taxonID	type	description	language	source
5F738E5FFF8E45169FFE7D06EA27FCDB.taxon	type_taxon	Type species Bulimulus (Bostryx) solutus Troschel, 1847 (original designation by monotypy).	en	Miranda, María José (2014): Bostryx tortoranus (Doering, 1879) species complex (Gastropoda: Stylommatophora: Bulimulidae), a review of taxonomy and distribution of endemic species from Argentina. Journal of Natural History 49 (17 - 18): 995-1022, DOI: 10.1080/00222933.2014.981313, URL: http://dx.doi.org/10.1080/00222933.2014.981313
5F738E5FFF8E45169FFE7D06EA27FCDB.taxon	discussion	Remarks This genus can be characterized as having a protoconch with axial costules or wrinkles, with numerous fine spiral grooves well marked and densely arranged. Penial retractor muscle inserted in terminal portion of phallic complex. Flagellum inner wall with diagonal folds converging at a central longitudinal fold. Epiphallus divided into proximal thin and wider distal portion. Penis divided by a thin area into proximal swollen portion 1 / 3 of total penis length and distal cylindrical portion, ¾ of total penis length. Penis proximal portion with inner penis gland and two different sculptures in the inner wall. Penial sheath muscular overlapping more than half portion of penis with retractor muscle inserted in its proximal end. Bursa copulatrix duct divided into proximal thin and wider distal portion.	en	Miranda, María José (2014): Bostryx tortoranus (Doering, 1879) species complex (Gastropoda: Stylommatophora: Bulimulidae), a review of taxonomy and distribution of endemic species from Argentina. Journal of Natural History 49 (17 - 18): 995-1022, DOI: 10.1080/00222933.2014.981313, URL: http://dx.doi.org/10.1080/00222933.2014.981313
5F738E5FFF8E45199F997F8FE8E0FC01.taxon	description	(Figures 1 – 4)	en	Miranda, María José (2014): Bostryx tortoranus (Doering, 1879) species complex (Gastropoda: Stylommatophora: Bulimulidae), a review of taxonomy and distribution of endemic species from Argentina. Journal of Natural History 49 (17 - 18): 995-1022, DOI: 10.1080/00222933.2014.981313, URL: http://dx.doi.org/10.1080/00222933.2014.981313
5F738E5FFF8E45199F997F8FE8E0FC01.taxon	materials_examined	Type material Holotype (MACN-In 380), Paratypes (MACN-In 380 – 1 (5 )). Type locality ‘ Baños de Villavil, provincia de Catamarca ’. Villavil is located in Catamarca province, Belén Department. Material examined Argentina, Prov. Catamarca, Dept. Belén: MACN-In 380, Baños de Villa Vil, 1933, Gómez M, leg.; MACN-In 380 – 1, Baños de Villa Vil, 1933, Gómez M, leg.; MLP 10171, Baños de Villa Vil, 1951, Carranza C; Dept. Santa María: IFML 16460, 10 km before La Hoyada, 26 ° 35 ′ 20 ″ S, 66 ° 21 ′ 40 ″ W, 2760 m, 25 November 2003, Cuezzo MG leg. Shell. Dextral, fusiform, slender, thin, of 5 ½ whorls slightly convex (Figure 1 A). Shell pale brown, with darker bands (Figure 1 A). Protoconch with thick, axial, elevated costules, more marked near suture, parallel to each other (Figure 1 B – E). Costules separated by regular narrow spaces. Spiral grooves, parallel, densely arranged, crossing them, thinner than costules (Figure 1 B – E). Spire high conic, with whorls increasing regularly in diameter (Figure 1 A). Body whorl tall in relation to total height of the shell, 70 % of the total length (Figure 1 A). Teleoconch with axial oblique ribs (Figure 1 A). Suture simple, slightly deep (Figure 1 A). Aperture elongated-ovate, tall, half of total shell length (Figure 1 A). Parietal space narrow, smooth (Figure 1 A). Peristome simple, not expanded (Figure 1 A). Umbilicus narrow, partially overlapped by the peristome (Figure 1 A).	en	Miranda, María José (2014): Bostryx tortoranus (Doering, 1879) species complex (Gastropoda: Stylommatophora: Bulimulidae), a review of taxonomy and distribution of endemic species from Argentina. Journal of Natural History 49 (17 - 18): 995-1022, DOI: 10.1080/00222933.2014.981313, URL: http://dx.doi.org/10.1080/00222933.2014.981313
