugal | Estrela m 1950 | 10. vii. [19] 79 Osella’ <handwritten> (2 ♂♂: cZan, 21 ♂♂: MCSNV); 1 ♂ (dissected): ‘ Portugal | Estrela 1950 | m 10 - VII- 79 | Osella’ <handwritten>, ‘ Lesteva [handwritten] | prope [handwritten] | lusitana [handwritten] | det. A. Zanetti 197 ’ <printed>, ‘ Lesteva | lusitanica Lohse | Det. Lohse’ <printed>, ‘ Coll. | G. A. Lohse | MHNG- 1994 ’ <printed> (MHNG); 1 ♂ (dissected; right antennomeres 10 - 11 missing): ‘ Portugal | Estrela 1950 m | 10. VII. 1979 | Osella’ <handwritten> (cSh); 1 ♂ (aedeagus mounted), 31 ♀♀ (two specimens without head and pronotum): ‘ Portugal | Estrela 1950 | m 10. VII. [19] 79 | Osella’ <handwritten> (1 ♀: cZan, 30 ♀♀: MCSNV); 2 ♂♂, 1 ♀: ‘ Espagne | Picos de | Europa’ <handwritten>, ‘ Covadonga | Lago Enol | 1600 m. 3. VI. 90 ’ <handwritten> (1 ♂, 1 ♀: cZan; 1 ♂: cSh); 1 ♀ (pronotum and mediobasal portion of left elytron damaged): ‘ Espagne | Picos de Europa | (Oviedo) ’ <handwritten>, ‘ Covadonga | Lago Enol | 1200 m 2. VI. [19] 90 ’ <handwritten> (cZ); 1 ♀: ‘ E Cord. Cantabrica | Pto. de Cubilla | 1400 m Spritzmoos | 08.06. [19] 91 [P.] Wunderle’ <printed>, ‘ Lesteva [handwritten] | det. P. Wunderle’ <printed>, ‘? coiffaiti’ <handwritten> (cW); 1 ♂ (aedeagus glued on the same card under the specimen): ‘ Pto. de Pajares, | Spain. | G. C. C. ’ <printed>, ‘ coiffaiti [handwritten] | det. Dr. G. A. Lohse 197 ’ <printed>, ‘ Coll. | G. A. Lohse | MHNG- 1994 ’ <printed> (MHNG); 1 ♀ (without left antennomeres 3 - 11 and right antenna, without apical part of the abdomen): ‘ SPAIN. Leon, | Pto. de Tarna, | 17 km. N. Riano, | 1490 m, 1. vii. 1965 ’ <printed>, ‘ Headwaters Rio Nalon. ’ <printed>, ‘ Among stones and | grit: fast stream. ’ <printed>, ‘ J. Balfour-Browne. | B. M. 1965 - 461. ’ <printed>, ‘ coiffaiti [handwritten] | det. Dr. G. A. Lohse 19 ’ <printed>, ‘ Coll. | G. A. Lohse | MHNG- 1994 ’ <printed> (MHNG). All paratypes with additional red printed labels: ‘ PARATYPE | Lesteva (s. str.) | osellai sp. n. | Zanetti A. & Shavrin A. V. des. 2020 ’.	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF80EE3149AE4A85FEC8F9FD.taxon	description	Description. Measurements (n = 50): HW: 0.67 – 0.80; HL: 0.52 – 0.60; OL: 0.18 – 0.25; LT: 0.10 – 0.12; AL (mean): 2.38; PL: 0.72 – 0.85; PWMax: 0.82 – 0.97; PWMin: 0.60 – 0.70; EL: 1.50 – 1.80; ESL: 1.35 – 1.60; EW: 1.27 – 1.57; MTbL: 1.02; MTrL: 0.47; AW: 1.22 – 1.57; AedL: 0.77 – 1.02; FL: 3.15 – 3.35; TL: 4.10 – 5.05 (holotype: 4.10). Body brown, with paler elytra and apical abdominal tergites in some specimens; antennae and legs yellow-brown; mouthparts and tarsi yellow. Punctation of head dense, moderately coarse and deep, sparser in middle, and slightly finer around eyes (some paratypes from Spain with oval impunctate portion between ocelli); punctation of neck as that of head, with impunctate longitudinal band in middle; punctation of pronotum usually coarser and deeper than that of head, sometimes deeper and coarser in medioapical third, finer and sparser in middle and mediobasal third (some paratypes from Estrela with indistinct impunctate longitudinal band); scutellum with several fine punctures; punctation of elytra similar to that on pronotum, but