5F738E5FFF8E45199F997F8FE8E0FC01.taxon	description	Measurements. Type material measurements in Table 1. Range of variability of the species: masd = 5.68 – 9.46; misd = 5.54 – 9.46; th = 12.89 – 20.63; bwh = 9.64 – 14.60; sh = 4.03 – 7.97; ah = 7.05 – 10.02; mad = 4.28 – 6.30. External morphology. Lateral groove from genital orifice towards mantle collar, well marked. Foot elongate, basal sole homogeneous, not divided. Digestive system. Jaw arched with 12 plaques, without sculpture. Central plaque rectangular divided into two minor plaques. Lateral plaques rectangular in shape, regular in size. Radula narrow and long. Central tooth small, triangular, tricuspid (Figure 2 A). First lateral tooth bicuspid, similar in size and shape to central tooth (Figure 2 A). Marginal teeth bicuspid or tricuspid (Figure 2 B, C). Pallial system (one specimen dissected). Kidney triangular, wider than long, slightly longer than pericardial cavity with inner longitudinal lamellae in contact between them. Pericardial cavity shorter than kidney. Primary ureter bordering kidney along its length. Secondary ureter parallel to rectum, opening at proximal portion of rectum. Distally the secondary ureter splitting into adrectal and abrectal branches. Interramus zone, rectangular, excavated. Rectum opening at mantle collar. Main pulmonary vein, parallel to rectum, with distal portion thinner than proximal portion. Minor veins well marked and thin. Reproductive system (one specimen dissected). Ovotestis embedded in digestive gland. Albumen gland bean-shaped (Figure 3 A, B). Hermaphroditic duct divided into three parts, central part convoluted seminal vesicle (Figure 3 A, B). Proximal and distal hermaphroditic duct portion thin and elongated, and distal portion inserting into half portion of albumen gland (Figure 3 A, B). Fertilization pouch – spermathecal complex, long and finger-shaped (Figure 3 B). Spermoviduct oviducal portion long, transversely sacculated (Figure 3 A). Distally, spermoviduct splitting into free oviduct and vas deferens (Figure 3 A). Free oviduct shorter and thinner than vagina (Figure 3 A). Bursa copulatrix sac rounded, small, with long duct, reaching distal portion of albumen gland, with broader distal portion (Figure 3 A). Bursa copulatrix duct with two different inner wall sculptures, proximal longitudinal straight folds, distal zigzag folds. Vagina cylindrical, 1 / 3 of penis length (Figure 3 A), inner wall with longitudinal parallel thick straight folds. Penial complex formed by penis, epiphallus and flagellum. Penial retractor muscle short, inserted terminally in flagellum (Figure 3 C, D). Flagellum thin, short and cylindrical (Figure 3 C, D) with inner folds diagonal with respect to central fold, longitudinal, thin fold extending towards epiphallus (Figure 3 D). Epiphallus longer than flagellum, progressively increasing its width towards penis (Figure 3 C, D). Inner epiphallus wall with straight, parallel folds in proximal portion, zigzag folds in distal portion (Figure 3 D). Penis as long as epiphallus, divided into two regions by a thinner middle section (Figure 3 C, D). Proximal penis swollen, internally wall sculpture consisting of thin straight folds followed by smooth area. Distal penis cylindrical with inner straight longitudinal folds area (Figure 3 D). Inner penial papilla in proximal penis portion, elongated, triangular with terminal pore (Figure 3 D). Penis sheath muscular, folded upon itself in its proximal portion, overlapping the distal portion of penis (Figure 3 D). Retractor muscle, not divided, inserting on upper end of penis sheath (Figure 3 A). Vas deferens thin, inserting at basal penis sheath (Figure 3 A). Atrium short (Figure 3 A), inner wall with zigzag folds.	en	Miranda, María José (2014): Bostryx tortoranus (Doering, 1879) species complex (Gastropoda: Stylommatophora: Bulimulidae), a review of taxonomy and distribution of endemic species from Argentina. Journal of Natural History 49 (17 - 18): 995-1022, DOI: 10.1080/00222933.2014.981313, URL: http://dx.doi.org/10.1080/00222933.2014.981313
5F738E5FFF8E45199F997F8FE8E0FC01.taxon	discussion	Remarks Bostryx rudisculptus was originally described in Bulimulus (Scansicochlea) Pilsbry, by Parodiz (1956), based on shell characters. However, strikingly in 1957 Parodiz did not list Bulimulus (Scansicochlea) rudisculptus as a species present in Argentina. Later Miquel (1995) relocated B. rudisculptus in Bostryx, the genus in which it is currently maintained (Cuezzo et al. 2013). The protoconch ’ s sculpture and the reproductive system organization confirm its classification in Bostryx. Bostryx rudisculptus has a similar range to Bostryx catamarcanus (Parodiz 1956), a species which can be clearly differentiated by its shell shape and size. The shell of B. rudisculptus is smaller in height and diameter as well as aperture (in height and diameter); the teleoconch has ribs and darker bands. In B. catamarcanus in contrast, the teleoconch has costules and uniform coloration. Bostryx rudisculptus is one of the smaller Argentinean species in the genus Bostryx, together with Bostryx cordillerae (Strobel), Bostryx costellatus (Hylton Scott) and Bostryx reedi (Parodiz). Bostryx rudisculptus differs from B. cordillerae and B. reedi by its shell that has ribs and a taller aperture. Moreover it differs from B. reedi because of its slender shell, without parietal callus and narrower umbilicus. Bostryx rudisculptus differs from B. costellatus by its more slender shell, with narrower umbilicus, thinner walls and a teleoconch with densely arranged ribs, whereas in B. costellatus there are thicker and higher ribs, and these are more widely spaced.	en	Miranda, María José (2014): Bostryx tortoranus (Doering, 1879) species complex (Gastropoda: Stylommatophora: Bulimulidae), a review of taxonomy and distribution of endemic species from Argentina. Journal of Natural History 49 (17 - 18): 995-1022, DOI: 10.1080/00222933.2014.981313, URL: http://dx.doi.org/10.1080/00222933.2014.981313
5F738E5FFF8E45199F997F8FE8E0FC01.taxon	distribution	Distribution and habitat Bostryx rudisculptus is endemic to Catamarca province (26 – 27 ° S, 66 – 67 ° W), and known from Belén and Santa María Departments. It inhabits High Monte ecoregion (Figure 4) ranging in altitude between 1630 and 2760 m. It is found under rocks in contact with moist soil.	en	Miranda, María José (2014): Bostryx tortoranus (Doering, 1879) species complex (Gastropoda: Stylommatophora: Bulimulidae), a review of taxonomy and distribution of endemic species from Argentina. Journal of Natural History 49 (17 - 18): 995-1022, DOI: 10.1080/00222933.2014.981313, URL: http://dx.doi.org/10.1080/00222933.2014.981313
5F738E5FFF8145039FED7FC5ED68FBC8.taxon	description	(Figures 4 – 8)	en	Miranda, María José (2014): Bostryx tortoranus (Doering, 1879) species complex (Gastropoda: Stylommatophora: Bulimulidae), a review of taxonomy and distribution of endemic species from Argentina. Journal of Natural History 49 (17 - 18): 995-1022, DOI: 10.1080/00222933.2014.981313, URL: http://dx.doi.org/10.1080/00222933.2014.981313