sparser, slightly finer and denser in parascutellar portion and finer and sparser along suture; abdominal tergites with fine, moderately dense, and sometimes indistinct punctation. Body glossy, mostly without microsculpture; basal portion of neck with coarse, dense, isodiametric sculpture, basal portion of scutellum with fine transverse meshes, abdominal tergites with distinct transverse microsculpture on tergites IV – VI and finer, isodiametric microreticulation on tergites VII – VIII. Forebody covered with dense and moderately long semierect setation; abdomen with fine and dense setae. Habitus as in Figs. 29 – 30. Head 1.2 – 1.3 times as broad as long, distinctly elevated in middle; middle portion separated from lateral portions by deep longitudinal depression, markedly deeper between middle length of eyes; eyes large and convex, about twice as long as temples (lateral view), gradually narrowing toward neck; ocelli large, distance between ocelli subequal to distance between ocellus and posterior margin of eye. Antennae moderately short, reaching basal portion of elytra when reclined, with slightly thickened and moderately short antennomeres 4 – 10; basal antennomere about twice as long as wide, 2 about half as wide as basal antennomere, 3 slightly longer than 2, 4 slightly broader and longer than 3, 5 – 7 slightly longer than 4, 8 – 10 slightly shorter and broader than 7, apical antennomere about 1.3 times as long as 10, in apical third sharply narrowed toward apex. Pronotum convex, slightly broader than long and head, widest slightly before middle, from their widely rounded lateral margins gradually narrowed toward subparallel latero-basal margins, with obtuse hind angles; mediobasal third of some paratypes with distinct, transverse, moderately deep impression. Elytra distinctly convex, slightly longer than broad, twice as long as pronotum, slightly widened toward hind angles; apical margins widely rounded; mediobasal surface of elytra in some paratypes with indistinct depression. Metatarsus more than twice as long as metatibia, with basal metatarsomere about as long as combined length of tarsomeres 2 – 3 and about as long as apical tarsomere. Abdomen slightly narrower or about as broad as elytra, with paired tomentose spots in the middle of abdominal tergites IV and V, more transverse in tergite IV, smaller and round in tergite V, with distinct narrow palisade fringe on apical margin of abdominal tergite VII. Male (Fig. 29). Apical margins of abdominal tergite VIII slightly and sternite VIII deeply concave. Aedeagus with wide basal portion, gradually narrowed toward moderately narrow, long median lobe, in apical third sharply narrowed toward subacute apex; parameres narrow, slightly reaching apex of median lobe, with four short apical setae; internal sac moderately wide and long (Figs. 32, 34, 36). Lateral aspect of aedeagus as in Figs. 33, 35, 37. Female (Fig. 30). Apical margins of abdominal tergite VIII and sternite VIII rounded. Comparative notes. Based on the general body shape, coloration, robust antennomeres and moderately sparse and large punctation of the pronotum, L. osellai sp. n. is similar to L. lusitana, L. longoelytrata, and L. monticola. Regarding body size, the shape of the apical margins of the elytra, the length of the apical metatarsomere (not longer than basal metatarsomere), this species is similar to L. lusitana and L. monticola. It differs from them by the shorter antennomeres 8 – 10, a narrower median lobe, and shorter parameres.	