5F738E5FFF8145039FED7FC5ED68FBC8.taxon	materials_examined	Type material Bulimulus (Scansicochlea) martinezi: Holotype (MLP 11448), Paratypes (MLP 11011 (1), MLP 11012 (1), MLP 11445 (1), MLP 11446 (1), MLP 11447 (1), MACN-In 27208 (3 )). Bulimulus (Scansicochlea) cicheroi: Holotype (MLP 10996), Paratypes (MLP 10995 (1), MLP 10997 (1), MACN-In 27281 (1 )). Type locality ‘ Chancani, Sierra de Pocho, Córdoba ’. Chancani is located in western Cordoba province, Pocho Department. Material examined Argentina, Prov. Córdoba, Dept. Pocho: IFML 15608 (ex CWW 911), Sierra de Pocho, between Las Palmas and Chancani, 1050 m, 12 November 1967, Weyrauch W leg. and det.; IFML 15611 (ex CWW 1257), Sierra de Pocho, on the road between Las Palmas and Chancani, 1050 – 1250 m, 11 December 1969, Weyrauch W leg. and det.; IFML 15610 (ex CWW 2889), Sierra de Pocho, between Las Palmas and Chancani, north-west slope of Sierra de Pocho, 600 – 900 m, 2 December 1969, Weyrauch W leg. and det.; IFML 15609 (ex SMF 195593), Sierra de Pocho, route between Las Palmas and Chancani, 1050 – 1250 m, 2 December 1969, Weyrauch W leg. and det.; IFML 15506 A, Chancani, from Las Palmas to El Cadillo, 869 m, 31 ° 22 ′ 01 ″ S, 65 ° 24 ′ 74 ″ W, 18 March 2006, Cuezzo MG and Salas Oroño E leg.; IFML 15507 A, Chancani, from Las Palmas to El Cadillo, before the third tunnel, 921 m, 31 ° 22 ′ 17 ″ S, 65 ° 24 ′ 53 ″ W, 18 March 2006, Cuezzo MG and Salas Oroño E leg.; IFML 15508 A, Chancani, from Las Palmas to El Cadillo, 1440 m, 31 ° 22 ′ 60 ″ S, 65 ° 23 ′ 30 ″ W, 18 March 2006, Cuezzo MG and Salas Oroño E leg.; IBN 826, Sierra de Pocho, 1021 m, 31 ° 22.273 ′ S, 65 ° 21.080 ′ W, 3 December 2012, Miranda MJ and Romero F leg.; IBN 829, Sierra de Pocho, Quebrada de La Mermela, 1085 m, 32 ° 22.485 ′ S, 65 ° 23.477 ′ W, Miranda MJ leg.; MACN-In 25870, Chancani, 1944, Castellanos A. leg.; MACN-In 27208, Sierra de Pocho, between Las Palmas and Chancani, Martínez A leg.; MACN-In 27431 A, Sierra de Pocho, between Las Palmas and Chancani, November 1966, Cichero M leg.; MACN-In 36892 A, Sierra de Pocho, between Las Palmas and Chancani; MLP 11011, Sierra de Pocho, between Las Palmas and Chancani; MLP 11012, Sierra de Pocho, between Las Palmas and Chancani, January 1965, Martínez A leg.; MLP 11446, Sierra de Pocho, between Las Palmas and Chancani, January 1965, Martínez A leg.; MLP 11447, Sierra de Pocho, between Las Palmas and Chancani, January 1965, Martínez A leg.; MLP 11448, Pampa de Pocho, Chancani, January 1965, Martínez A leg. Prov. San Luis, Dept. Junín: MACN-In 27281, Quebrada de Cautana, 3 December 1966, Cichero JA leg.; MLP 10995, Quebrada de Cautana; MLP 10996, Quebrada de Cautana, Cichero JA leg.; MLP 10997, Quebrada de Cautana, Cichero JA leg., MLP 11441, Quebrada de Cautana, 3 December 1966, Cichero JA leg. Shell. Dextral, fusiform, slender, thin, with 6 to 6 ¼, flat to slightly convex whorls (Figure 5 A – C). Shell uniform yellowish or with darker bands from the third whorl in some specimens. First whorl of the protoconch with thick, axial, elevated costules, parallel to each other and spiral grooves with the same thickness, densely arranged (Figure 6 A, B). Second whorl of protoconch with axial, elevated costules, thicker than the spiral grooves and more densely arranged than the first whorl (Figure 6 A, C, D). Spire high conic, with whorls increasing regularly in diameter (Figure 5 A – C). Body whorl tall in relation to total height of the shell, 80 % of the total length (Figure 5 A – C). Teleoconch with axial oblique elevated costules, more evident from fourth whorl (Figure 5 A – C). Spiral lines crossing them in some specimens. Suture simple, slightly deep (Figure 5 A – C). Aperture elongated-ovate, short, 40 % of total shell length (Figure 5 A – C). Parietal space narrow, smooth (Figure 5 A – C). Peristome simple, not expanded (Figure 5 A – C). Umbilicus narrow (Figure 5 A – C).	en	Miranda, María José (2014): Bostryx tortoranus (Doering, 1879) species complex (Gastropoda: Stylommatophora: Bulimulidae), a review of taxonomy and distribution of endemic species from Argentina. Journal of Natural History 49 (17 - 18): 995-1022, DOI: 10.1080/00222933.2014.981313, URL: http://dx.doi.org/10.1080/00222933.2014.981313
5F738E5FFF8145039FED7FC5ED68FBC8.taxon	description	Measurements. Type material measurements in Table 1. Range of variability of the species: masd = 7.89 – 11.0; misd = 7.27 – 9.70; th = 18.30 – 24.60; bwh = 13.80 – 17.70; sh = 5.40 – 8.80; ah = 9.70 – 12.70; mad = 4.98 – 6.97. External morphology. Animal body homogeneously pale brown, with lateral groove from genital orifice towards mantle collar, well marked. Foot elongate, basal sole homogeneous, not divided. Digestive system. Jaw arched with 14 plaques and fine transverse striae. Central plaque triangular divided into two minor triangular plaques. Lateral plaques rectangular in shape, irregular in size. Central tooth triangular, tricuspid, with rounded tip, smaller than the lateral teeth (Figure 7 A). First lateral tooth bicuspid, similar in size and shape to central tooth (Figure 7 A). Marginal teeth similar to lateral teeth, bicuspid with a sharply pointed ectocone (Figure 7 B, C). Pallial system (six specimens dissected). Kidney triangular, 1 / 3 of lung roof length. Secondary ureter opening at proximal portion of roof length. Rest of system idem to B. rudisculptus. Reproductive system (six specimens dissected). Free oviduct longer and thinner than vagina (Figure 8 A). Bursa copulatrix duct long, reaching distal portion of albumen gland (Figure 8 A). Bursa copulatrix duct inner wall with longitudinal zigzag folds. Vagina inner wall with longitudinal parallel thinner straight folds. Epiphallus length 2 / 3 of penis length (Figure 8 A, B). Proximal portion of inner wall of penis with straight folds followed by smooth area and distal portion with relaxed zigzag folds or straight longitudinal folds area (Figure 8 B). Rest of reproductive system idem to B. rudisculptus.	en	Miranda, María José (2014): Bostryx tortoranus (Doering, 1879) species complex (Gastropoda: Stylommatophora: Bulimulidae), a review of taxonomy and distribution of endemic species from Argentina. Journal of Natural History 49 (17 - 18): 995-1022, DOI: 10.1080/00222933.2014.981313, URL: http://dx.doi.org/10.1080/00222933.2014.981313
5F738E5FFF8145039FED7FC5ED68FBC8.taxon	discussion	Remarks Bostryx martinezi was originally described by Hylton Scott (1965) in Bulimulus (Scansicochlea) Pilsbry, based on the shell morphology, radula and jaw. In 1978, Breure published the first anatomical study of B. martinezi and relocated it in the genus Bostryx Troschel, 1847. Finally, B. martinezi was listed in synonymy with Bostryx tortoranus (Doering, 1879) by Miquel (1995) and Cuezzo et al. (2013). Based on the conchological, radular and anatomical observations performed, diagnostic characters discussed later allow the removal of B. martinezi from the synonymy with B. tortoranus. Additionally, the following change is proposed: Bulimulus (Scansicochlea) cicheroi Hylton Scott, 1967 is transferred from synonymy with B. tortoranus to synonymy with B. martinezi. This new synonymy is established based on similarities in shell morphology and the fact that shell measurements of B. cicheroi fit the range of those of B. martinezi. Moreover, the anatomical description of B. cicheroi performed by Hylton Scott (1967) is coincident with the anatomy of B. martinezi. Bostryx martinezi differs from Bostryx peristomatus (Doering, 1879), a species also found in Sierra de Pocho, because B. martinezi is smaller than B. peristomatus (in shell height and diameter) with narrower umbilicus without carina, shorter aperture and peristome simple. Moreover, in B. martinezi the teleoconch sculpture consists of axial costules whereas B. peristomatus has thicker axial ribs and a finely granulate appearance to the last whorls. Regarding morphological characters, B. martinezi differs from B. peristomatus because of its shorter kidney with respect to pulmonary roof length, the ureter secondary aperture in the proximal portion of the pulmonary roof, and the free oviduct longer than the vagina.	