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF80EE3149AE4A85FEC8F9FD.taxon	distribution	Distribution. The species is known from several localities in Serra da Estrela, Portugal, and the Cantabrian mountains, Spain (Fig. 8). Natural history. Specimens were collected at elevations from 1200 to 1950 m a. s. l. Detailed ecological data are unknown.	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF80EE3149AE4A85FEC8F9FD.taxon	etymology	Etymology. Patronymic, the species is named to honour our colleague, G. Osella (Verona, Italy), collector of the type material.	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF81EE3649AE4CAEFD68F8F9.taxon	description	(Figs. 31, 38 – 46)	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF81EE3649AE4CAEFD68F8F9.taxon	materials_examined	Type material examined. Paratypes of Lesteva fageli Lohse, 1960: 1 ♀: ‘ Espagne: Estremadura | Guadalupe | (canal scierie) | V — 1958, G. Fagel’ <printed>, ‘ fageli sp. n. [handwritten] | det. Dr. G. A. Lohse’ <printed>, ‘ Para- | type’ <red, printed>, ‘ Coll. | G. A. Lohse | MHNG- 1994 ’ <printed>, ‘ PARATYPE | Lesteva | fageli sp. n. | Det. Lohse 1960 ’ <yellow, printed>, ‘ MHNG | ENTO | 00087326 ’ <printed> (MHNG); 1 ♂ [dissected]: same labels, but: ‘ MHNG | ENTO | 00087327 ’ <printed> (MHNG); 1 ♂: same labels, but: ‘ MHNG | ENTO | 00087328 ’ <printed> (MHNG). All paratypes with additional printed label: ‘ Lesteva (str.) pubescens Mannerheim, 1830 | Shavrin A. V. det. 2021 ’. Material examined. PORTUGAL: CASTELO BRANCO: 3 ♂♂: Serra de Estrela. H. Franz leg. (NHMW); 1 ♂: Estrela (MHNG); 3 ♀♀: same data. H. Franz leg. (MHNG); SPAIN: GALICIA: 1 ♂, 1 ♀: Pontevedra Galice. 18 - 30.07.1959. H. Coiffait leg. (MHNH); 1 ♂: Lugo, Linares. 1000 m a. s. l. 27.05.1996. W. Starke leg. (cA); ASTURIAS: 1 ♂ [identified by G. A. Lohse as fageli], 2 ♀♀: ‘ Hispania la Granja’ (BMNH); SORIA: 2 ♂♂: Sierra de Urbión, forest W of Santa Inés. 1700 m a. s. l. 23.07.1996. D. Wrase & J. P. Zaballos leg. (cSch); CANTABRIA: 1 ♀: Espinama, 27.07.1972 (MHNG); 1 ♂, 4 ♀♀: same locality, Valle de Salvoron. 30.07.1972. (MHNG); 3 ♂♂: Poblagion (BMNH); CASTILLA Y LEÓN: 1 ♀: Sierra de Gredos. 16.07.1959. H. Coiffait leg. (MHNH); 1 ♂: Sierra de Gredos, Novacepeda. 01.06.1984. (cZan); 3 ♀♀: Puerto de Béjar. 800 - 1400 m a. s. l. 05 - 06.1957. G. Fagel leg. (MHNG); ARAGÓN: 1 ♂: Teruel (IRSN); CATALUñA: 1 ♂: Col de Aras (Span. Pyren.). 31.05.1971. G. Lohse leg. (MHNG); 1 ♂, 4 ♀♀: Barcelona (MHNG); 1 ♂: Sierra de Montseny, Tordera River, 5 km E San-Bernat. 02 - 12.5.1981. P. Zwick leg. (MHNG); 4 ♂, 6 ♀♀: Montseny (MHNG); MADRID: 1 ♀: Escorial (NMPC); 1 ♂: same locality. 30.07.1879. L. Bleuse leg. (NMPC); 1 ♂: ‘ Guadarrama | Eppelsh. ’, ‘ L. pandellei J. Bohač det. 1980 ’ (NMPC); 2 ♂♂, 2 ♀♀: Sistema Central Mts., Pinilla de Valle, 40 ° 91 ’ N - 3 ° 82 ’ W. 28.04.2014. A. V. Shavrin & A. Anichtchenko leg. (cSh); 2 ♀♀: Puerto de la Morcuera. 