en	Miranda, María José (2014): Bostryx tortoranus (Doering, 1879) species complex (Gastropoda: Stylommatophora: Bulimulidae), a review of taxonomy and distribution of endemic species from Argentina. Journal of Natural History 49 (17 - 18): 995-1022, DOI: 10.1080/00222933.2014.981313, URL: http://dx.doi.org/10.1080/00222933.2014.981313
5F738E5FFF8145039FED7FC5ED68FBC8.taxon	distribution	Distribution and habitat Bostryx martinezi is found from Cordoba to San Luis (31 – 32 ° S, 65 ° W), in the Dry Chaco ecoregions (Figure 4) and within an altitudinal range between 600 and 1440 m (both records from Sierra de Pocho). It is found on rocks on the side of rivers.	en	Miranda, María José (2014): Bostryx tortoranus (Doering, 1879) species complex (Gastropoda: Stylommatophora: Bulimulidae), a review of taxonomy and distribution of endemic species from Argentina. Journal of Natural History 49 (17 - 18): 995-1022, DOI: 10.1080/00222933.2014.981313, URL: http://dx.doi.org/10.1080/00222933.2014.981313
5F738E5FFF9B45049FD7789CE832FADD.taxon	description	(Figures 4, 9 – 12)	en	Miranda, María José (2014): Bostryx tortoranus (Doering, 1879) species complex (Gastropoda: Stylommatophora: Bulimulidae), a review of taxonomy and distribution of endemic species from Argentina. Journal of Natural History 49 (17 - 18): 995-1022, DOI: 10.1080/00222933.2014.981313, URL: http://dx.doi.org/10.1080/00222933.2014.981313
5F738E5FFF9B45049FD7789CE832FADD.taxon	materials_examined	Type material Bulimulus (Bulimulus) tortoranus: Syntypes (ZMB 34718 (3 )) ex Doering, according to Breure (2013). Bulimulus (Scansicochlea) gladysae: Holotype (MLP 13345), Paratypes (MLP 11007 (1), MLP 11442 (1), MLP 11006 (1), MLP 11444 (1), MLP 11443 (1), MACN-In 27282 (4 )); Bulimulus (Scansicochlea) hyltonscottae: Holotype (MACN-In 6516), Paratypes (MACN-In 6516 – 1 (4 )); Bulimulus jujuyensis: Holotype (MACN-In 32686); Bulimulus (Bulimulus) monticola: Syntypes (ZMB 34734 (2 )); Bulimulus sporadicus gracilis: Hylton Scott, 1948 Holotype (MLP 11015), Paratypes (MLP 11014 (4 )). Type locality ‘ … quebradas húmedas de la sierra de Pocho (Totoras, Yatan, Cerro Salado) ’. The mountain system ‘ Sierra de Pocho ’ is located in Cordoba Province, Pocho Department. Material examined Argentina, Prov. Catamarca, Dept. Andalgalá: MLP 11006, Foot of Cuesta de la Chilca, 1951; MLP 11007, Foot of Cuesta de la Chilca; MLP 13345, Cuesta de la Chilca, Birabén G leg.; MLP 11442, Cuesta de la Chilca, March 1950; MLP 11443, Foot of Cuesta de la Chilca; MLP 11444, Foot of Cuesta de la Chilca; MLP 11149, Foot of Cuesta de la Chilca; MACN-In 27282, Cuesta de la Chilca, March 1950, Birabén Scott G leg. Prov. Córdoba, Dept. Pocho, IFML 16359, Sierra de Pocho, route to Las Palmas, 31 ° 20.774 ″ S, 65 ° 5.396 ′ W, 944 m, 4 December 2012, Miranda MJ leg. Prov. Jujuy: MACN-In 19461, Humahuaca river, January 1931, Budín leg. Dept. Ledesma: IFML 158, in route from Tilcara to Garganta del Diablo, 2800 m, 12 September 1962, Willink A leg.; IFML 15607 (ex CWW 2657), in route from Tilcara to Garganta del Diablo, 2800 m, 18 January 1967, Weyrauch W leg.; IFML 14877 A, Cerro Negro, Tilcara, 3000 m, 26 January 2007, Cuezzo MG and Domínguez E leg.; IFML 15530, Cerro Negro, 23 ° 33 ′ 57.8 ″ S, 65 ° 22 ′ 14.0 ″ W, 3143 m, 21 November 2010, Cuezzo MG leg.; IFML 15001 A, Cerro Negro, 23 ° 34 ′ 13.8 ″ S, 65 ° 22 ′ 39.7 ″ W, 2806 m, 21 November 2010, Cuezzo MG leg.; MLP 8071, Fraile Pintado, Birabén M leg. Dept. Tilcara: MACN-In 19590 – 1, Maimará, March 1931, Budín leg.; MACN-In 30619 A, Cerro Pucará, February 1968, de la Puente R leg.; MACN-In 19506, Tilcara; MACN-In 32686, Tilcara, January 1908, Ambrosetti H leg.; MACN-In 19507, Tilcara, February 1931, Casanova and Gatto leg.; MLP 8075, Tilcara, 1945, Hylton Scott MI leg.; MLP 8070, Tilcara, February 1944, Hylton Scott MI leg. and det.; MLP 1526 – 1, Tilcara, February 1944, Hylton Scott MI leg. and det. Dept. Tumbaya: MLP 8078, Tumbaya, Weyrauch W leg. Prov. Salta, Dept. Cachi: IFML 15505 A, Route 40, from Cachi to Cafayate, 2160 m, 25 ° 12 ′ 37 ″ S, 66 ° 11 ′ 45 ″ W, Cuezzo MG leg.; IFML 15612, Los Cardones National Park, Pampa TinTin, 3200 m, 25 ° 08 ′ 08 ″ S, 66 ° 0 ′ 50 ″ W, Delhey leg. Dept. Cafayate: IFML 15248, between Cafayate and Colalao del Valle, 26 ° 08 ′ 11 ″ S, 65 ° 57 ′ 58 ″ W, 2156 m, Cuezzo MG leg. Prov. San Luis, Dept. Ayacucho: MACN-In 6516, El Zapallar, 1947, Hylton Scott MI and Birabén M leg.; MACN-In 6516 – 1, El Zapallar, 1947, Hylton Scott MI and Birabén M. Prov. Tucumán, Dept. Tafí del Valle: IFML 15502 A, between Amaicha and El Infiernillo, after Ampimpa, 26 ° 38 ′ 06 ″ S, 65 ° 50 ′ 05 ″ W, 21 December 2001; IFML 15503 A, Route 307, between Amaicha and El Infiernillo, 26 ° 40 ′ 14 ″ S, 65 ° 49 ′ 13 ″ W, 2756 m, 21 December 2001, Ituarte C leg.; IFML 15504 A, Ruinas de Quilmes, 3 January 2004, Domínguez E leg.; IFML 15282, 5 km after Colalao del Valle, on the side of route, 26 ° 15 ′ 07 ″ S, 65 ° 56 ′ 42 ″ W, Cuezzo MG leg.; IFML 15281, Route 307 between Amaicha and El Infiernillo, 26 ° 40 ′ 49 ″ S, 65 ° 48 ′ 46 ″ W, 2850 m, 26 November 2002, Cuezzo MG leg.; IFML 15510 A, Route 307, between Tafí del Valle and Amaicha, 26 ° 38 ′ 18 ″ S, 65 ° 49 ′ 30 ″ W, 2650 m, 26 November 2002, Cuezzo MG leg.	en	Miranda, María José (2014): Bostryx tortoranus (Doering, 1879) species complex (Gastropoda: Stylommatophora: Bulimulidae), a review of taxonomy and distribution of endemic species from Argentina. Journal of Natural History 49 (17 - 18): 995-1022, DOI: 10.1080/00222933.2014.981313, URL: http://dx.doi.org/10.1080/00222933.2014.981313
5F738E5FFF9B45049FD7789CE832FADD.taxon	description	Shell. Dextral, elongated-ovate, of 5 ½ to 6 ½ whorls slightly convex (Figure 9 A – C). Shell whitish to yellowish brown. Protoconch with thick, axial, elevated costules slightly wavy, discontinuous, and parallel to each other (Figure 10 A – C). Costules separated by regular narrow spaces (Figure 10 A – C). Spiral grooves thinner than costules, parallel, more densely arranged, crossing them (Figure 10 A – C). Spire high conic, with whorls increasing regularly in diameter (Figure 9 A – C). Body whorl tall in relation to the total height of the shell, 75 % of the total length (Figure 9 A – C). Teleoconch with axial, elevated oblique ribs (Figure 9 A – C), irregularly arranged. Suture simple, deep (Figure 9 A – C). Aperture elongated-ovate, tall, half the total shell length (Figure 9 A – C). Parietal space narrow, smooth (Figure 9 A – C). Peristome simple not expanded (Figure 9 A – C). Umbilicus rimate (Figure 9 A – C). Measurements. Type material measurements in Table 1. Range of variability of the species: masd = 8.93 – 11.98; misd = 8.70 – 11.0; th = 18.80 – 29.87; bwh = 14.70 – 22.11; sh = 5.60 – 8.90; ah = 10.0 – 15.35; mad = 6.10 – 8.90. External morphology. Animal body homogeneously pale yellowish, with lateral groove from genital orifice towards mantle collar, well marked. Foot elongate, basal sole homogeneous, not divided. Digestive system. Jaw arched with nine to 11 plaques, without sculpture. Central plaque triangular not divided. Lateral plaques rectangular in shape, the two next to central plaque similar in size, following lateral plaques smaller than the central plaque. Central tooth small, triangular, tricuspid, with rounded tip (Figure 11 A, B). First lateral tooth