11.10.2004. de la Rosa leg. (cSh); 1 ♂: Sierra de Estrela, Manteigas. 1000 - 1100 m a. s. l. H. Franz leg. (MHNH); CASTILLA LA MANCHA: 2 ♂♂: Sierra de Alcaraz, 15 km NNE Riópar, 38 ° 34 ’ 01 N 02 ° 20 ’ 03 W 1120 m a. s. l. 10.04.2003. P. Wunderle leg. (cW); VALENCIA: 1 ex.: ‘ Buñol Valencia Moróder’ (MHNH); ANDALUSÍA: 5 ♂♂, 7 ♀♀: Despenaperros, Arroyo de los Arcos, 38 ° 38 ‘ N - 3 ° 51 ‘ W. 01 - 02.05.2014. A. V. Shavrin leg. (cSh); 1 ♂: ‘ Pnte. Las Herrerias S. Cazorla (Jaén) Mateau-Cobos coll. ’ (MHNH); 1 ♀: ‘ Fte. Bermejo. Jaén’, ‘ ruela S. Cazorla Mateau-Cobos coll. ’ (MHNH); 1 ♂: Sierra Nevada, Capileira. 1600 m a. s. l. 22.03.1994. V. Assing leg. (cSh); 9 ♂♂, 4 ♀♀: same data, but P. Wunderle leg. (cW, cSh); 1 ♂: Mt. Mulhacen. 2000 m. a. s. l. 21.05.1967. M. E. Bacchus & B. Levey leg. (MHNG). LOCALITY NOT SPECIFIED: 1 ♂: ‘ Miguelon’ (MHNH). Additional material. ANDORRA: 1 ♂: Puerto de Envalira. 2200 - 2300 m a. s. l. 30.05.1994. M. Schülke & Grünberg leg. (cSch); FRANCE: 1 ♂: ‘ Gavarnie 25.5. [19] 31 ’ (MHNH); 2 ♂♂: ‘ Gavarnie, H. Pyr. 12 - 31, vii. 14. G. C. C. ’ (BMNH); 2 ♂♂: ‘ Lourdes’ (IRSN); 8 ♂♂, 7 ♀♀: Por d`Boucharo. 30.06.1976. (MHNG); 14 ♂♂, 6 ♀♀: Ht. Canigou. 02.07.1976. (MHNG); 4 ♂♂, 3 ♀♀: Sahorre d. Vernet. 04.07.1976. (MHNG); 1 ♀: Ariège, Auzat. 19.07.1972 (MHNG); 1 ex.: ‘ Pyr. Or. La Vall. H. Coiffait 24.3. [19] 48 ’ (MHNH); 1 ♂, 1 ♀, 1 ex.: Banyuls-sur-Mer (IRSN, MHNH); 1 ♂: Gard Fressac. 28.10.1982. I. Löbl leg. (MHNG); 8 ♂♂, 2 ♀♀: ‘ Col de Aras Pyren. or. ’ (MHNG).	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
B43EBA76FF81EE3649AE4CAEFD68F8F9.taxon	discussion	Remarks. Lesteva fageli was originally described based on several type specimens from Cantabria (“ Puente del Cerezo ”) and central Spain (“ Rio Guadalupejo ”). Lohse compared it with L. fontinalis and L. pubescens. He distinguished it from the latter by the darker body and smaller aedeagus (“ 0,64 mm gegenüber 0,75 mm bei pubescens ”). During the study of types of L. fageli, we found that these specimens are morphologically conspecific with L. pubescens, a variable species regarding coloration, body size, and the proportions of the forebody and aedeagus (Figs. 38 – 46). Thus, we synonymize L. fageli with L. pubescens. Regarding the presence of stout pubescence on the median lobe, this species is similar to L. pourtoyi Jarrige, 1972, a species originally described from “ Cirque de Mahourat (Pyr [énées]. - Atl [antiues].) ”, but the aedeagus of this species is evidently narrower (Fig. 5 in Jarrige (1972 )) and is furnished with longer pubescence. According to Tronquet (2014), L. pourtoyi is also known from Spain (“ pourtoyi Jarrige, 1972: 158. - Pyr. occ. et cent. Esp. (chaîne Cantabrique ” )), but this record needs confirmation. Habitus of L. pubescens as in Fig. 31; aedeagus as in Figs. 38 – 46. Lesteva pubescens is widely distributed in Europe (Schülke & Smetana 2015) and had already been reported from Spain (Lohse 1960). It is here recorded from Portugal for the first time.	en	Shavrin, Alexey V., Zanetti, Adriano (2021): Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula. Zootaxa 4966 (4): 401-427, DOI: https://doi.org/10.11646/zootaxa.4966.4.1