tricuspid, similar in size and shape to central tooth (Figure 11 A). Following lateral teeth bicuspid, ectocones developed (Figure 11 C). Tooth lateral from 23 teeth of the tooth central bicuspid with serrate ectocone (splitting into two little cones) (Figure 11 C). Marginal teeth bicuspid with a rounded ectocone (Figure 11 D). Pallial system (10 specimens dissected). Secondary ureter opening at half rectum length. Rest of system idem to B. rudisculptus. Reproductive system (10 specimens dissected). Free oviduct longer and thinner than vagina (Figure 12 A). Bursa copulatrix duct long reaching middle portion of albumen gland (Figure 12 A). Bursa copulatrix duct with inner zigzag longitudinal folds. Vagina cylindrical, 1 / 3 of penis length (Figure 12 A), inner wall with straight longitudinal parallel folds. Epiphallus the same length as the penis (Figure 12 A, B). Proximal penis swollen, with inner zigzag folds followed by smooth area, distal portion cylindrical with inner zigzag folds (Figure 12 B). Rest of reproductive system idem to B. rudisculptus.	en	Miranda, María José (2014): Bostryx tortoranus (Doering, 1879) species complex (Gastropoda: Stylommatophora: Bulimulidae), a review of taxonomy and distribution of endemic species from Argentina. Journal of Natural History 49 (17 - 18): 995-1022, DOI: 10.1080/00222933.2014.981313, URL: http://dx.doi.org/10.1080/00222933.2014.981313
5F738E5FFF9B45049FD7789CE832FADD.taxon	discussion	Remarks Bostryx tortoranus was originally described by Doering (1879) in Bulimulus (Bulimulus) Leach, 1815, based on shell and jaw characters. Subsequently Pilsbry (1895 – 1896) classified it in Bulimulus (Bostryx-Lissoacme) Pilsbry, 1896 due to the absence of sculpture on the protoconch. Later, Parodiz (1946) confirmed that the protoconch of this species had a spiral striation and classified it as B. tortoranus in Peronaeus Albers, 1850. Finally, Breure (1979) relocated B. tortoranus to Bostryx, a placement currently maintained (Miquel 1995; Cuezzo et al. 2013). The examination of the type material and numerous other specimens in the Argentinean museums indicates that this species shows high shell variability in form, size and coloration. In agreement with Miquel (1995) and Cuezzo et al. (2013), Bulimulus jujuyensis, Bulimulus hyltonscottae and Bulimulus gladysae are also considered here as synonyms of B. tortoranus.	en	Miranda, María José (2014): Bostryx tortoranus (Doering, 1879) species complex (Gastropoda: Stylommatophora: Bulimulidae), a review of taxonomy and distribution of endemic species from Argentina. Journal of Natural History 49 (17 - 18): 995-1022, DOI: 10.1080/00222933.2014.981313, URL: http://dx.doi.org/10.1080/00222933.2014.981313
5F738E5FFF9B45049FD7789CE832FADD.taxon	distribution	Distribution and habitat Bostryx tortoranus is found latitudinally from Jujuy to San Luis provinces (23 – 32 ° S) and longitudinally from Jujuy to Catamarca (65 – 67 ° W). A distributional break between 27 ° and 31 ° S in the distribution of B. tortoranus has been registered. This species inhabits a variety of xerophilic environments of High Monte, Dry Chaco and Central Andean Puna ecoregions (Figure 4). It is common at altitudes from 944 to 3200 m, with the lowest locality corresponding to Pocho Department in Cordoba Province (Dry Chaco) and the highest locality including Cachi Department in Salta Province (High Monte). It is usually found inside crevices between rocks or on the vegetation.	en	Miranda, María José (2014): Bostryx tortoranus (Doering, 1879) species complex (Gastropoda: Stylommatophora: Bulimulidae), a review of taxonomy and distribution of endemic species from Argentina. Journal of Natural History 49 (17 - 18): 995-1022, DOI: 10.1080/00222933.2014.981313, URL: http://dx.doi.org/10.1080/00222933.2014.981313
